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<document id="4D05A4E936DC5230C5E6ADDCE07AFC55" ID-DOI="10.5281/zenodo.185475" ID-GBIF-Dataset="137d587d-196f-4390-a0ab-132dd8ef8601" ID-ISSN="1175-5326" ID-Zenodo-Dep="185475" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459988028658" checkinUser="plazi" docAuthor="Tong, Yanfeng &amp; Li, Shuqiang" docDate="2009" docId="777F8C78FFCCFFA130EEF8B197354199" docLanguage="en" docName="zt01988p032.pdf" docOrigin="Zootaxa 1988" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Belisana exian Tong &amp; Li, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="19" masterDocId="8B46F400FFCDFFA33079FFD0961C4209" masterDocTitle="Six new cave-dwelling pholcid spiders (Araneae: Pholcidae) from Hainan Island, with two newly recorded genera from China" masterLastPageNumber="32" masterPageNumber="17" pageNumber="18" updateTime="1698232438384" updateUser="plazi">
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<mods:title id="04B6ADBBB3A4BCC8F15585295A109F7E">Six new cave-dwelling pholcid spiders (Araneae: Pholcidae) from Hainan Island, with two newly recorded genera from China</mods:title>
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<mods:namePart id="B596CCBFF2AA08124653765F1B87196C">Tong, Yanfeng</mods:namePart>
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<mods:namePart id="AD756B7B885E8DAD38F2864744B3AE2F">Li, Shuqiang</mods:namePart>
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<mods:date id="A37A36F2502C077DC531C3A7D35CF93D">2009</mods:date>
<mods:detail id="E5E60251A51E3F910771860AC05771A8" type="volume">
<mods:number id="FEB428C07ABD95274E1A57A38F802F2D">1988</mods:number>
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<treatment id="777F8C78FFCCFFA130EEF8B197354199" ID-DOI="http://doi.org/10.5281/zenodo.6216781" ID-GBIF-Taxon="119379577" ID-Zenodo-Dep="6216781" LSID="urn:lsid:plazi:treatment:777F8C78FFCCFFA130EEF8B197354199" httpUri="http://treatment.plazi.org/id/777F8C78FFCCFFA130EEF8B197354199" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
<subSubSection id="B7CC6EE5FFCCFFA230EEF8B197494597" pageId="1" pageNumber="18" type="nomenclature">
<paragraph id="FF693D6EFFCCFFA230EEF8B197B34573" blockId="1.[151,431,1888,1950]" box="[151,431,1888,1915]" pageId="1" pageNumber="18">
<heading id="A4218A02FFCCFFA230EEF8B197B34573" bold="true" box="[151,431,1888,1915]" fontSize="11" level="1" pageId="1" pageNumber="18" reason="1">
<emphasis id="CDA2E17CFFCCFFA230EEF8B197B34573" bold="true" box="[151,431,1888,1915]" pageId="1" pageNumber="18">
<taxonomicName id="38D646EDFFCCFFA230EEF8B197564572" ID-CoL="5WGWT" box="[151,330,1889,1915]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="exian" status="sp. nov.">
<emphasis id="CDA2E17CFFCCFFA230EEF8B197564572" bold="true" box="[151,330,1889,1915]" italics="true" pageId="1" pageNumber="18">Belisana exian</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6915C07FFCCFFA2312BF8B097B34573" box="[338,431,1888,1914]" pageId="1" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FF693D6EFFCCFFA230EEF85497494597" blockId="1.[151,431,1888,1950]" box="[151,341,1924,1950]" pageId="1" pageNumber="18">
<figureCitation id="67ED21EBFFCCFFA230EEF85496E34597" box="[151,255,1924,1950]" captionStart="FIGURES 1 23" captionStartId="3.[151,273,1901,1925]" captionTargetBox="[217,1367,200,1882]" captionTargetId="figure@3.[217,1369,197,1884]" captionTargetPageId="3" captionText="FIGURES 1 23. Belisana desciscens sp. nov., 1 3; Belisana exian sp. nov., 4 6; Belisana maogan sp. nov., 7 9; Belisana yalong sp. nov., 10 12; Modisimus culicinus (Simon, 1893), 13 15; Pholcus exceptus sp. nov., 16 19; Holocneminus huangdi sp. nov., 20 23. (1, 4, 7, 10, 13, 16, 20, male habitus, dorsal view; 2, 5, 8, 11, 14, 17, 21, male habitus, lateral view; 3, 6, 9, 12, 15, 18, 23, female abdomen, ventral view; 19, 22, male sternum, ventral view.)" httpUri="https://zenodo.org/record/185476/files/figure.png" pageId="1" pageNumber="18">Figs 46</figureCitation>
,
<figureCitation id="67ED21EBFFCCFFA23172F85497494597" box="[267,341,1924,1950]" captionStart="FIGURES 30 34" captionStartId="5.[151,271,1951,1975]" captionTargetBox="[239,1350,205,1913]" captionTargetId="figure@5.[234,1351,194,1927]" captionTargetPageId="5" captionText="FIGURES 30 34. Belisana exian sp. nov. 30 31. Male left palp, prolateral (30) and retrolateral (31) views; 32. Male chelicerae, frontal view; 33 34. Cleared epigynum, ventral (33) and dorsal (34) views. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/185478/files/figure.png" pageId="1" pageNumber="18">3034</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCCFFA130EEF80097E142D1" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18" type="materials_examined">
<paragraph id="FF693D6EFFCCFFA130EEF80097E142D1" blockId="1.[151,1435,2000,2027]" lastBlockId="2.[151,1437,152,912]" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
<emphasis id="CDA2E17CFFCCFFA230EEF800974D45E3" bold="true" box="[151,337,2000,2026]" pageId="1" pageNumber="18">
<typeStatus id="206D83CCFFCCFFA230EEF80096C945E3" box="[151,213,2000,2026]" pageId="1" pageNumber="18">Type</typeStatus>
material:
</emphasis>
<typeStatus id="206D83CCFFCCFFA23123F80197D445E2" box="[346,456,2001,2027]" pageId="1" pageNumber="18" type="holotype">Holotype</typeStatus>
male,
<specimenCount id="E9D0F6E7FFCCFFA23260F801947645E2" box="[537,618,2001,2027]" pageId="1" pageNumber="18" type="male">1 male</specimenCount>
and
<specimenCount id="E9D0F6E7FFCCFFA232DDF801950B45E2" box="[676,791,2001,2027]" pageId="1" pageNumber="18" type="female">7 females</specimenCount>
<typeStatus id="206D83CCFFCCFFA23366F801958845E2" box="[799,916,2001,2027]" pageId="1" pageNumber="18" type="paratype">paratypes</typeStatus>
,
<collectingCountry id="87C17DFEFFCCFFA233D8F801921F45E2" box="[929,1027,2001,2027]" name="China" pageId="1" pageNumber="18">CHINA</collectingCountry>
: Hainan Province, Dongfang City, Donghe Town, Nanlang Village, E´xian Cave (
<geoCoordinate id="9AE25BA9FFCFFFA132B2FF48956E42BB" box="[715,882,152,178]" direction="north" orientation="latitude" pageId="2" pageNumber="19" precision="1" value="19.0071">19°00.426´N</geoCoordinate>
,
<geoCoordinate id="9AE25BA9FFCFFFA13306FF48922D42BB" box="[895,1073,152,178]" direction="east" orientation="longitude" pageId="2" pageNumber="19" precision="1" value="109.08908">109°05.345´E</geoCoordinate>
),
<date id="8B681BAEFFCFFFA13430FF48931842BB" box="[1097,1284,152,178]" pageId="2" pageNumber="19" value="2005-03-29">29 March 2005</date>
, Y. Song, X. Han, Y. Tong and G. Deng leg.
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCFFFA130BCFF35928042F6" box="[197,1180,229,255]" pageId="2" pageNumber="19" type="etymology">
<paragraph id="FF693D6EFFCFFFA130BCFF35928042F6" blockId="2.[151,1437,152,912]" box="[197,1180,229,255]" pageId="2" pageNumber="19">
<emphasis id="CDA2E17CFFCFFFA130BCFF35974A42F6" bold="true" box="[197,342,229,255]" pageId="2" pageNumber="19">Etymology:</emphasis>
The specific name is a noun in apposition, taken from the
<typeStatus id="206D83CCFFCFFFA1347CFF35922442F6" box="[1029,1080,229,255]" pageId="2" pageNumber="19">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCFFFA130BCFEDB93034345" pageId="2" pageNumber="19" type="diagnosis">
<paragraph id="FF693D6EFFCFFFA130BCFEDB93034345" blockId="2.[151,1437,152,912]" pageId="2" pageNumber="19">
<emphasis id="CDA2E17CFFCFFFA130BCFEDB975B432C" bold="true" box="[197,327,267,293]" pageId="2" pageNumber="19">Diagnosis:</emphasis>
The new species is similar to
<taxonomicName id="38D646EDFFCFFFA132D2FEDC95B2432F" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[683,942,268,294]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="gedeh">
<emphasis id="CDA2E17CFFCFFFA132D2FEDC950E432C" box="[683,786,268,293]" italics="true" pageId="2" pageNumber="19">B. gedeh</emphasis>
<bibRefCitation id="9B47409FFFCFFFA13360FEDC95B2432F" author="Huber" box="[793,942,268,294]" pageId="2" pageNumber="31" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." type="journal article" year="2005">Huber, 2005</bibRefCitation>
</taxonomicName>
, but can be distinguished by the shapes of procursus, the narrowly separated cheliceral apophyses and the position of female epigynal pocket.
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCFFFA130BCFE8897354199" pageId="2" pageNumber="19" type="description">
<paragraph id="FF693D6EFFCFFFA130BCFE88978E40C6" blockId="2.[151,1437,152,912]" pageId="2" pageNumber="19">
<emphasis id="CDA2E17CFFCFFFA130BCFE88977F437B" bold="true" box="[197,355,344,370]" pageId="2" pageNumber="19">Description:</emphasis>
Male (
<typeStatus id="206D83CCFFCFFFA131C3FE899436437A" box="[442,554,345,371]" pageId="2" pageNumber="19" type="holotype">holotype</typeStatus>
). Total length 1.26 (1.36 with clypeus), carapace width 0.56. Leg 1: 8.33 (2.02+0.22+2.13+2.87+1.09), tibia 2: 1.48, tibia 3: 0.98, tibia 4: 1.36; tibia 1 L/d: 35. Habitus as in figs 4 and 5. Whole body yellowish white; carapace and opisthosoma both without marks. Ocular area not elevated, thoracic furrow absent; distance PME-PME 0.11; diameter PME 0.04; distance PME-ALE 0.01; AME absent. Clypeus unmodified. Sternum slightly wider than long (0.41/0.35). Chelicerae as in fig. 32, with pair of curved apophyses, tips 0.24 apart. Palps as in figs 30 and 31; trochanter with short retrolateral apophysis, femur with indistinct hump proximo-dorsally, procursus complicated distally, with ventral membranous flap, bulb with hooked apophysis and simple embolus. Retrolateral trichobothrium of tibia 1 at 30%; legs without spines, vertical hairs, and curved hairs; tarsus 1 apparently with about 15 pseudosegments, but only about 10 quite distinct distally.
</paragraph>
<paragraph id="FF693D6EFFCFFFA130BCFD0C96CE4115" blockId="2.[151,1437,152,912]" pageId="2" pageNumber="19">
Variation. Tibia
<quantity id="382E908BFFCFFFA131FDFD0C97AE40FF" box="[388,434,732,758]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="2" pageNumber="19" unit="in" value="1.0">1 in</quantity>
other male: 2.37. Distance between tips of distal cheliceral apophyses in other male: 0.23.
</paragraph>
<paragraph id="FF693D6EFFCFFFA130BCFCF897354199" blockId="2.[151,1437,152,912]" pageId="2" pageNumber="19">
<emphasis id="CDA2E17CFFCFFFA130BCFCF89737414B" bold="true" box="[197,299,808,834]" pageId="2" pageNumber="19">Females</emphasis>
(
<typeStatus id="206D83CCFFCFFFA13142FCF997AE414A" box="[315,434,809,835]" pageId="2" pageNumber="19" type="paratype">paratypes</typeStatus>
). In general similar to male. Tibia 1 (n = 7): 1.82.12 (mean: 1.94). Epigynum simple and flat externally, with internal structures visible through cuticle (
<figureCitation id="67ED21EBFFCFFFA133E7FC8095F64163" box="[926,1002,848,874]" captionStart="FIGURES 1 23" captionStartId="3.[151,273,1901,1925]" captionTargetBox="[217,1367,200,1882]" captionTargetId="figure@3.[217,1369,197,1884]" captionTargetPageId="3" captionText="FIGURES 1 23. Belisana desciscens sp. nov., 1 3; Belisana exian sp. nov., 4 6; Belisana maogan sp. nov., 7 9; Belisana yalong sp. nov., 10 12; Modisimus culicinus (Simon, 1893), 13 15; Pholcus exceptus sp. nov., 16 19; Holocneminus huangdi sp. nov., 20 23. (1, 4, 7, 10, 13, 16, 20, male habitus, dorsal view; 2, 5, 8, 11, 14, 17, 21, male habitus, lateral view; 3, 6, 9, 12, 15, 18, 23, female abdomen, ventral view; 19, 22, male sternum, ventral view.)" httpUri="https://zenodo.org/record/185476/files/figure.png" pageId="2" pageNumber="19">Figs 6</figureCitation>
,
<figureCitation id="67ED21EBFFCFFFA1338FFC80920A4163" box="[1014,1046,848,874]" captionStart="FIGURES 30 34" captionStartId="5.[151,271,1951,1975]" captionTargetBox="[239,1350,205,1913]" captionTargetId="figure@5.[234,1351,194,1927]" captionTargetPageId="5" captionText="FIGURES 30 34. Belisana exian sp. nov. 30 31. Male left palp, prolateral (30) and retrolateral (31) views; 32. Male chelicerae, frontal view; 33 34. Cleared epigynum, ventral (33) and dorsal (34) views. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/185478/files/figure.png" pageId="2" pageNumber="19">33</figureCitation>
), pockets 0.15 apart. Dorsal view as in fig. 34.
</paragraph>
</subSubSection>
</treatment>
</document>