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<document id="5C2966516FAD0FCFB13952BFFEA942C8" ID-DOI="10.5281/zenodo.185475" ID-GBIF-Dataset="137d587d-196f-4390-a0ab-132dd8ef8601" ID-ISSN="1175-5326" ID-Zenodo-Dep="185475" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459988028658" checkinUser="plazi" docAuthor="Tong, Yanfeng &amp; Li, Shuqiang" docDate="2009" docId="777F8C78FFCAFFAB30EEFA8E95CB43C9" docLanguage="en" docName="zt01988p032.pdf" docOrigin="Zootaxa 1988" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Holocneminus huangdi Tong &amp; Li, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="25" masterDocId="8B46F400FFCDFFA33079FFD0961C4209" masterDocTitle="Six new cave-dwelling pholcid spiders (Araneae: Pholcidae) from Hainan Island, with two newly recorded genera from China" masterLastPageNumber="32" masterPageNumber="17" pageNumber="24" updateTime="1698232438384" updateUser="plazi">
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<mods:title id="AF438721A95A19389D44EBC6B774FC7F">Six new cave-dwelling pholcid spiders (Araneae: Pholcidae) from Hainan Island, with two newly recorded genera from China</mods:title>
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<mods:namePart id="D62E46BBF73F66B847A59DF78027D11A">Tong, Yanfeng</mods:namePart>
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<mods:namePart id="E1AA024861E958AB2FEF60CA78A147FD">Li, Shuqiang</mods:namePart>
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<mods:title id="05B9134E3D03EFBA8911ADC74EBA6C8C">Zootaxa</mods:title>
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<mods:date id="3E2B16F95311520AC4B442C1AEA361C2">2009</mods:date>
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<mods:number id="B22FB9C9342B032BD206A31FC29FDB72">1988</mods:number>
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<treatment id="777F8C78FFCAFFAB30EEFA8E95CB43C9" ID-DOI="http://doi.org/10.5281/zenodo.6216789" ID-GBIF-Taxon="119379574" ID-Zenodo-Dep="6216789" LSID="urn:lsid:plazi:treatment:777F8C78FFCAFFAB30EEFA8E95CB43C9" httpUri="http://treatment.plazi.org/id/777F8C78FFCAFFAB30EEFA8E95CB43C9" lastPageId="8" lastPageNumber="25" pageId="7" pageNumber="24">
<subSubSection id="B7CC6EE5FFCAFFA430EEFA8E976E4792" pageId="7" pageNumber="24" type="nomenclature">
<paragraph id="FF693D6EFFCAFFA430EEFA8E94054771" blockId="7.[151,537,1374,1435]" box="[151,537,1374,1400]" pageId="7" pageNumber="24">
<heading id="A4218A02FFCAFFA430EEFA8E94054771" bold="true" box="[151,537,1374,1400]" fontSize="11" level="1" pageId="7" pageNumber="24" reason="1">
<emphasis id="CDA2E17CFFCAFFA430EEFA8E94054771" bold="true" box="[151,537,1374,1400]" pageId="7" pageNumber="24">
<taxonomicName id="38D646EDFFCAFFA430EEFA8E97A94771" ID-CoL="3M8SF" box="[151,437,1374,1400]" class="Arachnida" family="Pholcidae" genus="Holocneminus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="huangdi" status="sp. nov.">
<emphasis id="CDA2E17CFFCAFFA430EEFA8E97A94771" bold="true" box="[151,437,1374,1400]" italics="true" pageId="7" pageNumber="24">Holocneminus huangdi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6915C07FFCAFFA431C5FA8E94054771" box="[444,537,1374,1400]" pageId="7" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FF693D6EFFCAFFA430EEFA51976E4792" blockId="7.[151,537,1374,1435]" box="[151,370,1409,1435]" pageId="7" pageNumber="24">
<figureCitation id="67ED21EBFFCAFFA430EEFA5197004792" box="[151,284,1409,1435]" captionStart="FIGURES 1 23" captionStartId="3.[151,273,1901,1925]" captionTargetBox="[217,1367,200,1882]" captionTargetId="figure@3.[217,1369,197,1884]" captionTargetPageId="3" captionText="FIGURES 1 23. Belisana desciscens sp. nov., 1 3; Belisana exian sp. nov., 4 6; Belisana maogan sp. nov., 7 9; Belisana yalong sp. nov., 10 12; Modisimus culicinus (Simon, 1893), 13 15; Pholcus exceptus sp. nov., 16 19; Holocneminus huangdi sp. nov., 20 23. (1, 4, 7, 10, 13, 16, 20, male habitus, dorsal view; 2, 5, 8, 11, 14, 17, 21, male habitus, lateral view; 3, 6, 9, 12, 15, 18, 23, female abdomen, ventral view; 19, 22, male sternum, ventral view.)" httpUri="https://zenodo.org/record/185476/files/figure.png" pageId="7" pageNumber="24">Figs 2023</figureCitation>
,
<figureCitation id="67ED21EBFFCAFFA43150FA51976E4792" box="[297,370,1409,1435]" captionStart="FIGURES 48 53" captionStartId="10.[151,269,1916,1940]" captionTargetBox="[246,1348,194,1891]" captionTargetId="figure@10.[239,1349,194,1892]" captionTargetPageId="10" captionText="FIGURES 48 53. Holocneminus huangdi sp. nov. 48 49. Male left palp, prolateral (48) and retrolateral (49) views; 50. Male chelicerae, frontal view; 51. Male left procursus, ventral view; 52 53. Cleared epigynum, ventral (52) and dorsal (53) views. Scale lines: 0.2 mm." httpUri="https://zenodo.org/record/185481/files/figure.png" pageId="7" pageNumber="24">4853</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCAFFA430EEFA1E92214455" pageId="7" pageNumber="24" type="materials_examined">
<paragraph id="FF693D6EFFCAFFA430EEFA1E92214455" blockId="7.[151,1437,1486,2015]" pageId="7" pageNumber="24">
<emphasis id="CDA2E17CFFCAFFA430EEFA1E974947E1" bold="true" box="[151,341,1486,1512]" pageId="7" pageNumber="24">
<typeStatus id="206D83CCFFCAFFA430EEFA1E96CA47E1" box="[151,214,1486,1512]" pageId="7" pageNumber="24">Type</typeStatus>
material:
</emphasis>
<typeStatus id="206D83CCFFCAFFA43124FA1E97D147E1" box="[349,461,1486,1512]" pageId="7" pageNumber="24" type="holotype">Holotype</typeStatus>
male,
<specimenCount id="E9D0F6E7FFCAFFA43259FA1E949147E1" box="[544,653,1486,1512]" pageId="7" pageNumber="24" type="male">14 males</specimenCount>
and
<specimenCount id="E9D0F6E7FFCAFFA432B3FA1E955247E1" box="[714,846,1486,1512]" pageId="7" pageNumber="24" type="female">13 females</specimenCount>
<typeStatus id="206D83CCFFCAFFA4332FFA1E95D147E1" box="[854,973,1486,1512]" pageId="7" pageNumber="24" type="paratype">paratypes</typeStatus>
,
<collectingCountry id="87C17DFEFFCAFFA433A3FA1E922247E1" box="[986,1086,1486,1512]" name="China" pageId="7" pageNumber="24">CHINA</collectingCountry>
: Hainan Province, Dongfang City,
<taxonomicName id="38D646EDFFCAFFA430AEFA2597204406" box="[215,316,1525,1551]" class="Arachnida" family="Pholcidae" genus="Holocneminus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="27" phylum="Arthropoda" rank="species" species="huangdi" status="sp. nov.">Huangdi</taxonomicName>
Cave (
<geoCoordinate id="9AE25BA9FFCAFFA431EBFA25942B4406" box="[402,567,1525,1551]" direction="north" orientation="latitude" pageId="7" pageNumber="24" precision="1" value="18.962">18°57.720´N</geoCoordinate>
,
<geoCoordinate id="9AE25BA9FFCAFFA4323DFA2594E84406" box="[580,756,1525,1551]" direction="east" orientation="longitude" pageId="7" pageNumber="24" precision="1" value="109.13955">109°08.373´E</geoCoordinate>
),
<date id="8B681BAEFFCAFFA43372FA2595DF4406" box="[779,963,1525,1551]" pageId="7" pageNumber="24" value="2005-03-20">20 March 2005</date>
, Y. Song, X. Han, Y. Tong and G. Deng leg.;
<specimenCount id="E9D0F6E7FFCAFFA430A1F9CC974F443F" box="[216,339,1564,1590]" pageId="7" pageNumber="24" type="female">3 females</specimenCount>
<typeStatus id="206D83CCFFCAFFA43125F9CC97C5443F" box="[348,473,1564,1590]" pageId="7" pageNumber="24" type="paratype">paratypes</typeStatus>
,
<collectingCountry id="87C17DFEFFCAFFA4319EF9CC9453443F" box="[487,591,1564,1590]" name="China" pageId="7" pageNumber="24">CHINA</collectingCountry>
: Hainan Province, Changjiang County, Qicha Town, Baoyao Cave (
<geoCoordinate id="9AE25BA9FFCAFFA430E6F992975F4455" box="[159,323,1602,1628]" direction="north" orientation="latitude" pageId="7" pageNumber="24" precision="1" value="19.101534">19°06.092´N</geoCoordinate>
,
<geoCoordinate id="9AE25BA9FFCAFFA43136F99297E14455" box="[335,509,1602,1628]" direction="east" orientation="longitude" pageId="7" pageNumber="24" precision="1" value="109.020134">109°01.208´E</geoCoordinate>
),
<date id="8B681BAEFFCAFFA4326AF99294D44455" box="[531,712,1602,1628]" pageId="7" pageNumber="24" value="2005-04-02" valueMax="2005-04-03" valueMin="2005-04-02">23 April 2005</date>
, collector same as for
<typeStatus id="206D83CCFFCAFFA433B6F99292254455" box="[975,1081,1602,1628]" pageId="7" pageNumber="24" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCAFFA430BCF9B89280448A" box="[197,1180,1640,1667]" pageId="7" pageNumber="24" type="etymology">
<paragraph id="FF693D6EFFCAFFA430BCF9B89280448A" blockId="7.[151,1437,1486,2015]" box="[197,1180,1640,1667]" pageId="7" pageNumber="24">
<emphasis id="CDA2E17CFFCAFFA430BCF9B8974A448B" bold="true" box="[197,342,1640,1666]" pageId="7" pageNumber="24">Etymology:</emphasis>
The specific name is a noun in apposition, taken from the
<typeStatus id="206D83CCFFCAFFA4347CF9B99224448A" box="[1029,1080,1641,1667]" pageId="7" pageNumber="24">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCAFFA430BCF95F92BC44FE" pageId="7" pageNumber="24" type="diagnosis">
<paragraph id="FF693D6EFFCAFFA430BCF95F92BC44FE" blockId="7.[151,1437,1486,2015]" pageId="7" pageNumber="24">
<emphasis id="CDA2E17CFFCAFFA430BCF95F975044A0" bold="true" box="[197,332,1679,1705]" pageId="7" pageNumber="24">Diagnosis:</emphasis>
The new species can be easily distinguished from
<taxonomicName id="38D646EDFFCAFFA433C7F940975444D9" authority="Berland, 1942 (Beatty 2008)" authorityName="Berland, 1942 (Beatty" authorityYear="2008" class="Arachnida" family="Pholcidae" genus="Holocneminus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="piritarsis">
<emphasis id="CDA2E17CFFCAFFA433C7F94092FA44A0" box="[958,1254,1680,1705]" italics="true" pageId="7" pageNumber="24">Holocneminus piritarsis</emphasis>
<bibRefCitation id="9B47409FFFCAFFA43496F940938044A3" author="Berland" box="[1263,1436,1680,1706]" pageId="7" pageNumber="31" refString="Berland, L. (1942) Polynesian spiders. Occasional Papers of the Bernice Pauahi Bishop Museum, 17, 1 - 24." type="journal article" year="1942">Berland, 1942</bibRefCitation>
(
<bibRefCitation id="9B47409FFFCAFFA430E6F966975C44D9" author="Beatty" box="[159,320,1718,1744]" pageId="7" pageNumber="31" refString="Beatty, J. A., Berry, J. W. &amp; Huber, B. A. (2008) The pholcid spiders of Micronesia and Polynesia (Araneae, Pholcidae). Journal of Arachnology, 36, 1 - 25." type="journal article" year="2008">Beatty 2008</bibRefCitation>
)
</taxonomicName>
by the absence of two-horned projection on clypeus, the shape of palpal bulb and the complicated procursus in male; by the presence of a short scape on epigynum in female.
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCAFFAB30BCF8D395CB43C9" lastPageId="8" lastPageNumber="25" pageId="7" pageNumber="24" type="description">
<paragraph id="FF693D6EFFCAFFAB30BCF8D392D442F6" blockId="7.[151,1437,1486,2015]" lastBlockId="8.[151,1437,152,448]" lastPageId="8" lastPageNumber="25" pageId="7" pageNumber="24">
<emphasis id="CDA2E17CFFCAFFA430BCF8D397794514" bold="true" box="[197,357,1795,1821]" pageId="7" pageNumber="24">Description:</emphasis>
Male (
<typeStatus id="206D83CCFFCAFFA431C5F8D494324517" box="[444,558,1796,1822]" pageId="7" pageNumber="24" type="holotype">holotype</typeStatus>
). Total length 1.96 (2.2 with clypeus), carapace width 0.96. Leg 1: 8.02 (2.12+0.31+2.16+2.68+0.75), tibia 2: 1.78, tibia 3: 1.5, tibia 4: 1.97; tibia 1 L/d: 23. Habitus as in figs 20 and 21. Carapace yellow with wide median brown mark. Ocular area yellow, but brown laterally, thoracic groove distinct; sternum yellowish, shape as in fig. 22. Opisthosoma pale gray, with some spots dorsally and laterally. Distance PME-PME 0.1; diameter PME 0.09; distance PME-ALE 0.03; diameter AME 0.03. Chelicerae as in fig. 50, with stridulatory files, and a pair of strongly sclerotized frontal apophyses and two pairs of small cones proximolaterally. Palps as in figs 48, 49 and 51; bulbus with distinctive internal duct; femur conspicuously enlarged; procursus relatively huge and complicated. Retrolateral trichobothrium of tibia 1 at 54%; Legs without spines, curved and vertical hairs; tarsus 1 with about 6 pseudosegments.
</paragraph>
<paragraph id="FF693D6EFFC5FFAB30BCFEDC95DB432F" blockId="8.[151,1437,152,448]" box="[197,967,268,294]" pageId="8" pageNumber="25">
Variation. Tibia
<quantity id="382E908BFFC5FFAB31FAFEDC97AC432F" box="[387,432,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="25" unit="in" value="1.0">1 in</quantity>
other males (n = 14): 1.982.34 (mean: 2.19)
</paragraph>
<paragraph id="FF693D6EFFC5FFAB30BCFEE295CB43C9" blockId="8.[151,1437,152,448]" pageId="8" pageNumber="25">
<emphasis id="CDA2E17CFFC5FFAB30BCFEE297284345" bold="true" box="[197,308,306,332]" pageId="8" pageNumber="25">Females.</emphasis>
In general similar to male. Tibia
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females from
<taxonomicName id="38D646EDFFC5FFAB33DDFEE292164345" box="[932,1034,306,332]" class="Arachnida" family="Pholcidae" genus="Holocneminus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="27" phylum="Arthropoda" rank="species" species="huangdi" status="sp. nov.">Huangdi</taxonomicName>
Cave (n = 31): 1.872.53 (mean: 2.19); from Baoyou Cave (n = 3): 1.96, 2.03 (leg I lost in one specimen). Epigynum as in figs 23 and 52, with a short scape pointing posteriorly. Lateral surface of scape with pair of pockets, possibly to accommodate the male cheliceral apophyses during copulation. Dorsal view as in fig. 53.
</paragraph>
</subSubSection>
</treatment>
</document>