484 lines
64 KiB
XML
484 lines
64 KiB
XML
<document id="08F81752F81C825DE350188F5520F049" ENCODING="UTF-8" ID-GBIF-Dataset="97f581b3-4a31-4414-8753-97a66197d5ee" ModsDocID="z01612p025" checkinTime="1247241928304" checkinUser="thomas" docAuthor="Piet Verburg & Roger Bills" docDate="2007" docId="7635B225A6FB71EE3226D879731AADE8" docLanguage="en" docName="2007_Verburg_Bills_gg1.xml" docOrigin="Zootaxa 1612" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:0C7EE498-1D50-4856-8AF2-5F59D36B029F" docTitle="Neolamprologus walteri Verburg & Bills, 2007, sp. nov." docType="treatment" docVersion="11" lastPageNumber="35" masterDocId="1A2F48BA902D92E218EEB22288BA2C71" masterDocTitle="Two new cichild species Neolamprologus (Teleostei: Cichlidae) from Lake Tanganyika, East Africa." masterLastPageNumber="44" masterPageNumber="25" pageNumber="28" updateTime="1701314756014" updateUser="plazi">
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<mods:titleInfo id="CDD1796457989A926CC93377104A8952">
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<mods:title id="BD615A552DCC809B76547F65D59AA64E">Two new cichild species Neolamprologus (Teleostei: Cichlidae) from Lake Tanganyika, East Africa.</mods:title>
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</mods:titleInfo>
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<mods:name id="48B17CAECE26432934F6BFEF369BEF51" type="personal">
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<mods:roleTerm id="58AB83256BE7BD2CE4C8EA8FD335B0F5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0C0129297DBE20ADFE17C70F4FF27C16">Piet Verburg</mods:namePart>
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</mods:name>
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<mods:name id="3D82254555A1B68E727E1707E87A4B72" type="personal">
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<mods:roleTerm id="8E284E9EEF84660354B5D907C904F738">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6391E6EF2ED51CFA6F25D8C52EDE6274">Roger Bills</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="12C6D19181129333CD33238BD7FD5242">text</mods:typeOfResource>
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<mods:title id="20A5EC73AE7796B35D1581B65013DE45">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AB5850C20940297FEF20FBBFFE843E85">
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<mods:date id="94B7F3C028BD9EE41B897EA39F1806A4">2007</mods:date>
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<mods:detail id="55160E6176A1B0E035E8143AEDCFE621" type="volume">
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<mods:number id="904DA62A7749AA06AF93D4066A76EB07">1612</mods:number>
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<mods:start id="B74134F2D4E0B2C0A1802D09F09BC49A">25</mods:start>
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<mods:end id="12BF932900EC870F0BCD883B374451AD">44</mods:end>
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<mods:location id="7649A3750FAD977719B6A0BD32BB1FD5">
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<mods:url id="4102245311D2FC02F4FD39149F02D8B8">http://www.zoobank.org/urn:lsid:zoobank.org:pub:0C7EE498-1D50-4856-8AF2-5F59D36B029F</mods:url>
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</mods:location>
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<mods:classification id="BF36C9993AEE7D86B75D493795934FEE">journal article</mods:classification>
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<mods:identifier id="A85E77BC419F60FF62053C7FB652161A" type="Plazi-Custom">z01612p025</mods:identifier>
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<treatment id="7635B225A6FB71EE3226D879731AADE8" ID-DOI="http://doi.org/10.5281/zenodo.6244586" ID-GBIF-Taxon="100122889" ID-Zenodo-Dep="6244586" LSID="urn:lsid:plazi:treatment:7635B225A6FB71EE3226D879731AADE8" httpUri="http://treatment.plazi.org/id/7635B225A6FB71EE3226D879731AADE8" lastPageNumber="35" pageNumber="28">
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<subSubSection id="AAD2CD0D6397422A2D4FF75E9DBA8D8B" type="nomenclature">
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<paragraph id="B965A96FC8CCBC99A293ACDFF4B5298E" pageNumber="28">
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<taxonomicName id="7CF0A7C6292405AC14848A626B7425C4" ID-CoL="46K4B" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
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<taxonomicNameLabel id="A6F4721ED6CA7AFEF8A5D1849724181C" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="2794381B3553A2CC4FA1C83FA722B7D3" type="description">
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<paragraph id="2D0E447D04121814EB50A101947F6BCA" pageNumber="28">(Figs. 2 and 3. See also photographs in Brichard [1989, pp. 344 and 377] and Konings [1988, p. 93])</paragraph>
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</subSubSection>
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<subSubSection id="379231122E07086E7F88DBBD75946D1F" type="nomenclature">
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<paragraph id="FE5A3E1CAC5C1897DAD344E622AEF2EE" pageNumber="28">
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<materialsCitation id="C050A5681B21E5837BACE9EFBCA05889" ID-GBIF-Occurrence="923899930" country="Tanzania">
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Type material.
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<typeStatus id="15C8DF60FCE9096EE09E4F9CAB89702F">Holotype</typeStatus>
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:
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<collectionCode id="9F53E813EDE5E40D6FAB20744E3186C3" collectionName="SAIAB">SAIAB</collectionCode>
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56090, male, 48.4mm SL, Tembo Rock,
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<collectingCountry id="86E64D30D1306813F0740B417DB62CD9">Tanzania</collectingCountry>
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(
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<geoCoordinate id="203EA7FB583B0DF2F54B04D214117491" direction="south" orientation="latitude" value="-4.8869443">04°53'13" S</geoCoordinate>
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,
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<geoCoordinate id="BD12BC58A2FED2E6E2C1FAEA5B469927" direction="east" orientation="longitude" value="29.612501">29°36'45" E</geoCoordinate>
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), 02/10/1997.
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</materialsCitation>
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Type localities in Figs. 1 and 4.
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</paragraph>
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<paragraph id="2329922C4879C4B7E3EDF76D98C81C4F" pageNumber="28">
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<typeStatus id="393C92D0A36E505439AC50B5F24CF55F">Paratypes</typeStatus>
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: The material was collected from two localities 4 km apart: Tembo Rock (2 km north of Cape Bangwe) and Muzungu Beach on the southern side of Cape Bangwe (Fig. 4).
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<materialsCitation id="23D77AC2791BA87CA89DB1DAFCAFA5C5" ID-GBIF-Occurrence="923899936" country="Tanzania">
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<collectionCode id="AA5121963A084582A9CC0920B1D873B6" collectionName="SAIAB">SAIAB</collectionCode>
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58257 (17), 19.2-55.2 mm SL, Tembo Rock,
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<collectingCountry id="AA6E2C0BDDB6A7C4B274A7A7A11AE64B">Tanzania</collectingCountry>
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(
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<geoCoordinate id="9045FE9C3F5FC08F38ADE9AE3F61960F" direction="south" orientation="latitude" value="-4.8869443">04°53'13" S</geoCoordinate>
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,
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<geoCoordinate id="E529A1F46C6D961D14ACE5F993A8208F" direction="east" orientation="longitude" value="29.612501">29°36'45" E</geoCoordinate>
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), 02/10/1997
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</materialsCitation>
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;
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<materialsCitation id="B48DF079B41D2DEF0283F4C3171258B6" ID-GBIF-Occurrence="923899932" country="Tanzania">
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<collectionCode id="2BDA79E82D5AFA76494F1B431DC19106" collectionName="SAIAB">SAIAB</collectionCode>
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57912 (14), 40.5-56.7 mm SL, Muzungu Beach near Kigoma,
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<collectingCountry id="1A36CA06A7CB4911CCDBA8CC5D5F1C9D">Tanzania</collectingCountry>
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(
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||
<geoCoordinate id="3543FF6B3D186D640EB9927B66596418" direction="south" orientation="latitude" value="-4.9180555">04°55'05" S</geoCoordinate>
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||
,
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<geoCoordinate id="47D0DF55F75CA6F8E3C85A7924D02B3A" direction="east" orientation="longitude" value="29.596111">29°35'46" E</geoCoordinate>
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), 08/05/1998
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</materialsCitation>
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;
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<materialsCitation id="AF7CD39B41D2FE3CE33C8FD5BE3EE47D" ID-GBIF-Occurrence="923899931" country="Tanzania">
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<collectionCode id="CE547FAD18B495D2D40A33F260CC2F5F" collectionName="SAIAB">SAIAB</collectionCode>
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56215 (5), 14.2-41.2 mm SL, Tembo Rock,
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<collectingCountry id="555CD9A92D4B2D319DD19280D960DE98">Tanzania</collectingCountry>
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(
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<geoCoordinate id="8D13F4857336D19A82385917055F2314" direction="south" orientation="latitude" value="-4.8869443">04°53'13" S</geoCoordinate>
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,
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||
<geoCoordinate id="00E6E84FCBE7FA68A3BAFB5A751CB8CE" direction="east" orientation="longitude" value="29.612501">29°36'45" E</geoCoordinate>
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||
), 08/10/1997
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</materialsCitation>
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;
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<materialsCitation id="329D857CC54C8DA91EEDB0B6D4073272" ID-GBIF-Occurrence="923899928" country="Tanzania">
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<collectionCode id="3AB2C04FA0B68A163B3498AFEB84C0E2" collectionName="SAIAB">SAIAB</collectionCode>
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56117 (6), 21.8- 32.0 mm SL, Tembo Rock,
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<collectingCountry id="7AAB015316EE535677F0D2C7557E137E">Tanzania</collectingCountry>
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(
|
||
<geoCoordinate id="C657C7EA8E7EDD9B7B1B96F233AB2E4F" direction="south" orientation="latitude" value="-4.8869443">04°53'13" S</geoCoordinate>
|
||
,
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||
<geoCoordinate id="A90B4BEAB932358E265E9BA5808452FE" direction="east" orientation="longitude" value="29.612501">29°36'45" E</geoCoordinate>
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||
), 07/10/1997
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||
</materialsCitation>
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||
;
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<materialsCitation id="ABD20B5BCDC7219757965674CEF059A1" ID-GBIF-Occurrence="923899929" country="Tanzania">
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<collectionCode id="C1897137D2D3D96E19D3E4488DA8165B" collectionName="SAIAB">SAIAB</collectionCode>
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56107 (18), 16.1-48.8 mm SL, Tembo Rock,
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<collectingCountry id="3DF70250AFE011094BC3A6232345A8D1">Tanzania</collectingCountry>
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04/10/1997
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</materialsCitation>
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||
;
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<materialsCitation id="FCD6BD3FF9B8D3DC0097DF807811D274" ID-GBIF-Occurrence="923899934" country="Tanzania">
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||
<collectionCode id="9DD29A67AEECED67205DE340C27AB09C" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
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98-056-P-l (1), 56.7 mm SL, Muzungu Beach near Kigoma,
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<collectingCountry id="FA7759135985DC230CC69C2FAB4B777A">Tanzania</collectingCountry>
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||
(
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||
<geoCoordinate id="3B1A06A34AF08BB84F6A7E1A1E890ADE" direction="south" orientation="latitude" value="-4.9180555">04°55'05" S</geoCoordinate>
|
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,
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||
<geoCoordinate id="2FADECCBE71D538D96792C7F5E1091CC" direction="east" orientation="longitude" value="29.596111">29°35'46" E</geoCoordinate>
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||
), 08/05/1998
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||
</materialsCitation>
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||
.
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</paragraph>
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<paragraph id="F11267744AAF914F391306A1C7628D84" pageNumber="28">
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||
<materialsCitation id="D193D3EB33D3F6AD84E263C92ADD61EE" ID-GBIF-Occurrence="923899935" country="Tanzania">
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Topotypes:
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<collectionCode id="135DC67A2133A1FA1AF1F2241A10DFD0" collectionName="GMNH">GMNH</collectionCode>
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4419 (26), Tembo Rock,
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<collectingCountry id="6380C281407325BDBC5CCCE76A9F30F8">Tanzania</collectingCountry>
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(
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<geoCoordinate id="86FD062FDDCFE77E18C48F8DF0BDDCCB" direction="south" orientation="latitude" value="-4.8869443">04°53'13" S</geoCoordinate>
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||
,
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||
<geoCoordinate id="B103E47DB2A8493E7BB3F14AEBFCB951" direction="east" orientation="longitude" value="29.612501">29°36'45" E</geoCoordinate>
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), 07/7/1998.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A3986B896270A1E38E36B516B660F557" type="diagnosis">
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<paragraph id="F321D69955218A5A26EF5D6C310DEFB9" lastPageNumber="29" pageNumber="28">
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Diagnosis. The species is very similar to
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<taxonomicName id="E3BCFF18407D70070955721E631FFA81" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
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, and
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<taxonomicName id="133C6EDE7D8DCF75C69F393414E07175" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
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<taxonomicNameLabel id="03A6BEBD71347BDCF69F28F37360C676" rank="species">sp. nov.</taxonomicNameLabel>
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Most differences were found with
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<taxonomicName id="1EF8BD83D74FAD4CC30D0A8291C16AE7" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
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.
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<taxonomicName id="7139A77C5567328E1FA9477710941F49" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
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can be distinguished from
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<taxonomicName id="55E9735B61FD1BBA8C2E076898499E09" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
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,
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<taxonomicName id="95C079A3CD5F727F7D1E026613B8D682" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="crassus">N. crassus</taxonomicName>
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,
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<taxonomicName id="B1BBFB7A7921D51CC85D2D6BD7B92D32" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">N. gracilis</taxonomicName>
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,
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<taxonomicName id="588B4DF60B170916B29FA5C524D27627" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="splendens">N. splendens</taxonomicName>
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,
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<taxonomicName id="434AAEDCA8F4DA977AB3806AAFF99E99" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="pulcher">N. pulcher</taxonomicName>
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,
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<taxonomicName id="F68A07D50973DD887F9762491F6A3AC4" LSID-ZBK="urn:lsid:zoobank.org:act:D9635626-82B6-4514-A745-22C5F6887D5F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neolamprologus helianthus Büscher 1997:701" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="helianthus">N. helianthus</taxonomicName>
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and
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<taxonomicName id="24BC25126817E5605E660689A8906CE4" LSID-ZBK="urn:lsid:zoobank.org:act:7BB28A57-F0B7-4522-A82C-C511181015B1" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neolamprologus marunguensis Büscher 1989:739" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="marunguensis">N. marunguensis</taxonomicName>
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by a small interorbital width (14.8-22.1 % vs. 22.5-29.1 % HL).
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<taxonomicName id="AEE22469287F48A1A51917C13C03DA78" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
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can be distinguished from
|
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<taxonomicName id="809D21E7364D450CF71C011EEC9CA4AD" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
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,
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<taxonomicName id="C48B66CF6EA4198EC0B79C837E969900" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="crassus">N. crassus</taxonomicName>
|
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,
|
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<taxonomicName id="2DF9A011A036AAD5D64AE2984BCE0F14" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">N. gracilis</taxonomicName>
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,
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<taxonomicName id="54EBE13CEEE8C7A79C5F01D1EA0C28BC" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="splendens">N. splendens</taxonomicName>
|
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,
|
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<taxonomicName id="617578D5A80861E4958D81684067693E" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="pulcher">N. pulcher</taxonomicName>
|
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and
|
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<taxonomicName id="836BDB2FD41F513B73335547F8B8D8BE" LSID-ZBK="urn:lsid:zoobank.org:act:7BB28A57-F0B7-4522-A82C-C511181015B1" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neolamprologus marunguensis Büscher 1989:739" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="marunguensis">N. marunguensis</taxonomicName>
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by a small preorbital depth (11.8-16.9 % vs. 17.0-22.3 % HL).
|
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<taxonomicName id="F36628B3134A749E060307EE7A2A28E9" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
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differs from
|
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<taxonomicName id="7A17149691304758AB645F44D9D18FCF" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="olivaceous">N. olivaceous</taxonomicName>
|
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,
|
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<taxonomicName id="BAF331101D8C3EEC14A721636E5F4DF1" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
|
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,
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<taxonomicName id="BF33E5F9B6458695850CDE53F3EA1481" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="splendens">N. splendens</taxonomicName>
|
||
,
|
||
<taxonomicName id="7E2696E31022ED86E3A6E3BF6CC6C0DB" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="pulcher">N. pulcher</taxonomicName>
|
||
,
|
||
<taxonomicName id="5FD8BDA5EB0B1CDC3E06BF962C71222F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
and
|
||
<taxonomicName id="F70CFE593EE16A3D98C1F6A7B996BE5E" LSID-ZBK="urn:lsid:zoobank.org:act:D9635626-82B6-4514-A745-22C5F6887D5F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neolamprologus helianthus Büscher 1997:701" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="helianthus">N. helianthus</taxonomicName>
|
||
by the absence of markings on the operculum. With 33-35 scales in the longitudinal series,
|
||
<taxonomicName id="6F8D79D38DEADC352ACBD42F7424CB0E" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="6DCDE384E9D4354260A077572C74CFEF" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">N. gracilis</taxonomicName>
|
||
,
|
||
<taxonomicName id="5CE01FD778411A90444983AC188BA736" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="splendens">N. splendens</taxonomicName>
|
||
and
|
||
<taxonomicName id="67C5700283F8AB74272E1988D46CD14A" LSID-ZBK="urn:lsid:zoobank.org:act:D9635626-82B6-4514-A745-22C5F6887D5F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neolamprologus helianthus Büscher 1997:701" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="helianthus">N. helianthus</taxonomicName>
|
||
(36-38 scales). With 6-9 gill rakers,
|
||
<taxonomicName id="9B047046679B1E8EAAC1FB6F5A7850A8" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="EBA16DA9DEF939A984F6055F20EF82FB" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
and
|
||
<taxonomicName id="2064D8208BF0B94A9D9C5B19C97AAC5F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">N. gracilis</taxonomicName>
|
||
(10-18 gill rakers).
|
||
<taxonomicName id="DA75D62FBBE6FCF66179F2540FCC29E9" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
differs from all species in the
|
||
<taxonomicName id="3A453EFDBAD88D28684A5DF3E45A8053" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
|
||
complex except
|
||
<taxonomicName id="DDD97FD840F24FFDCA8AFF1DE024BBE2" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
by the absence of scales on the occiput; from
|
||
<taxonomicName id="35593235C7CA8197C41701597E95D685" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="splendens">N. splendens</taxonomicName>
|
||
,
|
||
<taxonomicName id="6B02671BD426C53180A6C695CA6F1921" LSID-ZBK="urn:lsid:zoobank.org:act:7BB28A57-F0B7-4522-A82C-C511181015B1" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neolamprologus marunguensis Büscher 1989:739" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="marunguensis">N. marunguensis</taxonomicName>
|
||
and
|
||
<taxonomicName id="5B5EED098155E3E42F0AA8246AD7AD0F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
|
||
by the absence of scales on the paired fins and by the absence of ctenoid scales on the dorsal and anal fins; from
|
||
<taxonomicName id="5ADD154EDB230E26B85331AE3D6BD2D9" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
|
||
,
|
||
<taxonomicName id="C5B5BDD0D359B4FCB7BC3B15EA727322" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
,
|
||
<taxonomicName id="9726A451DFD74FD890738650201DFEC3" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="olivaceous">N. olivaceous</taxonomicName>
|
||
and
|
||
<taxonomicName id="84FF481D8E3FA43ABE30C63A61DCD31D" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="pulcher">N. pulcher</taxonomicName>
|
||
by the presence of cephalic pits; from
|
||
<taxonomicName id="5C86DF379A3763F5BDA3334D98C897A3" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="olivaceous">N. olivaceous</taxonomicName>
|
||
and
|
||
<taxonomicName id="9DB123ED5339373229B4135FD224498E" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="pulcher">N. pulcher</taxonomicName>
|
||
by the absence of conspicuous spots on scales and from
|
||
<taxonomicName id="91ABFFF05EFD8727EA95D945D8E9EAD3" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
,
|
||
<taxonomicName id="59B659CF25769E7993B2C1749772093F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="pulcher">N. pulcher</taxonomicName>
|
||
,
|
||
<taxonomicName id="B36089F3735EF45F4D6710916E2F92D1" LSID-ZBK="urn:lsid:zoobank.org:act:D9635626-82B6-4514-A745-22C5F6887D5F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neolamprologus helianthus Büscher 1997:701" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="helianthus">N. helianthus</taxonomicName>
|
||
,
|
||
<taxonomicName id="6F053C0C205EB76EE79358F7EC7AAC41" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="crassus">N. crassus</taxonomicName>
|
||
,
|
||
<taxonomicName id="D5ACC1EBC5C4ED1FF1FE6C3098757EC3" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="splendens">N. splendens</taxonomicName>
|
||
and
|
||
<taxonomicName id="7B381CD7D2CEBB07F791DD60B2FF31F4" LSID-ZBK="urn:lsid:zoobank.org:act:7BB28A57-F0B7-4522-A82C-C511181015B1" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neolamprologus marunguensis Büscher 1989:739" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="marunguensis">N. marunguensis</taxonomicName>
|
||
by a smaller caudal peduncle depth (11.3-13.2 % vs. 13.4-15.3 % SL).
|
||
<taxonomicName id="39E742598B8BAEC3396339AB24BF8705" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="542B6170BF4F199886975B870143CA13" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
|
||
and
|
||
<taxonomicName id="F9D62D4743B9D4CA0E177E064D2F0EB3" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="gracilis">N. gracilis</taxonomicName>
|
||
by having less scales between the upper and lower lateral lines (2 vs. 3) and by having less scales around the caudal peduncle (16 vs. 19).
|
||
<taxonomicName id="AD957253815203A14717F514F488ECBB" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
further differs from
|
||
<taxonomicName id="47C77AEE9EB9A8254D74AFA61E053230" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
|
||
by the absence of bars on the body, by a smaller predorsal fin length (29.2-34.0 % vs. 34.6-36.8 % SL) and by having more pectoral fin rays (13 vs. 12).
|
||
<taxonomicName id="939F086017691F934EE6E8E448873F79" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="C3A7E55FCB12DB7AE602AD3BDF9402B4" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
<taxonomicNameLabel id="3A835703D282EC9C8015187433EDC751" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in the linear relations between cheek depth and head length and between body depth and standard length (significantly different intercepts, Ancova, P <0.002, Fig. 5) with larger cheek depth and larger body depth for specific specimen size.
|
||
<taxonomicName id="651FF12EA72734732AD271AB053C4ED2" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
further differs from
|
||
<taxonomicName id="A31F93646A20527566CE8A4D09D0E7CA" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
<taxonomicNameLabel id="7AAD931D57E9A207DBDA04FC216A7806" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in habitat preference (rubble substrate with fine sediment vs. large boulders with sand) and by having more pronounced markings on the dorsal and caudal fins (Fig. 2).
|
||
<taxonomicName id="9496588AB594DE250C29696D202DAEF5" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="3DBFF2C6C4993FC1634E2A3C5E8CD53A" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
<taxonomicNameLabel id="6AE768898A3B2BCF3E0A01AF7B61EE7A" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by fewer canal scales in the lower lateral line (median 6, 54 % of all scales in the lower lateral line vs. median 9.5, 77 %, P <0.003, t test). Preorbital depth (POD) was significantly smaller (Ancova, P = 0.002 for POD as % of head length and P <0.001 for POD as % of standard length, Fig. 5) in
|
||
<taxonomicName id="7BCACA43777DC60FC7F64A70FD0B2256" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
(11.8-16.9 % HL) than in all four examined type specimens of
|
||
<taxonomicName id="3A874AAD3EB2FD7ECBF83939BEA3A1A2" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
(17.0-18.5 % HL. The four examined type specimens include all type specimens of
|
||
<taxonomicName id="D468E2D8639AE788D4901C7856487CC6" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
that are not suspected to be non-specific with the holotype; pers. comm. Jos Snoeks).
|
||
<taxonomicName id="C8F846A053167EE838FA06E1E17D205E" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
differs further from
|
||
<taxonomicName id="0C6F2EA10F711B39F42389FCE4976D04" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
by a smaller maximum standard length (57 mm vs 66 mm) and other associated size measures (two of the four type specimens of
|
||
<taxonomicName id="DA6B3C98BE3B36416270659C2A489F55" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
, including the holotype, are larger in body depth and head length than the largest
|
||
<taxonomicName id="F60CFF38A4B853EC435C5157D0347D24" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
type specimen), by having drab grey-brown juveniles with bluish dorsal and anal fins (compared with bright yellow-orange fins of the juveniles of
|
||
<taxonomicName id="5452DA7C5FE521FE9BEA44E5EDA47CEA" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
; Konings, 1998), by having a distinct pattern on the unpaired fins (compared with no or vague markings in
|
||
<taxonomicName id="6DDED9606510247456E8E47F8141DC5D" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
. Compare photographs of
|
||
<taxonomicName id="B4FDAE15180483A1C008F62792F2DB49" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
[pp. 344 and 377 in Brichard, 1989] with
|
||
<taxonomicName id="ED0E65AE03C7D0F636CE04EEB6BF8F91" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
[page 339 in Brichard, 1989] and the description by Brichard [1989, p. 343]: “the unpaired fins [of
|
||
<taxonomicName id="6036943302EED786B64F97285DD1076F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
], which are devoid of any markings”), and perhaps by having more dorsal soft rays (8-10 vs. 7 for holotype of
|
||
<taxonomicName id="306CFE376489CD350EC79C92A286AEDC" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
). The juveniles of
|
||
<taxonomicName id="40D804D75A9879A7EE03F9747C952D37" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
with bluish fins and greybrown body differ strongly from juveniles of the undescribed form
|
||
<taxonomicName id="748559353F2127D7139C53AE8A443019" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="sp. 'cygnus'">N. sp. ‘cygnus’</taxonomicName>
|
||
(found at the south east end of the lake) which are bright blue and yellow (Konings, 1998). Juveniles of
|
||
<taxonomicName id="3F68BC5F7CCF776C4E0686C3DBC0D861" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
and
|
||
<taxonomicName id="C6BFD0B4282F62B71D7A63CC6D72BE6F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="sp. 'cygnus'">N. sp. ‘cygnus’</taxonomicName>
|
||
do not have the vertical bars on the body of the juveniles of
|
||
<taxonomicName id="0DB641B096750E7219ABA2693E80C29B" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
.
|
||
<taxonomicName id="EB74C73A7EE43827775327A1304F1ED8" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
had on average more canines in the upper (8) and lower jaws (6), than found in all other species, except
|
||
<taxonomicName id="C76DC95D73B958554A9607BA1C266E90" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
<taxonomicNameLabel id="590CA976BFBBCD3F6333E7FDCBDDA1AF" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="E7B5F5D6F551A37ADF92A578C2A20FEC" LSID-ZBK="urn:lsid:zoobank.org:act:7BB28A57-F0B7-4522-A82C-C511181015B1" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Neolamprologus marunguensis Büscher 1989:739" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="marunguensis">N. marunguensis</taxonomicName>
|
||
and
|
||
<taxonomicName id="C8140CE0BC35D125D39C65AE02D72138" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
(in 2 of the 4 investigated type specimens). There were fewer canines in the holotype of
|
||
<taxonomicName id="13C9F48EAAB5F8DBA06F763B3DFA406C" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
, 3 and 6 (in upper and lower jaw respectively) and 6 and 4 respectively in paratype 80-31-P7 (the holotype and paratype of
|
||
<taxonomicName id="D85C6B70378E0957D9E0A0701BAC0D28" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="28" phylum="Chordata" rank="species" species="falcicula">N. falcicula</taxonomicName>
|
||
may have lost canines).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3A92782411564ECED6D7D5DE84C601E3" type="description">
|
||
<paragraph id="4B0DFC3C687CCAF749795AD61F7E0333" pageNumber="29">Description. Counts and measurements for the type series are given in Table 1. Range of standard length (SL) of investigated specimens: males 34.1-56.7 mm (18 specimens), females 32.5-49.6 mm (11 specimens). No sexual dimorphism. Meristics, morphometric measurements and colour patterns showed no differences between the populations of Tembo Rock and Muzungu beach.</paragraph>
|
||
<paragraph id="89430F94F8DDE8D17C2BD3E269AB8D46" pageNumber="29">Four-8 canines in upper jaw (mode 8, 69%) and 4-7 canines in lower jaw (mode 6, 72 % of specimens, Table 1) on anterior part of the dental arcade, with two outer canines on both jaws enlarged and the middle two smallest (Fig. 6). Dentigerous arm of premaxilla straight. Upper jaw tooth rows 5-6 anteriorly reducing to a single row half way along the jaw, outer row teeth (including the canines) 32-38. Lower jaw with 5-6 rows of teeth reducing to a single row approximately 2/3 along jaw, outer row teeth 27. Mouth terminal, isognath.</paragraph>
|
||
<paragraph id="51C82929DFE9E1121066DB2E1E147492" pageNumber="31">Six-11 gill rakers along lower limb of first gill arch (mode 7, Fig. 6). Gill rakers were slender, concentrated near the posterior end of the lower arch and decreasing in size towards anterior end of the arch. Marked with small dark spots, sometimes 1 or 2 reduced at lower end of arch, sometimes one or several bifid.</paragraph>
|
||
<paragraph id="B4EE5A578C6164545C5E02E2D1D46EB7" pageNumber="31">Scales absent on cheek, preoperculum and between eye and dorsal margin of preoperculum. Opercular scales 8-15, cycloid, variable in size but smaller than flank scales. Scales absent on occiput. Scales on nape cycloid, deeply embedded, varying from a few around the origin of dorsal fin to fully covered. Small scales between pectoral and pelvic fins 5-10, cycloid, difficult to distinguish. Small scales between upper lateral line and origin of dorsal fin cycloid, often covered by skin. Small cycloid scales on chest and belly. Urogenital area small ctenoid scales. Chest scales smallest. Scales on flank ctenoid. Dorsal fin with scaly sheath, few cycloid scales between soft rays, less between spines. Anal fin with scaly sheath, few cycloid scales between rays, more than on dorsal fin. No scales on pectoral and pelvic fins. Caudal fin small ctenoid scales, mostly on rays, covering about on average the anterior 75 % of tail. Larger specimens tended to have a larger part of tail covered with scales. Scales on outer rays range in adults up to past posterior end of middle rays.</paragraph>
|
||
<paragraph id="72B9219454A95FECC4836976F6D8D892" lastPageNumber="32" pageNumber="31">Thirty-three-35 scales in the longitudinal series. In the upper lateral line 13-26 canal scales (76-100 % of all scales in the upper lateral line, median 20, 91%). In the lower lateral line 2-11 canal scales (13-100 % of all scales in the lower lateral line, median 6, 54 %). Scales without canal segments in between canal scales were pitted. Anterior to canal scales in lower lateral line, pitted scales generally extended almost up to the operculum. A positive relationship between standard length and number of scales in upper lateral line, and number of canal scales in lower and upper lateral lines. Lower and upper lateral lines overlapped in specimens>51 mm.</paragraph>
|
||
<paragraph id="1924999780A4F281ADCD44F5B06A6E8D" pageNumber="32">First pelvic fin ray longest. Caudal fin lunate. First spines of dorsal and anal fins small. Anal filament on average 2.5 % longer than dorsal fin filament. Dorsal fin margin posterior to sixth spine straight.</paragraph>
|
||
<paragraph id="81BA09FAD436E549E5E40A3926D3EC18" pageNumber="32">Lower pharyngeal jaw about as wide as long (Fig. 7). Dentigerous area covered with slender, pointed, unicuspid teeth, anteriorly slightly recurved, posterior rows straight and enlarged.</paragraph>
|
||
<paragraph id="2B891E4FE7493F089E1815194711C75F" pageNumber="32">Post-lachrymal infraorbital bone series absent (Fig. 8). Free neuromast papillae on infraorbital lateral line system. Suborbital dermal depression forming a thin but distinct groove. A row of more or less contiguous cephalic pits with papillae runs from a pore posterodorsal of orbit to a pore anteroventral of orbit. Papillae clearly visible in some specimens, tiny or absent in others.</paragraph>
|
||
<paragraph id="2E6348A4D86EC5B8562EF6861BA37E2E" pageNumber="32">Many epidermal papillae around the pectoral, behind and on the pelvic fin, around the genital pore and posterior edges of operculum and preoperculum in specimens of Tembo Rock. Slightly or not at all present in specimens from Muzungu Beach.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4741480271183EB5400CDA32B62290E" type="description">
|
||
<paragraph id="A72E6F50C244A6DD84F250CF8B78498F" pageNumber="32">
|
||
Coloration. Body dark, grey to almost black, darker than
|
||
<taxonomicName id="BC852B5099164A41E63D6BA0916108D0" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="32" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
<taxonomicNameLabel id="D18741DB398506B8D10796D9520B5B2E" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Larger specimens are lighter. Caudal margins of scales brown, produces a checkered pattern on the body. Between first 4 spines of dorsal fin (these increasing in size from rostral to dorsally) the margin of membrane yellow (lappets). Blue eye and blue stripe under the eye. On rare occasions stressed individuals show bands on body as
|
||
<taxonomicName id="5F3F9A3D4B015807C75A4F495AFDBE37" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="32" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="96F059ECF1872FF59EE350A7529C85C1" pageNumber="33">No trace of opercular markings. Conspicuous black and white bands along dorsal margin of tail and posterior margin of dorsal fin (submarginal white or transparent). Vertical bands on dorsal and caudal fins. In preserved specimens vertical bands on fins less visible, no colour on iris or suborbital stripe.</paragraph>
|
||
<paragraph id="32A5034066484AE8268E79626C84ED40" pageNumber="33">Juveniles. Vertical bands on fins less clear or not visible in juveniles, especially on tail. Smallest young (up to ± 20 mm length) observed in natural habitat bluish fins and light vertical bars, with dark and light bars. In specimens of 14-25 mm body grey-brown. Emarginate tail, no filaments on caudal, dorsal and anal fins. Pelvic fin blackish, margin dorsal fin black. Tail 10 - 40 % covered with scales, no scales on other fins, operculum scaled, no scales on nape and occiput.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C713E79B52EC1B2FA3F57C5FDEEA624A" type="distribution">
|
||
<paragraph id="93D0B1FEBC646005C599C5AEC621BD56" pageNumber="33">
|
||
Distribution.
|
||
<taxonomicName id="0EDF0F208A56BD41E700373723295DA3" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
was found from near the Burundi border with Tanzania in the north, to Cape Kabogo in the south (
|
||
<geoCoordinate id="575C57325EC87A967DEC8A3D5B3B5AA3" direction="south" orientation="latitude" value="-4.5">4°30'S</geoCoordinate>
|
||
-
|
||
<geoCoordinate id="DBB5CA022BA0A1837B8D457A192EDF6D" direction="south" orientation="latitude" value="-5.4166665">5°25'S</geoCoordinate>
|
||
). No observation dives were made in Tanzanian waters beyond these limits, except near Kisoje (Fig. 1) where
|
||
<taxonomicName id="A88DB7EBF812E8C1C398A254E2565D63" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
was absent. In Zambian waters, at the south end of the lake,
|
||
<taxonomicName id="84A563787C264289F576236117055307" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
is absent. Around Kigoma,
|
||
<taxonomicName id="25F56577F382EC8BDCE5C446D97254D5" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
is by far the most common fish species in the littoral.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="492BFE121D30B737CDCE0AEF7394339C" type="biology_ecology">
|
||
<paragraph id="6C04A03CB39D77A79FEE58B5CA1221AC" pageNumber="33">
|
||
Habitat and biology. Fig. 9 shows
|
||
<taxonomicName id="BDCDB37AB07C9C96FEE33FC32E523AFC" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
in its habitat.
|
||
<taxonomicName id="763B9620883716F8170F336E0693A381" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
was observed syntopically with
|
||
<taxonomicName id="CF93E4FE69F9EB547C4A3878F91E30C0" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
and with
|
||
<taxonomicName id="21304D85D14A00CB676D7A518790D38F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
|
||
.
|
||
<taxonomicName id="D74D6F5DF63C02D0A7AEEA484731370B" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
and
|
||
<taxonomicName id="45329272E9336A9FE4D14DB0235F49C3" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
<taxonomicNameLabel id="8EAB366F6739055E0F2A5FF95718A614" rank="species">sp. nov.</taxonomicNameLabel>
|
||
were not found in exactly the same area, but were associated with alternative habitat types.
|
||
<taxonomicName id="25D9C0BAB3EB8BF41A7DEDBF0A860EBE" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
was found in bays and sheltered sites, exposed to weaker currents than the (only) site in which
|
||
<taxonomicName id="9A57CDD005FA219860A30DC9C1B074AB" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
<taxonomicNameLabel id="B6ABEB2A533ECE0F490EDCFF09353AE5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
was found. At the Bangwe peninsula south of Kigoma,
|
||
<taxonomicName id="2913BE87895D48A1121F62530E1EADCA" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
occurs on the north and south shores of the peninsula, while
|
||
<taxonomicName id="34AADFB18009A8C5D94E7FA20DC5E26A" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
<taxonomicNameLabel id="27CFE9B431CDD1763F25147ECACB731D" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is found on a more exposed site between these two localities on the peninsula (Fig. 4). The habitat of
|
||
<taxonomicName id="94F7888A0873AC494414B7FB742CCE51" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
features gentle slopes, with small stones or rubble, cemented by calcite, with fine sediment in spaces between rubble, and stones with stromatolite growth (Cohen et al., 1997).
|
||
<taxonomicName id="8AC3883949FA89313B12AE63AB5B0C87" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
is often associated with bivalve shell accumulations covering large areas (always
|
||
<taxonomicName id="486B2A66F32437F3EF19FFE5967C8283" class="Bivalvia" family="Iridinidae" genus="Pleiodon" higherTaxonomySource="CoL" kingdom="Animalia" order="Unionida" pageNumber="33" phylum="Mollusca" rank="species" species="spekii">Pleiodon spekii</taxonomicName>
|
||
[Pelecypoda: Mutelidae]). The shells (10 to 15 cm long) are cemented together and provide many small spaces for refuge and breeding.
|
||
</paragraph>
|
||
<paragraph id="2B8A47608C94B0585873D5392BAB84B4" pageNumber="33">
|
||
The two sites where the paratypes were collected differed in topography and in species community composition. At Muzungu Beach
|
||
<taxonomicName id="D42CFF96C7835E097ED87F2F225B10CF" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
occurred in dense aggregates on patches of small encrusted rubble on relatively steep slopes. The patches were generally several tens of square metres, and separated from each other by large rocks.
|
||
<taxonomicName id="BC3D028BEF0665266564CE4037C7529C" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
was primarily associated with the larger rocks, and generally did not occur on the rubble patches. The fish species assemblage in this habitat was relatively diverse, due to habitat heterogeneity with large boulders and deep water near the shore. At Tembo Rock rubble patches were larger,>100 m2 and sloped less steeply and large boulders were absent.
|
||
<taxonomicName id="BF4AB0357AEE748D1D619F03B37A38FA" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="brichardi">Neolamprologus brichardi</taxonomicName>
|
||
occurred here in large feeding aggregations, in the same area as
|
||
<taxonomicName id="4F8E99DA5131FAD55CE3C0A758E69012" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
. Such large feeding aggregations of
|
||
<taxonomicName id="555C275482F4D30236399A52331555C6" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
have been reported by Brichard (1989) and can consist of 100,000's of individuals. In this habitat the fish community consisted virtually only of
|
||
<taxonomicName id="F5DB65D79221BDE5CD31347DE7653D4A" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
and
|
||
<taxonomicName id="BE1445218C903D3927318B3F7C6796FD" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
, with a sharp segregation of the space occupied in the water column between the two species.
|
||
<taxonomicName id="EF52C81F89CCA56A73254E91C1C01168" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
occupied the lower water layer, near the bottom at an average 20 to 30 cm height (max c. 50 cm), with a high population density, and the less sedentary
|
||
<taxonomicName id="E9DDEE1058627B1B66407585C62A087C" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
foraged in about equal numbers immediately above, at an average c. 60 cm off the bottom, over the same area.
|
||
</paragraph>
|
||
<paragraph id="F303590C67CB71558A8E496808895803" pageNumber="33">
|
||
In sheltered areas around Kigoma with rubble on a sandy bottom,
|
||
<taxonomicName id="D376F70C3BE99D739C5FCDE7510BE63B" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
is the most common fish species and can occur in high densities.
|
||
<taxonomicName id="0B00325FF713B5BD5384AB1B5F685BB5" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
is a substrate spawner that forms large congregations and maintains small contiguous territories of about 1 m2 or less, each occupied by a number of fish including two or more adults, several subadults and juveniles. Other species of similar size are chased from their territory.
|
||
<taxonomicName id="96886E97C7BE6FB7355FFE94DB29BD59" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="33" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
colonies can cover hundreds of square metres at depths of 5 to 20 m with densities reaching 20 to 30 individuals per square metre. They excavate sediment next to and between rocks, thereby creating numerous crevices that function as refuges in the territory in which they hide during the day from approaching predators and spend the night. Crevices are shared by several individuals. Eggs were never observed and are probably deposited within these crevices.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="8570D6E946AF53284CC9B84B3311DDA5" type="biology_ecology">
|
||
<paragraph id="DB42C2FE12CC96D1E4C86896588BDA22" pageNumber="35">
|
||
Every individual
|
||
<taxonomicName id="7AB1373A377ADBD9A320A404843D1A0A" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
is strongly restricted to a small area and is rarely seen to travel more than 2 metres (>2 m seen only once in about 60 observation dives). The home range of an individual generally has a diameter of 20 to 200 cm, the larger the individual the larger the area. Although locally very abundant,
|
||
<taxonomicName id="16F3457A2A8DDCEC9FB06238709D8320" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
is probably a poor disperser, in view of its extreme philopatry.
|
||
</paragraph>
|
||
<paragraph id="686B53436ED9661555A9FCABB7A9D9C0" pageNumber="35">
|
||
<taxonomicName id="21A78A6228422A3D8B96B27240A9811F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
either forages on the substrate, ingesting sediment, or hovers at 10 to 50 cm off the bottom, while taking plankton. When
|
||
<taxonomicName id="61B0F9062E764B718F38D5E6EB0A10C6" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
forages in the water column it stays closer to the bottom than
|
||
<taxonomicName id="A0B9264E6B2CAACB62390953E34F7437" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
. The larger fish move higher from the substrate than the younger ones and are probably more dependent on a planktonic diet. The juveniles stay within 5 cm of the substrate.
|
||
<taxonomicName id="5411204090FB551AB4AF2F884F0C89D1" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
takes occasionally bites from the substrate surface for feeding, and sediment is egested through gills and mouth. When feeding on plankton,
|
||
<taxonomicName id="C677484577C8A13ACA77F56B3D0375DD" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
has a lower feeding rate than
|
||
<taxonomicName id="E1BB630DABC7D520A85E197706481ED5" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E6B5B0E9FC0A66982625D6348D8387E2" pageNumber="35">
|
||
<taxonomicName id="508F139B15AD2CBEC255149C0940FE88" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
is relatively wary, as
|
||
<taxonomicName id="6067869A7892172354E74B7D46555087" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="savoryi">N. savoryi</taxonomicName>
|
||
, and stays close to the bottom and dives for the nearest cover immediately when approached. Fry are generally very few, as expected in lamprologine substrate breeders. Young <1 cm in length are usually visibly present in the colonies, though always very near the substrate and are first to vanish into the bottom.
|
||
</paragraph>
|
||
<paragraph id="0E3F6ECA52AE3FA8FDC330C88F5B7EF5" pageNumber="35">
|
||
<taxonomicName id="5C4186A70EED6C13EC0DA1D5A63DA383" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">Neolamprologus walteri</taxonomicName>
|
||
were observed at a depth range of 2 to 30 m, but can probably occur deeper as well. They are most common at 7 to 15 m. There is a large depth overlap with
|
||
<taxonomicName id="49636BE1A4AD544FDB4D679F7CDAFD00" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
, but
|
||
<taxonomicName id="E04B1374F4C842676369BBF099BA580A" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
has lower densities in shallower depths than
|
||
<taxonomicName id="1F9AA92DC132E131786DB48455D748F2" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="brichardi">N. brichardi</taxonomicName>
|
||
.
|
||
<taxonomicName id="B87D9FA50C304EAFB72FF91A607DEBDC" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="savoryi">Neolamprologus savoryi</taxonomicName>
|
||
at the Bangwe peninsula is most often found at depths of 15 to 30 m, but also occurs shallower. Within its distribution in the lake
|
||
<taxonomicName id="E09DF5B3EC1DC7205BC56981363B2005" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
is present almost everywhere where there is some rocky cover, even in areas with an otherwise flat sandy bottom at 3 m depth, except at the site where
|
||
<taxonomicName id="B459F64A20C87DD0FECA989777E8734D" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
occurs. While territories of
|
||
<taxonomicName id="E65DF7D7F762788D767E41334EBC32E9" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="walteri">N. walteri</taxonomicName>
|
||
occurred as close as 4 m from the southern boundary of the
|
||
<taxonomicName id="B986E0064A0482D878B61A144A81564F" family="Cichlidae" genus="Neolamprologus" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageNumber="35" phylum="Chordata" rank="species" species="chitamwebwai">N. chitamwebwai</taxonomicName>
|
||
community (its northern boundary is sandy and contains neither species), and the species can therefore be considered sympatric, their distributions were never found to overlap and it appears that their distributions are determined by the type of substrate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="821D7105A7C48DCFDAA0ADB166D23953" type="etymology">
|
||
<paragraph id="2B68A538F26411BE8B5E1B0E24735A7B" pageNumber="35">Etymology. Named after Walter Dieckhoff, who first recognised it as an undescribed species (Konings, 1988). It has been referred to as ‘walteri’ in the aquarium trade for two decades and by Konings (1988).</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |