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<document id="D8434C59D99F3F3D87E05E7C5107BA40" ID-DOI="10.5281/zenodo.4504184" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504184" ID-ZooBank="F3A75556-83C3-47A9-97CE-78342127CAE5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612465776621" checkinUser="carolina" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="68722F5E58622E4DFC5C72D7689AFF0C" docLanguage="en" docName="RafflesBZool.62.83-88.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Horaeomorphus solodovnikovi Jałoszyński 2014, new species" docType="treatment" docUuid="7EEAB5E9-791C-49C7-B79F-60ACC07D81A7" docUuidSource="ZooBank" docVersion="5" lastPageNumber="85" masterDocId="944B572658622E4FFFD375406C2EFFF9" masterDocTitle="Oriental Horaeomorphus Schaufuss: a new species, synonymic notes and distributional synopsis (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="88" masterPageNumber="83" pageNumber="83" updateTime="1698866066355" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="3F04C3B5E943DAA2FBBF986BB8993688" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6F6F5B771C3ECD23940CA7F319C626B9">
<mods:title id="A84F9A149516FE3764AA670BC69F111F">Oriental Horaeomorphus Schaufuss: a new species, synonymic notes and distributional synopsis (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
</mods:titleInfo>
<mods:name id="3A5165D67AC2F920ACCF5467E95813C4" type="personal">
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<mods:namePart id="FA1EFF39ED3493A6F203585BDF9046D8">Jałoszyński, Paweł</mods:namePart>
</mods:name>
<mods:typeOfResource id="1A1A20F99E77F8C5CFCDDF2C879DD277">text</mods:typeOfResource>
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<mods:title id="17A8B1E0AB38EEA62DAFC013BB3F4BDF">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="F89BE698013890027D83AE6C36D67BD9">
<mods:date id="93CD1BA96612DD59A04B687D3C214F00">2014</mods:date>
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<mods:number id="EFCEAD90034796C0023A689C701E8F90">2014-03-07</mods:number>
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<mods:detail id="FFAC857F8A67EDD65CF11821DC3CB156" type="volume">
<mods:number id="30539C151388CD49D5F590FD9536BBEA">62</mods:number>
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<mods:start id="32A5816E0E994A2A8362B17B703E1998">83</mods:start>
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<mods:classification id="02536780CA00DE5703498CE949EF39ED">journal article</mods:classification>
<mods:identifier id="FDC6EE55EA97C333E5F7F9523B393CF5" type="DOI">10.5281/zenodo.4504184</mods:identifier>
<mods:identifier id="1C0689004210D9549741522A8F894CA4" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="84C6951FDA2DDCDE3A44A44CFD7E56C0" type="Zenodo-Dep">4504184</mods:identifier>
<mods:identifier id="528BDAB7F23EFEB9930EBB9F240B9988" type="ZooBank">F3A75556-83C3-47A9-97CE-78342127CAE5</mods:identifier>
</mods:mods>
<treatment id="68722F5E58622E4DFC5C72D7689AFF0C" LSID="urn:lsid:zoobank.org:act:7EEAB5E9-791C-49C7-B79F-60ACC07D81A7" httpUri="http://treatment.plazi.org/id/68722F5E58622E4DFC5C72D7689AFF0C" lastPageId="2" lastPageNumber="85" pageId="0" pageNumber="83">
<subSubSection id="A8C1CDC358622E4FFC5C72D76959F849" box="[911,1399,1943,1968]" pageId="0" pageNumber="83" type="nomenclature">
<paragraph id="E0649E4858622E4FFC5C72D76959F849" blockId="0.[911,1399,1943,2000]" box="[911,1399,1943,1968]" pageId="0" pageNumber="83">
<heading id="BB2C292458622E4FFC5C72D76959F849" bold="true" box="[911,1399,1943,1968]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="83" reason="2">
<emphasis id="D2AF425A58622E4FFC5C72D76959F849" bold="true" box="[911,1399,1943,1968]" pageId="0" pageNumber="83">
<taxonomicName id="27DBE5CB58622E4FFC5C72D768CDF856" ID-CoL="3MNLN" authority="Jałoszyński, 2014" authorityName="Jałoszyński" authorityYear="2014" box="[911,1251,1943,1967]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="solodovnikovi" status="sp. nov.">
<emphasis id="D2AF425A58622E4FFC5C72D768CDF856" bold="true" box="[911,1251,1943,1967]" italics="true" pageId="0" pageNumber="83">Horaeomorphus solodovnikovi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C99CFF2158622E4FFB2172D96959F849" box="[1266,1399,1945,1968]" pageId="0" pageNumber="83" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358622E4FFBC772F968DFF829" box="[1044,1265,1977,2000]" pageId="0" pageNumber="83" type="description">
<paragraph id="E0649E4858622E4FFBC772F968DFF829" blockId="0.[911,1399,1943,2000]" box="[1044,1265,1977,2000]" pageId="0" pageNumber="83">
(
<figureCitation id="78E082CD58622E4FFBCF72F9684CF829" box="[1052,1122,1977,2000]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (A–C); and Horaeomorphus wailimae (Lhoste), new combination, (D–G). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="0" pageNumber="83">Figs. 1</figureCitation>
A–C, 2A, B)
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358622E4EFC9072B76EC9FEBF" lastPageId="1" lastPageNumber="84" pageId="0" pageNumber="83" type="materials_examined">
<paragraph id="E0649E4858622E4FFC9072B769ECF7D0" blockId="0.[835,1475,2039,2089]" pageId="0" pageNumber="83">
<emphasis id="D2AF425A58622E4FFC9072B7682BF7F5" bold="true" box="[835,1029,2039,2060]" pageId="0" pageNumber="83">Material examined.</emphasis>
<materialsCitation id="50B3941558622E4FFBDA72B769ECF7D0" country="Laos" location="Muang Paxong" municipality="Male" pageId="0" pageNumber="83" specimenCount="1" stateProvince="Champasak" typeStatus="holotype">
<typeStatus id="3F6020EA58622E4FFBDA72B7684DF7F5" box="[1033,1123,2039,2060]" pageId="0" pageNumber="83" type="holotype">Holotype</typeStatus>
.
<collectingMunicipality id="0000043258622E4FFBB872B768B5F7F5" box="[1131,1179,2039,2060]" pageId="0" pageNumber="83">Male</collectingMunicipality>
(
<figureCitation id="78E082CD58622E4FFB7572B768C0F7F5" box="[1190,1262,2039,2060]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (A–C); and Horaeomorphus wailimae (Lhoste), new combination, (D–G). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="0" pageNumber="83">Fig. 1A</figureCitation>
), three labels: “
<collectingCountry id="98CCDED858622E4FFAAE72B76991F7F5" box="[1405,1471,2039,2060]" name="Laos" pageId="0" pageNumber="83">LAOS</collectingCountry>
:
<collectingRegion id="221F50AA58622E4FFC907D546FDDF7D0" box="[835,1011,2068,2089]" country="Laos" name="Champasak" pageId="0" pageNumber="83">Champasak prov.</collectingRegion>
: / Bolaven Plt. [i.e., Plateau],
<location id="E504C89358622E4FFAF57D546990F7D0" LSID="urn:lsid:plazi:treatment:68722F5E58622E4DFC5C72D7689AFF0C:E504C89358622E4FFAF57D546990F7D0" box="[1318,1470,2068,2089]" country="Laos" municipality="Male" name="Muang Paxong" pageId="0" pageNumber="83" stateProvince="Champasak">Muang Paxong</location>
,
</materialsCitation>
</paragraph>
<paragraph id="E0649E4858632E4EFFA275FE6EC9FEBF" blockId="1.[113,753,190,326]" pageId="1" pageNumber="84">
/
<materialsCitation id="50B3941558632E4EFF5175FE6ECAFEBF" collectingDate="2008-06-14" collectionCode="ZMUC" collectorName="A. Solodovnikov &amp; Pedersen &amp; Disturbed" elevation="1100" latitude="15.05285" location="Ban Hoyayteuy" longLatPrecision="1" longitude="106.28995" municipality="Mt. Phu Din" pageId="1" pageNumber="84" specimenCount="1" typeStatus="holotype">
<location id="E504C89358632E4EFF5175FE6D0EFF2A" LSID="urn:lsid:plazi:treatment:68722F5E58622E4DFC5C72D7689AFF0C:E504C89358632E4EFF5175FE6D0EFF2A" box="[130,288,190,211]" latitude="15.05285" longLatPrecision="1" longitude="106.28995" municipality="Mt. Phu Din" name="Ban Hoyayteuy" pageId="1" pageNumber="84">Ban Hoyayteuy</location>
,
<collectingMunicipality id="0000043258632E4EFEFE75FE6D81FF2A" box="[301,431,190,211]" pageId="1" pageNumber="84">Mt. Phu Din</collectingMunicipality>
/
<date id="9465B88858632E4EFE1B75FE6E78FF2A" box="[456,598,190,211]" pageId="1" pageNumber="84" value="2008-06-14" valueMax="2008-06-14" valueMin="2008-06-14">
<collectingDate id="8421416058632E4EFE1B75FE6E78FF2A" box="[456,598,190,211]" pageId="1" pageNumber="84" value="2008-06-14" valueMax="2008-06-14" valueMin="2008-06-14">14-14.vi.2008</collectingDate>
</date>
;
<quantity id="272333AD58632E4EFDB775FE6E93FF2A" box="[612,701,190,211]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="84" unit="m" value="1100.0">
<elevation id="6BF6797B58632E4EFDB775FE6E93FF2A" box="[612,701,190,211]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="84" unit="m" value="1100.0">1,100 m</elevation>
</quantity>
., /
<geoCoordinate id="85EFF88F58632E4EFD0C75FE6CF4FF09" degrees="15" direction="north" minutes="03.171" orientation="latitude" pageId="1" pageNumber="84" precision="1" value="15.05285">N 15°03.171'</geoCoordinate>
<geoCoordinate id="85EFF88F58632E4EFF0B759B6D4AFF09" box="[216,356,219,240]" degrees="106" direction="east" minutes="17.397" orientation="longitude" pageId="1" pageNumber="84" precision="1" value="106.28995">E 106°17.397'</geoCoordinate>
” [white, printed], “
<collectorName id="4D2EFB9E58632E4EFDCD759B6EECFF09" box="[542,706,219,240]" pageId="1" pageNumber="84">A. Solodovnikov</collectorName>
&amp; J. /
<collectorName id="4D2EFB9E58632E4EFFAE75B86CFAFEF4" box="[125,212,248,269]" pageId="1" pageNumber="84">Pedersen</collectorName>
leg.
<collectorName id="4D2EFB9E58632E4EFED175B86D4FFEF4" box="[258,353,248,269]" pageId="1" pageNumber="84">Disturbed</collectorName>
/ primary rainforest. /
<collectionCode id="86CA068D58632E4EFDE575B86E55FEF4" box="[566,635,248,269]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="84">ZMUC</collectionCode>
collection.” [white, printed], “
<emphasis id="D2AF425A58632E4EFECE74546EABFED0" box="[285,645,276,297]" italics="true" pageId="1" pageNumber="84">
<taxonomicName id="27DBE5CB58632E4EFECE74546DC0FED0" authorityName="Schaufuss" authorityYear="1889" box="[285,494,276,297]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="genus">HORAEOMORPHUS</taxonomicName>
/
<taxonomicName id="27DBE5CB58632E4EFDD374546EABFED0" authorityName="Jałoszyński" authorityYear="2014" box="[512,645,276,297]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="solodovnikovi">solodovnikovi</taxonomicName>
</emphasis>
m. / det. P. JAŁOSZYŃSKI, 2013 /
<typeStatus id="3F6020EA58632E4EFEB774716DDAFEBF" box="[356,500,305,326]" pageId="1" pageNumber="84" type="holotype">HOLOTYPUS</typeStatus>
” [red, printed] (
<collectionCode id="86CA068D58632E4EFD4174716EF3FEBF" box="[658,733,305,326]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="1" pageNumber="84">ZMUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358632E4DFFA2742F6D74FDAC" lastPageId="2" lastPageNumber="85" pageId="1" pageNumber="84" type="diagnosis">
<paragraph id="E0649E4858632E4EFFA2742F6E73FD9F" blockId="1.[113,753,367,614]" pageId="1" pageNumber="84">
<emphasis id="D2AF425A58632E4EFFA2742F6CC7FE7F" bold="true" box="[113,233,367,390]" pageId="1" pageNumber="84">Diagnosis.</emphasis>
The following set of characters is unique for this species: body length below
<quantity id="272333AD58632E4EFE1474CF6E29FE5F" box="[455,519,399,422]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="84" unit="mm" value="3.0">3 mm</quantity>
, pigmentation brown, pronotum strongly elongate, with three small ante-basal pits located in deep groove; metatrochanters in males strongly modified, each forming very long, slightly recurved rod-like projection with rounded apex; and aedeagus with symmetrical internal armature containing long median tubular structure much longer than half length of median lobe.
</paragraph>
<paragraph id="E0649E4858632E4EFFA277CF6E86FD1F" blockId="1.[113,752,655,742]" pageId="1" pageNumber="84">
<emphasis id="D2AF425A58632E4EFFA277CF6C9CFD5F" bold="true" box="[113,178,655,678]" pageId="1" pageNumber="84">Male.</emphasis>
Body (
<figureCitation id="78E082CD58632E4EFED677CF6D74FD5F" box="[261,346,655,678]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (A–C); and Horaeomorphus wailimae (Lhoste), new combination, (D–G). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="1" pageNumber="84">Fig. 1A</figureCitation>
) moderately large (length
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), slender, strongly convex, dark brown with slightly lighter legs and palps, vestiture slightly lighter than cuticle.
</paragraph>
<paragraph id="E0649E4858632E4EFFA2764F6FC1FECC" blockId="1.[113,752,783,1062]" lastBlockId="1.[813,1452,190,309]" pageId="1" pageNumber="84">
Head broadest at moderately large and moderately strongly convex eyes, length
<quantity id="272333AD58632E4EFE8D766F6DE8FCBF" box="[350,454,815,838]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="84" unit="mm" value="0.4">0.40 mm</quantity>
, width
<quantity id="272333AD58632E4EFDCD766F6EA9FCBF" box="[542,647,815,838]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="84" unit="mm" value="0.6">0.60 mm</quantity>
; tempora rounded and about as long as eye in dorsal view; vertex strongly transverse and weakly convex, with pair of small but distinct pits located near posterior margins of supraantennal tubercles; frons weakly convex; supraantennal tubercles strongly raised, well delimited from frons but confluent with vertex. Punctures on vertex and frons fine and inconspicuous; setae moderately long, sparse, suberect to erect. Antennae slender, longer than half length of body, length
<quantity id="272333AD58632E4EFA9175FE6989FF2C" box="[1346,1447,190,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="84" unit="mm" value="1.5">1.50 mm</quantity>
; relative lengths of antennomeres (shortest antennomere II as 1): 1.43: 1.00: 1.71: 1.29: 1.29: 1.14: 1.14: 1.14: 1.43: 1.43: 2.00.
</paragraph>
<paragraph id="E0649E4858632E4EFCFE741E692FFD4C" blockId="1.[813,1452,350,693]" pageId="1" pageNumber="84">
Pronotum oval, elongate, broadest between anterior third and fourth, length
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, width at base
<quantity id="272333AD58632E4EFB03743E691DFE6C" box="[1232,1331,382,405]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="84" unit="mm" value="0.5">0.50 mm</quantity>
, maximum width
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; distal part strongly narrowing caudad to moderately sharply marked posterior constriction demarcating narrow basal collar; hind angles obtuse and blunt; posterior margin nearly straight; posterior collar delimited from disc by deep, narrow and slightly convex anteriorly transverse groove connecting three small pits: median pit as small as width of groove and lateral pits only slightly larger. Punctures on disc as fine as those on frons and vertex; vestiture composed of sparse, moderately long and suberect setae.
</paragraph>
<paragraph id="E0649E4858632E4EFCFE779E6FD7FBCC" blockId="1.[813,1452,734,1077]" pageId="1" pageNumber="84">
Elytra strongly elongate, oval, more convex than pronotum, broadest between middle and anterior third, length
<quantity id="272333AD58632E4EFAA877BE6F74FCCC" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="84" unit="mm" value="1.5">1.50 mm</quantity>
, width
<quantity id="272333AD58632E4EFC67765E6833FCCC" box="[948,1053,798,821]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="1" pageNumber="84" unit="mm" value="0.98">0.98 mm</quantity>
, elytral index 1.54. Humeral calli weakly marked, delimited from adsutural region by very short, shallow and broad basal impressions; basal elytral foveae indiscernible; median part of elytra without adsutural or circumsutural impressions; apices of elytra separately rounded. Punctures more distinct than those on pronotum, fine but sharply marked and separated by spaces 2–3× as wide as puncture diameters; vestiture similar to that on pronotum but slightly longer.
</paragraph>
<caption id="B4A4CEC058632E4EFFA272F96DDDF7B8" ID-DOI="http://doi.org/10.5281/zenodo.4504186" ID-Zenodo-Dep="4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="1" pageNumber="84" startId="1.[113,148,1977,1998]" targetBox="[120,1444,1159,1947]" targetPageId="1">
<paragraph id="E0649E4858632E4EFFA272F96DDDF7B8" blockId="1.[113,1452,1977,2113]" pageId="1" pageNumber="84">
Fig. 1.
<taxonomicName id="27DBE5CB58632E4EFF6672F96DF3F837" authority="Jałoszyński, 2014" authorityName="Jałoszyński" authorityYear="2014" box="[181,477,1977,1998]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="solodovnikovi" status="sp. nov.">
<emphasis id="D2AF425A58632E4EFF6672F96DF3F837" box="[181,477,1977,1998]" italics="true" pageId="1" pageNumber="84">Horaeomorphus solodovnikovi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="C99CFF2158632E4EFE3A72F96E71F837" box="[489,607,1977,1998]" pageId="1" pageNumber="84" rank="species">new species</taxonomicNameLabel>
, (A–C); and
<taxonomicName id="27DBE5CB58632E4EFD0972F96801F837" authority="(Lhoste)" authorityName="Jałoszyński" authorityYear="2014" baseAuthorityName="Jałoszyński" baseAuthorityYear="1939" box="[730,1071,1977,1998]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="wailimae" status="comb. nov.">
<emphasis id="D2AF425A58632E4EFD0972F96FFBF837" box="[730,981,1977,1998]" italics="true" pageId="1" pageNumber="84">Horaeomorphus wailimae</emphasis>
(Lhoste)
</taxonomicName>
,
<taxonomicNameLabel id="C99CFF2158632E4EFBEB72F968CFF837" box="[1080,1249,1977,1998]" pageId="1" pageNumber="84" rank="species">new combination</taxonomicNameLabel>
, (D–G). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C).
</paragraph>
</caption>
<paragraph id="E0649E4858602E4DFF5B75FE6D0EFEEC" blockId="2.[136,774,190,277]" pageId="2" pageNumber="85">
Metatrochanters (
<figureCitation id="78E082CD58602E4DFE9575FE6DB7FF2C" box="[326,409,190,213]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (A–C); and Horaeomorphus wailimae (Lhoste), new combination, (D–G). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="2" pageNumber="85">Fig. 1B</figureCitation>
) strongly modified, developed as long and recurved rod-like projections nearly as long as half of metafemur.
</paragraph>
<paragraph id="E0649E4858602E4DFF5B747E6EB7FDEC" blockId="2.[136,775,318,533]" pageId="2" pageNumber="85">
Aedeagus (
<figureCitation id="78E082CD58602E4DFF2C747E6D75FEAC" box="[255,347,318,341]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (A–C); and Horaeomorphus wailimae (Lhoste), new combination, (D–G). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="2" pageNumber="85">Figs. 1C</figureCitation>
,
<figureCitation id="78E082CD58602E4DFEB4747E6D88FEAC" box="[359,422,318,341]" captionStart="Fig" captionStartId="2.[136,171,2062,2083]" captionTargetBox="[138,772,1364,2039]" captionTargetId="figure-739@2.[128,781,1350,2045]" captionTargetPageId="2" captionText="Fig. 2. Horaeomorphus solodovnikovi new species. Aedeagus in ventral (A) and lateral (B) views. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4504190" httpUri="https://zenodo.org/record/4504190/files/figure.png" pageId="2" pageNumber="85">2A, B</figureCitation>
)
<quantity id="272333AD58602E4DFE66747E6E3DFEAC" box="[437,531,318,341]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="2" pageNumber="85" unit="mm" value="0.73">0.73 mm</quantity>
in length; median lobe broadest in basal third, distal part subtriangular and rounded at apex, in lateral view slightly curved dorsally; internal armature remarkable, with large and symmetrical central complex containing strongly elongate tubular structure in middle; parameres slender, each with apical bunch of several setae of various lengths, without subapical setae.
</paragraph>
<paragraph id="E0649E4858602E4DFF5B777E6D74FDAC" blockId="2.[136,346,574,597]" box="[136,346,574,597]" pageId="2" pageNumber="85">
<emphasis id="D2AF425A58602E4DFF5B777E6CCCFDAC" bold="true" box="[136,226,574,597]" pageId="2" pageNumber="85">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358602E4DFF5B773E6E68FD2C" pageId="2" pageNumber="85" type="etymology">
<paragraph id="E0649E4858602E4DFF5B773E6E68FD2C" blockId="2.[136,775,638,725]" pageId="2" pageNumber="85">
<emphasis id="D2AF425A58602E4DFF5B773E6D2BFD6C" bold="true" box="[136,261,638,661]" pageId="2" pageNumber="85">Etymology.</emphasis>
This species is dedicated to Alexey Solodovnikov (ZMUC), a specialist on
<taxonomicName id="27DBE5CB58602E4DFE4377DE6E0DFD4C" box="[400,547,670,693]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
and one of collectors of the specimen fixed here as a
<typeStatus id="3F6020EA58602E4DFE3277FE6E6CFD2C" box="[481,578,702,725]" pageId="2" pageNumber="85" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358602E4DFF5B77BE6E99FCEC" box="[136,695,766,789]" pageId="2" pageNumber="85" type="distribution">
<paragraph id="E0649E4858602E4DFF5B77BE6E99FCEC" blockId="2.[136,695,766,789]" box="[136,695,766,789]" pageId="2" pageNumber="85">
<emphasis id="D2AF425A58602E4DFF5B77BE6D34FCEC" bold="true" box="[136,282,766,789]" pageId="2" pageNumber="85">Distribution.</emphasis>
Southern
<collectingCountry id="98CCDED858602E4DFE5977BE6DEEFCEC" box="[394,448,766,789]" name="Laos" pageId="2" pageNumber="85">Laos</collectingCountry>
,
<collectingRegion id="221F50AA58602E4DFE1E77BE6E9DFCEC" box="[461,691,766,789]" country="Laos" name="Champasak" pageId="2" pageNumber="85">Champasak Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358602E4DFF5B767E689AFF0C" pageId="2" pageNumber="85" type="discussion">
<paragraph id="E0649E4858602E4DFF5B767E689AFF0C" blockId="2.[136,776,830,1333]" lastBlockId="2.[835,1474,190,245]" pageId="2" pageNumber="85">
<emphasis id="D2AF425A58602E4DFF5B767E6CD9FCAC" bold="true" box="[136,247,830,853]" pageId="2" pageNumber="85">Remarks.</emphasis>
<taxonomicName id="27DBE5CB58602E4DFF2D767E6E66FCAC" authorityName="Jałoszyński" authorityYear="2014" box="[254,584,830,853]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="solodovnikovi">
<emphasis id="D2AF425A58602E4DFF2D767E6E66FCAC" box="[254,584,830,853]" italics="true" pageId="2" pageNumber="85">Horaeomorphus solodovnikovi</emphasis>
</taxonomicName>
shares the highly modified and recurved male metatrochanters with several congeners:
<taxonomicName id="27DBE5CB58602E4DFED2763E6E78FC6C" authority="Jaloszynski, 2006" authorityName="Jaloszynski" authorityYear="2006" box="[257,598,894,917]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="D2AF425A58602E4DFED2763E6DBBFC6C" box="[257,405,894,917]" italics="true" pageId="2" pageNumber="85">H. antennatus</emphasis>
Jałoszyński, 2006
</taxonomicName>
(from
<collectingCountry id="98CCDED858602E4DFD4D763E6F2AFC6C" box="[670,772,894,917]" name="Malaysia" pageId="2" pageNumber="85">Malaysia</collectingCountry>
:
<collectingRegion id="221F50AA58602E4DFF5B76DE6CF3FC4C" box="[136,221,926,949]" country="Malaysia" name="Pahang" pageId="2" pageNumber="85">Pahang</collectingRegion>
),
<taxonomicName id="27DBE5CB58602E4DFF2076DE6EA8FC4C" authority="Jaloszynski, 2006" authorityName="Jaloszynski" authorityYear="2006" box="[243,646,926,949]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="caverniventris">
<emphasis id="D2AF425A58602E4DFF2076DE6D9BFC4C" box="[243,437,926,949]" italics="true" pageId="2" pageNumber="85">H. caverniventris</emphasis>
Jałoszyński, 2006
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFD4A76DE6F2DFC4C" box="[665,771,926,949]" name="Malaysia" pageId="2" pageNumber="85">Malaysia</collectingCountry>
:
<collectingRegion id="221F50AA58602E4DFF5B76FE6CF5FC2C" box="[136,219,958,981]" country="Malaysia" name="Pahang" pageId="2" pageNumber="85">Pahang</collectingRegion>
),
<taxonomicName id="27DBE5CB58602E4DFF3C76FE6E14FC2C" authority="Franz, 1992" authorityName="Franz" authorityYear="1992" box="[239,570,958,981]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="punctatissimus">
<emphasis id="D2AF425A58602E4DFF3C76FE6D9FFC2C" box="[239,433,958,981]" italics="true" pageId="2" pageNumber="85">H. punctatissimus</emphasis>
<bibRefCitation id="844AE3B958602E4DFE6A76FE6E14FC2C" author="Franz H" box="[441,570,958,981]" pageId="2" pageNumber="85" pagination="69 - 76" refId="ref4325" refString="Franz H (1992) Scydmaeniden aus Malaysia und Sulawesi (Coleoptera: Scydmaenidae). Koleopterologische Rundschau, 62: 69 - 76." type="journal article" year="1992">Franz, 1992</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFD9876FE6E9CFC2C" box="[587,690,958,981]" name="Malaysia" pageId="2" pageNumber="85">Malaysia</collectingCountry>
:
<collectingRegion id="221F50AA58602E4DFD6D76FE6F2DFC2C" box="[702,771,958,981]" country="Malaysia" name="Sabah" pageId="2" pageNumber="85">Sabah</collectingRegion>
;
<collectingCountry id="98CCDED858602E4DFF5B769E6CDAFC0C" box="[136,244,990,1013]" name="Indonesia" pageId="2" pageNumber="85">Indonesia</collectingCountry>
:
<collectingRegion id="221F50AA58602E4DFED2769E6D73FC0C" box="[257,349,990,1013]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="85">Sumatra</collectingRegion>
, Siberut Is.),
<taxonomicName id="27DBE5CB58602E4DFE20769E6CEFFBEC" authority="Jaloszynski, 2006" authorityName="Jaloszynski" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="jeraianus">
<emphasis id="D2AF425A58602E4DFE20769E6E52FC0C" box="[499,636,990,1013]" italics="true" pageId="2" pageNumber="85">H. jeraianus</emphasis>
Jałoszyński, 2006
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFF0676BE6D68FBEC" box="[213,326,1022,1045]" name="Malaysia" pageId="2" pageNumber="85">Malaysia</collectingCountry>
:
<collectingRegion id="221F50AA58602E4DFE8476BE6D89FBEC" box="[343,423,1022,1045]" country="Malaysia" name="Kedah" pageId="2" pageNumber="85">Kedah</collectingRegion>
),
<taxonomicName id="27DBE5CB58602E4DFE6C76BE6CEEFBCC" authority="Jaloszynski, 2006" authorityName="Jaloszynski" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="punctifrons">
<emphasis id="D2AF425A58602E4DFE6C76BE6E42FBEC" box="[447,620,1022,1045]" italics="true" pageId="2" pageNumber="85">H. punctifrons</emphasis>
Jałoszyński, 2006
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFF00715E6D6FFBCC" box="[211,321,1054,1077]" name="Malaysia" pageId="2" pageNumber="85">Malaysia</collectingCountry>
:
<collectingRegion id="221F50AA58602E4DFE82715E6D87FBCC" box="[337,425,1054,1077]" country="Malaysia" name="Pahang" pageId="2" pageNumber="85">Pahang</collectingRegion>
),
<taxonomicName id="27DBE5CB58602E4DFE12715E6F28FBCC" authority="Jaloszynski, 2006" authorityName="Jaloszynski" authorityYear="2006" box="[449,774,1054,1077]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="jaechi">
<emphasis id="D2AF425A58602E4DFE12715E6E02FBCC" box="[449,556,1054,1077]" italics="true" pageId="2" pageNumber="85">H. jaechi</emphasis>
Jałoszyński, 2006
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFF43717E6CD2FBAC" box="[144,252,1086,1109]" name="Malaysia" pageId="2" pageNumber="85">Malaysia</collectingCountry>
:
<collectingCountry id="98CCDED858602E4DFED8717E6D41FBAC" box="[267,367,1086,1109]" name="Malaysia" pageId="2" pageNumber="85">Sarawak</collectingCountry>
),
<taxonomicName id="27DBE5CB58602E4DFE55717E6F29FBAC" authority="Jaloszynski, 2006" authorityName="Jaloszynski" authorityYear="2006" box="[390,775,1086,1109]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="tiomanensis">
<emphasis id="D2AF425A58602E4DFE55717E6E1FFBAC" box="[390,561,1086,1109]" italics="true" pageId="2" pageNumber="85">H. tiomanensis</emphasis>
Jałoszyński, 2006
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFF43711E6D2EFB8C" box="[144,256,1118,1141]" name="Malaysia" pageId="2" pageNumber="85">Malaysia</collectingCountry>
: Tioman Is.),
<taxonomicName id="27DBE5CB58602E4DFE7E711E6D0CFB6C" authority="Jaloszynski &amp; Nomura, 2007" authorityName="Jaloszynski &amp; Nomura" authorityYear="2007" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="endauensis">
<emphasis id="D2AF425A58602E4DFE7E711E6E7AFB8C" box="[429,596,1118,1141]" italics="true" pageId="2" pageNumber="85">H. endauensis</emphasis>
Jałoszyński &amp; Nomura, 2007
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFEE2713E6DB6FB6C" box="[305,408,1150,1173]" name="Malaysia" pageId="2" pageNumber="85">Malaysia</collectingCountry>
:
<collectingRegion id="221F50AA58602E4DFE71713E6DDBFB6C" box="[418,501,1150,1173]" country="Malaysia" name="Pahang" pageId="2" pageNumber="85">Pahang</collectingRegion>
),
<taxonomicName id="27DBE5CB58602E4DFDD4713E6CEEFB4C" authority="Jaloszynski, 2009" authorityName="Jaloszynski" authorityYear="2009" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="imitator">
<emphasis id="D2AF425A58602E4DFDD4713E6E50FB6C" box="[519,638,1150,1173]" italics="true" pageId="2" pageNumber="85">H. imitator</emphasis>
Jałoszyński, 2009
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFF0071DE6D77FB4C" box="[211,345,1182,1205]" name="Philippines" pageId="2" pageNumber="85">Philippines</collectingCountry>
: Mindanao),
<taxonomicName id="27DBE5CB58602E4DFE2671DE6CEFFB2C" authority="Jaloszynski, 2004" authorityName="Jaloszynski" authorityYear="2004" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="blattnyi">
<emphasis id="D2AF425A58602E4DFE2671DE6E5CFB4C" box="[501,626,1182,1205]" italics="true" pageId="2" pageNumber="85">H. blattnyi</emphasis>
Jałoszyński, 2004
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFF1A71FE6D65FB2C" box="[201,331,1214,1237]" name="Philippines" pageId="2" pageNumber="85">Philippines</collectingCountry>
: Luzon),
<taxonomicName id="27DBE5CB58602E4DFE6971FE6C93FB0C" authority="Jaloszynski, 2002" authorityName="Jaloszynski" authorityYear="2002" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="sakishimanus">
<emphasis id="D2AF425A58602E4DFE6971FE6E5AFB2C" box="[442,628,1214,1237]" italics="true" pageId="2" pageNumber="85">H. sakishimanus</emphasis>
<bibRefCitation id="844AE3B958602E4DFD5371FE6C93FB0C" author="Jaloszynski P" pageId="2" pageNumber="85" pagination="223 - 232" refId="ref4413" refString="Jaloszynski P (2002) First record of the genus Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae) from Japan, with description of a new species. Bulletin of National Science Museum Tokyo, Series A, 28 (4): 223 - 232." type="journal article" year="2002">Jałoszyński, 2002</bibRefCitation>
</taxonomicName>
(
<collectingCountry id="98CCDED858602E4DFF1E719E6D21FB0C" box="[205,271,1246,1269]" name="Japan" pageId="2" pageNumber="85">Japan</collectingCountry>
: Iriomote-jima and Ishigaki-jima islands), and
<taxonomicName id="27DBE5CB58602E4DFF5B71BE6DC5FAEC" authority="(Franz, 1986) (Fiji)" baseAuthorityName="Franz" baseAuthorityYear="1986" box="[136,491,1278,1301]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="calcarifer">
<emphasis id="D2AF425A58602E4DFF5B71BE6D3CFAEC" box="[136,274,1278,1301]" italics="true" pageId="2" pageNumber="85">H. calcarifer</emphasis>
(
<bibRefCitation id="844AE3B958602E4DFEF371BE6D8CFAEC" author="Franz H" box="[288,418,1278,1301]" pageId="2" pageNumber="85" pagination="147 - 178" refId="ref4302" refString="Franz H (1986) Zweiter Beitrag zur Kenntnis der Scydmaenidenfauna der Fiji-Inseln. Entomologische Blatter, 82: 147 - 178." type="journal article" year="1986">Franz, 1986</bibRefCitation>
) (
<collectingCountry id="98CCDED858602E4DFE6B71BE6DCDFAEC" box="[440,483,1278,1301]" name="Fiji" pageId="2" pageNumber="85">Fiji</collectingCountry>
)
</taxonomicName>
. None of these species has the aedeagus with a long tubular median part of the internal armature, and this character allows for an unambiguous identification of
<emphasis id="D2AF425A58602E4DFC25759E689AFF0C" box="[1014,1204,222,245]" italics="true" pageId="2" pageNumber="85">
<taxonomicName id="27DBE5CB58602E4DFC25759E689EFF0C" authorityName="Jałoszyński" authorityYear="2014" box="[1014,1200,222,245]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="solodovnikovi">H. solodovnikovi</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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