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<mods:title id="95886F8777FB7B63789891E2E73D62BD">Adding the West-African riverine component: Revision of the Recent freshwater snails belonging to Pseudocleopatra Thiele, 1928 (Caenogastropoda, Cerithioidea, Paludomidae)</mods:title>
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<emphasis id="D2DAEAECF637C511FF6C6643748DCDAA" bold="true" box="[151,484,1404,1431]" italics="true" pageId="14" pageNumber="315">Pseudocleopatra dartevellei</emphasis>
<bibRefCitation id="843F4B0FF637C511FE1766427792CDAB" author="Mandahl-Barth, G." box="[492,763,1404,1431]" pageId="14" pageNumber="315" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
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<figureCitation id="78952A7BF637C511FF6C669F7598CD87" box="[151,241,1440,1466]" captionStart="FIGURE 8" captionStartId="11.[151,250,926,951]" captionTargetBox="[481,1112,201,877]" captionTargetId="figure@11.[466,1121,184,877]" captionTargetPageId="11" captionText="FIGURE 8. Opercula of Pseudocleopatra Thiele, 1928 species. A. Lectotype of P. togoensis Thiele, 1928, ZMB 190048b. B. Paratype of Pseudocleopatra voltana Mandahl-Barth, 1973, Mogeyenga village, DBL 2591. C. Probable paratype of Pseudocleopatra dartevellei Mandahl-Barth, 1973, Matadi, DBL 2583. D. Pseudocleopatra broecki (Putzeys, 1899) n. comb., MCZ 171520. E. Lectotype of Pseudocleopatra broecki (Putzeys, 1899) n. comb., MRAC 47360. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3460148" httpUri="https://zenodo.org/record/3460148/files/figure.png" pageId="14" pageNumber="315">Figs 8C</figureCitation>
,
<figureCitation id="78952A7BF637C511FF0666A0742ACD87" box="[253,323,1439,1466]" captionStart-0="FIGURE 15" captionStart-1="FIGURE 16" captionStart-2="FIGURE 17" captionStart-3="FIGURE 18" captionStartId-0="16.[151,250,1903,1928]" captionStartId-1="17.[151,250,1938,1963]" captionStartId-2="18.[151,250,990,1015]" captionStartId-3="18.[151,250,1708,1733]" captionTargetBox-0="[456,1135,1225,1869]" captionTargetBox-1="[151,1431,1020,1918]" captionTargetBox-2="[214,1374,174,971]" captionTargetBox-3="[482,1105,1102,1689]" captionTargetId-0="figure@16.[441,1146,1214,1884]" captionTargetId-1="figure@17.[170,1417,1033,1918]" captionTargetId-2="figure@18.[214,1374,174,971]" captionTargetId-3="figure@18.[482,1105,1102,1689]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="18" captionText-0="FIGURE 15. Shells of Pseudocleopatra dartevellei Mandahl-Barth, 1973. A. Holotype, MRAC 799161. B. Probable paratype, Matadi; MRAC 260702. CD. Probable paratypes, Matadi, MRAC 260702, growth series. Scale bar: 5 mm." captionText-1="FIGURE 16. Radula of Pseudocleopatra dartevellei Mandahl-Barth, 1973 (Morph a), probable paratype, Île des Princes, MRAC 146342. A. Radula ribbon, scale bar: 10 μm. B. Lateral and central teeth, scale bar: 10 μm. C. Cutting edge of lateral and central teeth, scale bar: 10 μm. D. Central, lateral and marginal teeth, scale bar: 10 μm." captionText-2="FIGURE 17. Radula of Pseudocleopatra dartevellei Mandahl-Barth, 1973 (Morph b), probable paratype, Île des Princes, MRAC 146342. A. Radula ribbon, scale bar: 10 μm. B. Lateral and central teeth, scale bar: 10 μm. C. Cutting edge of lateral and central teeth, scale bar: 2 μm. D. Central, lateral and marginal teeth, scale bar: 10 μm." captionText-3="FIGURE 18. Known occurrence records of Pseudocleopatra dartevellei Mandahl-Barth, 1973 in the Congo River system. The asterisk indicates the type locality; solid symbols indicate samples with alcohol material, open symbols those of dry material. The arrow indicates the approximate location of the Inga dams." figureDoi-0="http://doi.org/10.5281/zenodo.3460164" figureDoi-1="http://doi.org/10.5281/zenodo.3460166" figureDoi-2="http://doi.org/10.5281/zenodo.3460168" figureDoi-3="http://doi.org/10.5281/zenodo.3460170" httpUri-0="https://zenodo.org/record/3460164/files/figure.png" httpUri-1="https://zenodo.org/record/3460166/files/figure.png" httpUri-2="https://zenodo.org/record/3460168/files/figure.png" httpUri-3="https://zenodo.org/record/3460170/files/figure.png" pageId="14" pageNumber="315">1518</figureCitation>
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<emphasis id="D2DAEAECF637C511FF6C66DA74DFCDC0" box="[151,438,1509,1533]" italics="true" pageId="14" pageNumber="315">Pseudocleopatra dartevellei</emphasis>
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, fig. 3df, i; Mandahl-Barth (1974: 355);
<bibRefCitation id="843F4B0FF637C511FB5066DA70F1CDC0" author="Brown, D. S." box="[1195,1432,1509,1533]" pageId="14" pageNumber="315" refId="ref13993" refString="Brown, D. S. (1980) Freshwater snails of Africa and their medical importance. Taylor &amp; Francis, London, 487 pp." type="book" year="1980">Brown (1980: 112113</bibRefCitation>
, fig. 64a);
<bibRefCitation id="843F4B0FF637C511FED7653C77BECE21" author="Danish Bilharziasis Laboratory" box="[300,727,1539,1564]" pageId="14" pageNumber="315" refId="ref14192" refString="Danish Bilharziasis Laboratory (1982) Guide de terrain des gasteropodes d'eau douce Africains. 5: Afrique centrale. Danish Bilharziasis Laboratory, Charlottenlund, 55 pp." type="book" year="1982">Danish Bilharziasis Laboratory (1982: 25</bibRefCitation>
, 50, fig. 77);
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, 539, 570, fig. 62a);
<bibRefCitation id="843F4B0FF637C511FAD7653C7445CE07" author="Glaubrecht, M." pageId="14" pageNumber="315" refId="ref14270" refString="Glaubrecht, M. (1996) Evolutionsokologie und Systematik am Beispiel von Suss- und Brackwasserschnecken (Mollusca: Caenogastropoda: Cerithioidea): Ontogenese-Strategien, palaontologische Befunde und historische Zoogeographie. Backhuys Publishers, Leiden, xvi + 499 pp., 25 pls." type="book" year="1996">Glaubrecht (1996: 27</bibRefCitation>
, 144, 146148, 307308, 493, fig. 25, pl. 2, fig. 5, pl. 21, figs 34);
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, fig.
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Graf
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<typeStatus id="3F15885CF637C511FE8665F07481CED4" box="[381,488,1743,1769]" pageId="14" pageNumber="315" type="holotype">Holotype</typeStatus>
,
<specimenCode id="B0089E85F637C511FE0F65F077CCCED4" box="[500,677,1743,1769]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="14" pageNumber="315" type="Museum">MRAC 799161</specimenCode>
, leg.
<collectorName id="4D5B5328F637C511FD1A65EF7681CED4" box="[737,1000,1743,1770]" pageId="14" pageNumber="315">E. Dartevelle. Probable</collectorName>
<typeStatus id="3F15885CF637C511FC0B65EF7132CED7" box="[1008,1115,1744,1770]" pageId="14" pageNumber="315" type="paratype">paratypes</typeStatus>
(not explicitly mentioned in the original description): all from
<collectingCountry id="98B9766EF637C511FDF565CC76D5CF30" box="[526,956,1779,1805]" name="Democratic Republic of the Congo" pageId="14" pageNumber="315">the Democratic Republic of the Congo</collectingCountry>
,
<location id="E5716025F637C511FC3C65CB71B0CF30" LSID="urn:lsid:plazi:treatment:680787E8F637C50CFF6C664371CACA02:E5716025F637C511FC3C65CB71B0CF30" box="[967,1241,1779,1806]" country="Democratic Republic of the Congo" county="Matadi" latitude="-5.881111" longLatPrecision="21" longitude="13.029166" municipality="River" name="Kongo Central Province" pageId="14" pageNumber="315">Kongo Central Province</location>
: “Île des Princes” (=
<location id="E5716025F637C511FF4064297401CF0C" LSID="urn:lsid:plazi:treatment:680787E8F637C50CFF6C664371CACA02:E5716025F637C511FF4064297401CF0C" box="[187,360,1814,1841]" country="Democratic Republic of the Congo" county="Matadi" latitude="-5.881111" longLatPrecision="21" longitude="13.029166" municipality="River" name="Ile des Princes" pageId="14" pageNumber="315">Île des Princes</location>
, ca.
<geoCoordinate id="859A5039F637C511FE5A6429774DCF0C" box="[417,548,1814,1841]" degrees="5" direction="south" minutes="53" orientation="latitude" pageId="14" pageNumber="315" precision="15" seconds="00" value="-5.883333">5°5300” S</geoCoordinate>
,
<geoCoordinate id="859A5039F637C511FDCF642877AECF0F" box="[564,711,1815,1842]" degrees="13" direction="east" minutes="06" orientation="longitude" pageId="14" pageNumber="315" precision="15" seconds="00" value="13.1">13°0600” E</geoCoordinate>
),
<specimenCode id="B0089E85F637C511FD24642876FACF0C" box="[735,915,1815,1841]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="14" pageNumber="315" type="Museum">MRAC 146342</specimenCode>
,
<specimenCode id="B0089E85F637C511FC596428713FCF0C" box="[930,1110,1815,1841]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="14" pageNumber="315" type="Museum">MRAC 146371</specimenCode>
,&gt;
<specimenCount id="F6A8FD77F637C511FB8F6429707BCF0C" box="[1140,1298,1814,1841]" pageId="14" pageNumber="315" type="generic">25 specimens</specimenCount>
</materialsCitation>
;
<materialsCitation id="50C63CA3F637C511FADA642870F1CF68" ID-GBIF-Occurrence="2423596343" collectingDate="02-07-26" collectionCode="MRAC" collectorName="E. Dartevelle. Probable" country="Republic of the Congo" county="Matadi" latitude="-5.881111" location="Chaudron d'Enfer" longLatPrecision="21" longitude="13.029166" municipality="River" pageId="14" pageNumber="315" specimenCode="MRAC 106951, DBL 2583, MRAC 107737" specimenCount="33" typeStatus="holotype">
“Chaudron dEnfer prés Matadi” (=
<location id="E5716025F637C511FE536404771BCF68" LSID="urn:lsid:plazi:treatment:680787E8F637C50CFF6C664371CACA02:E5716025F637C511FE536404771BCF68" box="[424,626,1851,1877]" country="Republic of the Congo" county="Matadi" latitude="-5.881111" longLatPrecision="21" longitude="13.029166" municipality="River" name="Chaudron d'Enfer" pageId="14" pageNumber="315">Chaudron dEnfer</location>
near
<location id="E5716025F637C511FD5664047796CF68" LSID="urn:lsid:plazi:treatment:680787E8F637C50CFF6C664371CACA02:E5716025F637C511FD5664047796CF68" box="[685,767,1851,1877]" country="Republic of the Congo" county="Matadi" latitude="-5.881111" longLatPrecision="21" longitude="13.029166" municipality="River" name="Matadi" pageId="14" pageNumber="315">Matadi</location>
,
<geoCoordinate id="859A5039F637C511FCF1640576EECF68" box="[778,903,1850,1877]" degrees="5" direction="south" minutes="49" orientation="latitude" pageId="14" pageNumber="315" precision="15" seconds="48" value="-5.83">5°4948” S</geoCoordinate>
,
<geoCoordinate id="859A5039F637C511FC686405714BCF6B" box="[915,1058,1850,1878]" degrees="13" direction="east" minutes="25" orientation="longitude" pageId="14" pageNumber="315" precision="15" seconds="54" value="13.431667">13°2554” E</geoCoordinate>
),
<specimenCode id="B0089E85F637C511FBCE6404718DCF68" box="[1077,1252,1850,1877]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="14" pageNumber="315" type="Museum">MRAC 106951</specimenCode>
,&gt;
<specimenCount id="F6A8FD77F637C511FB05640570F1CF68" box="[1278,1432,1850,1877]" pageId="14" pageNumber="315" type="generic">25 specimens</specimenCount>
</materialsCitation>
;
<materialsCitation id="50C63CA3F637C511FF6C646071C5CF44" ID-GBIF-Occurrence="2423596348" box="[151,1196,1886,1914]" collectingDate="02-07-26" collectionCode="MRAC" collectorName="E. Dartevelle. Probable" country="Republic of the Congo" county="Matadi" latitude="-5.881111" location="River" longLatPrecision="21" longitude="13.029166" municipality="River" pageId="14" pageNumber="315" specimenCode="DBL 2583" specimenCount="3" typeStatus="holotype">
“Matadi” (=
<collectingCountry id="98B9766EF637C511FED26460741DCF44" box="[297,372,1887,1913]" name="Democratic Republic of the Congo" pageId="14" pageNumber="315">Congo</collectingCountry>
<location id="E5716025F637C511FE80646074D0CF44" LSID="urn:lsid:plazi:treatment:680787E8F637C50CFF6C664371CACA02:E5716025F637C511FE80646074D0CF44" box="[379,441,1887,1913]" country="Republic of the Congo" county="Matadi" latitude="-5.881111" longLatPrecision="21" longitude="13.029166" municipality="River" name="River" pageId="14" pageNumber="315">River</location>
at
<location id="E5716025F637C511FE2564607759CF44" LSID="urn:lsid:plazi:treatment:680787E8F637C50CFF6C664371CACA02:E5716025F637C511FE2564607759CF44" box="[478,560,1887,1913]" country="Republic of the Congo" county="Matadi" latitude="-5.881111" longLatPrecision="21" longitude="13.029166" municipality="River" name="Matadi" pageId="14" pageNumber="315">Matadi</location>
, ca.
<geoCoordinate id="859A5039F637C511FD9F6461778DCF44" box="[612,740,1886,1913]" degrees="5" direction="south" minutes="49" orientation="latitude" pageId="14" pageNumber="315" precision="15" seconds="15" value="-5.820833">5°4915” S</geoCoordinate>
,
<geoCoordinate id="859A5039F637C511FD0A646076EBCF47" box="[753,898,1887,1914]" degrees="13" direction="east" minutes="27" orientation="longitude" pageId="14" pageNumber="315" precision="15" seconds="00" value="13.45">13°2700” E</geoCoordinate>
),
<specimenCode id="B0089E85F637C511FC63645F717BCF44" box="[920,1042,1886,1914]" collectionCode="DBL" pageId="14" pageNumber="315">DBL 2583</specimenCode>
,
<specimenCount id="F6A8FD77F637C511FBE5646071C5CF44" box="[1054,1196,1887,1913]" pageId="14" pageNumber="315" type="generic">3 specimens</specimenCount>
</materialsCitation>
;
<materialsCitation id="50C63CA3F637C511FB4364607129CFA0" ID-GBIF-Occurrence="2423596351" collectingDate="02-07-26" collectionCode="MRAC" collectorName="E. Dartevelle. Probable" country="Republic of the Congo" county="Matadi" latitude="-5.881111" location="River" longLatPrecision="21" longitude="13.029166" municipality="River" pageId="14" pageNumber="315" specimenCode="MRAC 107737" specimenCount="5" typeStatus="holotype">
<collectingCountry id="98B9766EF637C511FB386460707DCF44" box="[1219,1300,1887,1913]" name="Democratic Republic of the Congo" pageId="14" pageNumber="315">Congo</collectingCountry>
: Matadi” (=
<collectingCountry id="98B9766EF637C511FF6C64BC758BCFA0" box="[151,226,1923,1949]" name="Democratic Republic of the Congo" pageId="14" pageNumber="315">Congo</collectingCountry>
<location id="E5716025F637C511FF1364BC744FCFA0" LSID="urn:lsid:plazi:treatment:680787E8F637C50CFF6C664371CACA02:E5716025F637C511FF1364BC744FCFA0" box="[232,294,1923,1949]" country="Republic of the Congo" county="Matadi" latitude="-5.881111" longLatPrecision="21" longitude="13.029166" municipality="River" name="River" pageId="14" pageNumber="315">River</location>
at
<location id="E5716025F637C511FEB364BC74F3CFA0" LSID="urn:lsid:plazi:treatment:680787E8F637C50CFF6C664371CACA02:E5716025F637C511FEB364BC74F3CFA0" box="[328,410,1923,1949]" country="Republic of the Congo" county="Matadi" latitude="-5.881111" longLatPrecision="21" longitude="13.029166" municipality="River" name="Matadi" pageId="14" pageNumber="315">Matadi</location>
, ca.
<geoCoordinate id="859A5039F637C511FE3064BD7723CFA0" box="[459,586,1922,1949]" degrees="5" direction="south" minutes="49" orientation="latitude" pageId="14" pageNumber="315" precision="15" seconds="15" value="-5.820833">5°4915” S</geoCoordinate>
,
<geoCoordinate id="859A5039F637C511FDAD64BC778CCFA3" box="[598,741,1923,1950]" degrees="13" direction="east" minutes="27" orientation="longitude" pageId="14" pageNumber="315" precision="15" seconds="00" value="13.45">13°2700” E</geoCoordinate>
),
<specimenCode id="B0089E85F637C511FD0264BC76C0CFA0" box="[761,937,1923,1949]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="14" pageNumber="315" type="Museum">MRAC 107737</specimenCode>
,
<specimenCount id="F6A8FD77F637C511FC4F64BD7129CFA0" box="[948,1088,1922,1949]" pageId="14" pageNumber="315" type="generic">5 specimens</specimenCount>
</materialsCitation>
;
<materialsCitation id="50C63CA3F637C511FBB064BC7127CFFC" ID-GBIF-Occurrence="2423596344" collectingDate="02-07-26" collectionCode="MRAC" collectorName="E. Dartevelle. Probable" country="Republic of the Congo" county="Matadi" latitude="-5.881111" location="River" longLatPrecision="21" longitude="13.029166" municipality="River" pageId="14" pageNumber="315" specimenCode="MRAC 146372, MRAC 146426" specimenCount="22" typeStatus="holotype">
“Matadi, ilot” (=
<collectingCountry id="98B9766EF637C511FAF664BC7031CFA0" box="[1293,1368,1923,1949]" name="Democratic Republic of the Congo" pageId="14" pageNumber="315">Congo</collectingCountry>
<collectingMunicipality id="0075AC84F637C511FAA564BC70F5CFA0" box="[1374,1436,1923,1949]" pageId="14" pageNumber="315">River</collectingMunicipality>
at
<collectingCounty id="09704E72F637C511FF496498746DCFFC" box="[178,260,1959,1985]" pageId="14" pageNumber="315">Matadi</collectingCounty>
, small island, ca.
<geoCoordinate id="859A5039F637C511FE3164997721CFFC" box="[458,584,1958,1985]" degrees="5" direction="south" minutes="49" orientation="latitude" pageId="14" pageNumber="315" precision="15" seconds="15" value="-5.820833">5°4915” S</geoCoordinate>
,
<geoCoordinate id="859A5039F637C511FDAE6498778DCFFF" box="[597,740,1959,1986]" degrees="13" direction="east" minutes="27" orientation="longitude" pageId="14" pageNumber="315" precision="15" seconds="00" value="13.45">13°2700” E</geoCoordinate>
),
<collectionCode id="86BFAE3BF637C511FD0364987623CFFC" box="[760,842,1959,1985]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="14" pageNumber="315" type="Museum">MRAC</collectionCode>
<date id="9410103EF637C511FCAA649876C1CFFC" box="[849,936,1959,1985]" pageId="14" pageNumber="315" value="02-07-26">
<collectingDate id="8454E9D6F637C511FCAA649876C1CFFC" box="[849,936,1959,1985]" pageId="14" pageNumber="315" value="02-07-26">260702</collectingDate>
</date>
,
<specimenCount id="F6A8FD77F637C511FC4864987127CFFC" box="[947,1102,1959,1985]" pageId="14" pageNumber="315" type="generic">13 specimens</specimenCount>
</materialsCitation>
;
<materialsCitation id="50C63CA3F637C511FBA2649976A6CFD8" ID-GBIF-Occurrence="2423596347" collectingDate="02-07-26" collectionCode="MRAC" collectorName="E. Dartevelle. Probable" country="Republic of the Congo" county="Matadi" latitude="-5.881111" location="River" longLatPrecision="21" longitude="13.029166" municipality="River" pageId="14" pageNumber="315" specimenCode="MRAC 146372, MRAC 146426" specimenCount="22" typeStatus="holotype">
“Île Selonga” (= Île Selonga,
<geoCoordinate id="859A5039F637C511FF6C64F5747FCFD8" box="[151,278,1994,2021]" degrees="5" direction="south" minutes="52" orientation="latitude" pageId="14" pageNumber="315" precision="15" seconds="52" value="-5.881111">5°5252” S</geoCoordinate>
,
<geoCoordinate id="859A5039F637C511FED864F574DACFDB" box="[291,435,1994,2022]" degrees="13" direction="east" minutes="01" orientation="longitude" pageId="14" pageNumber="315" precision="15" seconds="45" value="13.029166">13°0145” E</geoCoordinate>
),
<specimenCode id="B0089E85F637C511FE3C64F47711CFD8" box="[455,632,1995,2021]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="14" pageNumber="315" type="Museum">MRAC 146372</specimenCode>
,
<specimenCode id="B0089E85F637C511FD7864F4765ACFD8" box="[643,819,1995,2021]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="14" pageNumber="315" type="Museum">MRAC 146426</specimenCode>
,
<specimenCount id="F6A8FD77F637C511FCC564F476A2CFD8" box="[830,971,1995,2021]" pageId="14" pageNumber="315" type="generic">9 specimens</specimenCount>
.
</materialsCitation>
</paragraph>
<caption id="B4D16676F636C510FF6C60D477C9CC71" ID-DOI="http://doi.org/10.5281/zenodo.3460156" ID-Zenodo-Dep="3460156" httpUri="https://zenodo.org/record/3460156/files/figure.png" pageId="15" pageNumber="316" startId="15.[151,250,1003,1028]" targetBox="[253,1334,174,982]" targetPageId="15">
<paragraph id="E01136FEF636C510FF6C60D477C9CC71" blockId="15.[151,1437,1003,1100]" pageId="15" pageNumber="316">
<emphasis id="D2DAEAECF636C510FF6C60D47476CC39" bold="true" box="[151,287,1003,1028]" pageId="15" pageNumber="316">FIGURE 12.</emphasis>
<taxonomicName id="27AE4D7DF636C510FEDE60D47404CC39" box="[293,365,1003,1028]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="15" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
of
<taxonomicName id="27AE4D7DF636C510FE7560D47604CC39" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" box="[398,877,1003,1028]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="15" pageNumber="316" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF636C510FE7560D477E2CC3E" box="[398,651,1003,1027]" italics="true" pageId="15" pageNumber="316">Pseudocleopatra voltana</emphasis>
<bibRefCitation id="843F4B0FF636C510FD6A60D47604CC39" author="Mandahl-Barth, G." box="[657,877,1003,1028]" pageId="15" pageNumber="316" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
</taxonomicName>
, paratype, Apaso village, DBL 2592.
<emphasis id="D2DAEAECF636C510FB1360D47069CC39" bold="true" box="[1256,1280,1003,1028]" pageId="15" pageNumber="316">A.</emphasis>
<taxonomicName id="27AE4D7DF636C510FAFD60D47027CC39" box="[1286,1358,1003,1028]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="15" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
ribbon, scale bar: 25 μm.
<emphasis id="D2DAEAECF636C510FEB6672F740DCC15" bold="true" box="[333,356,1040,1064]" pageId="15" pageNumber="316">B.</emphasis>
Lateral and central teeth; scale bar: 5 μm.
<emphasis id="D2DAEAECF636C510FCE26730765BCC15" bold="true" box="[793,818,1039,1064]" pageId="15" pageNumber="316">C.</emphasis>
Cutting edge of lateral and central teeth; scale bar: 3 μm.
<emphasis id="D2DAEAECF636C510FA7F672F70F5CC15" bold="true" box="[1412,1436,1040,1064]" pageId="15" pageNumber="316">D.</emphasis>
Central, lateral and marginal teeth; scale bar: 5 μm.
</paragraph>
</caption>
<caption id="B4D16676F636C510FF6C64A4771ECFC0" ID-DOI="http://doi.org/10.5281/zenodo.3460158" ID-Zenodo-Dep="3460158" httpUri="https://zenodo.org/record/3460158/files/figure.png" pageId="15" pageNumber="316" startId="15.[151,250,1947,1972]" targetBox="[250,1337,1116,1928]" targetPageId="15">
<paragraph id="E01136FEF636C510FF6C64A4771ECFC0" blockId="15.[151,1437,1947,2045]" pageId="15" pageNumber="316">
<emphasis id="D2DAEAECF636C510FF6C64A47476CF89" bold="true" box="[151,287,1947,1972]" pageId="15" pageNumber="316">FIGURE 13.</emphasis>
<taxonomicName id="27AE4D7DF636C510FEDD64A47407CF89" box="[294,366,1947,1972]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="15" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
of
<taxonomicName id="27AE4D7DF636C510FE7464A37606CF89" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" box="[399,879,1947,1972]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="15" pageNumber="316" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF636C510FE7464A377E5CF89" box="[399,652,1948,1972]" italics="true" pageId="15" pageNumber="316">Pseudocleopatra voltana</emphasis>
<bibRefCitation id="843F4B0FF636C510FD6964A47606CF89" author="Mandahl-Barth, G." box="[658,879,1947,1972]" pageId="15" pageNumber="316" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
</taxonomicName>
, paratype, Daboya, DBL 2594.
<emphasis id="D2DAEAECF636C510FB5564A471AECF89" bold="true" box="[1198,1223,1947,1972]" pageId="15" pageNumber="316">A.</emphasis>
<taxonomicName id="27AE4D7DF636C510FB3664A4707CCF89" box="[1229,1301,1947,1972]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="15" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
ribbon, scale bar: 25 μm.
<emphasis id="D2DAEAECF636C510FEE864FF7443CFE5" bold="true" box="[275,298,1984,2008]" pageId="15" pageNumber="316">B.</emphasis>
Lateral and central teeth; scale bar: 20 μm.
<emphasis id="D2DAEAECF636C510FD1C64807669CFE5" bold="true" box="[743,768,1983,2008]" pageId="15" pageNumber="316">C.</emphasis>
Cutting edge of lateral and central teeth; scale bar: 6 μm.
<emphasis id="D2DAEAECF636C510FAB764FF700CCFE5" bold="true" box="[1356,1381,1984,2008]" pageId="15" pageNumber="316">D.</emphasis>
Central, lateral and marginal teeth; scale bar: 8 μm.
</paragraph>
</caption>
<caption id="B4D16676F629C50FFF6C60117416CBB2" ID-DOI="http://doi.org/10.5281/zenodo.3460162" ID-Zenodo-Dep="3460162" httpUri="https://zenodo.org/record/3460162/files/figure.png" pageId="16" pageNumber="317" startId="16.[151,250,814,839]" targetBox="[486,1101,174,795]" targetPageId="16">
<paragraph id="E01136FEF629C50FFF6C60117416CBB2" blockId="16.[151,1437,814,911]" pageId="16" pageNumber="317">
<emphasis id="D2DAEAECF629C50FFF6C60117449CB7A" bold="true" box="[151,288,814,839]" pageId="16" pageNumber="317">FIGURE 14.</emphasis>
Known occurrence records of
<taxonomicName id="27AE4D7DF629C50FFDA760107155CB7A" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" box="[604,1084,814,839]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="16" pageNumber="317" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF629C50FFDA760107633CB7A" box="[604,858,815,839]" italics="true" pageId="16" pageNumber="317">Pseudocleopatra voltana</emphasis>
<bibRefCitation id="843F4B0FF629C50FFC9A60117155CB7A" author="Mandahl-Barth, G." box="[865,1084,814,839]" pageId="16" pageNumber="317" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
</taxonomicName>
in the Volta River system. The asterisk indicates the type locality; open symbols indicate samples of dry material. The arrow indicates the approximate location of the Akosombo dam.
</paragraph>
</caption>
<paragraph id="E01136FEF629C50FFF3C60DE71FFCCB6" blockId="16.[151,1436,993,1164]" pageId="16" pageNumber="317">
<emphasis id="D2DAEAECF629C50FFF3C60DE741CCBC1" bold="true" box="[199,373,993,1020]" pageId="16" pageNumber="317">
Shell (
<figureCitation id="78952A7BF629C50FFEE960DE740ECBC1" box="[274,359,993,1020]" captionStart="FIGURE 15" captionStartId="16.[151,250,1903,1928]" captionTargetBox="[456,1135,1225,1869]" captionTargetId="figure@16.[441,1146,1214,1884]" captionTargetPageId="16" captionText="FIGURE 15. Shells of Pseudocleopatra dartevellei Mandahl-Barth, 1973. A. Holotype, MRAC 799161. B. Probable paratype, Matadi; MRAC 260702. CD. Probable paratypes, Matadi, MRAC 260702, growth series. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3460164" httpUri="https://zenodo.org/record/3460164/files/figure.png" pageId="16" pageNumber="317">Fig. 15</figureCitation>
).
</emphasis>
The shell of
<taxonomicName id="27AE4D7DF629C50FFDF160DD77CBCBC1" authorityName="Mandahl-Barth" authorityYear="1973" box="[522,674,994,1020]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="16" pageNumber="317" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF629C50FFDF160DD7772CBC6" box="[522,539,994,1019]" italics="true" pageId="16" pageNumber="317">P</emphasis>
.
<emphasis id="D2DAEAECF629C50FFDD160DD77CBCBC1" box="[554,674,994,1020]" italics="true" pageId="16" pageNumber="317">dartevellei</emphasis>
</taxonomicName>
is yellowish brown to olive and has an ovate to conical shape, with up to six whorls (usually with eroded spire, then only about three whorls remaining). The whorls are rather convex, with very fine growth lines crossed by indistinct spiral lines. The aperture is holostome and ovate, higher than wide and slightly drawn out at the upper angle; outer margin sinuous in lateral view, columella curved and the basal margin evenly rounded (somewhat produced in the most mature specimens). Umbilicus closed.
</paragraph>
</subSubSection>
<caption id="B4D16676F629C50FFF6C64507188CF96" ID-DOI="http://doi.org/10.5281/zenodo.3460164" ID-Zenodo-Dep="3460164" httpUri="https://zenodo.org/record/3460164/files/figure.png" pageId="16" pageNumber="317" startId="16.[151,250,1903,1928]" targetBox="[456,1135,1225,1869]" targetPageId="16">
<paragraph id="E01136FEF629C50FFF6C64507188CF96" blockId="16.[151,1437,1903,1964]" pageId="16" pageNumber="317">
<emphasis id="D2DAEAECF629C50FFF6C64507476CFB5" bold="true" box="[151,287,1903,1928]" pageId="16" pageNumber="317">FIGURE 15.</emphasis>
Shells of
<taxonomicName id="27AE4D7DF629C50FFE7E644F76EFCFB5" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" box="[389,902,1903,1928]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="16" pageNumber="317" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF629C50FFE7E644F77CBCFB5" box="[389,674,1904,1928]" italics="true" pageId="16" pageNumber="317">Pseudocleopatra dartevellei</emphasis>
<bibRefCitation id="843F4B0FF629C50FFD52645076EFCFB5" author="Mandahl-Barth, G." box="[681,902,1903,1928]" pageId="16" pageNumber="317" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
</taxonomicName>
.
<emphasis id="D2DAEAECF629C50FFC74645076C1CFB5" bold="true" box="[911,936,1903,1928]" pageId="16" pageNumber="317">A.</emphasis>
Holotype, MRAC 799161.
<emphasis id="D2DAEAECF629C50FFB3A644F71B1CFB5" bold="true" box="[1217,1240,1904,1928]" pageId="16" pageNumber="317">B.</emphasis>
Probable paratype, Matadi; MRAC 260702.
<emphasis id="D2DAEAECF629C50FFE6D64AC74A7CF91" bold="true" box="[406,462,1939,1964]" pageId="16" pageNumber="317">CD.</emphasis>
Probable paratypes, Matadi, MRAC 260702, growth series. Scale bar: 5 mm.
</paragraph>
</caption>
<subSubSection id="A8B46575F628C50CFF3C63A67050C95A" lastPageId="19" lastPageNumber="320" pageId="17" pageNumber="318" type="description">
<paragraph id="E01136FEF628C50EFF3C63A67446C8C6" blockId="17.[151,1437,152,647]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFF3C63A6748CC88E" bold="true" box="[199,485,152,179]" pageId="17" pageNumber="318">
Measurements (
<emphasis id="D2DAEAECF628C50EFE7363A674F1C88E" bold="true" box="[392,408,153,179]" italics="true" pageId="17" pageNumber="318">n</emphasis>
= 80).
</emphasis>
h: 4.21 ±
<quantity id="27569B1BF628C50EFDA463A677ADC88E" box="[607,708,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="17" pageNumber="318" unit="mm" value="1.67">1.67 mm</quantity>
(range:
<quantity id="27569B1BF628C50EFCDF63A676D8C88E" box="[804,945,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="5.6499999999999995" metricValueMax="9.6" metricValueMin="1.7" pageId="17" pageNumber="318" unit="mm" value="5.6499999999999995" valueMax="9.6" valueMin="1.7">1.79.6 mm</quantity>
); w: 3.22 ±
<quantity id="27569B1BF628C50EFBC663A671CAC88E" box="[1085,1187,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.16" pageId="17" pageNumber="318" unit="mm" value="1.16">1.16 mm</quantity>
(range:
<quantity id="27569B1BF628C50EFAF863A670E6C88E" box="[1283,1423,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="6.8" metricValueMin="1.4" pageId="17" pageNumber="318" unit="mm" value="4.1" valueMax="6.8" valueMin="1.4">1.46.8 mm</quantity>
); bw: 3.66 ±
<quantity id="27569B1BF628C50EFEE0638274E9C8EA" box="[283,384,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" pageId="17" pageNumber="318" unit="mm" value="1.44">1.44 mm</quantity>
(range:
<quantity id="27569B1BF628C50EFE1B63827705C8EA" box="[480,620,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" metricValueMax="8.3" metricValueMin="1.1" pageId="17" pageNumber="318" unit="mm" value="4.7" valueMax="8.3" valueMin="1.1">1.18.3 mm</quantity>
); la: 2.42 ±
<quantity id="27569B1BF628C50EFD0C63827635C8EA" box="[759,860,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" pageId="17" pageNumber="318" unit="mm" value="1.04">1.04 mm</quantity>
(range:
<quantity id="27569B1BF628C50EFC4763837121C8EA" box="[956,1096,188,215]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="5.8" metricValueMin="0.9" pageId="17" pageNumber="318" unit="mm" value="3.35" valueMax="5.8" valueMin="0.9">0.95.8 mm</quantity>
); wa: 1.79 ±
<quantity id="27569B1BF628C50EFB246382702CC8EA" box="[1247,1349,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="17" pageNumber="318" unit="mm" value="0.68">0.68 mm</quantity>
(range:
<quantity id="27569B1BF628C50EFF6C63DE744BC8C6" box="[151,290,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="3.9" metricValueMin="0.8" pageId="17" pageNumber="318" unit="mm" value="2.35" valueMax="3.9" valueMin="0.8">0.83.9 mm</quantity>
).
</paragraph>
<paragraph id="E01136FEF628C50EFF3C623B71DEC922" blockId="17.[151,1437,152,647]" box="[199,1207,260,288]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFF3C623B7414C922" bold="true" box="[199,381,260,287]" pageId="17" pageNumber="318">
Ratios (
<emphasis id="D2DAEAECF628C50EFEDF623A745DC922" bold="true" box="[292,308,261,287]" italics="true" pageId="17" pageNumber="318">n</emphasis>
= 80).
</emphasis>
h/w: 1.30 ± 0.09 (range: 1.111.62); h/la: 1.77 ± 0.15 (range: 1.482.20).
</paragraph>
<paragraph id="E01136FEF628C50EFF3C6217760DC95A" blockId="17.[151,1437,152,647]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFF3C621774AEC97E" bold="true" box="[199,455,296,323]" pageId="17" pageNumber="318">
Operculum (
<figureCitation id="78952A7BF628C50EFEA5621774D1C97E" box="[350,440,296,323]" captionStart="FIGURE 8" captionStartId="11.[151,250,926,951]" captionTargetBox="[481,1112,201,877]" captionTargetId="figure@11.[466,1121,184,877]" captionTargetPageId="11" captionText="FIGURE 8. Opercula of Pseudocleopatra Thiele, 1928 species. A. Lectotype of P. togoensis Thiele, 1928, ZMB 190048b. B. Paratype of Pseudocleopatra voltana Mandahl-Barth, 1973, Mogeyenga village, DBL 2591. C. Probable paratype of Pseudocleopatra dartevellei Mandahl-Barth, 1973, Matadi, DBL 2583. D. Pseudocleopatra broecki (Putzeys, 1899) n. comb., MCZ 171520. E. Lectotype of Pseudocleopatra broecki (Putzeys, 1899) n. comb., MRAC 47360. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.3460148" httpUri="https://zenodo.org/record/3460148/files/figure.png" pageId="17" pageNumber="318">Fig. 8C</figureCitation>
).
</emphasis>
The operculum is corneous, with a paucispiral, rather large nucleus that can reach up to ⅔ of the maximum diameter and possesses a concentric margin.
</paragraph>
<paragraph id="E01136FEF628C50EFF3C624E7467CABA" blockId="17.[151,1437,152,647]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFF3C624E7743C9B6" bold="true" box="[199,554,368,395]" pageId="17" pageNumber="318">
<taxonomicName id="27AE4D7DF628C50EFF3C624E7477C9B6" box="[199,286,369,395]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="17" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
(
<figureCitation id="78952A7BF628C50EFED4624F74DBC9B6" box="[303,434,368,395]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStartId-0="17.[151,250,1938,1963]" captionStartId-1="18.[151,250,990,1015]" captionTargetBox-0="[151,1431,1020,1918]" captionTargetBox-1="[214,1374,174,971]" captionTargetId-0="figure@17.[170,1417,1033,1918]" captionTargetId-1="figure@18.[214,1374,174,971]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURE 16. Radula of Pseudocleopatra dartevellei Mandahl-Barth, 1973 (Morph a), probable paratype, Île des Princes, MRAC 146342. A. Radula ribbon, scale bar: 10 μm. B. Lateral and central teeth, scale bar: 10 μm. C. Cutting edge of lateral and central teeth, scale bar: 10 μm. D. Central, lateral and marginal teeth, scale bar: 10 μm." captionText-1="FIGURE 17. Radula of Pseudocleopatra dartevellei Mandahl-Barth, 1973 (Morph b), probable paratype, Île des Princes, MRAC 146342. A. Radula ribbon, scale bar: 10 μm. B. Lateral and central teeth, scale bar: 10 μm. C. Cutting edge of lateral and central teeth, scale bar: 2 μm. D. Central, lateral and marginal teeth, scale bar: 10 μm." figureDoi-0="http://doi.org/10.5281/zenodo.3460166" figureDoi-1="http://doi.org/10.5281/zenodo.3460168" httpUri-0="https://zenodo.org/record/3460166/files/figure.png" httpUri-1="https://zenodo.org/record/3460168/files/figure.png" pageId="17" pageNumber="318">Figs 1617</figureCitation>
,
<tableCitation id="AD2C0345F628C50EFE3B624E7772C9B6" box="[448,539,368,395]" captionStart="TABLE 2" captionStartId="17.[151,239,694,718]" captionTargetPageId="17" captionText="TABLE 2. Number of denticles on the different radula teeth of examined samples of Pseudocleopatra Thiele, 1928 and Cleopatra Troschel, 1857 species. Data for Cleopatra bulimoides (Olivier, 1804) from Glaubrecht (1996) and data for P." httpUri="http://table.plazi.org/id/B4D16676F628C50EFF6C618977F2CB2A" pageId="17" pageNumber="318" tableUuid="B4D16676F628C50EFF6C618977F2CB2A">Table 2</tableCitation>
).
</emphasis>
Two distinct radula
<typeStatus id="3F15885CF628C50EFCED624E7638C9B6" box="[790,849,369,395]" pageId="17" pageNumber="318">types</typeStatus>
were found referred to as “Morph a” (
<figureCitation id="78952A7BF628C50EFAF4624E700AC9B1" box="[1295,1379,369,396]" captionStart="FIGURE 16" captionStartId="17.[151,250,1938,1963]" captionTargetBox="[151,1431,1020,1918]" captionTargetId="figure@17.[170,1417,1033,1918]" captionTargetPageId="17" captionText="FIGURE 16. Radula of Pseudocleopatra dartevellei Mandahl-Barth, 1973 (Morph a), probable paratype, Île des Princes, MRAC 146342. A. Radula ribbon, scale bar: 10 μm. B. Lateral and central teeth, scale bar: 10 μm. C. Cutting edge of lateral and central teeth, scale bar: 10 μm. D. Central, lateral and marginal teeth, scale bar: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.3460166" httpUri="https://zenodo.org/record/3460166/files/figure.png" pageId="17" pageNumber="318">Fig. 16</figureCitation>
) and “Morph b” (
<figureCitation id="78952A7BF628C50EFEE462AA741BC992" box="[287,370,405,431]" captionStart="FIGURE 17" captionStartId="18.[151,250,990,1015]" captionTargetBox="[214,1374,174,971]" captionTargetId="figure@18.[214,1374,174,971]" captionTargetPageId="18" captionText="FIGURE 17. Radula of Pseudocleopatra dartevellei Mandahl-Barth, 1973 (Morph b), probable paratype, Île des Princes, MRAC 146342. A. Radula ribbon, scale bar: 10 μm. B. Lateral and central teeth, scale bar: 10 μm. C. Cutting edge of lateral and central teeth, scale bar: 2 μm. D. Central, lateral and marginal teeth, scale bar: 10 μm." figureDoi="http://doi.org/10.5281/zenodo.3460168" httpUri="https://zenodo.org/record/3460168/files/figure.png" pageId="17" pageNumber="318">Fig. 17</figureCitation>
). Morph a represents a typical paludomid radula. The rachidian tooth is squarish, with a straight upper edge. The cutting edge tapers to a distinct V-shaped tip and bears 56/1/68 denticles. The mesocone is very prominent and triangular in shape. It is about twice as long as wide and also wider than the lateral denticles, which decrease in size towards the margin of the tooth. The lateral teeth have very short and sinuate lateral extensions and possess 68/1/35 denticles. Their mesocone is very prominent, being twice as long as the flanking denticles; these decrease in size towards the outer margin of the teeth. The marginal teeth bear 1318 denticles of approximately equal size.
</paragraph>
<caption id="B4D16676F628C50EFF6C618977F2CB2A" ID-Table-UUID="B4D16676F628C50EFF6C618977F2CB2A" httpUri="http://table.plazi.org/id/B4D16676F628C50EFF6C618977F2CB2A" pageId="17" pageNumber="318" targetBox="[166,1392,802,1016]" targetIsTable="true" targetPageId="17">
<paragraph id="E01136FEF628C50EFF6C618977F2CB2A" blockId="17.[151,1437,694,791]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFF6C61897463CAF3" bold="true" box="[151,266,694,718]" pageId="17" pageNumber="318">TABLE 2.</emphasis>
Number of denticles on the different radula teeth of examined samples of
<taxonomicName id="27AE4D7DF628C50EFBD761887007CAF2" authority="Thiele, 1928" authorityName="Thiele" authorityYear="1928" box="[1068,1390,695,719]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="17" pageNumber="318" phylum="Mollusca" rank="genus">
<emphasis id="D2DAEAECF628C50EFBD761887188CAF2" box="[1068,1249,695,719]" italics="true" pageId="17" pageNumber="318">Pseudocleopatra</emphasis>
<bibRefCitation id="843F4B0FF628C50EFB1C61887007CAF2" author="Thiele, J." box="[1255,1390,695,719]" pageId="17" pageNumber="318" pagination="394 - 402" refId="ref16913" refString="Thiele, J. (1928) Revision des Systems der Hydrobiiden und Melaniiden. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 55, 394 - 402, pl. 8." type="journal article" year="1928">Thiele, 1928</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="27AE4D7DF628C50EFF6C61E474C1CACF" authority="Troschel, 1857" authorityName="Troschel" authorityYear="1857" box="[151,424,730,755]" class="Gastropoda" family="Thiaridae" genus="Cleopatra" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="318" phylum="Mollusca" rank="genus">
<emphasis id="D2DAEAECF628C50EFF6C61E4746ACACE" box="[151,259,731,755]" italics="true" pageId="17" pageNumber="318">Cleopatra</emphasis>
Troschel, 1857
</taxonomicName>
species. Data for
<taxonomicName id="27AE4D7DF628C50EFD9261E4769ECACE" authority="(Olivier, 1804)" authorityName="Olivier" authorityYear="1804" baseAuthorityName="Olivier" baseAuthorityYear="1804" box="[617,1015,731,755]" class="Gastropoda" family="Thiaridae" genus="Cleopatra" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="318" phylum="Mollusca" rank="species" species="bulimoides">
<emphasis id="D2DAEAECF628C50EFD9261E47638CACE" box="[617,849,731,755]" italics="true" pageId="17" pageNumber="318">Cleopatra bulimoides</emphasis>
(
<bibRefCitation id="843F4B0FF628C50EFCA461E47686CACE" author="Olivier, G. A." box="[863,1007,731,755]" pageId="17" pageNumber="318" refId="ref16156" refString="Olivier, G. A. (1804) Voyage dans l'Empire Othoman, l'Egypte et la Perse, fait par ordre du gouvernement, pendant les six premieres annees de la Republique. Tome Second. Agasse, Paris, ii + 466 pp." type="book" year="1804">Olivier, 1804</bibRefCitation>
)
</taxonomicName>
from
<bibRefCitation id="843F4B0FF628C50EFBCC61E47195CACF" author="Glaubrecht, M." box="[1079,1276,730,755]" pageId="17" pageNumber="318" refId="ref14270" refString="Glaubrecht, M. (1996) Evolutionsokologie und Systematik am Beispiel von Suss- und Brackwasserschnecken (Mollusca: Caenogastropoda: Cerithioidea): Ontogenese-Strategien, palaontologische Befunde und historische Zoogeographie. Backhuys Publishers, Leiden, xvi + 499 pp., 25 pls." type="book" year="1996">Glaubrecht (1996)</bibRefCitation>
and data for P.
<taxonomicName id="27AE4D7DF628C50EFF6C61C074EBCB2A" authority="Putzeys, 1899" authorityName="Putzeys" authorityYear="1899" box="[151,386,767,791]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="17" pageNumber="318" phylum="Mollusca" rank="species" species="broecki">
<emphasis id="D2DAEAECF628C50EFF6C61C0758FCB2A" box="[151,230,767,791]" italics="true" pageId="17" pageNumber="318">broecki</emphasis>
<bibRefCitation id="843F4B0FF628C50EFF1661C074EBCB2A" author="Putzeys, S." box="[237,386,767,791]" pageId="17" pageNumber="318" pagination="55 - 60" refId="ref16286" refString="Putzeys, S. (1899) Diagnoses de quelques coquilles et d'un sous-genre nouveau provenant de l'Etat independant du Congo. Annales de la Societe Royale malacologique de Belgique. Bulletins des Seances, 34, 55 - 60." type="journal article" year="1899">Putzeys, 1899</bibRefCitation>
</taxonomicName>
from
<bibRefCitation id="843F4B0FF628C50EFE3F61C077FECB2A" author="Glaubrecht, M." box="[452,663,767,791]" pageId="17" pageNumber="318" pagination="283 - 293" refId="ref14370" refString="Glaubrecht, M. (2010 a) The enigmatic Cleopatra broecki Putzeys, 1899 of the Congo River system in Africa-re-transfer from Potadomoides Leloup, 1953 (Caenogastropoda, Cerithioidea, Paludomidae). Zoosystematics and Evolution, 86, 283 - 293. https: // doi. org / 10.1002 / zoos. 201000011" type="journal article" year="2010">Glaubrecht (2010a)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="E01136FEF628C50EFF5D601D7184CBCA" pageId="17" pageNumber="318">
<table id="92AEC45EF6283AE0FF5D601D7019CBC5" box="[166,1392,802,1016]" gridcols="7" gridrows="6" pageId="17" pageNumber="318">
<tr id="5E9E34BCF6283AE0FF5D601D7019CB06" box="[166,1392,802,827]" gridrow="0" pageId="17" pageNumber="318">
<th id="1D4F5DC0F6283AE0FF5D601D744CCB06" box="[166,293,802,827]" gridcol="0" gridrow="0" pageId="17" pageNumber="318">
<taxonomicName id="27AE4D7DF628C50EFF5D601D7587CB06" box="[166,238,802,827]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="17" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
teeth
</th>
<th id="1D4F5DC0F6283AE0FE8B601D7487CB06" box="[368,494,802,827]" gridcol="1" gridrow="0" pageId="17" pageNumber="318">
<taxonomicName id="27AE4D7DF628C50EFE8B601C7487CB06" authorityName="Thiele" authorityYear="1928" box="[368,494,803,827]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="17" pageNumber="318" phylum="Mollusca" rank="species" species="togoensis">
<emphasis id="D2DAEAECF628C50EFE8B601C7487CB06" bold="true" box="[368,494,803,827]" italics="true" pageId="17" pageNumber="318">
P.
<emphasis id="D2DAEAECF628C50EFE76601C7487CB06" bold="true" box="[397,494,803,827]" italics="true" pageId="17" pageNumber="318">togoensis</emphasis>
</emphasis>
</taxonomicName>
</th>
<th id="1D4F5DC0F6283AE0FDD2601D77FBCB06" box="[553,658,802,827]" gridcol="2" gridrow="0" pageId="17" pageNumber="318">
<taxonomicName id="27AE4D7DF628C50EFDD2601C77FBCB06" authorityName="Mandahl-Barth" authorityYear="1973" box="[553,658,802,827]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="17" pageNumber="318" phylum="Mollusca" rank="species" species="voltana">
<emphasis id="D2DAEAECF628C50EFDD2601C77FBCB06" bold="true" box="[553,658,802,827]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFDD2601C7751CB07" bold="true" box="[553,568,803,826]" italics="true" pageId="17" pageNumber="318">P</emphasis>
.
<emphasis id="D2DAEAECF628C50EFDBE601D77FBCB06" bold="true" box="[581,658,802,827]" italics="true" pageId="17" pageNumber="318">voltana</emphasis>
</emphasis>
</taxonomicName>
</th>
<th id="1D4F5DC0F6283AE0FD29601D7632CB06" box="[722,859,802,827]" gridcol="3" gridrow="0" pageId="17" pageNumber="318">
<taxonomicName id="27AE4D7DF628C50EFD29601C7632CB06" authorityName="Mandahl-Barth" authorityYear="1973" box="[722,859,802,827]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="17" pageNumber="318" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF628C50EFD29601C7632CB06" bold="true" box="[722,859,802,827]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFD29601C7788CB07" bold="true" box="[722,737,803,826]" italics="true" pageId="17" pageNumber="318">P</emphasis>
.
<emphasis id="D2DAEAECF628C50EFD14601D7632CB06" bold="true" box="[751,859,802,827]" italics="true" pageId="17" pageNumber="318">dartevellei</emphasis>
</emphasis>
</taxonomicName>
</th>
<th id="1D4F5DC0F6283AE0FC68601D7175CB06" box="[915,1052,802,827]" gridcol="4" gridrow="0" pageId="17" pageNumber="318">
<taxonomicName id="27AE4D7DF628C50EFC68601C7175CB06" authorityName="Mandahl-Barth" authorityYear="1973" box="[915,1052,802,827]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="17" pageNumber="318" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF628C50EFC68601C7175CB06" bold="true" box="[915,1052,802,827]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFC68601C76CBCB07" bold="true" box="[915,930,803,826]" italics="true" pageId="17" pageNumber="318">P</emphasis>
.
<emphasis id="D2DAEAECF628C50EFC4B601D7175CB06" bold="true" box="[944,1052,802,827]" italics="true" pageId="17" pageNumber="318">dartevellei</emphasis>
</emphasis>
</taxonomicName>
</th>
<th id="1D4F5DC0F6283AE0FBAF601D71D7CB06" box="[1108,1214,802,827]" gridcol="5" gridrow="0" pageId="17" pageNumber="318">
<taxonomicName id="27AE4D7DF628C50EFBAF601C71D7CB06" authorityName="Putzeys" authorityYear="1899" box="[1108,1214,802,827]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="17" pageNumber="318" phylum="Mollusca" rank="species" species="broecki">
<emphasis id="D2DAEAECF628C50EFBAF601C71D7CB06" bold="true" box="[1108,1214,802,827]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFBAF601C710ACB07" bold="true" box="[1108,1123,803,826]" italics="true" pageId="17" pageNumber="318">P</emphasis>
.
<emphasis id="D2DAEAECF628C50EFB8A601D71D7CB06" bold="true" box="[1137,1214,802,827]" italics="true" pageId="17" pageNumber="318">broecki</emphasis>
</emphasis>
</taxonomicName>
</th>
<th id="1D4F5DC0F6283AE0FB1B601D7019CB06" box="[1248,1392,802,827]" gridcol="6" gridrow="0" pageId="17" pageNumber="318">
<taxonomicName id="27AE4D7DF628C50EFB1B601C7019CB06" baseAuthorityName="Olivier" baseAuthorityYear="1804" box="[1248,1392,802,827]" class="Gastropoda" family="Thiaridae" genus="Cleopatra" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="17" pageNumber="318" phylum="Mollusca" rank="species" species="bulimoides">
<emphasis id="D2DAEAECF628C50EFB1B601C7019CB06" bold="true" box="[1248,1392,802,827]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFB1B601C7198CB06" bold="true" box="[1248,1265,803,827]" italics="true" pageId="17" pageNumber="318">C</emphasis>
.
<emphasis id="D2DAEAECF628C50EFB05601D7019CB06" bold="true" box="[1278,1392,802,827]" italics="true" pageId="17" pageNumber="318">bulimoides</emphasis>
</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="5E9E34BCF6283AE0FF5D60767019CB5C" box="[166,1392,841,865]" gridrow="1" pageId="17" pageNumber="318" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1">
<td id="1D4F5DC0F6283AE0FD2960767632CB5C" box="[722,859,841,865]" gridcol="3" gridrow="1" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFD296076765BCB5C" bold="true" box="[722,818,841,865]" pageId="17" pageNumber="318">Morph a</emphasis>
</td>
<td id="1D4F5DC0F6283AE0FC6860767175CB5C" box="[915,1052,841,865]" gridcol="4" gridrow="1" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFC686076769DCB5C" bold="true" box="[915,1012,841,865]" pageId="17" pageNumber="318">Morph b</emphasis>
</td>
</tr>
<tr id="5E9E34BCF6283AE0FF5D60527019CBBB" box="[166,1392,877,902]" gridrow="2" pageId="17" pageNumber="318" rowspan-0="1">
<td id="1D4F5DC0F6283AE0FE8B60527487CBBB" box="[368,494,877,902]" gridcol="1" gridrow="2" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFE8B605074CECBBB" bold="true" box="[368,423,878,902]" italics="true" pageId="17" pageNumber="318">n = 7</emphasis>
</td>
<td id="1D4F5DC0F6283AE0FDD2605277FBCBBB" box="[553,658,877,902]" gridcol="2" gridrow="2" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFDD260507736CBBB" bold="true" box="[553,607,877,902]" italics="true" pageId="17" pageNumber="318">n = 2</emphasis>
</td>
<td id="1D4F5DC0F6283AE0FD2960527632CBBB" box="[722,859,877,902]" gridcol="3" gridrow="2" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFD2960507660CBBB" bold="true" box="[722,777,877,902]" italics="true" pageId="17" pageNumber="318">n = 2</emphasis>
</td>
<td id="1D4F5DC0F6283AE0FC6860527175CBBB" box="[915,1052,877,902]" gridcol="4" gridrow="2" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFC68605076A3CBBB" bold="true" box="[915,970,877,902]" italics="true" pageId="17" pageNumber="318">n = 8</emphasis>
</td>
<td id="1D4F5DC0F6283AE0FBAF605271D7CBBB" box="[1108,1214,877,902]" gridcol="5" gridrow="2" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFBAF605071E2CBBB" bold="true" box="[1108,1163,877,902]" italics="true" pageId="17" pageNumber="318">n = 2</emphasis>
</td>
<td id="1D4F5DC0F6283AE0FB1B60527019CBBB" box="[1248,1392,877,902]" gridcol="6" gridrow="2" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFB1B6050707FCBBB" bold="true" box="[1248,1302,877,902]" italics="true" pageId="17" pageNumber="318">n = 2</emphasis>
</td>
</tr>
<tr id="5E9E34BCF6283AE0FF5D60AC7019CB90" box="[166,1392,915,941]" gridrow="3" pageId="17" pageNumber="318">
<th id="1D4F5DC0F6283AE0FF5D60AC744CCB90" box="[166,293,915,941]" gridcol="0" gridrow="3" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFF5D60AC747CCB91" bold="true" box="[166,277,915,940]" pageId="17" pageNumber="318">Rachidian</emphasis>
</th>
<td id="1D4F5DC0F6283AE0FE8B60AC7487CB90" box="[368,494,915,941]" gridcol="1" gridrow="3" pageId="17" pageNumber="318">56/1/56</td>
<td id="1D4F5DC0F6283AE0FDD260AC77FBCB90" box="[553,658,915,941]" gridcol="2" gridrow="3" pageId="17" pageNumber="318">4-5/1/4-5</td>
<td id="1D4F5DC0F6283AE0FD2960AC7632CB90" box="[722,859,915,941]" gridcol="3" gridrow="3" pageId="17" pageNumber="318">56/1/68</td>
<td id="1D4F5DC0F6283AE0FC6860AC7175CB90" box="[915,1052,915,941]" gridcol="4" gridrow="3" pageId="17" pageNumber="318">45/1/45</td>
<td id="1D4F5DC0F6283AE0FBAF60AC71D7CB90" box="[1108,1214,915,941]" gridcol="5" gridrow="3" pageId="17" pageNumber="318">7/1/7</td>
<td id="1D4F5DC0F6283AE0FB1B60AC7019CB90" box="[1248,1392,915,941]" gridcol="6" gridrow="3" pageId="17" pageNumber="318">4/1/34</td>
</tr>
<tr id="5E9E34BCF6283AE0FF5D60867019CBEF" box="[166,1392,953,978]" gridrow="4" pageId="17" pageNumber="318">
<th id="1D4F5DC0F6283AE0FF5D6086744CCBEF" box="[166,293,953,978]" gridcol="0" gridrow="4" pageId="17" pageNumber="318">Laterals</th>
<td id="1D4F5DC0F6283AE0FE8B60867487CBEF" box="[368,494,953,978]" gridcol="1" gridrow="4" pageId="17" pageNumber="318">24/1/47</td>
<td id="1D4F5DC0F6283AE0FDD2608677FBCBEF" box="[553,658,953,978]" gridcol="2" gridrow="4" pageId="17" pageNumber="318">1215</td>
<td id="1D4F5DC0F6283AE0FD2960867632CBEF" box="[722,859,953,978]" gridcol="3" gridrow="4" pageId="17" pageNumber="318">68/1/35</td>
<td id="1D4F5DC0F6283AE0FC6860867175CBEF" box="[915,1052,953,978]" gridcol="4" gridrow="4" pageId="17" pageNumber="318">34/1/34</td>
<td id="1D4F5DC0F6283AE0FBAF608671D7CBEF" box="[1108,1214,953,978]" gridcol="5" gridrow="4" pageId="17" pageNumber="318">23/1/&gt;9</td>
<td id="1D4F5DC0F6283AE0FB1B60867019CBEF" box="[1248,1392,953,978]" gridcol="6" gridrow="4" pageId="17" pageNumber="318">45/1/34</td>
</tr>
<tr id="5E9E34BCF6283AE0FF5D60E07019CBC5" box="[166,1392,991,1016]" gridrow="5" pageId="17" pageNumber="318">
<th id="1D4F5DC0F6283AE0FF5D60E0744CCBC5" box="[166,293,991,1016]" gridcol="0" gridrow="5" pageId="17" pageNumber="318">Marginals</th>
<td id="1D4F5DC0F6283AE0FE8B60E07487CBC5" box="[368,494,991,1016]" gridcol="1" gridrow="5" pageId="17" pageNumber="318">67</td>
<td id="1D4F5DC0F6283AE0FDD260E077FBCBC5" box="[553,658,991,1016]" gridcol="2" gridrow="5" pageId="17" pageNumber="318">810</td>
<td id="1D4F5DC0F6283AE0FD2960E07632CBC5" box="[722,859,991,1016]" gridcol="3" gridrow="5" pageId="17" pageNumber="318">1318</td>
<td id="1D4F5DC0F6283AE0FC6860E07175CBC5" box="[915,1052,991,1016]" gridcol="4" gridrow="5" pageId="17" pageNumber="318">0</td>
<td id="1D4F5DC0F6283AE0FBAF60E071D7CBC5" box="[1108,1214,991,1016]" gridcol="5" gridrow="5" pageId="17" pageNumber="318">1820</td>
<td id="1D4F5DC0F6283AE0FB1B60E07019CBC5" box="[1248,1392,991,1016]" gridcol="6" gridrow="5" pageId="17" pageNumber="318">7</td>
</tr>
</table>
</paragraph>
<caption id="B4D16676F628C50EFF6C64AD715ACFCE" ID-DOI="http://doi.org/10.5281/zenodo.3460166" ID-Zenodo-Dep="3460166" httpUri="https://zenodo.org/record/3460166/files/figure.png" pageId="17" pageNumber="318" startId="17.[151,250,1938,1963]" targetBox="[151,1431,1020,1918]" targetPageId="17">
<paragraph id="E01136FEF628C50EFF6C64AD715ACFCE" blockId="17.[151,1437,1938,2035]" pageId="17" pageNumber="318">
<emphasis id="D2DAEAECF628C50EFF6C64AD744DCF96" bold="true" box="[151,292,1938,1963]" pageId="17" pageNumber="318">FIGURE 16.</emphasis>
<taxonomicName id="27AE4D7DF628C50EFED564AD741FCF96" box="[302,374,1938,1963]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="17" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
of
<taxonomicName id="27AE4D7DF628C50EFE6464AC76C3CF96" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" box="[415,938,1938,1963]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="17" pageNumber="318" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF628C50EFE6464AC77A8CF96" box="[415,705,1939,1963]" italics="true" pageId="17" pageNumber="318">Pseudocleopatra dartevellei</emphasis>
<bibRefCitation id="843F4B0FF628C50EFD3764AD76C3CF96" author="Mandahl-Barth, G." box="[716,938,1938,1963]" pageId="17" pageNumber="318" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
</taxonomicName>
(Morph a), probable paratype, Île des Princes, MRAC 146342.
<emphasis id="D2DAEAECF628C50EFEB864897432CFF2" bold="true" box="[323,347,1974,1999]" pageId="17" pageNumber="318">A.</emphasis>
<taxonomicName id="27AE4D7DF628C50EFE98648974C2CFF2" box="[355,427,1974,1999]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="17" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
ribbon, scale bar: 10 μm.
<emphasis id="D2DAEAECF628C50EFD41648877B8CFF2" bold="true" box="[698,721,1975,1999]" pageId="17" pageNumber="318">B.</emphasis>
Lateral and central teeth, scale bar: 10 μm.
<emphasis id="D2DAEAECF628C50EFB6D648971C6CFF2" bold="true" box="[1174,1199,1974,1999]" pageId="17" pageNumber="318">C.</emphasis>
Cutting edge of lateral and central teeth, scale bar: 10 μm.
<emphasis id="D2DAEAECF628C50EFDFB64E47771CFCE" bold="true" box="[512,536,2011,2035]" pageId="17" pageNumber="318">D.</emphasis>
Central, lateral and marginal teeth, scale bar: 10 μm.
</paragraph>
</caption>
<caption id="B4D16676F62BC50DFF6C60E1714FCC02" ID-DOI="http://doi.org/10.5281/zenodo.3460168" ID-Zenodo-Dep="3460168" httpUri="https://zenodo.org/record/3460168/files/figure.png" pageId="18" pageNumber="319" startId="18.[151,250,990,1015]" targetBox="[214,1374,174,971]" targetPageId="18">
<paragraph id="E01136FEF62BC50DFF6C60E1714FCC02" blockId="18.[151,1437,990,1087]" pageId="18" pageNumber="319">
<emphasis id="D2DAEAECF62BC50DFF6C60E1744DCBCA" bold="true" box="[151,292,990,1015]" pageId="18" pageNumber="319">FIGURE 17.</emphasis>
<taxonomicName id="27AE4D7DF62BC50DFED560E1741FCBCA" box="[302,374,990,1015]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="18" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
of
<taxonomicName id="27AE4D7DF62BC50DFE6460E076C0CBCA" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" box="[415,937,990,1015]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="18" pageNumber="319" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF62BC50DFE6460E077A8CBCA" box="[415,705,991,1015]" italics="true" pageId="18" pageNumber="319">Pseudocleopatra dartevellei</emphasis>
<bibRefCitation id="843F4B0FF62BC50DFD3060E176C0CBCA" author="Mandahl-Barth, G." box="[715,937,990,1015]" pageId="18" pageNumber="319" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
</taxonomicName>
(Morph b), probable paratype, Île des Princes, MRAC 146342.
<emphasis id="D2DAEAECF62BC50DFEB8673D7432CC26" bold="true" box="[323,347,1026,1051]" pageId="18" pageNumber="319">A.</emphasis>
<taxonomicName id="27AE4D7DF62BC50DFE98673D74C2CC26" box="[355,427,1026,1051]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="18" pageNumber="304" phylum="Bryophyta" rank="genus">Radula</taxonomicName>
ribbon, scale bar: 10 μm.
<emphasis id="D2DAEAECF62BC50DFD41673C77B8CC26" bold="true" box="[698,721,1027,1051]" pageId="18" pageNumber="319">B.</emphasis>
Lateral and central teeth, scale bar: 10 μm.
<emphasis id="D2DAEAECF62BC50DFB6D673D71C6CC26" bold="true" box="[1174,1199,1026,1051]" pageId="18" pageNumber="319">C.</emphasis>
Cutting edge of lateral and central teeth, scale bar: 2 μm.
<emphasis id="D2DAEAECF62BC50DFE0867187765CC02" bold="true" box="[499,524,1063,1087]" pageId="18" pageNumber="319">D.</emphasis>
Central, lateral and marginal teeth, scale bar: 10 μm.
</paragraph>
</caption>
<caption id="B4D16676F62BC50DFF6C65937673CF30" ID-DOI="http://doi.org/10.5281/zenodo.3460170" ID-Zenodo-Dep="3460170" httpUri="https://zenodo.org/record/3460170/files/figure.png" pageId="18" pageNumber="319" startId="18.[151,250,1708,1733]" targetBox="[482,1105,1102,1689]" targetPageId="18">
<paragraph id="E01136FEF62BC50DFF6C65937673CF30" blockId="18.[151,1437,1708,1805]" pageId="18" pageNumber="319">
<emphasis id="D2DAEAECF62BC50DFF6C65937449CEF8" bold="true" box="[151,288,1708,1733]" pageId="18" pageNumber="319">FIGURE 18.</emphasis>
Known occurrence records of
<taxonomicName id="27AE4D7DF62BC50DFDAC6592713FCEF8" authority="Mandahl-Barth, 1973" authorityName="Mandahl-Barth" authorityYear="1973" box="[599,1110,1708,1733]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="18" pageNumber="319" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF62BC50DFDAC6592761CCEF8" box="[599,885,1709,1733]" italics="true" pageId="18" pageNumber="319">Pseudocleopatra dartevellei</emphasis>
<bibRefCitation id="843F4B0FF62BC50DFC876593713FCEF8" author="Mandahl-Barth, G." box="[892,1110,1708,1733]" pageId="18" pageNumber="319" pagination="277 - 286" refId="ref15611" refString="Mandahl-Barth, G. (1973) Descriptions of new species of African freshwater mollusks. Proceedings of the Malacological Society of London, 40, 277 - 286." type="journal article" year="1973">Mandahl-Barth, 1973</bibRefCitation>
</taxonomicName>
in the Congo River system. The asterisk indicates the type locality; solid symbols indicate samples with alcohol material, open symbols those of dry material. The arrow indicates the approximate location of the Inga dams.
</paragraph>
</caption>
<paragraph id="E01136FEF62BC50CFF3C64607050C95A" blockId="18.[151,1436,1887,2022]" lastBlockId="19.[151,1437,153,575]" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319">Morph b differs significantly from morph a and possibly represents a teratological specimen. The rachidian tooth has a more triangular shape and a V-shaped upper edge. Its cutting edge also tapers to a V-shaped tip. Morph b differs from Morph a most significantly with regard to the denticles on the rachidian. The cutting edge bears 45/1/45 cuspid denticles. The mesocone is twice as long as the adjacent flanking denticles. Furthermore, three prominent denticles are present on each side of the lateral edges of the rachidian. The foremost of these denticles is twice as long as the next one, which in turn is slightly larger than the third one. The lateral teeth have short lateral extensions that are slightly curved. The cutting edge is V-shaped and bears 34/1/34 long and cuspid denticles. The mesocone is twice as long as the flanking denticles, which decrease in size towards the margin of the teeth. The marginal teeth are somewhat curved and stand in two serial rows. The inner marginal teeth have a sickle-shaped cutting edge, lacking any denticles, the outer marginal teeth bear 810 small denticles of about equal size.</paragraph>
</subSubSection>
<subSubSection id="A8B46575F62AC50CFF3C624F7464C992" pageId="19" pageNumber="320" type="distribution">
<paragraph id="E01136FEF62AC50CFF3C624F7464C992" blockId="19.[151,1437,153,575]" pageId="19" pageNumber="320">
<emphasis id="D2DAEAECF62AC50CFF3C624F74A5C9B6" bold="true" box="[199,460,368,395]" pageId="19" pageNumber="320">
Distribution (
<figureCitation id="78952A7BF62AC50CFE93624F74D4C9B6" box="[360,445,368,395]" captionStart="FIGURE 18" captionStartId="18.[151,250,1708,1733]" captionTargetBox="[482,1105,1102,1689]" captionTargetId="figure@18.[482,1105,1102,1689]" captionTargetPageId="18" captionText="FIGURE 18. Known occurrence records of Pseudocleopatra dartevellei Mandahl-Barth, 1973 in the Congo River system. The asterisk indicates the type locality; solid symbols indicate samples with alcohol material, open symbols those of dry material. The arrow indicates the approximate location of the Inga dams." figureDoi="http://doi.org/10.5281/zenodo.3460170" httpUri="https://zenodo.org/record/3460170/files/figure.png" pageId="19" pageNumber="320">Fig. 18</figureCitation>
).
</emphasis>
<taxonomicName id="27AE4D7DF62AC50CFE29624E7666C9B6" authorityName="Mandahl-Barth" authorityYear="1973" box="[466,783,369,395]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="19" pageNumber="320" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF62AC50CFE29624E7666C9B6" box="[466,783,369,395]" italics="true" pageId="19" pageNumber="320">Pseudocleopatra dartevellei</emphasis>
</taxonomicName>
is only known from the lower
<collectingCountry id="98B9766EF62AC50CFB92624E71DDC9B6" box="[1129,1204,369,395]" name="Democratic Republic of the Congo" pageId="19" pageNumber="320">Congo</collectingCountry>
River in the vicinity of Matadi.
</paragraph>
</subSubSection>
<subSubSection id="A8B46575F62AC50CFF3C628671CACA02" pageId="19" pageNumber="320" type="discussion">
<paragraph id="E01136FEF62AC50CFF3C628671CACA02" blockId="19.[151,1437,153,575]" pageId="19" pageNumber="320">
<emphasis id="D2DAEAECF62AC50CFF3C62867452C9EE" bold="true" box="[199,315,441,467]" pageId="19" pageNumber="320">Remarks.</emphasis>
The record of
<taxonomicName id="27AE4D7DF62AC50CFE2762857707C9EE" authorityName="Mandahl-Barth" authorityYear="1973" box="[476,622,441,467]" class="Gastropoda" family="Thiaridae" genus="Pseudocleopatra" kingdom="Animalia" pageId="19" pageNumber="320" phylum="Mollusca" rank="species" species="dartevellei">
<emphasis id="D2DAEAECF62AC50CFE2762857707C9EE" box="[476,622,441,467]" italics="true" pageId="19" pageNumber="320">P. dartevellei</emphasis>
</taxonomicName>
from the
<collectingRegion id="226AF81CF62AC50CFD236286763BC9EE" box="[728,850,441,467]" country="Sudan" name="An Nil al Abyad" pageId="19" pageNumber="320">White Nile</collectingRegion>
in
<collectingCountry id="98B9766EF62AC50CFC8F628676D3C9EE" box="[884,954,441,467]" name="Sudan" pageId="19" pageNumber="320">Sudan</collectingCountry>
(
<bibRefCitation id="843F4B0FF62AC50CFC3C62867132C9EE" author="Ahmed, A. A. E." box="[967,1115,441,467]" pageId="19" pageNumber="320" refId="ref13712" refString="Ahmed, A. A. E. (2014) Survey of the Snail fauna of the White Nile State, Sudan. Master Thesis, University of Khartoum, Khartoum, 53 pp." type="book" year="2014">Ahmed 2014</bibRefCitation>
) certainly belongs to another species. We interpret the aberrant radula of a probable
<typeStatus id="3F15885CF62AC50CFCF362E27601C9CA" box="[776,872,477,503]" pageId="19" pageNumber="320" type="paratype">paratype</typeStatus>
from Île des Princes (MRAC 146342) to be teratological as the shell was not different from other type specimens. Future studies should test this using molecular techniques. Nothing is known about the ecology of the species and new records do not exist.
</paragraph>
</subSubSection>
</treatment>
</document>