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<document id="38D9A8F071882BD987A252D87B5B5703" ID-DOI="10.5281/zenodo.4504184" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504184" ID-ZooBank="F3A75556-83C3-47A9-97CE-78342127CAE5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612465776621" checkinUser="carolina" docAuthor="Jałoszyński, Paweł" docDate="2014" docId="68722F5E58602E4CFC85741D6EDEFEEF" docLanguage="en" docName="RafflesBZool.62.83-88.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Horaeomorphus wailimae Jałoszyński 2014, new combination" docType="treatment" docVersion="5" lastPageNumber="85" masterDocId="944B572658622E4FFFD375406C2EFFF9" masterDocTitle="Oriental Horaeomorphus Schaufuss: a new species, synonymic notes and distributional synopsis (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="88" masterPageNumber="83" pageNumber="85" updateTime="1698866066355" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="86873D4CE738CD1F25038D8DA5F5D825">Oriental Horaeomorphus Schaufuss: a new species, synonymic notes and distributional synopsis (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="440C1DA543AD0CDE187FFF0BC3CA96D6">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="DD81EC5104C3F13BF2012916D60E1201">Raffles Bulletin of Zoology</mods:title>
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<subSubSection id="A8C1CDC358602E4DFC85741D6981FE8C" box="[854,1455,349,373]" pageId="2" pageNumber="85" type="nomenclature">
<paragraph id="E0649E4858602E4DFC85741D6981FE8C" blockId="2.[854,1455,349,405]" box="[854,1455,349,373]" pageId="2" pageNumber="85">
<heading id="BB2C292458602E4DFC85741D6981FE8C" bold="true" box="[854,1455,349,373]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="85" reason="2">
<emphasis id="D2AF425A58602E4DFC85741D6981FE8C" bold="true" box="[854,1455,349,373]" pageId="2" pageNumber="85">
<taxonomicName id="27DBE5CB58602E4DFC85741D68F0FE8C" ID-CoL="3MNM4" authority="(Lhoste)" authorityName="Jałoszyński" authorityYear="2014" baseAuthorityName="Jałoszyński" baseAuthorityYear="1939" box="[854,1246,349,373]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="wailimae" status="comb. nov.">
<emphasis id="D2AF425A58602E4DFC85741D6858FE8C" bold="true" box="[854,1142,349,373]" italics="true" pageId="2" pageNumber="85">Horaeomorphus wailimae</emphasis>
(Lhoste)
</taxonomicName>
,
<taxonomicNameLabel id="C99CFF2158602E4DFB38741E6981FE8C" box="[1259,1455,350,373]" pageId="2" pageNumber="85" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358602E4DFB93743E68EBFE6C" box="[1088,1221,382,405]" pageId="2" pageNumber="85" type="description">
<paragraph id="E0649E4858602E4DFB93743E68EBFE6C" blockId="2.[854,1455,349,405]" box="[1088,1221,382,405]" pageId="2" pageNumber="85">
(
<figureCitation id="78E082CD58602E4DFB9B743E68AAFE6C" box="[1096,1156,382,405]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (AC); and Horaeomorphus wailimae (Lhoste), new combination, (DG). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="2" pageNumber="85">Fig. 1</figureCitation>
DG)
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358602E4DFC9074FD6809FD62" pageId="2" pageNumber="85" type="reference_group">
<paragraph id="E0649E4858602E4DFC9074FD684EFE17" blockId="2.[835,1474,445,667]" pageId="2" pageNumber="85">
<treatmentCitationGroup id="C0CBB96658602E4DFC9074FD684EFE17" pageId="2" pageNumber="85">
<treatmentCitation id="617AB85958602E4DFC9074FD6978FE2B" author="Lhoste J" box="[835,1366,445,466]" page="1" pageId="2" pageNumber="85" year="1939">
<taxonomicName id="27DBE5CB58602E4DFC9074FD6978FE2B" ID-CoL="4ZVCM" authority="Lhoste, 1939: 1" authorityName="Lhoste" authorityPageNumber="1" authorityYear="1939" box="[835,1366,445,466]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="wailimae" subGenus="Cyrtoscydmus">
<emphasis id="D2AF425A58602E4DFC9074FD6F82FE2B" box="[835,940,445,466]" italics="true" pageId="2" pageNumber="85">Stenichnus</emphasis>
(
<emphasis id="D2AF425A58602E4DFC6E74FD6868FE2B" box="[957,1094,445,466]" italics="true" pageId="2" pageNumber="85">Cyrtoscydmus</emphasis>
)
<emphasis id="D2AF425A58602E4DFB8474FD689EFE2B" box="[1111,1200,445,466]" italics="true" pageId="2" pageNumber="85">wailimae</emphasis>
<bibRefCitation id="844AE3B958602E4DFB6A74FD6978FE2B" author="Lhoste J" box="[1209,1366,445,466]" pageId="2" pageNumber="85" pagination="1 - 10" refId="ref5050" refString="Lhoste J (1939) Etude de quelques Scydmaenidae des rives de l'Ocean Indien. (Coleoptera). 2 ieme note. Arbeiten uber Morphologische und Taxonomische Entomologie, 6: 1 - 10." type="journal article" year="1939">Lhoste, 1939: 1</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 12;
<treatmentCitation id="617AB85958602E4DFCBA7499683FFE17" author="Franz H" box="[873,1041,473,494]" page="546" pageId="2" pageNumber="85" year="1970">
<bibRefCitation id="844AE3B958602E4DFCBA7499683FFE17" author="Franz H" box="[873,1041,473,494]" pageId="2" pageNumber="85" pagination="535 - 578" refId="ref4125" refString="Franz H (1970) Zur Kenntnis der Scydmaeniden-Fauna von Singapore, Malakka und Indonesien (Coleoptera: Scydmaenidae). Beitrage zur Entomologie, 20: 535 - 578." type="journal article" year="1970">Franz, 1970: 546</bibRefCitation>
</treatmentCitation>
, fig. 11.
</treatmentCitationGroup>
</paragraph>
<paragraph id="E0649E4858602E4DFC9074B66FE1FDD1" blockId="2.[835,1474,445,667]" pageId="2" pageNumber="85">
<taxonomicName id="27DBE5CB58602E4DFC9074B66876FDF2" authority="Lhoste" authorityName="Lhoste" box="[835,1112,502,523]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="waihinae">
<emphasis id="D2AF425A58602E4DFC9074B66823FDF2" box="[835,1037,502,523]" italics="true" pageId="2" pageNumber="85">Stenichnus waihinae</emphasis>
Lhoste
</taxonomicName>
(misspelling in
<bibRefCitation id="844AE3B958602E4DFB2E74B6695BFDF2" author="Franz H" box="[1277,1397,502,523]" pageId="2" pageNumber="85" pagination="535 - 578" refId="ref4125" refString="Franz H (1970) Zur Kenntnis der Scydmaeniden-Fauna von Singapore, Malakka und Indonesien (Coleoptera: Scydmaenidae). Beitrage zur Entomologie, 20: 535 - 578." type="journal article" year="1970">Franz, 1970</bibRefCitation>
, legend to fig. 11).
</paragraph>
<paragraph id="E0649E4858602E4DFC907770680FFD9B" blockId="2.[835,1474,445,667]" pageId="2" pageNumber="85">
<treatmentCitationGroup id="C0CBB96658602E4DFC907770680FFD9B" pageId="2" pageNumber="85">
<treatmentCitation id="617AB85958602E4DFC90777069BBFDBC" author="Franz H" box="[835,1429,560,581]" page="124" pageId="2" pageNumber="85" year="1985">
<taxonomicName id="27DBE5CB58602E4DFC90777069BBFDBC" authority="Franz, 1985: 124" authorityName="Franz" authorityPageNumber="124" authorityYear="1985" box="[835,1429,560,581]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="heissi" subGenus="Pseudosyndicus">
<emphasis id="D2AF425A58602E4DFC90777068F2FDBC" box="[835,1244,560,581]" italics="true" pageId="2" pageNumber="85">Horaeomorphus (Pseudosyndicus) heissi</emphasis>
<bibRefCitation id="844AE3B958602E4DFB36777069BBFDBC" author="Franz H" box="[1253,1429,560,581]" pageId="2" pageNumber="85" pagination="91 - 128" refId="ref4270" refString="Franz H (1985) Neue und ungenugend bekannte Scydmaeniden (Coleoptera) aus Taiwan, Fukien und Thailand. Mitteilungen der Munchner Entomologische Gesellschaft, 74: 91 - 128." type="journal article" year="1985">Franz, 1985: 124</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 36,
<emphasis id="D2AF425A58602E4DFC5D770D680FFD9B" bold="true" box="[910,1057,589,610]" pageId="2" pageNumber="85">new synonym.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="E0649E4858602E4DFC9077296809FD62" blockId="2.[835,1474,445,667]" pageId="2" pageNumber="85">
<taxonomicName id="27DBE5CB58602E4DFC9077296870FD87" authority="Franz" authorityName="Franz" authorityYear="1985" box="[835,1118,617,638]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="heissi">
<emphasis id="D2AF425A58602E4DFC9077296830FD87" box="[835,1054,617,638]" italics="true" pageId="2" pageNumber="85">Horaeomorphus heissi</emphasis>
Franz
</taxonomicName>
; Jałoszyński, 2006: 39, figs. 1, 8, 22, 50, 83, 8689, 124.
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358602E4DFC907780696FFBEC" pageId="2" pageNumber="85" type="materials_examined">
<paragraph id="E0649E4858602E4DFC9077806FBEFC21" blockId="2.[835,1474,704,1045]" pageId="2" pageNumber="85">
<emphasis id="D2AF425A58602E4DFC9077806836FD2C" bold="true" box="[835,1048,704,725]" pageId="2" pageNumber="85">Material examined.</emphasis>
<typeStatus id="3F6020EA58602E4DFBF1778068A9FD2C" box="[1058,1159,704,725]" pageId="2" pageNumber="85" type="lectotype">Lectotype</typeStatus>
(here designated). Male, drymounted beetle (
<figureCitation id="78E082CD58602E4DFC3A779D6819FD0B" box="[1001,1079,733,754]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (AC); and Horaeomorphus wailimae (Lhoste), new combination, (DG). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="2" pageNumber="85">Fig. 1D</figureCitation>
) and genital preparation on microscope slide (
<figureCitation id="78E082CD58602E4DFC5777B96FFCFCF7" box="[900,978,761,782]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (AC); and Horaeomorphus wailimae (Lhoste), new combination, (DG). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="2" pageNumber="85">Fig. 1F</figureCitation>
), card-mounted specimen with five labels (
<figureCitation id="78E082CD58602E4DFA4A77B96F4AFCD2" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (AC); and Horaeomorphus wailimae (Lhoste), new combination, (DG). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="2" pageNumber="85">Fig. 1G</figureCitation>
): “Wai Lima Z. Sum. / Lampongs [i.e.,
<collectingRegion id="221F50AA58602E4DFAD576566949FCD2" box="[1286,1383,790,811]" country="Indonesia" name="Lampung" pageId="2" pageNumber="85">Lampung</collectingRegion>
] No 156 / Karny. XIXII.1921” [white, printed except for handwritten number], “
<taxonomicName id="27DBE5CB58602E4DFC6376106835FC9C" box="[944,1051,848,869]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">Stenichnus</taxonomicName>
/ wailimae / J. LHOSTE det. n. sp.” [white, handwritten and printed], “
<typeStatus id="3F6020EA58602E4DFB97762D6885FC7B" box="[1092,1195,877,898]" pageId="2" pageNumber="85" type="holotype">Holotypus</typeStatus>
” [red, printed], “Lhoste det.” [white, printed], “
<emphasis id="D2AF425A58602E4DFC2376C968E1FC67" box="[1008,1231,905,926]" italics="true" pageId="2" pageNumber="85">
<taxonomicName id="27DBE5CB58602E4DFC2376C968EFFC67" authorityName="Schaufuss" authorityYear="1889" box="[1008,1217,905,926]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">HORAEOMORPHUS</taxonomicName>
/
</emphasis>
<taxonomicName id="27DBE5CB58602E4DFB0576C969ECFC67" authority="(Lhoste, 1939)" authorityName="Jałoszyński" authorityYear="2014" baseAuthorityName="Lhoste" baseAuthorityYear="1939" box="[1238,1474,905,926]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="wailimae">
<emphasis id="D2AF425A58602E4DFB0576C96900FC67" box="[1238,1326,905,926]" italics="true" pageId="2" pageNumber="85">wailimae</emphasis>
(
<bibRefCitation id="844AE3B958602E4DFAE876C96995FC67" author="Lhoste J" box="[1339,1467,905,926]" pageId="2" pageNumber="85" pagination="1 - 10" refId="ref5050" refString="Lhoste J (1939) Etude de quelques Scydmaenidae des rives de l'Ocean Indien. (Coleoptera). 2 ieme note. Arbeiten uber Morphologische und Taxonomische Entomologie, 6: 1 - 10." type="journal article" year="1939">Lhoste, 1939</bibRefCitation>
)
</taxonomicName>
/
<typeStatus id="3F6020EA58602E4DFC8176E66FC5FC42" box="[850,1003,934,955]" pageId="2" pageNumber="85" type="lectotype">LECTOTYPUS</typeStatus>
/ det. P. JAŁOSZYŃSKI, 2013 [white, printed] (
<collectionCode id="86CA068D58602E4DFC9976836FAAFC21" box="[842,900,963,984]" pageId="2" pageNumber="85">SDEI</collectionCode>
).
</paragraph>
<paragraph id="E0649E4858602E4DFC9076A0696FFBEC" blockId="2.[835,1474,704,1045]" pageId="2" pageNumber="85">
<materialsCitation id="50B3941558602E4DFC9076A06913FBEC" collectionCode="SDEI" pageId="2" pageNumber="85" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paralectotype">
<typeStatus id="3F6020EA58602E4DFC9076A06FFCFC0C" box="[835,978,992,1013]" pageId="2" pageNumber="85" type="paralectotype">Paralectotypes</typeStatus>
.
<specimenCount id="F6DD55C158602E4DFC0D76A0680FFC0C" box="[990,1057,992,1013]" pageId="2" pageNumber="85" type="male">1 male</specimenCount>
and
<specimenCount id="F6DD55C158602E4DFB8776A06887FC0C" box="[1108,1193,992,1013]" pageId="2" pageNumber="85" type="female">1 female</specimenCount>
(
<figureCitation id="78E082CD58602E4DFB6A76A0692AFC0C" box="[1209,1284,992,1013]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (AC); and Horaeomorphus wailimae (Lhoste), new combination, (DG). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="2" pageNumber="85">Fig. 1E</figureCitation>
), same data except for “
<typeStatus id="3F6020EA58602E4DFCA371406FFCFBEC" box="[880,978,1024,1045]" pageId="2" pageNumber="85" type="paratype">Paratypus</typeStatus>
” and “
<typeStatus id="3F6020EA58602E4DFBC77140688BFBEC" box="[1044,1189,1024,1045]" pageId="2" pageNumber="85" type="paralectotype">Paralectotypus</typeStatus>
” labels (
<collectionCode id="86CA068D58602E4DFB287140691BFBEC" box="[1275,1333,1024,1045]" pageId="2" pageNumber="85">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A8C1CDC358602E4CFC90717E6EDEFEEF" lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85" type="discussion">
<paragraph id="E0649E4858602E4DFC90717E6822FB0C" blockId="2.[835,1475,1086,1269]" pageId="2" pageNumber="85">
<emphasis id="D2AF425A58602E4DFC90717E6F9BFBAC" bold="true" box="[835,949,1086,1109]" pageId="2" pageNumber="85">Remarks.</emphasis>
<bibRefCitation id="844AE3B958602E4DFC13717E6871FBAC" author="Lhoste J" box="[960,1119,1086,1109]" pageId="2" pageNumber="85" pagination="1 - 10" refId="ref5050" refString="Lhoste J (1939) Etude de quelques Scydmaenidae des rives de l'Ocean Indien. (Coleoptera). 2 ieme note. Arbeiten uber Morphologische und Taxonomische Entomologie, 6: 1 - 10." type="journal article" year="1939">Lhoste (1939)</bibRefCitation>
did not specify the number of individuals included in the type series, therefore the three specimens preserved at SDEI have the status of
<typeStatus id="3F6020EA58602E4DFA92713E698CFB6C" box="[1345,1442,1150,1173]" pageId="2" pageNumber="85" type="syntype">syntypes</typeStatus>
. In order to ensure the stability of nomenclature and provide a unique name-bearing type for
<taxonomicName id="27DBE5CB58602E4DFBAB71FE6960FB2C" authorityName="Jałoszyński" authorityYear="2014" baseAuthorityName="Lhoste" baseAuthorityYear="1939" box="[1144,1358,1214,1237]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="wailimae">
<emphasis id="D2AF425A58602E4DFBAB71FE6960FB2C" box="[1144,1358,1214,1237]" italics="true" pageId="2" pageNumber="85">Stenichnus wailimae</emphasis>
</taxonomicName>
a
<typeStatus id="3F6020EA58602E4DFAB071FE69EDFB2C" box="[1379,1475,1214,1237]" pageId="2" pageNumber="85" type="lectotype">lectotype</typeStatus>
is here designated.
</paragraph>
<paragraph id="E0649E4858602E4DFC90705E68F9F92C" blockId="2.[835,1474,1310,1749]" pageId="2" pageNumber="85">
<bibRefCitation id="844AE3B958602E4DFC90705E6FF5FACC" author="Franz H" box="[835,987,1310,1333]" pageId="2" pageNumber="85" pagination="535 - 578" refId="ref4125" refString="Franz H (1970) Zur Kenntnis der Scydmaeniden-Fauna von Singapore, Malakka und Indonesien (Coleoptera: Scydmaenidae). Beitrage zur Entomologie, 20: 535 - 578." type="journal article" year="1970">Franz (1970)</bibRefCitation>
redescribed and illustrated the aedeagus of
<taxonomicName id="27DBE5CB58602E4DFCB6707E6863FAAC" authorityName="Jałoszyński" authorityYear="2014" baseAuthorityName="Lhoste" baseAuthorityYear="1939" box="[869,1101,1342,1365]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="wailimae">
<emphasis id="D2AF425A58602E4DFCB6707E6863FAAC" box="[869,1101,1342,1365]" italics="true" pageId="2" pageNumber="85">Stenichnus wailimae</emphasis>
</taxonomicName>
and did not express any doubts concerning the generic placement of this species. Later
<bibRefCitation id="844AE3B958602E4DFA55701E6FA7FA6C" author="Franz H" pageId="2" pageNumber="85" pagination="91 - 128" refId="ref4270" refString="Franz H (1985) Neue und ungenugend bekannte Scydmaeniden (Coleoptera) aus Taiwan, Fukien und Thailand. Mitteilungen der Munchner Entomologische Gesellschaft, 74: 91 - 128." type="journal article" year="1985">Franz (1985)</bibRefCitation>
described
<taxonomicName id="27DBE5CB58602E4DFC24703E68CCFA6C" authorityName="Franz" authorityYear="1985" box="[1015,1250,1406,1429]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="heissi">
<emphasis id="D2AF425A58602E4DFC24703E68CCFA6C" box="[1015,1250,1406,1429]" italics="true" pageId="2" pageNumber="85">Horaeomorphus heissi</emphasis>
</taxonomicName>
from Sumatra, which was redescribed by
<bibRefCitation id="844AE3B958602E4DFBC970DE68DDFA4C" author="Jaloszynski P" box="[1050,1267,1438,1461]" pageId="2" pageNumber="85" pagination="19 - 66" refId="ref4590" refString="Jaloszynski P (2006 a) Revision of Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae) of East [sic] Malaysia, Singapore and Sunda Islands. Genus, 17 (1): 19 - 66." type="journal article" year="2006">Jałoszyński (2006a)</bibRefCitation>
. In the latter study, which is a comprehensive revision of
<taxonomicName id="27DBE5CB58602E4DFB0A70FE69A6FA2C" authorityName="Schaufuss" authorityYear="1889" box="[1241,1416,1470,1493]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="D2AF425A58602E4DFB0A70FE69A6FA2C" box="[1241,1416,1470,1493]" italics="true" pageId="2" pageNumber="85">Horaeomorphus</emphasis>
</taxonomicName>
from
<collectingCountry id="98CCDED858602E4DFC90709E6F89FA0C" box="[835,935,1502,1525]" name="Malaysia" pageId="2" pageNumber="85">Malaysia</collectingCountry>
,
<collectingCountry id="98CCDED858602E4DFC61709E680FFA0C" box="[946,1057,1502,1525]" name="Singapore" pageId="2" pageNumber="85">Singapore</collectingCountry>
,
<collectingCountry id="98CCDED858602E4DFBF8709E68BBFA0C" box="[1067,1173,1502,1525]" name="Indonesia" pageId="2" pageNumber="85">Indonesia</collectingCountry>
, and
<collectingCountry id="98CCDED858602E4DFB1F709E6939FA0C" box="[1228,1303,1502,1525]" name="Brunei" pageId="2" pageNumber="85">Brunei</collectingCountry>
, the
<typeStatus id="3F6020EA58602E4DFA9B709E698BFA0C" box="[1352,1445,1502,1525]" pageId="2" pageNumber="85" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="27DBE5CB58602E4DFC9070BE6831F9EC" authorityName="Jałoszyński" authorityYear="2014" baseAuthorityName="Lhoste" baseAuthorityYear="1939" box="[835,1055,1534,1557]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="wailimae">
<emphasis id="D2AF425A58602E4DFC9070BE6831F9EC" box="[835,1055,1534,1557]" italics="true" pageId="2" pageNumber="85">Stenichnus wailimae</emphasis>
</taxonomicName>
were not studied, as it seemed unlikely that Franz, who himself published a number of papers on
<taxonomicName id="27DBE5CB58602E4DFC90737E6F99F9AC" box="[835,951,1598,1621]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="D2AF425A58602E4DFC90737E6F99F9AC" box="[835,951,1598,1621]" italics="true" pageId="2" pageNumber="85">Stenichnus</emphasis>
</taxonomicName>
and
<taxonomicName id="27DBE5CB58602E4DFC21737E688FF9AC" authorityName="Schaufuss" authorityYear="1889" box="[1010,1185,1598,1621]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="D2AF425A58602E4DFC21737E688FF9AC" box="[1010,1185,1598,1621]" italics="true" pageId="2" pageNumber="85">Horaeomorphus</emphasis>
</taxonomicName>
, might have misidentified these genera. However, examination of the type series of
<taxonomicName id="27DBE5CB58602E4DFC90733E6808F96C" authorityName="Jałoszyński" authorityYear="2014" baseAuthorityName="Lhoste" baseAuthorityYear="1939" box="[835,1062,1662,1685]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="wailimae">
<emphasis id="D2AF425A58602E4DFC90733E6808F96C" box="[835,1062,1662,1685]" italics="true" pageId="2" pageNumber="85">Stenichnus wailimae</emphasis>
</taxonomicName>
carried out during the present study revealed that this species not only belongs to
<taxonomicName id="27DBE5CB58602E4DFAC373DE6995F94C" authorityName="Schaufuss" authorityYear="1889" box="[1296,1467,1694,1717]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="D2AF425A58602E4DFAC373DE6995F94C" box="[1296,1467,1694,1717]" italics="true" pageId="2" pageNumber="85">Horaeomorphus</emphasis>
</taxonomicName>
, but is also conspecific with
<taxonomicName id="27DBE5CB58602E4DFBA273FE68FEF92C" authorityName="Franz" authorityYear="1985" box="[1137,1232,1726,1749]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="heissi">
<emphasis id="D2AF425A58602E4DFBA273FE68FEF92C" box="[1137,1232,1726,1749]" italics="true" pageId="2" pageNumber="85">H. heissi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E0649E4858602E4CFC9073BE6EDEFEEF" blockId="2.[835,1475,1790,2101]" lastBlockId="3.[113,752,191,278]" lastPageId="3" lastPageNumber="86" pageId="2" pageNumber="85">
<taxonomicName id="27DBE5CB58602E4DFC9073BE685AF8EC" authorityName="Jałoszyński" authorityYear="2014" baseAuthorityName="Jałoszyński" baseAuthorityYear="1939" box="[835,1140,1790,1813]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="wailimae">
<emphasis id="D2AF425A58602E4DFC9073BE685AF8EC" box="[835,1140,1790,1813]" italics="true" pageId="2" pageNumber="85">Horaeomorphus wailimae</emphasis>
</taxonomicName>
(=
<taxonomicName id="27DBE5CB58602E4DFB7573BE6921F8EC" authorityName="Franz" authorityYear="1985" box="[1190,1295,1790,1813]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="heissi">
<emphasis id="D2AF425A58602E4DFB7573BE6921F8EC" box="[1190,1295,1790,1813]" italics="true" pageId="2" pageNumber="85">H. heissi</emphasis>
</taxonomicName>
) can be easily identified on the basis of a hook-like projection on the ventral surface of male metafemora (
<figureCitation id="78E082CD58602E4DFBA0727E68EBF8AC" box="[1139,1221,1854,1877]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (AC); and Horaeomorphus wailimae (Lhoste), new combination, (DG). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="2" pageNumber="85">Fig. 1D</figureCitation>
, marked with an arrow), missing in females (
<figureCitation id="78E082CD58602E4DFBC8721E6842F88C" box="[1051,1132,1886,1909]" captionStart="Fig" captionStartId="1.[113,148,1977,1998]" captionTargetBox="[120,1444,1159,1947]" captionTargetId="figure-329@1.[113,1454,1145,1953]" captionTargetPageId="1" captionText="Fig. 1. Horaeomorphus solodovnikovi, new species, (AC); and Horaeomorphus wailimae (Lhoste), new combination, (DG). Dorsal habitus of holotype male (A), lectotype male (D; arrow indicates femoral projection) and paralectotype female (E); right metatrochanter and femur in ventral view (B; arrow indicates the apex of trochanteral projection); internal armature of aedeagus in ventral view (C; arrow indicates median tubular structure); genital preparation of lectotype male (F); and original set of labels of lectotype male (G). Scale bars = 0.5 mm (A, D, E), 0.1 mm (B), 0.2 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.4504186" httpUri="https://zenodo.org/record/4504186/files/figure.png" pageId="2" pageNumber="85">Fig. 1E</figureCitation>
), non-modified metatrochanters in both sexes, three ante-basal pits on the pronotum, of which the middle one is anteriorly connected to a short longitudinal groove, and a remarkable and unique aedeagus (illustrated in
<bibRefCitation id="844AE3B958602E4DFCB3729E6800F80C" author="Jaloszynski P" box="[864,1070,2014,2037]" pageId="2" pageNumber="85" pagination="19 - 66" refId="ref4590" refString="Jaloszynski P (2006 a) Revision of Horaeomorphus Schaufuss (Coleoptera, Scydmaenidae) of East [sic] Malaysia, Singapore and Sunda Islands. Genus, 17 (1): 19 - 66." type="journal article" year="2006">Jałoszyński, 2006a</bibRefCitation>
, Figs. 8689). The only other known species of
<taxonomicName id="27DBE5CB58602E4DFC6572BE684BF7EC" authorityName="Schaufuss" authorityYear="1889" box="[950,1125,2046,2069]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="D2AF425A58602E4DFC6572BE684BF7EC" box="[950,1125,2046,2069]" italics="true" pageId="2" pageNumber="85">Horaeomorphus</emphasis>
</taxonomicName>
with the hook-like metafemoral projection is
<taxonomicName id="27DBE5CB58602E4DFC0B7D5E6979F7CC" authority="Jaloszynski, 2009" authorityName="Jaloszynski" authorityYear="2009" box="[984,1367,2078,2101]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="samosirensis">
<emphasis id="D2AF425A58602E4DFC0B7D5E68A6F7CC" box="[984,1160,2078,2101]" italics="true" pageId="2" pageNumber="85">H. samosirensis</emphasis>
Jałoszyński, 2009
</taxonomicName>
from the Samosir Island,
<collectingRegion id="221F50AA58612E4CFEF875FF6DFDFF2F" box="[299,467,191,214]" country="Indonesia" name="Sumatera Utara" pageId="3" pageNumber="86">North Sumatra</collectingRegion>
, but it has the pronotum nearly circular and nearly as broad as elytra, and not strongly elongate and much narrower than elytra, as in
<taxonomicName id="27DBE5CB58612E4CFDB575BF6EC4FEEF" authorityName="Jałoszyński" authorityYear="2014" baseAuthorityName="Jałoszyński" baseAuthorityYear="1939" box="[614,746,255,278]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="wailimae">
<emphasis id="D2AF425A58612E4CFDB575BF6EC4FEEF" box="[614,746,255,278]" italics="true" pageId="3" pageNumber="86">H. wailimae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>