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<document id="717912FE25B92C8620914C0859922134" ID-DOI="10.5281/zenodo.185475" ID-GBIF-Dataset="137d587d-196f-4390-a0ab-132dd8ef8601" ID-ISSN="1175-5326" ID-Zenodo-Dep="185475" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459988028658" checkinUser="plazi" docAuthor="Tong, Yanfeng &amp; Li, Shuqiang" docDate="2009" docId="777F8C78FFCAFFA430EEFF4794BF4647" docLanguage="en" docName="zt01988p032.pdf" docOrigin="Zootaxa 1988" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Belisana yalong Tong &amp; Li, 2009, sp. nov." docType="treatment" docVersion="9" lastPageNumber="24" masterDocId="8B46F400FFCDFFA33079FFD0961C4209" masterDocTitle="Six new cave-dwelling pholcid spiders (Araneae: Pholcidae) from Hainan Island, with two newly recorded genera from China" masterLastPageNumber="32" masterPageNumber="17" pageNumber="24" updateTime="1698232438384" updateUser="plazi">
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<mods:title id="D9306D7B4D76815F361C720DECFFFEC7">Six new cave-dwelling pholcid spiders (Araneae: Pholcidae) from Hainan Island, with two newly recorded genera from China</mods:title>
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<mods:namePart id="4AA94438B07CDA41570375808CA1AFC5">Tong, Yanfeng</mods:namePart>
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<mods:namePart id="CF49023FDB1DA076427F30FCA2996293">Li, Shuqiang</mods:namePart>
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<mods:date id="6F8E87F758633A8E14AA8119CADB2DCE">2009</mods:date>
<mods:detail id="5D0BF0A5EFB6EDF2F70813683F0B3833" type="volume">
<mods:number id="D3F452FA6537D2B817FA961FA12FF0A6">1988</mods:number>
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<treatment id="777F8C78FFCAFFA430EEFF4794BF4647" ID-DOI="http://doi.org/10.5281/zenodo.6216785" ID-GBIF-Taxon="119379579" ID-Zenodo-Dep="6216785" LSID="urn:lsid:plazi:treatment:777F8C78FFCAFFA430EEFF4794BF4647" httpUri="http://treatment.plazi.org/id/777F8C78FFCAFFA430EEFF4794BF4647" lastPageNumber="24" pageId="7" pageNumber="24">
<subSubSection id="B7CC6EE5FFCAFFA430EEFF47976E42DD" pageId="7" pageNumber="24" type="nomenclature">
<paragraph id="FF693D6EFFCAFFA430EEFF4797A242B8" blockId="7.[151,446,151,212]" box="[151,446,151,177]" pageId="7" pageNumber="24">
<heading id="A4218A02FFCAFFA430EEFF4797A242B8" bold="true" box="[151,446,151,177]" fontSize="11" level="1" pageId="7" pageNumber="24" reason="1">
<emphasis id="CDA2E17CFFCAFFA430EEFF4797A242B8" bold="true" box="[151,446,151,177]" pageId="7" pageNumber="24">
<taxonomicName id="38D646EDFFCAFFA430EEFF47974542B8" ID-CoL="68GYL" box="[151,345,151,177]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="yalong" status="sp. nov.">
<emphasis id="CDA2E17CFFCAFFA430EEFF47974542B8" bold="true" box="[151,345,151,177]" italics="true" pageId="7" pageNumber="24">Belisana yalong</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D6915C07FFCAFFA43118FF4797A242B8" box="[353,446,151,177]" pageId="7" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FF693D6EFFCAFFA430EEFF6A976E42DD" blockId="7.[151,446,151,212]" box="[151,370,186,212]" pageId="7" pageNumber="24">
<figureCitation id="67ED21EBFFCAFFA430EEFF6A970042DD" box="[151,284,186,212]" captionStart="FIGURES 1 23" captionStartId="3.[151,273,1901,1925]" captionTargetBox="[217,1367,200,1882]" captionTargetId="figure@3.[217,1369,197,1884]" captionTargetPageId="3" captionText="FIGURES 1 23. Belisana desciscens sp. nov., 1 3; Belisana exian sp. nov., 4 6; Belisana maogan sp. nov., 7 9; Belisana yalong sp. nov., 10 12; Modisimus culicinus (Simon, 1893), 13 15; Pholcus exceptus sp. nov., 16 19; Holocneminus huangdi sp. nov., 20 23. (1, 4, 7, 10, 13, 16, 20, male habitus, dorsal view; 2, 5, 8, 11, 14, 17, 21, male habitus, lateral view; 3, 6, 9, 12, 15, 18, 23, female abdomen, ventral view; 19, 22, male sternum, ventral view.)" httpUri="https://zenodo.org/record/185476/files/figure.png" pageId="7" pageNumber="24">Figs 1012</figureCitation>
,
<figureCitation id="67ED21EBFFCAFFA43150FF6A976E42DD" box="[297,370,186,212]" captionStart="FIGURES 42 47" captionStartId="9.[151,270,1943,1967]" captionTargetBox="[237,1350,211,1920]" captionTargetId="figure@9.[235,1353,194,1929]" captionTargetPageId="9" captionText="FIGURES 42 47. Belisana yalong sp. nov. 42 43. Male left palp, prolateral (42) and retrolateral (43) views; 44. Male left procursus, prolateral view; 45. Male chelicerae, frontal view; 46 47. Cleared epigynum, ventral (46) and dorsal (47) views. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/185480/files/figure.png" pageId="7" pageNumber="24">4247</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCAFFA430EEFED794904366" pageId="7" pageNumber="24" type="materials_examined">
<paragraph id="FF693D6EFFCAFFA430EEFED794904366" blockId="7.[151,1437,263,1102]" pageId="7" pageNumber="24">
<emphasis id="CDA2E17CFFCAFFA430EEFED797524328" bold="true" box="[151,334,263,289]" pageId="7" pageNumber="24">
<typeStatus id="206D83CCFFCAFFA430EEFED796CE4328" box="[151,210,263,289]" pageId="7" pageNumber="24">Type</typeStatus>
material:
</emphasis>
<typeStatus id="206D83CCFFCAFFA43121FED897D9432B" box="[344,453,264,290]" pageId="7" pageNumber="24" type="holotype">Holotype</typeStatus>
male,
<specimenCount id="E9D0F6E7FFCAFFA4326FFED8946C432B" box="[534,624,264,290]" pageId="7" pageNumber="24" type="male">2 males</specimenCount>
and
<specimenCount id="E9D0F6E7FFCAFFA432D2FED89501432B" box="[683,797,264,290]" pageId="7" pageNumber="24" type="female">5 females</specimenCount>
<typeStatus id="206D83CCFFCAFFA4335CFED89585432B" box="[805,921,264,290]" pageId="7" pageNumber="24" type="paratype">paratypes</typeStatus>
,
<collectingCountry id="87C17DFEFFCAFFA433DCFED8921B432B" box="[933,1031,264,290]" name="China" pageId="7" pageNumber="24">CHINA</collectingCountry>
: Hainan Province, Dongfang City, Donghe Town,
<taxonomicName id="38D646EDFFCAFFA43134FEFE97BD4341" box="[333,417,302,328]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="26" phylum="Arthropoda" rank="species" species="yalong" status="sp. nov.">Yalong</taxonomicName>
Village, a cave without special name (
<geoCoordinate id="9AE25BA9FFCAFFA43313FEFE92134341" box="[874,1039,302,328]" direction="north" orientation="latitude" pageId="7" pageNumber="24" precision="1" value="18.98225">18°58.935´N</geoCoordinate>
,
<geoCoordinate id="9AE25BA9FFCAFFA43462FEFE92D64341" box="[1051,1226,302,328]" direction="east" orientation="longitude" pageId="7" pageNumber="24" precision="1" value="108.8866">108°53.196´E</geoCoordinate>
),
<date id="8B681BAEFFCAFFA43499FEFE938B4341" box="[1248,1431,302,328]" pageId="7" pageNumber="24" value="2005-03-31">31 March 2005</date>
, Y. Song, X. Han, Y. Tong and G. Deng leg.
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCAFFA430BCFEAB9280439F" box="[197,1180,379,406]" pageId="7" pageNumber="24" type="etymology">
<paragraph id="FF693D6EFFCAFFA430BCFEAB9280439F" blockId="7.[151,1437,263,1102]" box="[197,1180,379,406]" pageId="7" pageNumber="24">
<emphasis id="CDA2E17CFFCAFFA430BCFEAB974A439C" bold="true" box="[197,342,379,405]" pageId="7" pageNumber="24">Etymology:</emphasis>
The specific name is a noun in apposition, taken from the
<typeStatus id="206D83CCFFCAFFA4347CFEAC9224439F" box="[1029,1080,380,406]" pageId="7" pageNumber="24">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCAFFA430BCFE72935443EA" pageId="7" pageNumber="24" type="diagnosis">
<paragraph id="FF693D6EFFCAFFA430BCFE72935443EA" blockId="7.[151,1437,263,1102]" pageId="7" pageNumber="24">
<emphasis id="CDA2E17CFFCAFFA430BCFE72975343B5" bold="true" box="[197,335,418,444]" pageId="7" pageNumber="24">Diagnosis:</emphasis>
The new species is similar to
<taxonomicName id="38D646EDFFCAFFA432ABFE72921943B5" authority="Huber, 2005" authorityName="Huber" authorityYear="2005" box="[722,1029,418,444]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="24" phylum="Arthropoda" rank="species" species="ketambe">
<emphasis id="CDA2E17CFFCAFFA432ABFE72954043B2" box="[722,860,418,443]" italics="true" pageId="7" pageNumber="24">B. ketambe</emphasis>
<bibRefCitation id="9B47409FFFCAFFA4331FFE72921943B5" author="Huber" box="[870,1029,418,444]" pageId="7" pageNumber="31" refString="Huber, B. A. (2005) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana Thorell 1898 (Araneae, Pholcidae). Zoologica, 155, 1 - 126." type="journal article" year="2005">Huber, 2005</bibRefCitation>
</taxonomicName>
, but can be distinguished by the cheliceral distal apophyses (tips close together), the shape of procursus and the long bulbal apophyses.
</paragraph>
</subSubSection>
<subSubSection id="B7CC6EE5FFCAFFA430BCFE3F94BF4647" pageId="7" pageNumber="24" type="description">
<paragraph id="FF693D6EFFCAFFA430BCFE3F927D4185" blockId="7.[151,1437,263,1102]" pageId="7" pageNumber="24">
<emphasis id="CDA2E17CFFCAFFA430BCFE3F97794000" bold="true" box="[197,357,495,521]" pageId="7" pageNumber="24">Description:</emphasis>
Male (
<typeStatus id="206D83CCFFCAFFA431C5FE2094324003" box="[444,558,496,522]" pageId="7" pageNumber="24" type="holotype">holotype</typeStatus>
). Total length 1.08 (1.14 with clypeus), carapace width 0.5. Leg 1: 5.58 (1.46+0.18+1.48+1.7+0.76), tibia 2: 0.98, tibia 3: 0.46, tibia 4: 0.76; tibia 1 L/d: 25. Habitus as in figs 10 and 11. Carapace yellowish white, with indistinctive pair of brownish marks laterally, and a narrow median line on ocular area. Sternum whitish. Legs yellowish white, without darker rings. Opisthosoma whitish gray, without marks. Ocular area not elevated, thoracic furrow absent; distance PME-PME 0.04; diameter PME 0.06; distance PME-ALE 0.02; AME absent. Clypeus unmodified. Sternum slightly wider than long (0.4/0.36). Chelicerae as in fig. 45, with pair of curved apophyses, tips 0.13 apart. Palps as in figs 42, 43 and 44; trochanter with short retrolateral apophysis, femur with indistinct hump proximo-dorsally, procursus complicated distally, with ventral membranous flap, bulb with hooked apophysis and simple embolus. Retrolateral trichobothrium of tibia 1 at 36%; legs without spines, vertical hairs, and curved hairs; tarsus 1 apparently with about 20 pseudosegments, but only about 10 quite distinct distally.
</paragraph>
<paragraph id="FF693D6EFFCAFFA430BCFC4997C141D3" blockId="7.[151,1437,263,1102]" pageId="7" pageNumber="24">
Variation. Tibia
<quantity id="382E908BFFCAFFA431FFFC4997A841BA" box="[390,436,921,947]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="24" unit="in" value="1.0">1 in</quantity>
other two males: 1.32, 1.45. Distance between tips of distal cheliceral apophyses in other two males: 0.11, 0.12.
</paragraph>
<paragraph id="FF693D6EFFCAFFA430BCFC3694BF4647" blockId="7.[151,1437,263,1102]" pageId="7" pageNumber="24">
<emphasis id="CDA2E17CFFCAFFA430BCFC36972C4609" bold="true" box="[197,304,998,1024]" pageId="7" pageNumber="24">Females</emphasis>
(
<typeStatus id="206D83CCFFCAFFA43138FC3697A34609" box="[321,447,998,1024]" pageId="7" pageNumber="24" type="paratype">paratypes</typeStatus>
). In general similar to male. Tibia 1 (n = 5): 1.061.22 (mean: 1.13). Epigynum simple and flat externally, with indistinctive internal dark structures visible through cuticle (
<figureCitation id="67ED21EBFFCAFFA43486FBDD9342462E" box="[1279,1374,1037,1063]" captionStart="FIGURES 1 23" captionStartId="3.[151,273,1901,1925]" captionTargetBox="[217,1367,200,1882]" captionTargetId="figure@3.[217,1369,197,1884]" captionTargetPageId="3" captionText="FIGURES 1 23. Belisana desciscens sp. nov., 1 3; Belisana exian sp. nov., 4 6; Belisana maogan sp. nov., 7 9; Belisana yalong sp. nov., 10 12; Modisimus culicinus (Simon, 1893), 13 15; Pholcus exceptus sp. nov., 16 19; Holocneminus huangdi sp. nov., 20 23. (1, 4, 7, 10, 13, 16, 20, male habitus, dorsal view; 2, 5, 8, 11, 14, 17, 21, male habitus, lateral view; 3, 6, 9, 12, 15, 18, 23, female abdomen, ventral view; 19, 22, male sternum, ventral view.)" httpUri="https://zenodo.org/record/185476/files/figure.png" pageId="7" pageNumber="24">Figs 12</figureCitation>
,
<figureCitation id="67ED21EBFFCAFFA43515FBDD9392462E" box="[1388,1422,1037,1063]" captionStart="FIGURES 42 47" captionStartId="9.[151,270,1943,1967]" captionTargetBox="[237,1350,211,1920]" captionTargetId="figure@9.[235,1353,194,1929]" captionTargetPageId="9" captionText="FIGURES 42 47. Belisana yalong sp. nov. 42 43. Male left palp, prolateral (42) and retrolateral (43) views; 44. Male left procursus, prolateral view; 45. Male chelicerae, frontal view; 46 47. Cleared epigynum, ventral (46) and dorsal (47) views. Scale lines: 0.1 mm." httpUri="https://zenodo.org/record/185480/files/figure.png" pageId="7" pageNumber="24">46</figureCitation>
), pockets 0.18 apart. Dorsal view as in fig. 47.
</paragraph>
</subSubSection>
</treatment>
</document>