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<mods:title>Comparative morphology and key to Amydetinae genera, with description of three new firefly species (Coleoptera, Lampyridae)</mods:title>
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<mods:namePart>Campello, Lucas</mods:namePart>
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<mods:namePart>Vaz, Stephanie</mods:namePart>
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<mods:affiliation>Programa de Pos-graduacao em Biodiversidade e Biologia Evolutiva-Laboratorio de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 1 - 107, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:namePart>Mermudes, Jose R. M.</mods:namePart>
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<mods:affiliation>Programa de Pos-graduacao em Biodiversidade e Biologia Evolutiva-Laboratorio de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 1 - 107, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:namePart>Ferreira, Andre L. D.</mods:namePart>
<mods:affiliation>Programa de Pos-graduacao em Biodiversidade e Biologia Evolutiva-Laboratorio de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 1 - 107, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<mods:namePart>Silveira, Luiz F. L.</mods:namePart>
<mods:affiliation>Laboratorio de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, A 1 - 107, Bloco A, Av. Carlos Chagas Filho, 373, Cidade Universitaria, Ilha do Fundao, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<taxonomicName LSID="77759220-95F4-5BE3-8B30-CA7A98B6521B" authority="Silveira &amp; Mermudes, 2013" authorityName="Silveira &amp; Mermudes" authorityYear="2013" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">Memoan Silveira &amp; Mermudes, 2013</taxonomicName>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Memoan conani sp. nov., male habitus A ventral B lateral C dorsal. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1114.77692.figure2" httpUri="https://binary.pensoft.net/fig/720410" pageId="0" pageNumber="131">Figs 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Memoan conani sp. nov., pronotum A dorsal B ventral C lateral D posterior. Scale bar: 1 mm (A-D). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows). L: length; H: height." figureDoi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Memoan conani sp. nov., elytra A dorsal B lateral C ventral; right wing D dorsal; proleg E lateral; mesoleg F lateral; metaleg G lateral. Scale bar: 1 mm (A-G)." figureDoi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Memoan conani sp. nov., abdomen A ventral B lateral C dorsal. Scale bar: 1 mm (A-C)." figureDoi="10.3897/zookeys.1114.77692.figure7" httpUri="https://binary.pensoft.net/fig/720415" pageId="0" pageNumber="131">, 7</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Memoan conani sp. nov., aedeagus A dorsal B lateral C ventral; sternum IX D ventral; sternite VIII E ventral; pygidium F dorsal. Scale bars: 200 um (A-D); 2 mm (E, F)." figureDoi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131">, 8</figureCitation>
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<taxonomicName authorityName="Silveira &amp; Mermudes" authorityYear="2013" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">Memoan</taxonomicName>
Silveira &amp; Mermudes, 2013; Silveira and Mer mudes 2014a: 204;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3835.3.2" author="Silveira, LFL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="131" pagination="325 - 337" refId="B26" refString="Silveira, LFL, Mermudes, JRM, 2014b. Ybytyramoan, a new genus of fireflies (Coleoptera: Lampyridae, Lampyrinae, Photinini) endemic to the Brazilian Atlantic Rainforest, with description of three new species. Zootaxa 3835 (3): 325 - 337, DOI: https://doi.org/10.11646/zootaxa.3835.3.2" title="Ybytyramoan, a new genus of fireflies (Coleoptera: Lampyridae, Lampyrinae, Photinini) endemic to the Brazilian Atlantic Rainforest, with description of three new species." url="https://doi.org/10.11646/zootaxa.3835.3.2" volume="3835" year="2014 b">Silveira and Mermudes 2014b</bibRefCitation>
: 325;
<bibRefCitation DOI="https://doi.org/10.1590/1678-476620151053359366" author="Silveira, LFL" journalOrPublisher="Serie Zoologia" pageId="0" pageNumber="131" pagination="359 - 366" refId="B27" refString="Silveira, LFL, Bocakova, M, Mermudes, JRM, 2015. Taxonomy of the Brazilian firefly Cladodes illigeri (Coleoptera, Lampyridae): Morphology and new records. Iheringia. Serie Zoologia 105 (3): 359 - 366, DOI: https://doi.org/10.1590/1678-476620151053359366" title="Taxonomy of the Brazilian firefly Cladodes illigeri (Coleoptera, Lampyridae): Morphology and new records. Iheringia." url="https://doi.org/10.1590/1678-476620151053359366" volume="105" year="2015">Silveira et al. 2015</bibRefCitation>
: 359;
<bibRefCitation DOI="https://doi.org/10.1080/14772000.2016.1153006" author="Silveira, LFL" journalOrPublisher="Systematics and Biodiversity" pageId="0" pageNumber="131" pagination="371 - 384" refId="B28" refString="Silveira, LFL, Khattar, G, Souto, P, Mermudes, JRM, Takiya, DM, Monteiro, RF, 2016a. Integrative taxonomy of new firefly taxa from the Atlantic Rainforest. Systematics and Biodiversity 14 (4): 371 - 384, DOI: https://doi.org/10.1080/14772000.2016.1153006" title="Integrative taxonomy of new firefly taxa from the Atlantic Rainforest." url="https://doi.org/10.1080/14772000.2016.1153006" volume="14" year="2016 a">Silveira et al. 2016a</bibRefCitation>
: 11;
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.558.6040" author="Silveira, LFL" journalOrPublisher="ZooKeys" pageId="0" pageNumber="131" pagination="55 - 75" refId="B29" refString="Silveira, LFL, Mermudes, JRM, Bocakova, M, 2016b. Systematic review of the firefly genus Scissicauda (Coleoptera, Lampyridae, Amydetinae) from Brazil. ZooKeys 558: 55 - 75, DOI: https://doi.org/10.3897/zookeys.558.6040" title="Systematic review of the firefly genus Scissicauda (Coleoptera, Lampyridae, Amydetinae) from Brazil." url="https://doi.org/10.3897/zookeys.558.6040" volume="558" year="2016 b">Silveira et al. 2016b</bibRefCitation>
: 56;
<bibRefCitation DOI="https://doi.org/10.1016/j.jcz.2018.10.006" author="Souto, PM" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="131" pagination="1 - 13" refId="B33" refString="Souto, PM, Campello, L, Khattar, G, Mermudes, JRM, Monteiro, RF, Silveira, LFL, 2019. How to design a predatory firefly? Lessons from the Photurinae (Coleoptera: Lampyridae). Zoologischer Anzeiger 278: 1 - 13, DOI: https://doi.org/10.1016/j.jcz.2018.10.006" title="How to design a predatory firefly? Lessons from the Photurinae (Coleoptera: Lampyridae)." url="https://doi.org/10.1016/j.jcz.2018.10.006" volume="278" year="2019">Souto et al. 2019</bibRefCitation>
: 2;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4585.1.4" author="Campello-Goncalves, L" journalOrPublisher="Zootaxa" pageId="0" pageNumber="131" pagination="59 - 72" refId="B3" refString="Campello-Goncalves, L, Souto, PM, Mermudes, JRM, Silveira, LFL, 2019. Uanauna gen. nov., a new genus of fireflies endemic to the Brazilian Atlantic forest (Coleoptera: Lampyridae), with key to Brazilian genera of Lucidotina. Zootaxa 4585 (1): 59 - 72, DOI: https://doi.org/10.11646/zootaxa.4585.1.4" title="Uanauna gen. nov., a new genus of fireflies endemic to the Brazilian Atlantic forest (Coleoptera: Lampyridae), with key to Brazilian genera of Lucidotina." url="https://doi.org/10.11646/zootaxa.4585.1.4" volume="4585" year="2019">
<normalizedToken originalValue="Campello-Gonçalves">Campello-Goncalves</normalizedToken>
et al. 2019
</bibRefCitation>
: 59;
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixz024" author="Martin, GJ" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="131" refId="B10" refString="Martin, GJ, Stanger-Hall, KF, Branham, MA, Silveira, LFL, Lower, SE, Hall, DW, Bybee, SM, 2019. Higher-level phylogeny and reclassification of Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 3(6): e11. [15 pp] https://doi.org/10.1093/isd/ixz024" title="Higher-level phylogeny and reclassification of Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 3 (6): e 11. [15 pp]" url="https://doi.org/10.1093/isd/ixz024" year="2019">Martin et al. 2019</bibRefCitation>
: 3;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4571.4.8" author="Nunes, VCS" journalOrPublisher="Zootaxa" pageId="0" pageNumber="131" pagination="562 - 570" refId="B15" refString="Nunes, VCS, Souto, PM, Monteiro, RF, Silveira, LFL, 2019. A second species of Araucariocladus Silveira and Mermudes, with notes on the variation in antennomere numbers in this genus (Coleoptera: Lampyridae). Zootaxa 4571 (4): 562 - 570, DOI: https://doi.org/10.11646/zootaxa.4571.4.8" title="A second species of Araucariocladus Silveira and Mermudes, with notes on the variation in antennomere numbers in this genus (Coleoptera: Lampyridae)." url="https://doi.org/10.11646/zootaxa.4571.4.8" volume="4571" year="2019">Nunes et al. 2019</bibRefCitation>
: 562;
<bibRefCitation DOI="https://doi.org/10.1016/j.jcz.2020.02.006" author="Nunes, VCS" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="131" pagination="1 - 10" refId="B16" refString="Nunes, VCS, Souto, PM, Minelli, A, Stanger-Hall, KF, Silveira, LFL, 2020. Antennomere numbers in fireflies (Coleoptera: Lampyridae): unique patterns and tentative explanations. Zoologischer Anzeiger 286: 1 - 10, DOI: https://doi.org/10.1016/j.jcz.2020.02.006" title="Antennomere numbers in fireflies (Coleoptera: Lampyridae): unique patterns and tentative explanations." url="https://doi.org/10.1016/j.jcz.2020.02.006" volume="286" year="2020">Nunes et al. 2020</bibRefCitation>
: 4, 7, 8.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="new diagnosis">
<paragraph pageId="0" pageNumber="131">New diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="131">
Vertex straight (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3C</figureCitation>
); frons convex in lateral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3D</figureCitation>
); labrum connate to fronto-clypeus, frontoclypeo-labral suture obliterate (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3C</figureCitation>
); gular margins contiguous and straight (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3B</figureCitation>
); antenna serrate, with ten antennomeres (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3G, H</figureCitation>
); pronotum rectangular, with punctures contiguous or separated by 0.1
<normalizedToken originalValue="×">x</normalizedToken>
the puncture width, except for two raised tubercles on the posterior
<normalizedToken originalValue="½">1/2</normalizedToken>
of the disc with sides divergent posteriorly (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Memoan conani sp. nov., pronotum A dorsal B ventral C lateral D posterior. Scale bar: 1 mm (A-D). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows). L: length; H: height." figureDoi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131">4A</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Memoan conani sp. nov., elytra A dorsal B lateral C ventral; right wing D dorsal; proleg E lateral; mesoleg F lateral; metaleg G lateral. Scale bar: 1 mm (A-G)." figureDoi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131">6A</figureCitation>
); elytron slightly dehiscent (i.e., sinuose inner margin) (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Memoan conani sp. nov., elytra A dorsal B lateral C ventral; right wing D dorsal; proleg E lateral; mesoleg F lateral; metaleg G lateral. Scale bar: 1 mm (A-G)." figureDoi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131">6A-C</figureCitation>
); suture between mesanepisternum and mesoventrite visible (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">5C</figureCitation>
); metathoracic discrimen as long as 3/4 of sternite length (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">5C</figureCitation>
); lanterns occupying the entire area of sternite VI and VII (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Memoan conani sp. nov., male habitus A ventral B lateral C dorsal. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1114.77692.figure2" httpUri="https://binary.pensoft.net/fig/720410" pageId="0" pageNumber="131">2A</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Memoan conani sp. nov., abdomen A ventral B lateral C dorsal. Scale bar: 1 mm (A-C)." figureDoi="10.3897/zookeys.1114.77692.figure7" httpUri="https://binary.pensoft.net/fig/720415" pageId="0" pageNumber="131">7A</figureCitation>
); pygidium with posterior margin bisinuate (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Memoan conani sp. nov., aedeagus A dorsal B lateral C ventral; sternum IX D ventral; sternite VIII E ventral; pygidium F dorsal. Scale bars: 200 um (A-D); 2 mm (E, F)." figureDoi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131">8E</figureCitation>
); ventral plate of phallus deeply bilobate (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Memoan conani sp. nov., aedeagus A dorsal B lateral C ventral; sternum IX D ventral; sternite VIII E ventral; pygidium F dorsal. Scale bars: 200 um (A-D); 2 mm (E, F)." figureDoi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131">8C</figureCitation>
), phallobase symmetrical (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Memoan conani sp. nov., aedeagus A dorsal B lateral C ventral; sternum IX D ventral; sternite VIII E ventral; pygidium F dorsal. Scale bars: 200 um (A-D); 2 mm (E, F)." figureDoi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131">8A-C</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1114.77692.figure2" httpUri="https://binary.pensoft.net/fig/720410" pageId="0" pageNumber="131" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figure 2.</emphasis>
<taxonomicName authorityName="Campello, Vaz, Mermudes &amp; Silveira" authorityYear="2022" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan conani" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="conani">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan conani</emphasis>
</taxonomicName>
sp. nov., male habitus
<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
lateral
<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
dorsal. Scale bar: 1 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figure 3.</emphasis>
<taxonomicName authorityName="Campello, Vaz, Mermudes &amp; Silveira" authorityYear="2022" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan conani" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="conani">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan conani</emphasis>
</taxonomicName>
sp. nov., head
<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
frontal
<emphasis bold="true" pageId="0" pageNumber="131">D</emphasis>
lateral
<emphasis bold="true" pageId="0" pageNumber="131">E</emphasis>
posterior
<emphasis bold="true" pageId="0" pageNumber="131">F</emphasis>
occipital; antenna
<emphasis bold="true" pageId="0" pageNumber="131">G</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="131">H</emphasis>
lateral. Scale bar: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="131">A-H</emphasis>
). Note the gular margins contiguous (arrow).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="redescription">
<paragraph pageId="0" pageNumber="131">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">
Male.
<emphasis italics="true" pageId="0" pageNumber="131">Head</emphasis>
.
</emphasis>
Capsule 1.5
<normalizedToken originalValue="×">x</normalizedToken>
wider than long (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3A</figureCitation>
); vertex straight (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3C</figureCitation>
); eyes separated by 2/3 of head width in frontal view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3C</figureCitation>
) and each eye as wide as 2/3 of head width in dorsal view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3A</figureCitation>
); frons raised and convex in lateral view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3D</figureCitation>
); labrum connate to fronto-clypeus, frontoclypeo-labral suture obliterate (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3C</figureCitation>
); gular margins contiguous and straight (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3B</figureCitation>
); antennal insertion with outer margin straight, inner margin rounded, as wide as labrum (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3C</figureCitation>
). Antenna serrate, with 10 antennomeres (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3G, H</figureCitation>
; but may the apical antennomere may be subdivided; see Discussion), as long as 1/2 of body length (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Memoan conani sp. nov., male habitus A ventral B lateral C dorsal. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1114.77692.figure2" httpUri="https://binary.pensoft.net/fig/720410" pageId="0" pageNumber="131">2A-C</figureCitation>
); pedicel as long as wide (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3G</figureCitation>
); antennomere IV 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than III, V-IX subequal in length, X as long as IV (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3G, H</figureCitation>
). Maxillary palp with palpomeres I and III 2
<normalizedToken originalValue="×">x</normalizedToken>
wider than long (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3A-D</figureCitation>
), II and IV 1.5
<normalizedToken originalValue="×">x</normalizedToken>
wider than long. Labial palp with one or two palpomeres, apical palpomere digitiform (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3B, F</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3640.1.6" author="Silveira, LFL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="131" pagination="79 - 87" refId="B24" refString="Silveira, LFL, Mermudes, JR, 2013. Memoan ciceroi gen. et sp. n., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae). Zootaxa 3640 (1): 79 - 87, DOI: https://doi.org/10.11646/zootaxa.3640.1.6" title="Memoan ciceroi gen. et sp. n., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae)." url="https://doi.org/10.11646/zootaxa.3640.1.6" volume="3640" year="2013">Silveira and Mermudes 2013</bibRefCitation>
: fig. 5). Mentum completely longitudinally divided (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3B</figureCitation>
); gular margins contiguous and straight (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3B</figureCitation>
). Occipital foramen ellipsoid in posterior view (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Memoan conani sp. nov., head A dorsal B ventral C frontal D lateral E posterior F occipital; antenna G dorsal H lateral. Scale bar: 0.5 mm (A-H). Note the gular margins contiguous (arrow)." figureDoi="10.3897/zookeys.1114.77692.figure3" httpUri="https://binary.pensoft.net/fig/720411" pageId="0" pageNumber="131">3E, F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="131">
<emphasis italics="true" pageId="0" pageNumber="131">Thorax</emphasis>
.
</emphasis>
Pronotum rectangular (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Memoan conani sp. nov., pronotum A dorsal B ventral C lateral D posterior. Scale bar: 1 mm (A-D). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows). L: length; H: height." figureDoi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131">4A, B</figureCitation>
), 2
<normalizedToken originalValue="×">x</normalizedToken>
longer than head length in ventral view (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Memoan conani sp. nov., male habitus A ventral B lateral C dorsal. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1114.77692.figure2" httpUri="https://binary.pensoft.net/fig/720410" pageId="0" pageNumber="131">2A</figureCitation>
), 1.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide in dorsal view (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Memoan conani sp. nov., pronotum A dorsal B ventral C lateral D posterior. Scale bar: 1 mm (A-D). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows). L: length; H: height." figureDoi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131">4A</figureCitation>
), with punctures contiguous or separated by 0.1 puncture width, except for two elongate tubercles on the posterior 1/2 of the disc, sides of tubercles divergent posteriorly (Figs
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Memoan conani sp. nov., pronotum A dorsal B ventral C lateral D posterior. Scale bar: 1 mm (A-D). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows). L: length; H: height." figureDoi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131">4A</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Memoan conani sp. nov., elytra A dorsal B lateral C ventral; right wing D dorsal; proleg E lateral; mesoleg F lateral; metaleg G lateral. Scale bar: 1 mm (A-G)." figureDoi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131">6A</figureCitation>
); hypomeron 2.5
<normalizedToken originalValue="×">x</normalizedToken>
longer than tall (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Memoan conani sp. nov., pronotum A dorsal B ventral C lateral D posterior. Scale bar: 1 mm (A-D). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows). L: length; H: height." figureDoi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131">4C</figureCitation>
), with punctures contiguous or separated by 0.1
<normalizedToken originalValue="×">x</normalizedToken>
the puncture width (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Memoan conani sp. nov., pronotum A dorsal B ventral C lateral D posterior. Scale bar: 1 mm (A-D). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows). L: length; H: height." figureDoi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131">4C</figureCitation>
); prosternum 0.5
<normalizedToken originalValue="×">x</normalizedToken>
as wide as pronotum in ventral view (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Memoan conani sp. nov., pronotum A dorsal B ventral C lateral D posterior. Scale bar: 1 mm (A-D). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows). L: length; H: height." figureDoi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131">4B</figureCitation>
); proendosternite as long as 1/5 of prosternum width (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Memoan conani sp. nov., pronotum A dorsal B ventral C lateral D posterior. Scale bar: 1 mm (A-D). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows). L: length; H: height." figureDoi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131">4B</figureCitation>
). Mesocutellum with wide, irregular shaped punctures, contiguous or separated by 0.1
<normalizedToken originalValue="×">x</normalizedToken>
puncture width (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">5A</figureCitation>
). Elytron slightly dehiscent (i.e. sinuose inner margin), each subparallel-sided (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Memoan conani sp. nov., elytra A dorsal B lateral C ventral; right wing D dorsal; proleg E lateral; mesoleg F lateral; metaleg G lateral. Scale bar: 1 mm (A-G)." figureDoi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131">6A-C</figureCitation>
), almost 6
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide. Hind wing (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Memoan conani sp. nov., elytra A dorsal B lateral C ventral; right wing D dorsal; proleg E lateral; mesoleg F lateral; metaleg G lateral. Scale bar: 1 mm (A-G)." figureDoi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131">6D</figureCitation>
) with vein r4 poorly sclerotized apically; radial cell poorly sclerotized posteriorly, 2
<normalizedToken originalValue="×">x</normalizedToken>
to 3
<normalizedToken originalValue="×">x</normalizedToken>
wider than long; vein r3 present or absent, vein CuA1 and CuA3+4 present or absent, vein J evanescent as long as 1/4 AP3+4 length (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Memoan conani sp. nov., elytra A dorsal B lateral C ventral; right wing D dorsal; proleg E lateral; mesoleg F lateral; metaleg G lateral. Scale bar: 1 mm (A-G)." figureDoi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131">6D</figureCitation>
,
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3640.1.6" author="Silveira, LFL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="131" pagination="79 - 87" refId="B24" refString="Silveira, LFL, Mermudes, JR, 2013. Memoan ciceroi gen. et sp. n., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae). Zootaxa 3640 (1): 79 - 87, DOI: https://doi.org/10.11646/zootaxa.3640.1.6" title="Memoan ciceroi gen. et sp. n., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae)." url="https://doi.org/10.11646/zootaxa.3640.1.6" volume="3640" year="2013">Silveira and Mermudes 2013</bibRefCitation>
: fig. 8). Metanotum 1.5
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, posterior margin straight, allocrista distinct, well-sclerotized (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">5B</figureCitation>
). Mesoventrite sclerotized (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">5C</figureCitation>
), posterior margin rounded; suture between mesanepisternum and mesoventrite visible (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">5C</figureCitation>
); mesendosternum with irregular flap-like projections (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">5E</figureCitation>
); metaendosternum spatulate, diamond-shaped (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">5D</figureCitation>
); metathoracic discrimen as long as 3/4 of sternite length (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Memoan conani sp. nov., mesoscutellum A dorsal; metanotum B dorsal; pterothorax C dorsal; mesoendoesternite D dorsal; metaendoesternite E dorsal. Scale bars: 200 µm (A, D-E); 2 mm (B, C). Note the sutures between mesoventrite / mesanespisternum and mesanepisternum / mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows)." figureDoi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131">5C</figureCitation>
). Tibial spurs absent (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Memoan conani sp. nov., elytra A dorsal B lateral C ventral; right wing D dorsal; proleg E lateral; mesoleg F lateral; metaleg G lateral. Scale bar: 1 mm (A-G)." figureDoi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131">6E-G</figureCitation>
), procoxa distally constricted, femur as long as tibia, tarsus I&gt;V&gt;II&gt;IV&gt;III (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Memoan conani sp. nov., elytra A dorsal B lateral C ventral; right wing D dorsal; proleg E lateral; mesoleg F lateral; metaleg G lateral. Scale bar: 1 mm (A-G)." figureDoi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131">6E-G</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="131">
<emphasis italics="true" pageId="0" pageNumber="131">Abdomen</emphasis>
.
</emphasis>
Lanterns occupying the entire area of sternite VI and VII (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Memoan conani sp. nov., male habitus A ventral B lateral C dorsal. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1114.77692.figure2" httpUri="https://binary.pensoft.net/fig/720410" pageId="0" pageNumber="131">2A</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Memoan conani sp. nov., abdomen A ventral B lateral C dorsal. Scale bar: 1 mm (A-C)." figureDoi="10.3897/zookeys.1114.77692.figure7" httpUri="https://binary.pensoft.net/fig/720415" pageId="0" pageNumber="131">7A</figureCitation>
); sternite VIII 3
<normalizedToken originalValue="×">x</normalizedToken>
wider than long, lateral margins rounded, posterior margin mucronate (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Memoan conani sp. nov., aedeagus A dorsal B lateral C ventral; sternum IX D ventral; sternite VIII E ventral; pygidium F dorsal. Scale bars: 200 um (A-D); 2 mm (E, F)." figureDoi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131">8E</figureCitation>
). Sternite IX symmetrical (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Memoan conani sp. nov., aedeagus A dorsal B lateral C ventral; sternum IX D ventral; sternite VIII E ventral; pygidium F dorsal. Scale bars: 200 um (A-D); 2 mm (E, F)." figureDoi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131">8D</figureCitation>
), 3
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide, posterior margin rounded. Pygidium as long as wide (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Memoan conani sp. nov., aedeagus A dorsal B lateral C ventral; sternum IX D ventral; sternite VIII E ventral; pygidium F dorsal. Scale bars: 200 um (A-D); 2 mm (E, F)." figureDoi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131">8F</figureCitation>
), posterior margin bisinuate, posterolateral angles acute, median 1/3 extending slightly beyond posterolateral angles. Phallus with dorsal and ventral plates overlapping mesal concavity of phallobase (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Memoan conani sp. nov., aedeagus A dorsal B lateral C ventral; sternum IX D ventral; sternite VIII E ventral; pygidium F dorsal. Scale bars: 200 um (A-D); 2 mm (E, F)." figureDoi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131">8A-C</figureCitation>
); dorsal plate shorter than ventral plate, ventral plate deeply cleft medially, forming two lobes, apical lobes bent dorsad; parameres symmetrical and broadly rounded at apex in dorsal and lateral views, anterior margin rounded, apically separated from each other and 0.5
<normalizedToken originalValue="×">x</normalizedToken>
shorter than phallus; phallobase symmetrical (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Memoan conani sp. nov., aedeagus A dorsal B lateral C ventral; sternum IX D ventral; sternite VIII E ventral; pygidium F dorsal. Scale bars: 200 um (A-D); 2 mm (E, F)." figureDoi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131">8A-C</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1114.77692.figure4" httpUri="https://binary.pensoft.net/fig/720412" pageId="0" pageNumber="131" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figure 4.</emphasis>
<taxonomicName authorityName="Campello, Vaz, Mermudes &amp; Silveira" authorityYear="2022" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan conani" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="conani">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan conani</emphasis>
</taxonomicName>
sp. nov., pronotum
<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
lateral
<emphasis bold="true" pageId="0" pageNumber="131">D</emphasis>
posterior. Scale bar: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="131">A-D</emphasis>
). Note the L-shaped sclerotized raised tubercles, and proendosternite (arrows).
<emphasis bold="true" pageId="0" pageNumber="131">L</emphasis>
: length;
<emphasis bold="true" pageId="0" pageNumber="131">H</emphasis>
: height.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1114.77692.figure5" httpUri="https://binary.pensoft.net/fig/720413" pageId="0" pageNumber="131" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figure 5.</emphasis>
<taxonomicName authorityName="Campello, Vaz, Mermudes &amp; Silveira" authorityYear="2022" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan conani" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="conani">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan conani</emphasis>
</taxonomicName>
sp. nov., mesoscutellum
<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
dorsal; metanotum
<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
dorsal; pterothorax
<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
dorsal; mesoendoesternite
<emphasis bold="true" pageId="0" pageNumber="131">D</emphasis>
dorsal; metaendoesternite
<emphasis bold="true" pageId="0" pageNumber="131">E</emphasis>
dorsal. Scale bars: 200
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="131">A, D-E</emphasis>
); 2 mm (
<emphasis bold="true" pageId="0" pageNumber="131">B, C</emphasis>
). Note the sutures between mesoventrite/mesanespisternum and mesanepisternum/mesepimeron, the metathoracic discrimen, and irregular flap-like projections on mesendosternum (arrows).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1114.77692.figure6" httpUri="https://binary.pensoft.net/fig/720414" pageId="0" pageNumber="131" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figure 6.</emphasis>
<taxonomicName authorityName="Campello, Vaz, Mermudes &amp; Silveira" authorityYear="2022" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan conani" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="conani">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan conani</emphasis>
</taxonomicName>
sp. nov., elytra
<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
lateral
<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
ventral; right wing
<emphasis bold="true" pageId="0" pageNumber="131">D</emphasis>
dorsal; proleg
<emphasis bold="true" pageId="0" pageNumber="131">E</emphasis>
lateral; mesoleg
<emphasis bold="true" pageId="0" pageNumber="131">F</emphasis>
lateral; metaleg
<emphasis bold="true" pageId="0" pageNumber="131">G</emphasis>
lateral. Scale bar: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="131">A-G</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1114.77692.figure7" httpUri="https://binary.pensoft.net/fig/720415" pageId="0" pageNumber="131" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figure 7.</emphasis>
<taxonomicName authorityName="Campello, Vaz, Mermudes &amp; Silveira" authorityYear="2022" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan conani" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="conani">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan conani</emphasis>
</taxonomicName>
sp. nov., abdomen
<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
lateral
<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
dorsal. Scale bar: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="131">A-C</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1114.77692.figure8" httpUri="https://binary.pensoft.net/fig/720416" pageId="0" pageNumber="131" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figure 8.</emphasis>
<taxonomicName authorityName="Campello, Vaz, Mermudes &amp; Silveira" authorityYear="2022" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan conani" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="species" species="conani">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan conani</emphasis>
</taxonomicName>
sp. nov., aedeagus
<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
dorsal
<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
lateral
<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
ventral; sternum IX
<emphasis bold="true" pageId="0" pageNumber="131">D</emphasis>
ventral; sternite VIII
<emphasis bold="true" pageId="0" pageNumber="131">E</emphasis>
ventral; pygidium
<emphasis bold="true" pageId="0" pageNumber="131">F</emphasis>
dorsal. Scale bars: 200 um (
<emphasis bold="true" pageId="0" pageNumber="131">A-D</emphasis>
); 2 mm (
<emphasis bold="true" pageId="0" pageNumber="131">E, F</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Females and immature stages.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="remarks">
<paragraph pageId="0" pageNumber="131">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="131">
The discovery of a second species of
<taxonomicName authorityName="Silveira &amp; Mermudes" authorityYear="2013" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan</emphasis>
</taxonomicName>
(see below) called for an updated diagnosis for this genus.
<taxonomicName authorityName="Silveira &amp; Mermudes" authorityYear="2013" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan</emphasis>
</taxonomicName>
is distinguishable from other amydetine genus by their serrate antennae.
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.3640.1.6" author="Silveira, LFL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="131" pagination="79 - 87" refId="B24" refString="Silveira, LFL, Mermudes, JR, 2013. Memoan ciceroi gen. et sp. n., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae). Zootaxa 3640 (1): 79 - 87, DOI: https://doi.org/10.11646/zootaxa.3640.1.6" title="Memoan ciceroi gen. et sp. n., a remarkable new firefly genus and species from the Atlantic Rainforest (Coleoptera: Lampyridae)." url="https://doi.org/10.11646/zootaxa.3640.1.6" volume="3640" year="2013">Silveira and Mermudes (2013)</bibRefCitation>
pointed out that
<taxonomicName authorityName="Silveira &amp; Mermudes" authorityYear="2013" class="Insecta" family="Lampyridae" genus="Memoan" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Memoan" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Memoan</emphasis>
</taxonomicName>
shares the following traits with the other amydetine genera: continuous glow (present in
<taxonomicName authorityName="Hoffmannsegg in Illiger" authorityYear="1807" class="Insecta" family="Lampyridae" genus="Amydetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amydetes" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Amydetes</emphasis>
</taxonomicName>
spp. and
<taxonomicName authorityName="McDermott" authorityYear="1966" class="Insecta" family="Lampyridae" genus="Magnoculus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Magnoculus" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Magnoculus</emphasis>
</taxonomicName>
spp.); eyes ventrally close-set (present in some species of
<taxonomicName authorityName="McDermott" authorityYear="1966" class="Insecta" family="Lampyridae" genus="Magnoculus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Magnoculus" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Magnoculus</emphasis>
</taxonomicName>
;
<bibRefCitation author="Constantin, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="131" refId="B4" refString="Constantin, R, 2011. Contribution a l'etude du genre Magnoculus McDermott, 1964 (Coleoptera, Lampyridae) avec description de cinq especes nouvelles de Guyane. ACOREP-France, Coleopteres de Guyane, 52-59." title="Contribution a l'etude du genre Magnoculus McDermott, 1964 (Coleoptera, Lampyridae) avec description de cinq especes nouvelles de Guyane. ACOREP-France, Coleopteres de Guyane, 52 - 59." year="2011">Constantin 2011</bibRefCitation>
); labial and maxillary apical palpomere with rounded apex (present in
<taxonomicName authorityName="Hoffmannsegg in Illiger" authorityYear="1807" class="Insecta" family="Lampyridae" genus="Amydetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amydetes" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Amydetes</emphasis>
</taxonomicName>
spp. and
<taxonomicName authorityName="McDermott" authorityYear="1966" class="Insecta" family="Lampyridae" genus="Magnoculus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Magnoculus" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Magnoculus</emphasis>
</taxonomicName>
spp.); punctures wide and irregularly outlined on pronotum and scutellum, sometimes on hypomeron as well (widespread in
<taxonomicName authorityName="McDermott" authorityYear="1966" class="Insecta" family="Lampyridae" genus="Magnoculus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Magnoculus" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Magnoculus</emphasis>
</taxonomicName>
spp., absent from disc in
<taxonomicName authorityName="Hoffmannsegg in Illiger" authorityYear="1807" class="Insecta" family="Lampyridae" genus="Amydetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amydetes" order="Coleoptera" pageId="0" pageNumber="131" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Amydetes</emphasis>
</taxonomicName>
spp.). Recent phylogenetic studies based on molecular data corroborate a placement in
<taxonomicName authorityName="E. Olivier in Wytsman" authorityYear="1907" lsidName="" pageId="0" pageNumber="131" rank="subFamily" subFamily="Amydetinae">Amydetinae</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixz024" author="Martin, GJ" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="131" refId="B10" refString="Martin, GJ, Stanger-Hall, KF, Branham, MA, Silveira, LFL, Lower, SE, Hall, DW, Bybee, SM, 2019. Higher-level phylogeny and reclassification of Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 3(6): e11. [15 pp] https://doi.org/10.1093/isd/ixz024" title="Higher-level phylogeny and reclassification of Lampyridae (Coleoptera: Elateroidea). Insect Systematics and Diversity 3 (6): e 11. [15 pp]" url="https://doi.org/10.1093/isd/ixz024" year="2019">Martin et al. 2019</bibRefCitation>
). The evolutionary history of these traits is yet to be assessed.
</paragraph>
</subSubSection>
</treatment>
</document>