844 lines
92 KiB
XML
844 lines
92 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1180.107551" ID-Pensoft-Pub="1313-2970-1180-159" ID-Pensoft-UUID="165BC209F734524FB09936671B04125B" ID-ZooBank="50242204002B41F8B9B9BC2983CD7AC1" ModsDocID="1313-2970-1180-159" checkinTime="1695263980471" checkinUser="pensoft" docAuthor="Fuentes, Katlyn M., Baldwin, Carole C., Robertson, D. Ross, Lardizabal, Claudia C. & Tornabene, Luke" docDate="2023" docId="689886504CC25D18ABB4FFFD6CE13EBB" docLanguage="en" docName="ZooKeys 1180: 159-180" docOrigin="ZooKeys 1180" docPubDate="2023-09-20" docSource="http://dx.doi.org/10.3897/zookeys.1180.107551" docTitle="Varicus prometheus Fuentes & Baldwin & Robertson & Lardizábal & Tornabene 2023, sp. nov." docType="treatment" docUuid="E996CE37-FF77-4BA4-A550-039380D8ABBE" docUuidSource="ZooBank" docVersion="2" id="165BC209F734524FB09936671B04125B" lastPageNumber="159" masterDocId="165BC209F734524FB09936671B04125B" masterDocTitle="Two new species of Varicus from Caribbean deep reefs, with comments on the related genus Pinnichthys (Teleostei, Gobiidae, Gobiosomatini, Nes subgroup)" masterLastPageNumber="180" masterPageNumber="159" pageNumber="159" updateTime="1695264281519" updateUser="ExternalLinkService">
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<mods:title id="1276E4E7680C373190277DD9CF1F0CE8">Two new species of Varicus from Caribbean deep reefs, with comments on the related genus Pinnichthys (Teleostei, Gobiidae, Gobiosomatini, Nes subgroup)</mods:title>
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<mods:roleTerm id="8C220F85817F835B3147F87A9A48046D">Author</mods:roleTerm>
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<mods:namePart id="794F5730FF3F6D064DBAAF0F526CBC98">Fuentes, Katlyn M.</mods:namePart>
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<mods:affiliation id="D9FCB22D86F76E015C4904816A7C6001">School of Aquatic and Fishery Sciences and the Burke Museum of Natural History and Culture, University of Washington, 1122 NE Boat Street, Seattle, WA 98105, USA</mods:affiliation>
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<mods:namePart id="E39454587421ECE1909BC948CB339F44">Baldwin, Carole C.</mods:namePart>
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<mods:nameIdentifier id="0A8ED4FEA04691E1C7FBCAA5E28A0626" type="ORCID">https://orcid.org/0000-0002-2875-0474</mods:nameIdentifier>
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<mods:affiliation id="7F872DA59E6AE665E9E14B8E9AB20C46">Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, PO Box 37012, Washington, DC 20013 - 7012, USA</mods:affiliation>
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</mods:name>
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<mods:name id="9193ECC3A7F99B50B30F5A75F92511CF" type="personal">
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<mods:namePart id="4D7E1A09D68384209B2E524782D2325D">Robertson, D. Ross</mods:namePart>
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<mods:nameIdentifier id="3A36A44438A9D5ADC1642F8DC8CEC4D0" type="ORCID">https://orcid.org/0000-0003-3972-149X</mods:nameIdentifier>
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<mods:affiliation id="874BC1ABF7AA4C8FBF1253938951CCF1">Smithsonian Tropical Research Institute, Balboa, Panama</mods:affiliation>
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<mods:namePart id="AC5057F63BE9FA302DA2545662F4E5E7">Lardizabal, Claudia C.</mods:namePart>
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<mods:nameIdentifier id="48069BB9510EEBF82EE0E4A2541B05EC" type="ORCID">https://orcid.org/0000-0002-2802-7593</mods:nameIdentifier>
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<mods:affiliation id="FEE3B1172CCA8C11755A73A403E2A4CF">Instituto de Investigacion en Ciencias Biologicas y Ambientales del Norte de Honduras (IBIOANH), Departamento de Biologia, Universidad Nacional Autonoma de Honduras en el Valle de Sula, Zona el Playon, final del Blvd Micheletti, San Pedro Sula 21102, Cortes, Honduras</mods:affiliation>
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<mods:namePart id="F79A17D816B18AD836F7AC0DB7BDC8F8">Tornabene, Luke</mods:namePart>
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<mods:nameIdentifier id="D0A6089BA0DC180679B25D785B6CDF9F" type="ORCID">https://orcid.org/0000-0002-0673-2320</mods:nameIdentifier>
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<mods:affiliation id="B162A35E31187C833C996782D860A437">School of Aquatic and Fishery Sciences and the Burke Museum of Natural History and Culture, University of Washington, 1122 NE Boat Street, Seattle, WA 98105, USA</mods:affiliation>
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<mods:nameIdentifier id="EDA8D3407D64A06C4939BF9B8A1A6F24" type="email">luke.tornabene@gmail.com</mods:nameIdentifier>
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<mods:title id="CF6B78AF54F2E457A5D9E97B12FEEE6E">ZooKeys</mods:title>
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<mods:date id="1AA5155CB04C238604933CE94CAC007F">2023</mods:date>
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<mods:number id="CAADFD772A5082B0D2D57294918BC5EF">2023-09-20</mods:number>
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<mods:number id="4EC89EE006352F6CDE71C3E33AA6B85C">1180</mods:number>
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<mods:start id="2FC8BC38F34B938181D7BED16C867522">159</mods:start>
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<mods:url id="CF380616FCB163658D47DD06C4A660D4">http://dx.doi.org/10.3897/zookeys.1180.107551</mods:url>
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<mods:classification id="C679E584F6E8D8D756FF8799AC23F089">journal article</mods:classification>
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<mods:identifier id="FADEC084915E8A165E59E9C9DBDEF0BB" type="DOI">http://dx.doi.org/10.3897/zookeys.1180.107551</mods:identifier>
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<mods:identifier id="CDF29C467C0FFCD7DA413CF55610CE11" type="Pensoft-Pub">1313-2970-1180-159</mods:identifier>
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<mods:identifier id="993C14FB11CCFD45DA6664A2903F451B" type="ZooBank">50242204002B41F8B9B9BC2983CD7AC1</mods:identifier>
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<treatment id="689886504CC25D18ABB4FFFD6CE13EBB" LSID="urn:lsid:zoobank.org:act:E996CE37-FF77-4BA4-A550-039380D8ABBE" httpUri="http://treatment.plazi.org/id/689886504CC25D18ABB4FFFD6CE13EBB" lastPageNumber="159" pageId="0" pageNumber="159">
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<subSubSection id="68020C3477046FA7AD0B59066F8E3FB6" pageId="0" pageNumber="159" type="nomenclature">
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<paragraph id="CA2FC760454B4FD060599D4CD91EE863" pageId="0" pageNumber="159">
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<taxonomicName id="CDCC3940EB2E24298B458F4131365A44" LSID="https://zoobank.org/E996CE37-FF77-4BA4-A550-039380D8ABBE" authority="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene, 2023" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus" status="sp. nov.">Varicus prometheus</taxonomicName>
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<taxonomicNameLabel id="0A5B5662D0CB776C1FE04AF5B131DA2F" pageId="0" pageNumber="159">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="A5A11B381E7F3A303EFA572A7520685C" pageId="0" pageNumber="159" type="description">
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<paragraph id="858E0F9A2463F17242E8DBDC904D66E8" pageId="0" pageNumber="159">
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<figureCitation id="831F198E153F36378F6139CE45482223" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Varicus prometheus sp. nov. holotype, UW 158119, 30.5 mm SL, female, prior to preservation (top two photos) and preserved (bottom). Photos by Luke Tornabene." figureDoi="10.3897/zookeys.1180.107551.figure2" httpUri="https://binary.pensoft.net/fig/908647" pageId="0" pageNumber="159">Figs 2</figureCitation>
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<figureCitation id="0376CA406BBEE0831C17A7FD9F895D41" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Vertebral skeleton and anal-fin pterygiophores of Varicus prometheus sp. nov. Yellow numbers are the number of vertebrae." figureDoi="10.3897/zookeys.1180.107551.figure3" httpUri="https://binary.pensoft.net/fig/908648" pageId="0" pageNumber="159">, 3</figureCitation>
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<figureCitation id="4F4E50BBA6482D8B34529239F1DB77E3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Sensory papillae pattern for Varicus prometheus. Labels for papillae rows follow Sanzo (1911) as modified by Tornabene et al. (2016 b). Photo by L. Tornabene." figureDoi="10.3897/zookeys.1180.107551.figure4" httpUri="https://binary.pensoft.net/fig/908649" pageId="0" pageNumber="159">, 4 Promethean goby, Gobio prometeico (Spanish)</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E67FCB90C1C279A0F2B45DF647D128BE" pageId="0" pageNumber="159" type="type locality">
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<paragraph id="27A3EDB4333A2766F79E3856C622C0DB" pageId="0" pageNumber="159">Type locality.</paragraph>
|
||
<paragraph id="14426FF62253E4A9558047A4A64DC27A" pageId="0" pageNumber="159">Roatan, Honduras; western Caribbean.</paragraph>
|
||
</subSubSection>
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<subSubSection id="152E9868763F7E1CE150D38AE9786CC7" pageId="0" pageNumber="159" type="holotype">
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||
<paragraph id="A9D64BBEE610EAE47499D60FE715604F" pageId="0" pageNumber="159">Holotype.</paragraph>
|
||
<paragraph id="BC99DAE6BE78C0E831C6C9C531004C71" pageId="0" pageNumber="159">
|
||
Honduras • 1 female 30.5 mm SL; western Caribbean, island of Roatan, west End, off Half Moon Bay, sta. IDABEL18-02;
|
||
<geoCoordinate id="98DCB8FB4F1FBCD1DBB0422357F2A8F7" degrees="16.304" direction="north" orientation="latitude" precision="55" value="16.304">16.304°N</geoCoordinate>
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||
,
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||
<geoCoordinate id="1E0C9754A36059B5E1E90DCDF5C8C1AF" degrees="86.598" direction="west" orientation="longitude" precision="55" value="-86.598">86.598°W</geoCoordinate>
|
||
; 247 m depth; 5 June 2018; Luke Tornabene, D. Ross Robertson and Karl Stanley; 5% quinaldine-sulfate dispersed from Idabel submersible; UW 158119, DNA sample ROA18032.
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="432276F18CF4D4B97BE4DAA9A15BE4FE" pageId="0" pageNumber="159" type="generic placement">
|
||
<paragraph id="FBD55E2B2CC3B6EFDD1B14997998F6F4" pageId="0" pageNumber="159">Generic placement.</paragraph>
|
||
<paragraph id="A2489642A1802A49E52E1B9E097168BC" pageId="0" pageNumber="159">
|
||
In addition to molecular characters supporting the phylogenetic placement of this species, the following morphological characters support its inclusion in the genus
|
||
<taxonomicName id="7F25C66A1447E276F0E855C2234EBE72" authorityName="C.R.Robins & J.E.Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
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||
<emphasis id="67BF46772A478EE669C45E01A2A6DB90" italics="true" pageId="0" pageNumber="159">Varicus</emphasis>
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||
</taxonomicName>
|
||
(
|
||
<emphasis id="DA8CF1545945F9DD7B1306C2E8154037" italics="true" pageId="0" pageNumber="159">sensu</emphasis>
|
||
<bibRefCitation id="E868FE8E091F51DBA51669D93AB7151A" DOI="https://doi.org/10.1111/zoj.12394" author="Tornabene, L" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="159" pagination="764 - 812" refId="B21" refString="Tornabene, L, Van Tassell, JL, Gilmore, RG, Robertson, DR, Young, F, Baldwin, CC, 2016b. Molecular phylogeny, analysis of character evolution, and submersible collections enable a new classification for a diverse group of gobies (Teleostei: Gobiidae: Nes subgroup), including nine new species and four new genera. Zoological Journal of the Linnean Society 177 (4): 764 - 812, DOI: https://doi.org/10.1111/zoj.12394" title="Molecular phylogeny, analysis of character evolution, and submersible collections enable a new classification for a diverse group of gobies (Teleostei: Gobiidae: Nes subgroup), including nine new species and four new genera." url="https://doi.org/10.1111/zoj.12394" volume="177" year="2016 b">Tornabene et al. 2016b</bibRefCitation>
|
||
): dorsal-fin pterygiophore formula of 3-221110; 27 vertebrae (11 precaudal + 16 caudal, but see
|
||
<normalizedToken id="E1A82034C9E748D4AE0E0228A340BC08" originalValue="“Remarks”">"Remarks"</normalizedToken>
|
||
section below); one anal-fin pterygiophore inserted anterior to first haemal spine (but see
|
||
<normalizedToken id="A0EC96FDD18F3DB4D9D8E79F75D76886" originalValue="‘Remarks’">'Remarks'</normalizedToken>
|
||
section below); anal-fin rays I,8; head pores absent; pelvic fins completely separate, lacking both anterior frenum and membrane connecting bases of innermost pelvic-fin rays; fifth pelvic-fin ray unbranched.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2EC9C3754A91D4BBC875859504071AC9" pageId="0" pageNumber="159" type="diagnosis">
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||
<paragraph id="198DFCB15FB56B394E311899E4CCC01A" pageId="0" pageNumber="159">Diagnosis.</paragraph>
|
||
<paragraph id="D0213096967DC411D0830C169799766F" pageId="0" pageNumber="159">
|
||
<taxonomicName id="A50208472E7F5D0E264AA691B2A5D256" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus">
|
||
<emphasis id="C72BECD8AD6F96340F755C1655FF1B0A" italics="true" pageId="0" pageNumber="159">Varicus prometheus</emphasis>
|
||
</taxonomicName>
|
||
is distinguishable from all other
|
||
<taxonomicName id="FCF530A41FE909E7473892FE0D28480C" authorityName="C.R.Robins & J.E.Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="A21532477EBBEE1CD2C32267F684072F" italics="true" pageId="0" pageNumber="159">Varicus</emphasis>
|
||
</taxonomicName>
|
||
species by the following combination of characters: second dorsal fin I,9; anal fin I,8; pectoral fin 17; scales absent except for two modified basicaudal scales; pelvic rays 1-4 unbranched without fleshy tips; one anal-fin pterygiophore inserted anterior to first haemal spine (but see
|
||
<normalizedToken id="A380E45FEB811A80C02E0B09032457E1" originalValue="‘Remarks’">'Remarks'</normalizedToken>
|
||
section below); body with four incomplete brown saddles on pale to bright yellow background; pelvic and pectoral fins white to pale yellow in life, dorsal, anal, and caudal fins pale to bright yellow.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D139124FA21DE369C7D3EBAEDEFC0064" pageId="0" pageNumber="159" type="description">
|
||
<paragraph id="7E03C84C901D0D77EB0C345162D15992" pageId="0" pageNumber="159">Description.</paragraph>
|
||
<paragraph id="21BBA6E7F5AE7087FB881E16B9EC1E6E" pageId="0" pageNumber="159">
|
||
<emphasis id="3FBD293BBEE283F718A00416995671C9" bold="true" italics="true" pageId="0" pageNumber="159">General shape</emphasis>
|
||
: body robust, widest and deepest at posterior of head, trunk tapering in width and depth posteriorly, dorsal profile of head gently sloping from dorsum to tip of snout.
|
||
</paragraph>
|
||
<paragraph id="14288091D1E4447282AF8D1397324A2E" pageId="0" pageNumber="159">
|
||
<emphasis id="833119FF088FA7C19E69A831C6234D83" bold="true" italics="true" pageId="0" pageNumber="159">Morphometrics (%SL)</emphasis>
|
||
: head length 26.9; eye diameter 7.5; snout length 7.5; upper-jaw length 12.8; post-orbital length 13.1; predorsal length 36.7; body depth at 1st dorsal-fin origin 18.7; body depth at anal-fin origin 17.0; preanal length 61.0; body depth at caudal peduncle 10.5; caudal-peduncle length 12.1; pectoral-fin length 32.8; pelvic-fin length 25.6.
|
||
</paragraph>
|
||
<paragraph id="14C4884253E9FABAABAF28AE25E8348E" pageId="0" pageNumber="159">
|
||
<emphasis id="AC970DC31828AB18AB863E84DBEBDB9C" bold="true" italics="true" pageId="0" pageNumber="159">Median and paired fins</emphasis>
|
||
: first dorsal fin VII; second dorsal fin I,9; anal fin I,8; pectoral fin 17; pelvic fins I,5, fins well separated, lacking both anterior frenum and membrane connecting bases of innermost rays; 4th pelvic-fin ray longest, extending posteriorly to base of anal-fin spine; rays 1-4 all unbranched, no fleshy tips; 5th ray unbranched and <10% the length of the 4th ray; caudal fin shape undeterminable due to poor condition of specimen, branched caudal-fin rays 13, segmented caudal-fin rays 17.
|
||
</paragraph>
|
||
<caption id="78804BEDCFC2A80E3458D8C803A6F0C3" doi="10.3897/zookeys.1180.107551.figure2" httpUri="https://binary.pensoft.net/fig/908647" pageId="0" pageNumber="159" start="Figure 2" startId="F2">
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<paragraph id="E7A8C6D0E6C7AA6C6FA19B391065C2CE" pageId="0" pageNumber="159">
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||
<emphasis id="05CE955176697DE3F5111CF800732ECA" bold="true" pageId="0" pageNumber="159">Figure 2.</emphasis>
|
||
<taxonomicName id="90984CE4367D5F985B6C659BD00FC470" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus">
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||
<emphasis id="4B216C6FC61E833E0A910404CDBA6C20" italics="true" pageId="0" pageNumber="159">Varicus prometheus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. holotype, UW 158119, 30.5 mm SL, female, prior to preservation (top two photos) and preserved (bottom). Photos by Luke Tornabene.
|
||
</paragraph>
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||
</caption>
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||
<paragraph id="FA3364A9735A9C80C967AB153DF60AC8" pageId="0" pageNumber="159">
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<emphasis id="59136D23225C5383559E062C5A590D9C" bold="true" pageId="0" pageNumber="159">
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<emphasis id="5B40BBECCC5BD9EDE4DDD2E720D3F1C8" italics="true" pageId="0" pageNumber="159">Vertebral skeleton</emphasis>
|
||
(Fig.
|
||
<figureCitation id="0CE6908F7E7792602C8034F1B12EF2E3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Vertebral skeleton and anal-fin pterygiophores of Varicus prometheus sp. nov. Yellow numbers are the number of vertebrae." figureDoi="10.3897/zookeys.1180.107551.figure3" httpUri="https://binary.pensoft.net/fig/908648" pageId="0" pageNumber="159">3</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: 27 vertebrae, 11 precaudal vertebrae, 16 caudal vertebrae (see
|
||
<normalizedToken id="26D9A9AC1E557C49031BC88A7551560B" originalValue="‘Remarks’">'Remarks'</normalizedToken>
|
||
section below); dorsal-fin pterygiophore insertion pattern of 3-221110.
|
||
</paragraph>
|
||
<caption id="D0EF0FA1C1F6CF49021F9E81A5A9CEC8" doi="10.3897/zookeys.1180.107551.figure3" httpUri="https://binary.pensoft.net/fig/908648" pageId="0" pageNumber="159" start="Figure 3" startId="F3">
|
||
<paragraph id="9BBC16822B7D9D02BBC3B319174E3347" pageId="0" pageNumber="159">
|
||
<emphasis id="8342F9BD499ECB36A69F8B4D1EBA7E65" bold="true" pageId="0" pageNumber="159">Figure 3.</emphasis>
|
||
Vertebral skeleton and anal-fin pterygiophores of
|
||
<taxonomicName id="76358CF8F27168DA8981B6E021491564" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus">
|
||
<emphasis id="61B42AB76AACF24E0B1F4C7D9B5BBC87" italics="true" pageId="0" pageNumber="159">Varicus prometheus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Yellow numbers are the number of vertebrae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EF3AEEAFFDDF0AFF233953DD1ABF67C3" pageId="0" pageNumber="159">
|
||
<emphasis id="C295B8ACA81BEBA272DF041BEA1BAF12" bold="true" italics="true" pageId="0" pageNumber="159">Head</emphasis>
|
||
: jaw terminal, angled approximately 45 degrees from horizontal axis of the body, extending posteriorly to vertical through anterior margin of pupil; anterior nares on a short tube, posterior nares small, barely visible as an opening on a slightly raised rim immediately anterior to eye; eyes large, dorsolateral, extending to dorsal profile of head; operculum opening slightly wider than width of pectoral-fin base; teeth difficult to accurately assess due to condition of specimen, teeth in both upper and lower jaws in multiple rows, two to three rows anteriorly, with outer row in upper jaw slightly enlarged with recurved canines.
|
||
</paragraph>
|
||
<paragraph id="BB9D653F1FD5683731DF31811E1F55D0" pageId="0" pageNumber="159">
|
||
<emphasis id="1C26850C8595EEA17C056911AA39778E" bold="true" pageId="0" pageNumber="159">
|
||
<emphasis id="1C1A84E4C636CA49B2E4BE85482A8968" italics="true" pageId="0" pageNumber="159">Sensory papillae</emphasis>
|
||
(Fig.
|
||
<figureCitation id="401118AC150E56CDD778BF93F91A5017" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Sensory papillae pattern for Varicus prometheus. Labels for papillae rows follow Sanzo (1911) as modified by Tornabene et al. (2016 b). Photo by L. Tornabene." figureDoi="10.3897/zookeys.1180.107551.figure4" httpUri="https://binary.pensoft.net/fig/908649" pageId="0" pageNumber="159">4</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: sensory papillae on the sides of head of holotype are badly damaged, and the following description is a composite of both sides of the head. Five transverse rows extending from below eye, with two longitudinal rows; first longitudinal row (row
|
||
<normalizedToken id="10984FB564CAF5E26F9A7199779FD0FB" originalValue="‘d’">'d'</normalizedToken>
|
||
of
|
||
<bibRefCitation id="DFA6F6FB1EF76C80DF467BCC6400A845" author="Sanzo, L" journalOrPublisher="Mittheilungen aus der Zoologischen Station zu Neapal" pageId="0" pageNumber="159" pagination="251 - 328" refId="B16" refString="Sanzo, L, 1911. Distribuzione delle papille cutanee (organi ciatiformi) e suo valore sistematico nei Gobi. Mittheilungen aus der Zoologischen Station zu Neapal 20: 251 - 328" title="Distribuzione delle papille cutanee (organi ciatiformi) e suo valore sistematico nei Gobi." volume="20" year="1911">Sanzo 1911</bibRefCitation>
|
||
) at level just above angle of jaw row, extending between second and fourth transverse row; second longitudinal row (row
|
||
<normalizedToken id="8B9449439EB2DC3C88859BF2ADDDDAB8" originalValue="‘b’">'b'</normalizedToken>
|
||
of
|
||
<bibRefCitation id="43C8B0B3AD540440405B80DE3837D862" author="Sanzo, L" journalOrPublisher="Mittheilungen aus der Zoologischen Station zu Neapal" pageId="0" pageNumber="159" pagination="251 - 328" refId="B16" refString="Sanzo, L, 1911. Distribuzione delle papille cutanee (organi ciatiformi) e suo valore sistematico nei Gobi. Mittheilungen aus der Zoologischen Station zu Neapal 20: 251 - 328" title="Distribuzione delle papille cutanee (organi ciatiformi) e suo valore sistematico nei Gobi." volume="20" year="1911">Sanzo 1911</bibRefCitation>
|
||
) short, originating below posterior margin of eye immediately posterior to fourth transverse row; first transverse row originating at anteroventral corner of eye, second transverse row below originating at anterior margin of pupil, third transverse row originating below posterior margin of pupil, fourth transverse row (row 5i/5s of
|
||
<bibRefCitation id="9D13958A132F65E45ECD3BFF207CD68E" DOI="https://doi.org/10.1111/zoj.12394" author="Tornabene, L" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="159" pagination="764 - 812" refId="B21" refString="Tornabene, L, Van Tassell, JL, Gilmore, RG, Robertson, DR, Young, F, Baldwin, CC, 2016b. Molecular phylogeny, analysis of character evolution, and submersible collections enable a new classification for a diverse group of gobies (Teleostei: Gobiidae: Nes subgroup), including nine new species and four new genera. Zoological Journal of the Linnean Society 177 (4): 764 - 812, DOI: https://doi.org/10.1111/zoj.12394" title="Molecular phylogeny, analysis of character evolution, and submersible collections enable a new classification for a diverse group of gobies (Teleostei: Gobiidae: Nes subgroup), including nine new species and four new genera." url="https://doi.org/10.1111/zoj.12394" volume="177" year="2016 b">Tornabene et al. 2016b</bibRefCitation>
|
||
) originating below posterior margin of eye and extending below longitudinal as a single continuous row (5i/5s continuous versus separate), fifth transverse row diagonal, originating at posterior margin of eye, well above fourth row; several short vertical rows of papillae above operculum; a vertical row of four evenly space papillae near anterior margin of operculum; a short oblique row of papillae across middle of operculum; papillae row p not apparent except for a single papilla
|
||
<normalizedToken id="48E1D79D516C8E218E046DDAA8B76B15" originalValue="peʹ">peʹ</normalizedToken>
|
||
behind each eye; body with approximately 15 short vertical rows of papillae evenly spaced along lateral midline, beginning below origin of second dorsal fin and continuing onto caudal peduncle.
|
||
</paragraph>
|
||
<caption id="C3BF0570B751B95FAC837EC8968C1768" doi="10.3897/zookeys.1180.107551.figure4" httpUri="https://binary.pensoft.net/fig/908649" pageId="0" pageNumber="159" start="Figure 4" startId="F4">
|
||
<paragraph id="DFF9007BF04E4935BC2C89A175C15579" pageId="0" pageNumber="159">
|
||
<emphasis id="F8A49F972D3CBB525FDEC1B5E4EFFAE5" bold="true" pageId="0" pageNumber="159">Figure 4.</emphasis>
|
||
Sensory papillae pattern for
|
||
<taxonomicName id="7DE743D97AD127F710F7384E4CF88F0E" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus">
|
||
<emphasis id="28AC2E32C9FCAF83427719B7F23B70B3" italics="true" pageId="0" pageNumber="159">Varicus prometheus</emphasis>
|
||
</taxonomicName>
|
||
. Labels for papillae rows follow
|
||
<bibRefCitation id="61B5C21BB6FB4CFF39C48A45A72839FD" author="Sanzo, L" journalOrPublisher="Mittheilungen aus der Zoologischen Station zu Neapal" pageId="0" pageNumber="159" pagination="251 - 328" refId="B16" refString="Sanzo, L, 1911. Distribuzione delle papille cutanee (organi ciatiformi) e suo valore sistematico nei Gobi. Mittheilungen aus der Zoologischen Station zu Neapal 20: 251 - 328" title="Distribuzione delle papille cutanee (organi ciatiformi) e suo valore sistematico nei Gobi." volume="20" year="1911">Sanzo (1911)</bibRefCitation>
|
||
as modified by
|
||
<bibRefCitation id="775D776EC7F6C610BD4E30A3AE433F07" DOI="https://doi.org/10.1111/zoj.12394" author="Tornabene, L" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="159" pagination="764 - 812" refId="B21" refString="Tornabene, L, Van Tassell, JL, Gilmore, RG, Robertson, DR, Young, F, Baldwin, CC, 2016b. Molecular phylogeny, analysis of character evolution, and submersible collections enable a new classification for a diverse group of gobies (Teleostei: Gobiidae: Nes subgroup), including nine new species and four new genera. Zoological Journal of the Linnean Society 177 (4): 764 - 812, DOI: https://doi.org/10.1111/zoj.12394" title="Molecular phylogeny, analysis of character evolution, and submersible collections enable a new classification for a diverse group of gobies (Teleostei: Gobiidae: Nes subgroup), including nine new species and four new genera." url="https://doi.org/10.1111/zoj.12394" volume="177" year="2016 b">Tornabene et al. (2016b)</bibRefCitation>
|
||
. Photo by L. Tornabene.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="915E3EF4F6580D98F9A65EEE2AB28C77" pageId="0" pageNumber="159">
|
||
<emphasis id="D249B9BE3FBA023498F4A0E78CB5C3BC" bold="true" italics="true" pageId="0" pageNumber="159">Squamation</emphasis>
|
||
: scales on head and body absent except for two modified basicaudal scales with enlarged ctenii present at dorsal and ventral margins of caudal-fin base.
|
||
</paragraph>
|
||
<paragraph id="7842B435B8140E341F3B51E6C05EC2DF" pageId="0" pageNumber="159">
|
||
<emphasis id="5197985963F05595291568717C977EFE" bold="true" italics="true" pageId="0" pageNumber="159">Genitalia</emphasis>
|
||
: female with short, slightly rounded, conical papilla, wider at base and tapering to a slight point; male unknown.
|
||
</paragraph>
|
||
<paragraph id="A95196079C09B6D6A6843D373C75F115" pageId="0" pageNumber="159">
|
||
<emphasis id="DAF677C2C09D91E22208580090F8C196" bold="true" pageId="0" pageNumber="159">
|
||
<emphasis id="C0D9AF27A64A15101E3E239AF0B0CB4B" italics="true" pageId="0" pageNumber="159">Color before preservation</emphasis>
|
||
(Fig.
|
||
<figureCitation id="207FD42AD02D1E25D3974003DE9A2093" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Varicus prometheus sp. nov. holotype, UW 158119, 30.5 mm SL, female, prior to preservation (top two photos) and preserved (bottom). Photos by Luke Tornabene." figureDoi="10.3897/zookeys.1180.107551.figure2" httpUri="https://binary.pensoft.net/fig/908647" pageId="0" pageNumber="159">2</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: background color of head and body white to pale yellow, with upper portion of head and body slightly brighter yellow than posterior half of body (pigmentation of which is more visible when fish was photographed against white vs. black background). Head with areas of bright yellow pigment lightly speckled with small brown melanophores across upper jaw and snout; four dark bars formed by speckled brown melanophores; first two bars, slightly narrower than pupil diameter, extending ventrally from interorbital, one across the anterior eye and ending at posterior tip of jaw and the second crossing slightly posterior to pupil and ending mid-cheek; third bar originating well behind eye at dorsal midline and extending ventrally slightly onto middle of opercle; and fourth bar wider than first three, anterior to first dorsal fin, extending ventrally to approximately to just above pectoral-fin base. Eyes with green-yellow irises speckled with silvery dots and a thin silvery ring around the pupil. Body pale yellow to white with four dark saddles heavily speckled with brown melanophores; first saddle slightly narrower in width than eye diameter, located at middle of base of first dorsal fin extending ventrally to lateral midline where melanophores become less dense; second saddle about equal in width to eye diameter, located at anterior of second dorsal fin extending ventrally just short of lateral midline; third saddle slightly wider than eye diameter, located under rear of second dorsal fin base, extending ventrally to just beyond lateral midline; fourth saddle located laterally on the caudal peduncle at base of caudal fin, the front border rounded and the rear border straight, slightly larger than eye diameter. First dorsal fin bright yellow with a few brown melanophores along base immediately above saddle on body, few scattered melanophores on interradial membranes; second dorsal fin similarly colored as first. Caudal fin with a very wide oblique yellow bar down middle of fin rays. Anal fin bright yellow. Pectoral-fin base yellow with blotch of brown melanophores at center on the upper third of the fin base, extending to origin of rays. Pelvic fins pale.
|
||
</paragraph>
|
||
<paragraph id="DEDBF8ED6D71103F1EA4AD1E1FA37885" pageId="0" pageNumber="159">
|
||
<emphasis id="D552CD80C934097AF1E616DB69705AD7" bold="true" pageId="0" pageNumber="159">
|
||
<emphasis id="F0DE8E706A06C10EA0F6B73BE2C9EE67" italics="true" pageId="0" pageNumber="159">Color in preservation</emphasis>
|
||
(Fig.
|
||
<figureCitation id="509D6ABB2056E32D84FD80BD109102BB" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Varicus prometheus sp. nov. holotype, UW 158119, 30.5 mm SL, female, prior to preservation (top two photos) and preserved (bottom). Photos by Luke Tornabene." figureDoi="10.3897/zookeys.1180.107551.figure2" httpUri="https://binary.pensoft.net/fig/908647" pageId="0" pageNumber="159">2</figureCitation>
|
||
)
|
||
</emphasis>
|
||
: body and head uniformly pale yellow, fins translucent. Only remaining pigment consisting of light speckling of small melanophores on trunk at location where dark saddles occurred in life.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8565BDD146E5842CD9936286F93C940" pageId="0" pageNumber="159" type="habitat">
|
||
<paragraph id="F2BB40F59DC54548CC9C0EC9088ED737" pageId="0" pageNumber="159">Habitat.</paragraph>
|
||
<paragraph id="AE746C5A4EDDBB71AE5A888BC1507574" pageId="0" pageNumber="159">
|
||
The holotype was collected at from the deep-reef slope at 247 m on sand with
|
||
<taxonomicName id="B3A4AB866417F7FD33A7550CD12DCF6F" authorityName="Jean Vincent Felix Lamouroux" authorityYear="1812" class="Bryopsidophyceae" family="Halimedaceae" genus="Halimeda" higherTaxonomySource="IPNI" kingdom="Plantae" lsidName="Halimeda" order="Bryopsidales" pageId="0" pageNumber="159" phylum="Chlorophyta" rank="genus">
|
||
<emphasis id="F83DB8FD29F2FD485B59138C0AC1F774" italics="true" pageId="0" pageNumber="159">Halimeda</emphasis>
|
||
</taxonomicName>
|
||
rubble.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DFC34B9F77FFA54B31B432CA0948E31C" pageId="0" pageNumber="159" type="distribution">
|
||
<paragraph id="B752A389FA40CEDCCC7B9F403C394C35" pageId="0" pageNumber="159">Distribution.</paragraph>
|
||
<paragraph id="0808414F87068F473A999DB8DFD2B3DD" pageId="0" pageNumber="159">Known only from the type locality off Roatan, Honduras.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67235C360F9C7FD6A083F2647E3B1038" pageId="0" pageNumber="159" type="etymology">
|
||
<paragraph id="CF9D86145165422A7A67F7DEC74B3608" pageId="0" pageNumber="159">Etymology.</paragraph>
|
||
<paragraph id="4D4D441B13E9434C305057636BA3CC35" pageId="0" pageNumber="159">
|
||
The specific epithet
|
||
<taxonomicName id="00EA1DE6E31E3E815919521DAAED6A34" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus">
|
||
<emphasis id="57F11257CF2E507EFA299877E0D0E2EA" italics="true" pageId="0" pageNumber="159">Varicus prometheus</emphasis>
|
||
</taxonomicName>
|
||
is a patronym in reference to the Greek god Prometheus. In Greek mythology, as punishment from the god Zeus, Prometheus had his liver eaten out by an eagle, only to have the liver grow back overnight so it might be eaten again the next day. The name refers to the fact that the abdomen of the holotype of the new species was partially eaten by a hermit crab. The name is treated as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FC487787AB7AD5DF968D80FBBE0747F" pageId="0" pageNumber="159" type="remarks">
|
||
<paragraph id="5262949EE6453A5A07A56AC600FBC964" pageId="0" pageNumber="159">Remarks.</paragraph>
|
||
<paragraph id="4C27C553440A9AA67C6C0E474E877FE1" pageId="0" pageNumber="159">
|
||
<taxonomicName id="F1A37E47EDD9D4CA6D2AC8E6B89E74FB" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus">
|
||
<emphasis id="7386EB6453B2ABA55C56DB0DFD4E06B9" italics="true" pageId="0" pageNumber="159">Varicus prometheus</emphasis>
|
||
</taxonomicName>
|
||
most likely possesses 27 total vertebrae, like all other species in genus. However, our radiographs did not have enough resolution for us to determine whether the 12th vertebra had a haemal spine (if present, it is very reduced) rendering it the first caudal vertebra versus the last precaudal vertebra (Fig.
|
||
<figureCitation id="D5090C037BADE9642768653EC0EA92B0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Vertebral skeleton and anal-fin pterygiophores of Varicus prometheus sp. nov. Yellow numbers are the number of vertebrae." figureDoi="10.3897/zookeys.1180.107551.figure3" httpUri="https://binary.pensoft.net/fig/908648" pageId="0" pageNumber="159">3</figureCitation>
|
||
). Thus, the counts of precaudal and caudal vertebrae may be 11+17 or 12+16. Accordingly, it is unclear how many anal-fin pterygiophores are inserted anterior to the first haemal spine. All other species of
|
||
<taxonomicName id="31AF2F7885145919B7A126B9ACCCA23F" authorityName="C.R.Robins & J.E.Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="4E3CF318E1B281ABF372B92081058F2C" italics="true" pageId="0" pageNumber="159">Varicus</emphasis>
|
||
</taxonomicName>
|
||
have one anal-fin pterygiophore inserted anterior to the haemal arch that is present on the 12th vertebra (other species of
|
||
<taxonomicName id="E00F86F970EEABD188460DF314A57930" authorityName="Birdsong" authorityYear="1975" lsidName="" pageId="0" pageNumber="159" rank="tribe" tribe="Gobiosomatini">Gobiosomatini</taxonomicName>
|
||
have two pterygiophores inserted here, including the morphologically similar genus
|
||
<taxonomicName id="E142683C2C3BCFECC327AAB4F7E13995" authorityName="Ginsburg" authorityYear="1953" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="A61B36A7A3604435E5A09E88BF030631" italics="true" pageId="0" pageNumber="159">Psilotris</emphasis>
|
||
</taxonomicName>
|
||
Ginsburg, 1953). In some rare instances in
|
||
<taxonomicName id="167D647C95CD7E11B707D81751F4334C" authorityName="C.R.Robins & J.E.Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="92BC7CD7A4B023478CF0F8E9FA6876A0" italics="true" pageId="0" pageNumber="159">Varicus</emphasis>
|
||
</taxonomicName>
|
||
, the haemal arch on the 12th vertebra is very reduced and does not insert deeply into the space between the first two anal-fin pterygiophores, making it incorrectly appear on the x-ray as if there are two anal pterygiophores inserted before the
|
||
<normalizedToken id="0094223601BC05154FC486E1A4417A19" originalValue="“first”">"first"</normalizedToken>
|
||
haemal spine on the 13th vertebrae, rather than one pterygiophore inserted before the 12th vertebra (Fig.
|
||
<figureCitation id="2A9D5976B8805595D1404B9C56A12654" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Vertebral skeleton and anal-fin pterygiophores of Varicus prometheus sp. nov. Yellow numbers are the number of vertebrae." figureDoi="10.3897/zookeys.1180.107551.figure3" httpUri="https://binary.pensoft.net/fig/908648" pageId="0" pageNumber="159">3</figureCitation>
|
||
; also see
|
||
<bibRefCitation id="59C75C379A47FD2C56C3B4B7C779F917" DOI="https://doi.org/10.1111/zoj.12394" author="Tornabene, L" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="159" pagination="764 - 812" refId="B21" refString="Tornabene, L, Van Tassell, JL, Gilmore, RG, Robertson, DR, Young, F, Baldwin, CC, 2016b. Molecular phylogeny, analysis of character evolution, and submersible collections enable a new classification for a diverse group of gobies (Teleostei: Gobiidae: Nes subgroup), including nine new species and four new genera. Zoological Journal of the Linnean Society 177 (4): 764 - 812, DOI: https://doi.org/10.1111/zoj.12394" title="Molecular phylogeny, analysis of character evolution, and submersible collections enable a new classification for a diverse group of gobies (Teleostei: Gobiidae: Nes subgroup), including nine new species and four new genera." url="https://doi.org/10.1111/zoj.12394" volume="177" year="2016 b">Tornabene et al. 2016b</bibRefCitation>
|
||
, fig. 2). This is likely the case with
|
||
<taxonomicName id="9DDEF537261C332B2F91A29C99E28F81" lsidName="V. prometheus" pageId="0" pageNumber="159" rank="species" species="prometheus">
|
||
<emphasis id="A4C06809C67E6390E6AD557D83C27FED" italics="true" pageId="0" pageNumber="159">V. prometheus</emphasis>
|
||
</taxonomicName>
|
||
, and we tentatively consider the vertebral counts to be 11+16 with a single anal-fin pterygiophore inserted before the haemal arch.
|
||
</paragraph>
|
||
<paragraph id="17BF1610FBE51B38A29C300E5C3E044A" pageId="0" pageNumber="159">
|
||
<taxonomicName id="4EB658377FCD793FDDE667F6EF8F47AD" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus">
|
||
<emphasis id="7AA6E1452F55B43170EB2B67D4A5614D" italics="true" pageId="0" pageNumber="159">Varicus prometheus</emphasis>
|
||
</taxonomicName>
|
||
is easily distinguished from other species of
|
||
<taxonomicName id="83F940769B18D8DA3E31953E1B43EFDE" authorityName="C.R.Robins & J.E.Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="B23AECCFAECC3BA8CD7E7B6FFFB94E12" italics="true" pageId="0" pageNumber="159">Varicus</emphasis>
|
||
</taxonomicName>
|
||
and species in the morphologically similar genus
|
||
<taxonomicName id="D6D0567F5AD7D8BEBEF7060B4529EAD2" authorityName="Ginsburg" authorityYear="1953" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="0328C98807C4C4BB32951B5CFA39C274" italics="true" pageId="0" pageNumber="159">Psilotris</emphasis>
|
||
</taxonomicName>
|
||
(Table
|
||
<tableCitation id="90DFBC7CDF329A1E2A52777C3DA3E3FE" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Important taxonomic characters for the genera Varicus, Psilotris and Pinnichthys." httpUri="http://table.plazi.org/id/42DDABA574CA38025C21E7B792CB5EE6" pageId="0" pageNumber="159" tableUuid="42DDABA574CA38025C21E7B792CB5EE6">1</tableCitation>
|
||
) by the presence of four broad brown saddles on a uniformly yellow body. No other species of
|
||
<taxonomicName id="3574B9AD577EE3A9EE29EE313350916D" authorityName="C.R.Robins & J.E.Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="543AE13A4394D57740C6324EEF410FA5" italics="true" pageId="0" pageNumber="159">Varicus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4E6ADF68EA01C342FFDF3E94847BB6C1" authorityName="Ginsburg" authorityYear="1953" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="587305080B388D311886064B0FC93CC7" italics="true" pageId="0" pageNumber="159">Psilotris</emphasis>
|
||
</taxonomicName>
|
||
has such saddles, although a few species [e.g.,
|
||
<taxonomicName id="1C32EC4E94739FFB2279F79975872149" lsidName="V. adamsi" pageId="0" pageNumber="159" rank="species" species="adamsi">
|
||
<emphasis id="4C289F2E612BB5A9CBD4EFD587A4C49C" italics="true" pageId="0" pageNumber="159">V. adamsi</emphasis>
|
||
</taxonomicName>
|
||
Gilmore, Van Tassell & Tornabene, 2016;
|
||
<taxonomicName id="1D8EE47FA1EF907531AFA66144B8CB7D" lsidName="V. vespa" pageId="0" pageNumber="159" rank="species" species="vespa">
|
||
<emphasis id="81069E48A50A8230C6F1A53E0C7D0A39" italics="true" pageId="0" pageNumber="159">V. vespa</emphasis>
|
||
</taxonomicName>
|
||
(Hastings & Bortone, 1981);
|
||
<taxonomicName id="37305EA2DCFD6DA0E06B5AD49AD06331" lsidName="V. marilynae" pageId="0" pageNumber="159" rank="species" species="marilynae">
|
||
<emphasis id="DAD95A112DCBDBEDE931CB5DF5515D7E" italics="true" pageId="0" pageNumber="159">V. marilynae</emphasis>
|
||
</taxonomicName>
|
||
Gilmore, 1979] have complete or nearly complete vertical bars that are narrower than the saddles in
|
||
<taxonomicName id="31C360931C0F32501111FE7A1B5E33F2" lsidName="V. prometheus" pageId="0" pageNumber="159" rank="species" species="prometheus">
|
||
<emphasis id="6A518C13BF9FC9801DD896925DF6D35F" italics="true" pageId="0" pageNumber="159">V. prometheus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="45BC44088293310582978959F37EC407" lsidName="V. nigritus" pageId="0" pageNumber="159" rank="species" species="nigritus">
|
||
<emphasis id="F041771625C29BA22CE73E1AC57CB1AF" italics="true" pageId="0" pageNumber="159">V. nigritus</emphasis>
|
||
</taxonomicName>
|
||
has large dark blotches on the body that are as wide as the saddles in
|
||
<taxonomicName id="4A281E9195C9D0B9E1E8A81122554C5B" lsidName="V. prometheus" pageId="0" pageNumber="159" rank="species" species="prometheus">
|
||
<emphasis id="652698E62E1569BFF144C7EEE964E2D1" italics="true" pageId="0" pageNumber="159">V. prometheus</emphasis>
|
||
</taxonomicName>
|
||
but are round rather than saddle shaped (Fig.
|
||
<figureCitation id="0DB2651179BA3F80EBE86DA4C5DD2A72" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Live or fresh coloration of Varicus species (coloration unknown for V. benthonis and V. bucca) A V. adamsi, photo by L. Tornabene B V. cephalocellatus, photo by B. Brown C V. decorum, photo by B. Brown D V. lacerta, photo by C. Baldwin E V. marilynae, illustration by R. G. Gilmore F V. veliguttatus, photo by C. Baldwin G V. nigritus, illustration by R. G. Gilmor H V. vespa, photo by the crew of the R / V BELLOWS." figureDoi="10.3897/zookeys.1180.107551.figure5" httpUri="https://binary.pensoft.net/fig/908650" pageId="0" pageNumber="159">5</figureCitation>
|
||
).
|
||
<taxonomicName id="37129F26679D3CE10D0B7078E6B90451" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus">
|
||
<emphasis id="10705299C6A21AB4D8561B01868D1174" italics="true" pageId="0" pageNumber="159">Varicus prometheus</emphasis>
|
||
</taxonomicName>
|
||
lacks scales on the trunk, whereas all other
|
||
<taxonomicName id="29B7DE777B4D66EF6AF525B728998157" authorityName="C.R.Robins & J.E.Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="6D2AC96AF096995A96FC4E6A30CE1D61" italics="true" pageId="0" pageNumber="159">Varicus</emphasis>
|
||
</taxonomicName>
|
||
species except
|
||
<taxonomicName id="6DDE56304D8008055F2EAB8E7B37DAB2" lsidName="V. lacerta" pageId="0" pageNumber="159" rank="species" species="lacerta">
|
||
<emphasis id="DFCE3711365FDE45C42A8174D701CD5A" italics="true" pageId="0" pageNumber="159">V. lacerta</emphasis>
|
||
</taxonomicName>
|
||
Tornabene, Robertson & Baldwin, 2016, and
|
||
<taxonomicName id="3BCFC1F85E7953CC87E022D565DB1062" lsidName="V. decorum" pageId="0" pageNumber="159" rank="species" species="decorum">
|
||
<emphasis id="7EA4FFAA054ECC0224824EC445785690" italics="true" pageId="0" pageNumber="159">V. decorum</emphasis>
|
||
</taxonomicName>
|
||
Van Tassell, Baldwin & Tornabene, 2016, have many rows of scales on the trunk. Both
|
||
<taxonomicName id="2BCC00B349AE6D6180106EECB836BED0" lsidName="V. decorum" pageId="0" pageNumber="159" rank="species" species="decorum">
|
||
<emphasis id="3FC3ACA7479E3CF971E7EF26E38B0284" italics="true" pageId="0" pageNumber="159">V. decorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7AF30F392D3E6BD94E3290EB67AA5DAE" lsidName="V. prometheus" pageId="0" pageNumber="159" rank="species" species="prometheus">
|
||
<emphasis id="5FFCAAC916BAE0A5C2177286F9344CA7" italics="true" pageId="0" pageNumber="159">V. prometheus</emphasis>
|
||
</taxonomicName>
|
||
possess modified basicaudal scales, which are lacking in
|
||
<taxonomicName id="387CFA4BE71280830C970572C2FCFAF2" lsidName="V. lacerta" pageId="0" pageNumber="159" rank="species" species="lacerta">
|
||
<emphasis id="74917A21BBE5F12DE4DC06B817739AD9" italics="true" pageId="0" pageNumber="159">V. lacerta</emphasis>
|
||
</taxonomicName>
|
||
. The unbranched pelvic rays of
|
||
<taxonomicName id="DC888E93D1BA3BBAE2DA32D0E2C8BACB" lsidName="V. prometheus" pageId="0" pageNumber="159" rank="species" species="prometheus">
|
||
<emphasis id="5ABFA7CF53D937F782368B5E1B8DB91E" italics="true" pageId="0" pageNumber="159">V. prometheus</emphasis>
|
||
</taxonomicName>
|
||
and the presence of one anal-fin pterygiophore (versus 2) inserted before the haemal arch distinguish it from all species of
|
||
<taxonomicName id="0BE431A9F640A2B4FC499021DEFE3891" authorityName="Ginsburg" authorityYear="1953" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="B447D0CA4762C3399588BEC95745164F" italics="true" pageId="0" pageNumber="159">Psilotris</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="70FB2D9D18F4CC8CC5BB83D75E1441CB" doi="10.3897/zookeys.1180.107551.figure5" httpUri="https://binary.pensoft.net/fig/908650" pageId="0" pageNumber="159" start="Figure 5" startId="F5">
|
||
<paragraph id="FB2E99C79E3C6E857583DB833CF62C3A" pageId="0" pageNumber="159">
|
||
<emphasis id="08D86411330C9658FB4E0572495C48B5" bold="true" pageId="0" pageNumber="159">Figure 5.</emphasis>
|
||
Live or fresh coloration of
|
||
<taxonomicName id="91CA1C793E858667BB26985580F916D8" authorityName="C.R.Robins & J.E.Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="3B6531CB761FD2F49C58B4872EFCC080" italics="true" pageId="0" pageNumber="159">Varicus</emphasis>
|
||
</taxonomicName>
|
||
species (coloration unknown for
|
||
<taxonomicName id="E4CC2CEDC76F84B68C2D2E72BC47C90E" lsidName="V. benthonis" pageId="0" pageNumber="159" rank="species" species="benthonis">
|
||
<emphasis id="50F16C42D7B30B0E69DA04E36CE99BB7" italics="true" pageId="0" pageNumber="159">V. benthonis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="17D07CD484F36F1B6CED36D714063588" lsidName="V. bucca" pageId="0" pageNumber="159" rank="species" species="bucca">
|
||
<emphasis id="E7C7AAA435DB918FA6CDDCC9D26DC218" italics="true" pageId="0" pageNumber="159">V. bucca</emphasis>
|
||
</taxonomicName>
|
||
)
|
||
<emphasis id="CECA1695A70FB0BBD066C2CE89EC1147" bold="true" pageId="0" pageNumber="159">A</emphasis>
|
||
<taxonomicName id="C2AAB4895217F995EC087EAD4EBA5522" lsidName="V. adamsi" pageId="0" pageNumber="159" rank="species" species="adamsi">
|
||
<emphasis id="F3466C20D84241C1F1E39AD4CD0C426F" italics="true" pageId="0" pageNumber="159">V. adamsi</emphasis>
|
||
</taxonomicName>
|
||
, photo by L. Tornabene
|
||
<emphasis id="17DD1D4F45AE6B9BFD7E63EEDD498ECB" bold="true" pageId="0" pageNumber="159">B</emphasis>
|
||
<taxonomicName id="6D8B78FFF991C8D8D08CD0FAE4BCE00D" lsidName="V. cephalocellatus" pageId="0" pageNumber="159" rank="species" species="cephalocellatus">
|
||
<emphasis id="9A6726D25F097C45B480D923AC069769" italics="true" pageId="0" pageNumber="159">V. cephalocellatus</emphasis>
|
||
</taxonomicName>
|
||
, photo by B. Brown
|
||
<emphasis id="EA24C951D117EE887325F5E5B44AC7F2" bold="true" pageId="0" pageNumber="159">C</emphasis>
|
||
<taxonomicName id="1278AAD2A9C28D5E24659629BBF0380A" lsidName="V. decorum" pageId="0" pageNumber="159" rank="species" species="decorum">
|
||
<emphasis id="BA905EA12DE9B582D6389D0095B19B2E" italics="true" pageId="0" pageNumber="159">V. decorum</emphasis>
|
||
</taxonomicName>
|
||
, photo by B. Brown
|
||
<emphasis id="4E6045FBD09C4C1984189EB6FB242F59" bold="true" pageId="0" pageNumber="159">D</emphasis>
|
||
<taxonomicName id="CFFDB0A4613C4C408FD6A94E4198C701" lsidName="V. lacerta" pageId="0" pageNumber="159" rank="species" species="lacerta">
|
||
<emphasis id="560650F6E9638E89867EDDC80FD968C6" italics="true" pageId="0" pageNumber="159">V. lacerta</emphasis>
|
||
</taxonomicName>
|
||
, photo by C. Baldwin
|
||
<emphasis id="7F321F103D16A49CDD36E997DC00CABC" bold="true" pageId="0" pageNumber="159">E</emphasis>
|
||
<taxonomicName id="309B0FAC0C70B3268C43D63DEA23CE92" lsidName="V. marilynae" pageId="0" pageNumber="159" rank="species" species="marilynae">
|
||
<emphasis id="7A838CC8C8DF8E646ABC15ABB2B86EF7" italics="true" pageId="0" pageNumber="159">V. marilynae</emphasis>
|
||
</taxonomicName>
|
||
, illustration by R. G. Gilmore
|
||
<emphasis id="58DB6860A70229D7BF05764F5BD7BA56" bold="true" pageId="0" pageNumber="159">F</emphasis>
|
||
<taxonomicName id="FB897D3B761E09A89AA1DDF55C0C1BD9" lsidName="V. veliguttatus" pageId="0" pageNumber="159" rank="species" species="veliguttatus">
|
||
<emphasis id="7694B3F8AE6821EEC5DAE211112FC441" italics="true" pageId="0" pageNumber="159">V. veliguttatus</emphasis>
|
||
</taxonomicName>
|
||
, photo by C. Baldwin
|
||
<emphasis id="79AC2E975A65425AC90D0C70BF3D62F6" bold="true" pageId="0" pageNumber="159">G</emphasis>
|
||
<taxonomicName id="6E95FAB265177C35575DD2857775B428" lsidName="V. nigritus" pageId="0" pageNumber="159" rank="species" species="nigritus">
|
||
<emphasis id="4F320668264D2B0CE0BEBF7F8BBF69A2" italics="true" pageId="0" pageNumber="159">V. nigritus</emphasis>
|
||
</taxonomicName>
|
||
, illustration by R. G. Gilmor
|
||
<emphasis id="ED94067E33C8D2471794A36E0D8F7554" bold="true" pageId="0" pageNumber="159">H</emphasis>
|
||
<taxonomicName id="601DDBDB133014FD98EA8DEE117D6BC4" lsidName="V. vespa" pageId="0" pageNumber="159" rank="species" species="vespa">
|
||
<emphasis id="5E631D4B5A42388D26C0D75200E5BDCA" italics="true" pageId="0" pageNumber="159">V. vespa</emphasis>
|
||
</taxonomicName>
|
||
, photo by the crew of the R/V BELLOWS.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="42DDABA574CA38025C21E7B792CB5EE6" ID-Table-UUID="42DDABA574CA38025C21E7B792CB5EE6" httpUri="http://table.plazi.org/id/42DDABA574CA38025C21E7B792CB5EE6" pageId="0" pageNumber="159" start="Table 1" startId="T1">
|
||
<paragraph id="D6A4764B62C89FB53FB5FAC3E627C16E" pageId="0" pageNumber="159">
|
||
<emphasis id="AF6A8DB8825A3773E15321C2E76F9BED" bold="true" pageId="0" pageNumber="159">Table 1.</emphasis>
|
||
Important taxonomic characters for the genera
|
||
<taxonomicName id="691D0212E74FF2488EB470018E476F43" authorityName="C.R.Robins & J.E.Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="C9A50D0F7F077C9F75B9BC87B1001DA8" italics="true" pageId="0" pageNumber="159">Varicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="98F0921E1655DB684E715F660353D9BE" authorityName="Ginsburg" authorityYear="1953" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="CFF22F02EB0513420FF288A907931D35" italics="true" pageId="0" pageNumber="159">Psilotris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="8BCFE1223679C4951E5517E89E414817" authorityName="Van Tassell, Tornabene & Gilmorean Tassell" authorityYear="2016" family="Gobiidae" genus="Pinnichthys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pinnichthys" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="genus">
|
||
<emphasis id="E79E31AFCC93608B2E337AF733179BAE" italics="true" pageId="0" pageNumber="159">Pinnichthys</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FB2C6AB9427414D8DFAE034D8884A3B1" pageId="0" pageNumber="159">
|
||
<table id="5D61511313351895E016C6095421C3B8" pageId="0" pageNumber="159">
|
||
<tr id="7AE7296E7D60665D15AB2C662820383F" pageId="0" pageNumber="159">
|
||
<th id="E50366A1FF605444E0DAF7DD925FFAE4" colspan="1" pageId="0" pageNumber="159" rowspan="1">Species</th>
|
||
<th id="58271C560C56ADD65B6C9F8BA43D77C5" colspan="1" pageId="0" pageNumber="159" rowspan="1">Second dorsal</th>
|
||
<th id="938F2D26ABCACD6512926BC894403F2A" colspan="1" pageId="0" pageNumber="159" rowspan="1">Anal</th>
|
||
<th id="44E878CC4C3171FA041183042C06FE1C" colspan="1" pageId="0" pageNumber="159" rowspan="1">Pectoral</th>
|
||
<th id="462BF264D1A23E9A14467680328A6464" colspan="1" pageId="0" pageNumber="159" rowspan="1">AP</th>
|
||
<th id="5AC1133B4968683506868CF2E431A0CA" colspan="1" pageId="0" pageNumber="159" rowspan="1">Pelvic rays 1-4</th>
|
||
<th id="66E489879D99B84B59811C8D2B5AA89F" colspan="1" pageId="0" pageNumber="159" rowspan="1">Papillae rows 5i/5s</th>
|
||
<th id="B39DE8852D01FAA134905002296B341D" colspan="1" pageId="0" pageNumber="159" rowspan="1">Lateral scale rows</th>
|
||
<th id="10DC8DB59CFCA49B862AC45CE5965546" colspan="1" pageId="0" pageNumber="159" rowspan="1">Basicaudal Scales</th>
|
||
</tr>
|
||
<tr id="8E73E53F942004FA8ED187DDA3C6D336" pageId="0" pageNumber="159">
|
||
<td id="1407B5BE4DC8932C572F243B7B830018" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="8E8E9D5E1D94AD1C4822DC30CD6834A1" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus prometheus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prometheus">
|
||
<emphasis id="573721FE1FE30562FB5388E743AF942C" italics="true" pageId="0" pageNumber="159">Varicus prometheus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov
|
||
</td>
|
||
<td id="9147001D4F3463A87E82125318AA6F55" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="81C64A3FA7CA60DDA598FB7254B99283" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,8</td>
|
||
<td id="539F49DAF0512927CD3670C8075F4D6D" colspan="1" pageId="0" pageNumber="159" rowspan="1">17</td>
|
||
<td id="B65ED63BF52848158E4CD69985AE4075" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="BC486108AB394FCE8C6BABE33BEB4176" colspan="1" pageId="0" pageNumber="159" rowspan="1">unbranched</td>
|
||
<td id="F4DE0955DD108758D531687A1788BFE1" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="1D346D72047946A8A2B67D3B608C1021" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="95393EBAFFB8EA87569B2BBC4911D77F" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="E02B00EC5AB7BAAECEAA7513D6EA05F1" pageId="0" pageNumber="159">
|
||
<td id="35CF4CB9D5CA5496A733F19A909B4787" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="B8C2E184E317896BF0424AF2D336CABD" authorityName="Fuentes & Baldwin & Robertson & Lardizábal & Tornabene" authorityYear="2023" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus roatanensis" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="roatanensis">
|
||
<emphasis id="912763AAACDA2DE39A86FDDB841D9B6F" italics="true" pageId="0" pageNumber="159">Varicus roatanensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov
|
||
</td>
|
||
<td id="D98D8E92A608E918399994C738CE3201" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,8</td>
|
||
<td id="B5EE5CF1D7782DB78973DC9C903CDDB6" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,8</td>
|
||
<td id="ABDEA3E7D64908F59D323FDFCC0B1446" colspan="1" pageId="0" pageNumber="159" rowspan="1">17</td>
|
||
<td id="CCFD5F65F3E69C24753343395E49AED1" colspan="1" pageId="0" pageNumber="159" rowspan="1">?</td>
|
||
<td id="E225007D687C294F5EFED594B5B71E1F" colspan="1" pageId="0" pageNumber="159" rowspan="1">unbranched</td>
|
||
<td id="67A84257FC7B19DA18E9605122989C8C" colspan="1" pageId="0" pageNumber="159" rowspan="1">?</td>
|
||
<td id="D5B2A43A9BD2C6B6A7C069BB0ECD5ABE" colspan="1" pageId="0" pageNumber="159" rowspan="1">28</td>
|
||
<td id="61637B7E7B6127A709A6E68CE18B53D0" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="AA69664E32A5EF746C8651A9F40B99E6" pageId="0" pageNumber="159">
|
||
<td id="B81F4693CC8BC30D31D8A379720C5866" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="728B368B11338EF5B93D68E1364FF6B3" authorityName="Gilmore, Van Tassell & Tornabenean Tassell" authorityYear="2016" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus adamsi" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="adamsi">
|
||
<emphasis id="4084D2825BED2A09D4874C77DE047D34" italics="true" pageId="0" pageNumber="159">Varicus adamsi</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="32A34D5A05E572D9CC299B25C3314E10" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="8861EE50932EE72423A4FE844A541CB6" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,7-8</td>
|
||
<td id="E1A9C2620561ADF8140FEFC3918BD035" colspan="1" pageId="0" pageNumber="159" rowspan="1">18</td>
|
||
<td id="5D02B2974484EE7C7DB039CD3EAB369C" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="6C10EEEFDC22EDFEE69912FEA821F06A" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched - fleshy pads</td>
|
||
<td id="062E80B9EA2DE9C559A18B8B23F7CF97" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="CA88516FE9A9572B037C7735131EE5DB" colspan="1" pageId="0" pageNumber="159" rowspan="1">21-24</td>
|
||
<td id="DBB2A83A9B5C0B0CAE8676ACB2B08272" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="B60566BFB7C119CFE93D2F477A2273C0" pageId="0" pageNumber="159">
|
||
<td id="615E1BDE66F259B1768731F9F2AE3D8D" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="5B37986B54D102F8377F82AB92737E27" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus benthonis" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="benthonis">
|
||
<emphasis id="25EE5978B4445118A2388C8C815C98CD" italics="true" pageId="0" pageNumber="159">Varicus benthonis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="AB7FC0F2780F871438DB9B18B8D7103A" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,8</td>
|
||
<td id="44639C34B283B847FBD12EE12FE721BA" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,7</td>
|
||
<td id="86DB1B31BDC1A6AF600ADECF02856667" colspan="1" pageId="0" pageNumber="159" rowspan="1">16</td>
|
||
<td id="E419349AFC5E4F3D0A786C7B40962092" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="63D7036983826FC1E7BBA34B0B92694E" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="380430305DF9FCB5F3A24F03652A49DF" colspan="1" pageId="0" pageNumber="159" rowspan="1">separate</td>
|
||
<td id="9032473B9255F30647CACFC7681F99E1" colspan="1" pageId="0" pageNumber="159" rowspan="1">12</td>
|
||
<td id="B4532D4567A9826A278C9766468095E4" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="158E9AC27DC2CD1D9A98B618D5C22669" pageId="0" pageNumber="159">
|
||
<td id="5332C40B4EF6820A2AF47F0C467D1B6E" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="DBE663D80AB17EA5156BB4A752E5ADB7" authorityName="Robins & Bohlke" authorityYear="1961" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus bucca" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="bucca">
|
||
<emphasis id="29151AD69AE266BB8641376AE2B2350A" italics="true" pageId="0" pageNumber="159">Varicus bucca</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="78FBAD76371C4F2A5F9B3C789082E7BA" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="07D1E25D2D72308D384A3ADEC8E5EEC3" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,7-8</td>
|
||
<td id="5EAC1DF5D85B7E1FB54E6530B3947258" colspan="1" pageId="0" pageNumber="159" rowspan="1">16-19</td>
|
||
<td id="76B8A569C0FB8F9F8FEED8DDCFF4F2A1" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="1ED4C371EC040B9B1B76080BEFB74958" colspan="1" pageId="0" pageNumber="159" rowspan="1">unbranched or branched - fleshy pads</td>
|
||
<td id="70F3C014C5ED8FF8D85FE0644093B8E4" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="4494A6A76E60BD3814AFAA64730FF6A6" colspan="1" pageId="0" pageNumber="159" rowspan="1">27</td>
|
||
<td id="83445B2DB7F33C83851FB49C4B1EF7B9" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="1B8FA6B9FA8A88D1CE0C2875048D3B8B" pageId="0" pageNumber="159">
|
||
<td id="CDD9907587F83BC72651B86B7A738C0B" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="4356DF53A00DCB35F1B85E799F5A530A" authorityName="Gilmore, Van Tassell & Baldwinan Tassell" authorityYear="2016" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus cephalocellatus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="cephalocellatus">
|
||
<emphasis id="35EF8F6F2BC25F0B996F31CCEE24AF53" italics="true" pageId="0" pageNumber="159">Varicus cephalocellatus</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="19E36FCC06A4D278E65244AA39D6BFA5" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,10</td>
|
||
<td id="28C7247A3692B57C8BBDFB3E204AE8DC" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="61ADECAB2E68F87AA9611B8BE6F562C1" colspan="1" pageId="0" pageNumber="159" rowspan="1">19-20</td>
|
||
<td id="97357D389F8595281E09DAD818E7BE4B" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="F4CD5223B5731729007FEE0E3AB06041" colspan="1" pageId="0" pageNumber="159" rowspan="1">unbranched</td>
|
||
<td id="AC730A60C6265C17178786C4F08DEF46" colspan="1" pageId="0" pageNumber="159" rowspan="1">variable</td>
|
||
<td id="8BCF9C2F72A58E04221ADCF1B98CD497" colspan="1" pageId="0" pageNumber="159" rowspan="1">12-23</td>
|
||
<td id="DB3652B706A7AC09B0FF401C1DFA8A32" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="F04F1BFE5F0814A58E30653BFC82555B" pageId="0" pageNumber="159">
|
||
<td id="8D84CC6300246E0911B46899D6C138A9" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="E0A38A1B0D4ED08FB80556DF7037EDA2" authorityName="Van Tassell, Baldwin & Tornabenean Tassell" authorityYear="2016" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus decorum" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="decorum">
|
||
<emphasis id="7E65A3354E7FF8A94EDBC5689B455D10" italics="true" pageId="0" pageNumber="159">Varicus decorum</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="297AD630A0885929C36213A57B447C86" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="24394015095E9916C7210297CD75F425" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,7-8</td>
|
||
<td id="F53334BCA37A80C40542643A080053D1" colspan="1" pageId="0" pageNumber="159" rowspan="1">17</td>
|
||
<td id="BAC5925BC56E7A5E036AB2A0D42C10CA" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="EF5460BC03DAC82C380449DFBD1F22BE" colspan="1" pageId="0" pageNumber="159" rowspan="1">unbranched</td>
|
||
<td id="FC19049806B00EEC36435F61F9D9E807" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="8A883ECA14F2835F2A5AD2F8C92BD526" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="146C77B93F25EA15DCAEDC156F4CBD9D" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="6A04DD6539B1A8D7DDD9BFBB6220589D" pageId="0" pageNumber="159">
|
||
<td id="C49868DF09EF012910B0EE231650C045" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="DFFC380A65CE7571669F569F7521E56F" authorityName="Tornabene, Robertson & Baldwin" authorityYear="2016" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus lacerta" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="lacerta">
|
||
<emphasis id="836CF635A1982BD482B5211C2CC10E4E" italics="true" pageId="0" pageNumber="159">Varicus lacerta</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="C36C1F2952CC76017CCCC99C75CE0D39" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="587C2D65E79733AD82D8E4C289D4C7A4" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,7</td>
|
||
<td id="F93A8F68AD250682E1440E31697BD5AA" colspan="1" pageId="0" pageNumber="159" rowspan="1">18</td>
|
||
<td id="7F764F11550F9C03BCA434D95CD493FC" colspan="1" pageId="0" pageNumber="159" rowspan="1">s</td>
|
||
<td id="D4934784FBF1950A01273EEAE5FEC595" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched - feathery</td>
|
||
<td id="EC02C18F4A7DDB3847B11D7CDC63338A" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="009C784F884E5350084E400B5BDA317E" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="528D35E342021713661691B668E00874" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr id="0A78AE1AFD86616E7BFEC45958570DFB" pageId="0" pageNumber="159">
|
||
<td id="F5F33B7738AA2007D52A4FED9E9E39D8" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="EC49414CB7BA9014747F519EB8EAD62A" authorityName="Gilmore" authorityYear="1979" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus marilynae" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="marilynae">
|
||
<emphasis id="889AA8F29749AD84A6C936C7A6E060A8" italics="true" pageId="0" pageNumber="159">Varicus marilynae</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="9C3C05DD290376356F0E6C4E59A7BE15" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,8</td>
|
||
<td id="0ECB760400BBA04D92024E60BDE107CC" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,7</td>
|
||
<td id="8FBD10997332CA691D91288B11BD0E1F" colspan="1" pageId="0" pageNumber="159" rowspan="1">16-18</td>
|
||
<td id="D61BFE37D171F1D2A2C29BDD3A4AAF0A" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="308DAE1A25264D5FA635A83CA1540CFC" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched - fleshy pads</td>
|
||
<td id="C0210B170A763CBBF9907C5FF554DD65" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="E7551E426BC35F5D9C4624C00424D679" colspan="1" pageId="0" pageNumber="159" rowspan="1">18-19</td>
|
||
<td id="88621E651A63193825618AEB03399144" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="4524D1EDC8B4BB77833B40C67553E991" pageId="0" pageNumber="159">
|
||
<td id="321396E55B203FDCD159A5B522F0E152" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="D8C94D03B6FC465C4A77B5122DC43E71" authorityName="Gilmore, Van Tassell & Baldwinan Tassell" authorityYear="2016" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus nigritus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="nigritus">
|
||
<emphasis id="552C5B2BDEF1058B1FBE0DBE47DD5178" italics="true" pageId="0" pageNumber="159">Varicus nigritus</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="312002CF3C51363FAEB3A2F29697BD49" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="96B388833E87FA9E3300638EB075F6DD" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,8</td>
|
||
<td id="C07DEB1BE371D90F8A6D5DB29FAAE4B4" colspan="1" pageId="0" pageNumber="159" rowspan="1">17</td>
|
||
<td id="A330F5DDDC1D51C10BF3B12B89B4B5E0" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="3FD56A29CC366989EE9059F22CB93F33" colspan="1" pageId="0" pageNumber="159" rowspan="1">unbranched</td>
|
||
<td id="CD961A900F944FD3081C5A85BD8419E5" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="3609A1FCC6C8C7F44A99FD5BEC3DFCDC" colspan="1" pageId="0" pageNumber="159" rowspan="1">present (exact count unknown)</td>
|
||
<td id="7C97A5801F66D77F57000643740A1954" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="70BEE5F505C31B53781C26E32719E7C9" pageId="0" pageNumber="159">
|
||
<td id="7A4044918F55CD917AFB8BAD5DCEBE63" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="D5A37DCDEE457C8C8293C1E38CD421A6" authorityName="Van Tassell, Baldwin & Gilmorean Tassell" authorityYear="2016" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus veliguttatus" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="veliguttatus">
|
||
<emphasis id="65824402760EF6BA25FDE0F00D0CBE4B" italics="true" pageId="0" pageNumber="159">Varicus veliguttatus</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="B25527FB05039ABE5798DFD11342FF93" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,8</td>
|
||
<td id="76872FEA31953D4E30341CB54370C9F6" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,6-7</td>
|
||
<td id="5C445135541C32A853216D36937C27E9" colspan="1" pageId="0" pageNumber="159" rowspan="1">17-19</td>
|
||
<td id="99A6A6BC40017CC8D178C5F4F1A8765C" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="DEB97B7CC15D5041E84B7240ECADE96E" colspan="1" pageId="0" pageNumber="159" rowspan="1">unbranched</td>
|
||
<td id="72E7375AC7175B9DCC24B4E5CF314A65" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="B031FF456E3AFEDC26EA311572E7ACD5" colspan="1" pageId="0" pageNumber="159" rowspan="1">23-34</td>
|
||
<td id="125D06EE7CA4502594F07BF2E1007712" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="073A58A5C25F9AFD9909214412F5AE73" pageId="0" pageNumber="159">
|
||
<td id="4CA42603AA489F96D2C45843CBDF5400" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="F2680728711D317AE0B57A2534166B97" family="Gobiidae" genus="Varicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Varicus vespa" order="Perciformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="vespa">
|
||
<emphasis id="F0088A82FBD53806BD559FA18FD6C294" italics="true" pageId="0" pageNumber="159">Varicus vespa</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="8AE13F57C8B52C515CE80E7E3638F423" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="F3B0B618990C1FFF63EE988468B0F1F4" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,7 (rarely I,6 or I,8)</td>
|
||
<td id="C290F8A88FCADEBF7EA69B408663DAC1" colspan="1" pageId="0" pageNumber="159" rowspan="1">15-17</td>
|
||
<td id="52B3D791847414296DB731CE0B2D324B" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="DFE14C7C0D13912D9C7E512A8DC665F5" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched - fleshy pads</td>
|
||
<td id="723442314BCF6D9ED64EF9605E172762" colspan="1" pageId="0" pageNumber="159" rowspan="1">separate</td>
|
||
<td id="DFF3D0452CE997A81F2A190BB12DDBBD" colspan="1" pageId="0" pageNumber="159" rowspan="1">9-14</td>
|
||
<td id="59151840DCC129AC92292B05C1DE2F46" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="1743283CD6B19F6082F5F05C3AC16125" pageId="0" pageNumber="159">
|
||
<td id="9C5505336EDC8B43A2FB0B049311A7B5" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="F18D72C14A16E74A375ED8C2ED9024E5" authorityName="Ginsburg" authorityYear="1953" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris alepis" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="alepis">
|
||
<emphasis id="609F45912251BDB69650A7D4C58F21AC" italics="true" pageId="0" pageNumber="159">Psilotris alepis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="A8D312BD801295812CC71DB4CBE1B6D8" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9 (rarely I,8)</td>
|
||
<td id="76067804406493BC52909A0BBA66E712" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,7-8</td>
|
||
<td id="6CC3E91165F76AF498FC2CA91BFF1996" colspan="1" pageId="0" pageNumber="159" rowspan="1">15</td>
|
||
<td id="E58A9364833BD64125934E8E6B1D5524" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="6D5C9E0C7666538181460DD2FCB15290" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="B06674675F7525E9CA9667550CD61F50" colspan="1" pageId="0" pageNumber="159" rowspan="1">separate</td>
|
||
<td id="A01FADE14ABA552FB94735CF33EE1323" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="25DB101031189D5E8B4785A99CF5FC03" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr id="0A21CE9B3849F3A57DBBE74BF2FFD7DE" pageId="0" pageNumber="159">
|
||
<td id="B8555AB94EC6BCB9284E2BAA011A2FB0" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="AFD51CA4200877AAABAD4A3BCADEC734" authorityName="Greenfield" authorityYear="1993" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris boehlkei" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="boehlkei">
|
||
<emphasis id="2D6ECA4358231194EC3BD1536D152C0C" italics="true" pageId="0" pageNumber="159">Psilotris boehlkei</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="E8ED0433CF2701A3BAB2FBD4143AAFAF" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9-10</td>
|
||
<td id="8911F2EB2E3DBCE9CDE4A9F8E1B34650" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="61B37BB11B2893C6D83446BDB8A54527" colspan="1" pageId="0" pageNumber="159" rowspan="1">16-18</td>
|
||
<td id="820C91A4BCB5C1D7EFE92841B6CD7488" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="572DC5CAF5E0779BBC8380780007663C" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="9C43DA05CF1C12CDEEFB755C4AA9393D" colspan="1" pageId="0" pageNumber="159" rowspan="1">separate</td>
|
||
<td id="D8A443794D5CB30936916A99743ED011" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="3A12891FD172DA5C9F6333A308E45066" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr id="211C818113A66992032B1F4BF4C126EC" pageId="0" pageNumber="159">
|
||
<td id="E48810EAB098B04817D9104D4C47769B" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="A2B3AD51CD65F2970B919D9580134016" authorityName="Bohlke" authorityYear="1963" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris celsa" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="celsa">
|
||
<emphasis id="282D2F8B05C8791DB2E520593549CED4" italics="true" pageId="0" pageNumber="159">Psilotris celsa</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="48C75D99EB449BC8C399DD118362CE83" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9-10 (rarely I,8)</td>
|
||
<td id="7E3E91585F4C89C5D4A891FCB6A04F9A" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9-10 (rarely I,8)</td>
|
||
<td id="F4BEB393539516555037C997477A62F1" colspan="1" pageId="0" pageNumber="159" rowspan="1">16-17</td>
|
||
<td id="D16F1F38650EFEF20E49584B6F146A8A" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="724225B970082AAB8A4338117EF45515" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="C0844DD1FDB63B0D2062731AA4790A2C" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="5072BD0B0D956A0D2FD5552AD99A09A1" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="39E348576EDC40268425A1E61EB9280C" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr id="4D4A2E2687453DDD53D538CC166FA719" pageId="0" pageNumber="159">
|
||
<td id="A79B8AD3F5D9EC56356141B381A88432" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="309E98B3E4A066A02932E46EE8E771C7" authorityName="Greenfield, Findley & Johnson" authorityYear="1993" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris kaufmani" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="kaufmani">
|
||
<emphasis id="AEE62E5FF984E8BEA710B6EBE760BC8F" italics="true" pageId="0" pageNumber="159">Psilotris kaufmani</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="C269FEC0897CACAB5EA1E2320003D20B" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,10 (rarely I,9)</td>
|
||
<td id="F313BF188857445E214D2956446B13B6" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,10 (rarely I,9)</td>
|
||
<td id="734E55215376EA2F0837FC299B67E1CB" colspan="1" pageId="0" pageNumber="159" rowspan="1">16-19</td>
|
||
<td id="1FD0CF63658FAC3489E2C2B05C9E98B5" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="6AE104485272EB5759E2F28FBFB5B195" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="43D4FE348FA6B58C20ADDE49B43F8C40" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="B7D976B67C1AAED9FD27194150F48DCA" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="F5A88CAC94840FB448075650C759B158" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr id="44D8D7411B51BF962C7CC1287659ED16" pageId="0" pageNumber="159">
|
||
<td id="10E3761ABC38DAD4ECC4F4C9CB8559A2" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="E65AE8FF5CABB1B16E8CF927A344409E" authorityName="Van Tassell & Youngan Tassell" authorityYear="2016" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris laetarii" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="laetarii">
|
||
<emphasis id="5F24C52CDCCE030B1494DBC52000DE40" italics="true" pageId="0" pageNumber="159">Psilotris laetarii</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="F44D50D830AC16A62DA87706AA87491A" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9-10</td>
|
||
<td id="BE470B0B9A4DBF1D1AEBA9781D23E0F4" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,7-8</td>
|
||
<td id="A3C76411AFD6622404D820E635DD31C3" colspan="1" pageId="0" pageNumber="159" rowspan="1">15-17</td>
|
||
<td id="91C57B9BCCF8EFBA07A22C73C89DA2DB" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="AD8E2BC5EE653E6874E46EC128E7FA09" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="59D0C20872756D91710821D599B6E69E" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="4E5314ABB10529567C6636BEC38A237E" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="C952EFEAB1B4720415CA7E888F4CA176" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr id="F8C33C6F50DAF981EF4A9402221D5F33" pageId="0" pageNumber="159">
|
||
<td id="79282EBF14CF6128A199D03D7B3010A7" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="CC32DC71E085B9631C3EA243EA1A7C53" authorityName="Van Tassell, Tornabene & Baldwinan Tassell" authorityYear="2016" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris laurae" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="laurae">
|
||
<emphasis id="2E9B45BFCE61A30465392DFA62367BB0" italics="true" pageId="0" pageNumber="159">Psilotris laurae</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="F8E6A6548A222EA9C071FF7278263D2B" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="06DB5DDC036CFAC31B6601DAA79DBDA4" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,8</td>
|
||
<td id="38FAEEBFCB07F3149A124A0F7E6DDE42" colspan="1" pageId="0" pageNumber="159" rowspan="1">18</td>
|
||
<td id="E8DC593B69F8EA7BA684E226ECC030A5" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="495AF828AC14927C4278AEE150AF4154" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="E7E9C44ABB8A727F2B68B625276F751F" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="C6CB6A7420DBE4430971416BF89BA0FC" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="14C31F5CE7DFF33A2C227101EBCA4123" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr id="CE06F1F8E3AD5C85D3A9CF4885EB5853" pageId="0" pageNumber="159">
|
||
<td id="816DBBCAAC7A15A8A80D754381286C73" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="5E291CF47CDDA9C8E68C89E11B9333F9" authorityName="Tornabene & Baldwin" authorityYear="2019" class="Actinopterygii" family="Gobiidae" genus="Psilotris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psilotris vantasselli" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="vantasselli">
|
||
<emphasis id="0C42B48B7BD7B054CB485C2D8F3EC9FF" italics="true" pageId="0" pageNumber="159">Psilotris vantasselli</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="108C6E208BD13137FD8FAA242CCAB527" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,9</td>
|
||
<td id="88218CA4E87C64FC3DCDE1EE52B8FF4F" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,8</td>
|
||
<td id="9D004DE4B79D9FCD97BB04A2B6331973" colspan="1" pageId="0" pageNumber="159" rowspan="1">15</td>
|
||
<td id="9A0A2C1A8AB46D2CDB1FE59C458B84E6" colspan="1" pageId="0" pageNumber="159" rowspan="1">1</td>
|
||
<td id="EEB8BA53716E11B97AB902BC64AF21EF" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="E14E4DA818FF72168E3CD9892F249EC2" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="4D4CD8AD1221B3C9BB0E9162ACEA6578" colspan="1" pageId="0" pageNumber="159" rowspan="1">absent</td>
|
||
<td id="101AF35F17BFD13C7C9617231C2A813E" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="3D3D500599B7AA16EFE230D92E8E0728" pageId="0" pageNumber="159">
|
||
<td id="52A6ABD42041319219E982703E8DEB5A" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="EA1E98A7A6CD19CAE3998059959143C5" authorityName="Van Tassell & Tornabene" authorityYear="2016" family="Gobiidae" genus="Pinnichthys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pinnichthys aimoriensis" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="aimoriensis">
|
||
<emphasis id="6CE0A05055DA3C9B9AC6F6760E20CF81" italics="true" pageId="0" pageNumber="159">Pinnichthys aimoriensis</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="00E6E14B971AD264A3F67FF57EBE7F77" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,10-11</td>
|
||
<td id="45B8CE63D1200E6427E37036D8F94DB0" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,10</td>
|
||
<td id="B68A3FB75D0A48E023D3B0F4321CE7ED" colspan="1" pageId="0" pageNumber="159" rowspan="1">18-19</td>
|
||
<td id="391DDD0A1C628DB07B5A59B7B1FAF9BA" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="6DB097A4D77D6083DC9F708935D6EA33" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="1CB9B302CF8E3C3E2D5457C6987649DB" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="D1330815AE8FD20E378A431DEB1AE4F8" colspan="1" pageId="0" pageNumber="159" rowspan="1">40-47</td>
|
||
<td id="7EEFACD4E886BCD3A494779512A4A614" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="BBBAE02BBAFD9DA42F5CF4CDA95578AE" pageId="0" pageNumber="159">
|
||
<td id="61695877F8DB9E89FAD407C7DAEFA836" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="56E7424BD6A133FDFDCEC0FFD38B2B98" baseAuthorityName="Hastings & Findley" baseAuthorityYear="2013" family="Gobiidae" genus="Pinnichthys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pinnichthys bilix" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="bilix">
|
||
<emphasis id="660FD92DB9EB28BC246E31E42E21CBD2" italics="true" pageId="0" pageNumber="159">Pinnichthys bilix</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="4CC21207E729A7DC385106DB6F3BF256" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,11</td>
|
||
<td id="327DDB668238771E81EBE580FF7CA07E" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,10-11</td>
|
||
<td id="8BA88571F8C35F8955799FE6A6F55BA2" colspan="1" pageId="0" pageNumber="159" rowspan="1">19-20</td>
|
||
<td id="7C10FB38A5F97C026D6123628B80B05A" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="56984CF8095B883A39A3535A72FE30DF" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="A73BAFC280836F4DFB0017CD2CEAFD12" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="F6C8290C4AD0B0CAB4071E24EFD3357B" colspan="1" pageId="0" pageNumber="159" rowspan="1">30-35</td>
|
||
<td id="058D5F7FC42BEC5233B6D7AD0F5C5875" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="111030678D2C140F88A28B648CCB7CF5" pageId="0" pageNumber="159">
|
||
<td id="388557A50842BE9C5EC4A2E489679AA3" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="9759409631B942FF5D7E300FF8B07D4A" baseAuthorityName="Hastings & Findley" baseAuthorityYear="2015" family="Gobiidae" genus="Pinnichthys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pinnichthys prolata" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="prolata">
|
||
<emphasis id="3AB3AE112CBBD12B2558B4AB8F099337" italics="true" pageId="0" pageNumber="159">Pinnichthys prolata</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="A7638316E9EA6BE8F0FC0F9422C09168" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,10-11</td>
|
||
<td id="8D6DBD93015BD0EA0F9583457B3306E6" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,10-11</td>
|
||
<td id="E508EA2D5702BA8CE3994BC592568D15" colspan="1" pageId="0" pageNumber="159" rowspan="1">17-19</td>
|
||
<td id="595CEF6DD8F235A5ACC3F6AE215303C5" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="A03B7C0C67C18CF9EA18463ADD6140EC" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="D6F6B31932D7ABD79C5D668E444CFC1A" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="53521E71BFD63EE6F184396524CCCC6A" colspan="1" pageId="0" pageNumber="159" rowspan="1">34-37</td>
|
||
<td id="2315D03B03C49EBF723C2B05D8D02B94" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="7C1232766DBBB021060EBD96F5445523" pageId="0" pageNumber="159">
|
||
<td id="0DA121DC2457CDC585A8BA4A680F04EA" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="4C4A5363FE7F405E7CB6C8B4316B71BE" authorityName="Gilmore, Van Tassell & Tornabenean Tassell" authorityYear="2016" family="Gobiidae" genus="Pinnichthys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pinnichthys saurimimica" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="saurimimica">
|
||
<emphasis id="7BE397900019803E1E55C1EA9A378C45" italics="true" pageId="0" pageNumber="159">Pinnichthys saurimimica</emphasis>
|
||
</taxonomicName>
|
||
</td>
|
||
<td id="4645AE9C5E8A587C73BF84E7C44BC794" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,11</td>
|
||
<td id="F7809892FC60D5BD9D339C2DE77D47DD" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,11</td>
|
||
<td id="549CDA3B5E148C207A4E43CB0B5E41CC" colspan="1" pageId="0" pageNumber="159" rowspan="1">20</td>
|
||
<td id="0440E83DB326F6A098CE470E420E8D22" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="766C6A444C1E6D5BFD39B2E681E40BA0" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="A0D1E72D5A4AEEFDBBD535F8BC58EF3A" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="D69C3AE3F0C521194A967430FD99DB3E" colspan="1" pageId="0" pageNumber="159" rowspan="1">47-53</td>
|
||
<td id="139F4E1FC6F16E831AC2C82FBC8EE69A" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
<tr id="4A438E95659F3B0674301748003B59BF" pageId="0" pageNumber="159">
|
||
<td id="22BD344A799EFBF71F408ED7E4C61EC2" colspan="1" pageId="0" pageNumber="159" rowspan="1">
|
||
<taxonomicName id="588675CCDD6B44F8D842B23A973673B4" family="Gobiidae" genus="Pinnichthys" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Pinnichthys atrimela" order="Gobiiformes" pageId="0" pageNumber="159" phylum="Chordata" rank="species" species="atrimela">
|
||
<emphasis id="21487721FF74A58C26055C2757943EC2" italics="true" pageId="0" pageNumber="159">Pinnichthys atrimela</emphasis>
|
||
</taxonomicName>
|
||
(Pacific)
|
||
</td>
|
||
<td id="F4532349549519447832538E3E552425" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,11</td>
|
||
<td id="8573F0E0100091E13411A5B6F914AA26" colspan="1" pageId="0" pageNumber="159" rowspan="1">I,10</td>
|
||
<td id="42C4F1C6C73EB7DD89F7A14486466F0F" colspan="1" pageId="0" pageNumber="159" rowspan="1">20</td>
|
||
<td id="C9F9C60C6CB386F2BF15D1BDBEA8C8E6" colspan="1" pageId="0" pageNumber="159" rowspan="1">2</td>
|
||
<td id="941C57D4864E8B41B7EC9B3042318FDF" colspan="1" pageId="0" pageNumber="159" rowspan="1">branched</td>
|
||
<td id="F417F916D7A0F2484CDDC715E6BEE489" colspan="1" pageId="0" pageNumber="159" rowspan="1">connected</td>
|
||
<td id="84F27768D8719C9124D31ACCC156FA7A" colspan="1" pageId="0" pageNumber="159" rowspan="1">41</td>
|
||
<td id="1D464D833167129ABA1A9993C345210F" colspan="1" pageId="0" pageNumber="159" rowspan="1">present</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |