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<document id="6D3297673F4991CFE036FEFE9CF82F72" ID-CLB-Dataset="264236" ID-DOI="10.11646/zootaxa.5333.1.1" ID-GBIF-Dataset="d8e76b5e-3274-413a-afb8-0a55d8c205de" ID-ISSN="1175-5326" ID-Zenodo-Dep="8261139" ID-ZooBank="CC8CEE25-A7BD-48B3-B315-B67FB455748C" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1692349533096" checkinUser="plazi" docAuthor="Stonis, Jonas R., Diškus, Arūnas, Remeikis, Andrius, Orlovskytė, Svetlana, Solis, Alma, Paulavičiūtė, Brigita, Xu, Jiasheng &amp; Dai, Xiaohua" docDate="2023" docId="601087E8FFEEFF90989E5BC8FB659676" docLanguage="en" docName="zootaxa.5333.1.1.pdf" docOrigin="Zootaxa 5333 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5333.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Coptotriche sapaensis Diskus &amp; Stonis 2023, sp. nov." docType="treatment" docUuid="B6B226A0-C331-4838-86C3-7319D49BDC61" docUuidSource="ZooBank" docVersion="5" lastPageNumber="40" masterDocId="9C29FF90FFF3FFB798095932FFC6943D" masterDocTitle="Genera of Tischeriidae (Lepidoptera): a review of the global fauna, with descriptions of new taxa" masterLastPageNumber="131" masterPageNumber="1" pageNumber="30" updateTime="1699592551810" updateUser="ExternalLinkService">
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<mods:title id="9A0CE0695C244EF04994193236F30C66">Genera of Tischeriidae (Lepidoptera): a review of the global fauna, with descriptions of new taxa</mods:title>
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<mods:namePart id="A1C0B10D46EB530CBBC2127CFB7A9356">Diškus, Arūnas</mods:namePart>
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<mods:namePart id="8758966E6C48B3AB40ACA576A769BBF5">Remeikis, Andrius</mods:namePart>
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<mods:namePart id="63B952D4A6C93E9062DBEC18D8904FE1">Orlovskytė, Svetlana</mods:namePart>
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<mods:namePart id="4E0DF05837E2144B0063CF36DF9EF192">Solis, Alma</mods:namePart>
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<mods:namePart id="62C5810EF8AB5B2C0FA24B680E15952B">Paulavičiūtė, Brigita</mods:namePart>
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<mods:namePart id="36D32148B41F1D3AB5A1059649EF09DF">Xu, Jiasheng</mods:namePart>
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<mods:namePart id="468D76B1C0A8B2896D37BCEA32B7FC00">Dai, Xiaohua</mods:namePart>
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<heading id="B34E8192FFEEFFAA989E5BC8FD1A9729" bold="true" box="[151,732,762,789]" fontSize="11" level="2" pageId="29" pageNumber="30" reason="4">
<emphasis id="DACDEAECFFEEFFAA989E5BC8FD1A9729" bold="true" box="[151,732,762,789]" pageId="29" pageNumber="30">
<taxonomicName id="2FB94D7DFFEEFFAA989E5BC8FDB79729" ID-CoL="BS8Q5" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2023" box="[151,625,762,789]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="sapaensis" status="sp. nov.">
<emphasis id="DACDEAECFFEEFFAA989E5BC8FE669729" bold="true" box="[151,416,762,789]" italics="true" pageId="29" pageNumber="30">Coptotriche sapaensis</emphasis>
Diškus &amp; Stonis
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,
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urn:lsid:zoobank.org:act:
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(
<figureCitation id="70822A7BFFEEFFAA98965A70FEFD9761" box="[159,315,834,860]" captionStart-0="FIGURES 114120" captionStart-1="FIGURES 121125" captionStart-2="FIGURES 126130" captionStart-3="FIGURES 131135" captionStart-4="FIGURES 136142" captionStartId-0="34.[151,264,1933,1957]" captionStartId-1="35.[151,264,1920,1944]" captionStartId-2="36.[151,264,1922,1946]" captionStartId-3="37.[151,264,1924,1948]" captionStartId-4="38.[151,264,1924,1948]" captionTargetBox-0="[157,1425,189,1901]" captionTargetBox-1="[158,1424,193,1889]" captionTargetBox-2="[176,1407,198,1888]" captionTargetBox-3="[158,1425,188,1894]" captionTargetBox-4="[163,1423,193,1891]" captionTargetId-0="figure-25@34.[154,1434,181,1909]" captionTargetId-1="figure-20@35.[154,1434,181,1896]" captionTargetId-2="figure-25@36.[156,1432,181,1898]" captionTargetId-3="figure-20@37.[154,1433,181,1901]" captionTargetId-4="figure-25@38.[154,1433,181,1901]" captionTargetPageId-0="34" captionTargetPageId-1="35" captionTargetPageId-2="36" captionTargetPageId-3="37" captionTargetPageId-4="38" captionText-0="FIGURES 114120. Adults of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam. 114116, head; 117120 general view of holotype and paratypes (MfN)" captionText-1="FIGURES 121125. Male genitalia of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam, holotype, genitalia slide no. AD1111 (MfN). 121, socii and apex of phallus; 122, uncus and apical part of phallus; 123, general view; 124, 125, spines of phallus" captionText-2="FIGURES 126130. Male genitalia of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam, paratype, genitalia slide no. AD1106 (MfN). 126, 127, socii, tegumen and spined diaphragm; 128, valva and transtilla, ventral view; 129, valva, lateral view; 130, phallus" captionText-3="FIGURES 131135. Female genitalia of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam, paratype, genitalia slide no. AD1109 (MfN). 131, general view; 132, ovipositor lobes; 133, rounded accessory sac and proximal part of ductus spermathecae; 134, ovipositor lobes; 135, accessory sac and “neck” (slender proximal part) of corpus bursae" captionText-4="FIGURES 136142. Bionomics of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov. 136, 137, habitat, elevation 1920 m, 15 km NW of Sapa, 22°2058”N, 103°4616”E, Lao Cai Prov., northern Vietnam; 138142, leaf mines on unidentified Symplocaceae plant" figureDoi-0="http://doi.org/10.5281/zenodo.8261186" figureDoi-1="http://doi.org/10.5281/zenodo.8261188" figureDoi-2="http://doi.org/10.5281/zenodo.8261190" figureDoi-3="http://doi.org/10.5281/zenodo.8261192" figureDoi-4="http://doi.org/10.5281/zenodo.8261194" httpUri-0="https://zenodo.org/record/8261186/files/figure.png" httpUri-1="https://zenodo.org/record/8261188/files/figure.png" httpUri-2="https://zenodo.org/record/8261190/files/figure.png" httpUri-3="https://zenodo.org/record/8261192/files/figure.png" httpUri-4="https://zenodo.org/record/8261194/files/figure.png" pageId="29" pageNumber="30">Figs 114142</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A0A36575FFEEFFAA989E5AB8FDF3902D" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="E80636FEFFEEFFAA989E5AB8FDF3902D" blockId="29.[151,1437,906,2012]" pageId="29" pageNumber="30">
<emphasis id="DACDEAECFFEEFFAA989E5AB8FE819799" bold="true" box="[151,327,906,932]" pageId="29" pageNumber="30">Type material.</emphasis>
<materialsCitation id="58D13CA3FFEEFFAA99465AB8FF2297D1" ID-GBIF-Occurrence="4178999301" collectingDate="2015-02-20" collectingDateMax="2015-03" collectingDateMin="2015-02-20" collectionCode="MfN" collectorName="A. Diskus" country="Vietnam" elevation="1920" latitude="22.349445" location="Sapa" longLatPrecision="21" longitude="103.77112" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Lao Cai" typeStatus="holotype">
<typeStatus id="3702885CFFEEFFAA99465AB8FE7A9799" box="[335,444,906,932]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
: Ô,
<collectingCountry id="90AE766EFFEEFFAA99E55AB8FDB49799" box="[492,626,906,932]" name="Vietnam" pageId="29" pageNumber="30">VIETNAM</collectingCountry>
:
<collectingRegion id="2A7DF81CFFEEFFAA9A775AB9FCD99798" box="[638,799,906,933]" country="Vietnam" name="Lao Cai" pageId="29" pageNumber="30">Lao Cai Prov.</collectingRegion>
,
<quantity id="2F419B1BFFEEFFAA9B225AB8FCB29799" box="[811,884,906,933]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="km" value="15.0">15 km</quantity>
NW
<location id="ED666025FFEEFFAA9BBA5AB8FC2A9799" LSID="urn:lsid:plazi:treatment:601087E8FFEEFF90989E5BC8FB659676:ED666025FFEEFFAA9BBA5AB8FC2A9799" box="[947,1004,906,932]" country="Vietnam" latitude="22.349445" longLatPrecision="21" longitude="103.77112" name="Sapa" pageId="29" pageNumber="30" stateProvince="Lao Cai">Sapa</location>
,
<geoCoordinate id="8D8D5039FFEEFFAA9BF05AB8FB439798" box="[1017,1157,906,933]" degrees="22" direction="north" minutes="20" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="58" value="22.349445">22°2058”N</geoCoordinate>
,
<geoCoordinate id="8D8D5039FFEEFFAA9C9B5AB8FAEF9799" box="[1170,1321,906,932]" degrees="103" direction="east" minutes="46" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="16" value="103.77112">103°4616”E</geoCoordinate>
,
<elevation id="6394D1CDFFEEFFAA9D3F5AB8FF2997F4" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="29" pageNumber="30" unit="m" value="1920.0">
elevation
<quantity id="2F419B1BFFEEFFAA989E5A9CFF2997F4" box="[151,239,942,969]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="29" pageNumber="30" unit="m" value="1920.0">1920 m</quantity>
</elevation>
, feeding larva
<date id="9C07103EFFEEFFAA999E5A9CFDCD97F4" box="[407,523,942,969]" pageId="29" pageNumber="30" value="2015-02-20">
<collectingDate id="8C43E9D6FFEEFFAA999E5A9CFDCD97F4" box="[407,523,942,969]" pageId="29" pageNumber="30" value="2015-02-20">20.ii.2015</collectingDate>
</date>
, ex pupa
<date id="9C07103EFFEEFFAA9A715A9CFD1697F4" box="[632,720,942,969]" pageId="29" pageNumber="30" value="2015-03">
<collectingDate id="8C43E9D6FFEEFFAA9A715A9CFD1697F4" box="[632,720,942,969]" pageId="29" pageNumber="30" value="2015-03">iii.2015</collectingDate>
</date>
, field card no. 5203, leg.
<collectorName id="454C5328FFEEFFAA9BF85A9CFBA397F5" box="[1009,1125,942,968]" pageId="29" pageNumber="30">A. Diškus</collectorName>
, genitalia slide no. AD1111 (
<collectionCode id="8EA8AE3BFFEEFFAA98965AE0FF1A97D1" box="[159,220,978,1004]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="29" pageNumber="30" type="Museum">MfN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="58D13CA3FFEEFFAA98E75AE1FB8397D1" ID-GBIF-Occurrence="4178999317" box="[238,1093,978,1007]" collectingDate="2015-02-20" collectingDateMax="2015-03" collectingDateMin="2015-02-20" collectionCode="MfN" collectorName="A. Diskus" country="Vietnam" elevation="1920" latitude="22.349445" location="Sapa" longLatPrecision="21" longitude="103.77112" pageId="29" pageNumber="30" specimenCount="2" specimenCount-female="2" stateProvince="Lao Cai" typeStatus="paratype">
<typeStatus id="3702885CFFEEFFAA98E75AE1FE9997D0" box="[238,351,979,1005]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
: 1 Ô,
<specimenCount id="FEBFFD77FFEEFFAA99A85AE0FE0D97D0" box="[417,459,978,1005]" pageId="29" pageNumber="30" type="female">2 ♀</specimenCount>
same label data as holotype, genitalia slide no. AD1109
</materialsCitation>
<materialsCitation id="58D13CA3FFEEFFAA9C4C5AE0FDF7902D" ID-GBIF-Occurrence="4178999318" collectionCode="MfN" country="Vietnam" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai" typeStatus="paratype">
<specimenCount id="FEBFFD77FFEEFFAA9C4C5AE0FB9597D1" box="[1093,1107,978,1004]" pageId="29" pageNumber="30" type="female"></specimenCount>
(
<collectionCode id="8EA8AE3BFFEEFFAA9C685AE0FB5897D1" box="[1121,1182,978,1004]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="29" pageNumber="30" type="Museum">MfN</collectionCode>
); 1 Ô, same label data, genitalia slide no. AD1106Ô (GNU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A0A36575FFEEFFAA98CE5D28FB3E90D5" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="E80636FEFFEEFFAA98CE5D28FB3E90D5" blockId="29.[151,1437,906,2012]" pageId="29" pageNumber="30">
<emphasis id="DACDEAECFFEEFFAA98CE5D28FE849009" bold="true" box="[199,322,1050,1076]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
Externally, this new species is similar to and can be confused with some other bright ochre-colored
<taxonomicName id="2FB94D7DFFEEFFAA989E5D0CFEB29065" box="[151,372,1086,1112]" pageId="29" pageNumber="30">
<emphasis id="DACDEAECFFEEFFAA989E5D0CFEDB9065" box="[151,285,1086,1112]" italics="true" pageId="29" pageNumber="30">Coptotriche</emphasis>
species
</taxonomicName>
with distinctive black markings on the forewing apex, including the South East Asian
<taxonomicName id="2FB94D7DFFEEFFAA9D3B5D0DFE889041" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis NEPTI" authorityYear="2021" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="asiana">
<emphasis id="DACDEAECFFEEFFAA9D3B5D0DFA5B9065" box="[1330,1437,1087,1113]" italics="true" pageId="29" pageNumber="30">C. asiana</emphasis>
Diškus &amp; Stonis
</taxonomicName>
and
<taxonomicName id="2FB94D7DFFEEFFAA998C5D51FD5F9040" authority="Xu &amp; Dai. In" authorityName="Xu &amp; Dai. In" authorityYear="2021" box="[389,665,1122,1149]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="turpinia">
<emphasis id="DACDEAECFFEEFFAA998C5D51FE399041" box="[389,511,1123,1149]" italics="true" pageId="29" pageNumber="30">C. turpinia</emphasis>
Xu &amp; Dai. In
</taxonomicName>
the male genitalia, this new species can be easily differentiated from all congeneric species by the phallus with curved lateral clusters of large spines and unique morphology of phallus apex (see
<figureCitation id="70822A7BFFEEFFAA99045D98FEB790F9" box="[269,369,1194,1220]" captionStart="FIGURES 121125" captionStartId="35.[151,264,1920,1944]" captionTargetBox="[158,1424,193,1889]" captionTargetId="figure-20@35.[154,1434,181,1896]" captionTargetPageId="35" captionText="FIGURES 121125. Male genitalia of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam, holotype, genitalia slide no. AD1111 (MfN). 121, socii and apex of phallus; 122, uncus and apical part of phallus; 123, general view; 124, 125, spines of phallus" figureDoi="http://doi.org/10.5281/zenodo.8261188" httpUri="https://zenodo.org/record/8261188/files/figure.png" pageId="29" pageNumber="30">Fig. 122</figureCitation>
). In the female genitalia, the new species is characterized by the combination of a distinctly globular, heavily folded accessory sac, stout prela, and inconspicuous pectinations of corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="A0A36575FFEEFF9098CE5DC0FA5395E6" lastPageId="39" lastPageNumber="40" pageId="29" pageNumber="30" type="description">
<paragraph id="E80636FEFFEEFFAA98CE5DC0FDC09131" blockId="29.[151,1437,906,2012]" box="[199,518,1266,1292]" pageId="29" pageNumber="30">
<emphasis id="DACDEAECFFEEFFAA98CE5DC0FEB69131" bold="true" box="[199,368,1266,1292]" pageId="29" pageNumber="30">DNA barcode.</emphasis>
Unavailable.
</paragraph>
<paragraph id="E80636FEFFEEFFAA98CE5C24FD249281" blockId="29.[151,1437,906,2012]" pageId="29" pageNumber="30">
<emphasis id="DACDEAECFFEEFFAA98CE5C24FE9F910D" bold="true" box="[199,345,1302,1328]" pageId="29" pageNumber="30">Description.</emphasis>
<emphasis id="DACDEAECFFEEFFAA99695C24FE5C910D" box="[352,410,1302,1328]" italics="true" pageId="29" pageNumber="30">Male</emphasis>
(
<figureCitation id="70822A7BFFEEFFAA99A05C24FD80910D" box="[425,582,1302,1328]" captionStart="FIGURES 114120" captionStartId="34.[151,264,1933,1957]" captionTargetBox="[157,1425,189,1901]" captionTargetId="figure-25@34.[154,1434,181,1909]" captionTargetPageId="34" captionText="FIGURES 114120. Adults of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam. 114116, head; 117120 general view of holotype and paratypes (MfN)" figureDoi="http://doi.org/10.5281/zenodo.8261186" httpUri="https://zenodo.org/record/8261186/files/figure.png" pageId="29" pageNumber="30">Figs 114118</figureCitation>
). Forewing length
<quantity id="2F419B1BFFEEFFAA9B105C24FCB5910C" box="[793,883,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="29" pageNumber="30" unit="mm" value="4.5">4.5 mm</quantity>
; wingspan
<quantity id="2F419B1BFFEEFFAA9BF85C24FB86910C" box="[1009,1088,1302,1329]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="mm" value="10.0">10 mm</quantity>
(n = 1). Head: frons, palpi and pecten glossy yellow cream; frontal tuft comprised of golden cream lamellar scales and dark brown, long, piliformlike scales (
<figureCitation id="70822A7BFFEEFFAA991D5C6CFE6D9144" box="[276,427,1374,1401]" captionStart="FIGURES 114120" captionStartId="34.[151,264,1933,1957]" captionTargetBox="[157,1425,189,1901]" captionTargetId="figure-25@34.[154,1434,181,1909]" captionTargetPageId="34" captionText="FIGURES 114120. Adults of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam. 114116, head; 117120 general view of holotype and paratypes (MfN)" figureDoi="http://doi.org/10.5281/zenodo.8261186" httpUri="https://zenodo.org/record/8261186/files/figure.png" pageId="29" pageNumber="30">Figs 114, 115</figureCitation>
); collar ochreous yellow; antenna significantly longer than one half of the length of forewing; flagellum pale ochre-yellow; sensilla whitish cream, very fine, almost indistinctive. Thorax: bright ochreous yellow, with dark brown scales medially; tegula ochreous yellow. Forewing bright ochreous yellow, with some brown-black to black scales which scattered along costal margin and especially along dorsal margin and apex, and form a small irregular spot on tornus; fringe intense yellow-ochre, ochre-grey on tornus; fringe line absent; forewing underside with ochreous brown scales, except for the large, yellowish cream area below the fold. Hindwing dark brown-grey on upper side and underside; fringe ochre-brown. Legs ochre-yellow, densely covered with dark grey-brown scales on upper side. Abdomen dark brown with a very, strong golden gloss and some purple irridescence on upper side, pale brown, golden glossy on underside, yellow cream laterally; genital segments medium large, yellow cream; anal tuft dorsal, small, comprised of lamellar grey scales.
</paragraph>
<paragraph id="E80636FEFFEEFFAA98CE5FF4FAAB9339" blockId="29.[151,1437,906,2012]" pageId="29" pageNumber="30">
<emphasis id="DACDEAECFFEEFFAA98CE5FF4FEDD92DD" box="[199,283,1734,1760]" italics="true" pageId="29" pageNumber="30">Female</emphasis>
(
<figureCitation id="70822A7BFFEEFFAA99225FF4FE0A92DD" box="[299,460,1734,1760]" captionStart="FIGURES 114120" captionStartId="34.[151,264,1933,1957]" captionTargetBox="[157,1425,189,1901]" captionTargetId="figure-25@34.[154,1434,181,1909]" captionTargetPageId="34" captionText="FIGURES 114120. Adults of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam. 114116, head; 117120 general view of holotype and paratypes (MfN)" figureDoi="http://doi.org/10.5281/zenodo.8261186" httpUri="https://zenodo.org/record/8261186/files/figure.png" pageId="29" pageNumber="30">Figs 119, 120</figureCitation>
). Forewing length 5.0
<quantity id="2F419B1BFFEEFFAA9ADF5FF4FCE992DC" box="[726,815,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="29" pageNumber="30" unit="mm" value="5.1">5.1 mm</quantity>
; wingspan
<quantity id="2F419B1BFFEEFFAA9BB95FF4FB9492DC" box="[944,1106,1734,1761]" metricMagnitude="-2" metricUnit="m" metricValue="1.1149999999999998" metricValueMax="1.1199999999999999" metricValueMin="1.1099999999999999" pageId="29" pageNumber="30" unit="mm" value="11.149999999999999" valueMax="11.2" valueMin="11.1">11.111.2 mm</quantity>
(n = 2). Similar to male, but without antennal sensilla, anal tuft yellowish cream; ovipositor not protruding. Otherwise, identical with male.
</paragraph>
<paragraph id="E80636FEFFEEFFAA98CE5E3CFD7893E1" blockId="29.[151,1437,906,2012]" pageId="29" pageNumber="30">
<emphasis id="DACDEAECFFEEFFAA98CE5E3CFEAB9315" box="[199,365,1806,1832]" italics="true" pageId="29" pageNumber="30">Male genitalia</emphasis>
(
<figureCitation id="70822A7BFFEEFFAA99745E3CFDDC9315" box="[381,538,1806,1832]" captionStart-0="FIGURES 121125" captionStart-1="FIGURES 126130" captionStartId-0="35.[151,264,1920,1944]" captionStartId-1="36.[151,264,1922,1946]" captionTargetBox-0="[158,1424,193,1889]" captionTargetBox-1="[176,1407,198,1888]" captionTargetId-0="figure-20@35.[154,1434,181,1896]" captionTargetId-1="figure-25@36.[156,1432,181,1898]" captionTargetPageId-0="35" captionTargetPageId-1="36" captionText-0="FIGURES 121125. Male genitalia of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam, holotype, genitalia slide no. AD1111 (MfN). 121, socii and apex of phallus; 122, uncus and apical part of phallus; 123, general view; 124, 125, spines of phallus" captionText-1="FIGURES 126130. Male genitalia of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam, paratype, genitalia slide no. AD1106 (MfN). 126, 127, socii, tegumen and spined diaphragm; 128, valva and transtilla, ventral view; 129, valva, lateral view; 130, phallus" figureDoi-0="http://doi.org/10.5281/zenodo.8261188" figureDoi-1="http://doi.org/10.5281/zenodo.8261190" httpUri-0="https://zenodo.org/record/8261188/files/figure.png" httpUri-1="https://zenodo.org/record/8261190/files/figure.png" pageId="29" pageNumber="30">Figs 121130</figureCitation>
) with capsule about 730 µm long, 380385 µm wide. Uncus comprised of two large, triangular lobes. Socii relatively large, weakly paired, membranous. Tegumen very, short medially, with long lateral arms. Diaphragm covered with numerous tiny spines. Valva ca. 700 µm long, wide in lateral view. Transtilla with a long transverse bar and short sublateral processes. Anellus mostly membranous, indistinctive. Vinculum short, triangular. Phallus about 770 µm long, apically tulip-shaped, with three distinctive rod-like thickenings and lateral clusters of large, outwardly curved spines.
</paragraph>
<caption id="BCC66676FFEDFFA9989E5E51FB6D93CC" ID-DOI="http://doi.org/10.5281/zenodo.8261178" ID-Zenodo-Dep="8261178" httpUri="https://zenodo.org/record/8261178/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,264,1891,1915]" targetBox="[162,1412,175,1872]" targetPageId="30" targetType="figure">
<paragraph id="E80636FEFFEDFFA9989E5E51FB6D93CC" blockId="30.[151,1436,1891,2033]" pageId="30" pageNumber="31">
<emphasis id="DACDEAECFFEDFFA9989E5E51FE929346" bold="true" box="[151,340,1891,1915]" pageId="30" pageNumber="31">FIGURES 8087.</emphasis>
Male genital capsule of
<taxonomicName id="2FB94D7DFFEDFFA99A425E51FD029346" authorityName="Walsingham" authorityYear="1890" box="[587,708,1891,1915]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="DACDEAECFFEDFFA99A425E51FD029346" box="[587,708,1891,1915]" italics="true" pageId="30" pageNumber="31">Coptotriche</emphasis>
</taxonomicName>
. 80,
<taxonomicName id="2FB94D7DFFEDFFA99AFF5E56FBF59346" authority="Diskus &amp; Stonis, Far" authorityName="Diskus &amp; Stonis, Far" authorityYear="2014" box="[758,1075,1891,1916]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="DACDEAECFFEDFFA99AFF5E56FC9F9346" box="[758,857,1892,1915]" italics="true" pageId="30" pageNumber="31">C. minuta</emphasis>
Diškus &amp; Stonis, Far
</taxonomicName>
Eastern Russia (
<bibRefCitation id="8C284B0FFFEDFFA99CD25E51FA579341" author="Stonis, J. R. &amp; Diskus, A. &amp; Rociene, A. &amp; Sruoga, V. &amp; Davis, D. R." box="[1243,1425,1891,1916]" pageId="30" pageNumber="31" pagination="141 - 155" refId="ref51667" refString="Stonis, J. R., Diskus, A., Rociene, A., Sruoga, V. &amp; Davis, D. R. (2014) New and little known Coptotriche and Tischeria species (Lepidoptera: Tischeriidae) from Primorskiy Kray, Russian Far East. Zootaxa, 3884 (2), 141 - 155. https: // doi. org / 10.11646 / zootaxa. 3884.2.3" type="journal article" year="2014">
Stonis
<emphasis id="DACDEAECFFEDFFA99D2B5E56FA939346" box="[1314,1365,1891,1915]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2014
</bibRefCitation>
); 81,
<taxonomicName id="2FB94D7DFFEDFFA998B65EB3FE3893A4" authority="(Clemens)" baseAuthorityName="Clemens" box="[191,510,1921,1945]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="citripenipennella">
<emphasis id="DACDEAECFFEDFFA998B65EB3FE4D93A4" box="[191,395,1921,1945]" italics="true" pageId="30" pageNumber="31">C. citripenipennella</emphasis>
(Clemens)
</taxonomicName>
, USA, slide no. 28961 BMNH; 82,
<taxonomicName id="2FB94D7DFFEDFFA99B7A5EB3FB4B93A4" authority="(Duponchel)" baseAuthorityName="Duponchel" box="[883,1165,1921,1945]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="gaunacella">
<emphasis id="DACDEAECFFEDFFA99B7A5EB3FBC593A4" box="[883,1027,1921,1945]" italics="true" pageId="30" pageNumber="31">C. gaunacella</emphasis>
(Duponchel)
</taxonomicName>
, Turkmenistan, a genitalia mount in glycerin (MfN); 83,
<taxonomicName id="2FB94D7DFFEDFFA999C35EADFCC2938B" authority="Puplesis &amp; Diskus" authorityName="Puplesis &amp; Diskus" authorityYear="2003" box="[458,772,1950,1974]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="africana">
<emphasis id="DACDEAECFFEDFFA999C35EADFDF8938B" box="[458,574,1950,1974]" italics="true" pageId="30" pageNumber="31">C. africana</emphasis>
Puplesis &amp; Diškus
</taxonomicName>
, South Africa (
<bibRefCitation id="8C284B0FFFEDFFA99B975EACFBA4938B" author="Stonis, J. R. &amp; Diskus, A. &amp; Mey, W." box="[926,1122,1950,1974]" pageId="30" pageNumber="31" refId="ref50640" refString="Stonis, J. R., Diskus, A. &amp; Mey, W. (2019 a) Tischeriidae (Lepidoptera: Tischerioidea). In: Mey, W. &amp; Kruger, M. (Eds.), The Lepidoptera fauna of a crater valley in the Great escarpment of South Africa: The Asante Sana Project. Esperiana Memoir. Vol. 8. Esperiana-Verlag, Deutschland, pp. 85 - 98. [http: // www. esperiana. net / esperiana. html]" type="url" year="2019">
Stonis
<emphasis id="DACDEAECFFEDFFA99BEF5EADFBDC938B" box="[998,1050,1950,1974]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2019a
</bibRefCitation>
); 84,
<taxonomicName id="2FB94D7DFFEDFFA99C925EADFAA6938B" authority="(Braun)" baseAuthorityName="Braun" baseAuthorityYear="1972" box="[1179,1376,1950,1974]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="DACDEAECFFEDFFA99C925EADFACE938B" box="[1179,1288,1950,1974]" italics="true" pageId="30" pageNumber="31">C. confusa</emphasis>
(Braun)
</taxonomicName>
, USA (
<bibRefCitation id="8C284B0FFFEDFFA998975E89FEA193E9" author="Stonis, J. R. &amp; Remeikis, A. &amp; Diskus, A. &amp; Davis, D. R. &amp; Solis, M. A." box="[158,359,1979,2004]" pageId="30" pageNumber="31" pagination="482 - 505" refId="ref52106" refString="Stonis, J. R., Remeikis, A., Diskus, A., Davis, D. R. &amp; Solis, M. A. (2020 c) American Tischeriidae (Lepidoptera) from the collection of the National Museum of Natural History, Washington D. C. Proceedings of the Entomological Society of Washington, 122 (2), 482 - 505. https: // doi. org / 10.4289 / 0013 - 8797.122.2.482" type="journal article" year="2020">
Stonis
<emphasis id="DACDEAECFFEDFFA998EE5E8EFEDB93EE" box="[231,285,1979,2003]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2020c
</bibRefCitation>
); 85,
<taxonomicName id="2FB94D7DFFEDFFA999AA5E8EFCC993EE" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis L" authorityYear="2544" box="[419,783,1979,2004]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="parvisacculata">
<emphasis id="DACDEAECFFEDFFA999AA5E8EFD9F93EE" box="[419,601,1979,2003]" italics="true" pageId="30" pageNumber="31">C. parvisacculata</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<taxonomicName id="2FB94D7DFFEDFFA99B105E89FB9993EE" authority="(Stonis et al. 2019 b)" baseAuthorityName="Stonis" baseAuthorityYear="2019" box="[793,1119,1979,2003]" class="Magnoliopsida" family="Rosaceae" genus="Argentina" kingdom="Plantae" order="Rosales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="genus">
Argentina (
<bibRefCitation id="8C284B0FFFEDFFA99B875E89FB9E93EE" author="Stonis, J. R. &amp; Diskus, A. &amp; Remeikis, A. &amp; Katinas, L. &amp; Cumbicus Torres, N. &amp; Schuster, J. &amp; Puplesyte-Chambers, J." box="[910,1112,1979,2003]" pageId="30" pageNumber="31" pagination="1 - 32" refId="ref51357" refString="Stonis, J. R., Diskus, A., Remeikis, A., Katinas, L., Cumbicus Torres, N., Schuster, J. &amp; Puplesyte-Chambers, J. (2019 b) Diagnostics of new species of Neotropical Tischeriidae (Lepidoptera), with the first record of Coptotriche Walshingham from South America. Zootaxa, 4691 (1), 1 - 32. https: // doi. org / 10.11646 / zootaxa. 4691.1.1" type="journal article" year="2019">
Stonis
<emphasis id="DACDEAECFFEDFFA99BDE5E8EFBCA93EE" box="[983,1036,1979,2003]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2019b
</bibRefCitation>
)
</taxonomicName>
; 86,
<taxonomicName id="2FB94D7DFFEDFFA99C9D5E8EFA5E93EE" authority="(Rebel)" baseAuthorityName="Rebel" box="[1172,1432,1979,2003]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="longiciliatella">
<emphasis id="DACDEAECFFEDFFA99C9D5E8EFA8793EE" box="[1172,1345,1979,2003]" italics="true" pageId="30" pageNumber="31">C. longiciliatella</emphasis>
(Rebel)
</taxonomicName>
, Tenerife, Canary Is., slide no. Diškus088 ZMUC; 87,
<taxonomicName id="2FB94D7DFFEDFFA99AB15EEBFCB493CC" authority="Diskus" authorityName="Diskus" authorityYear="1998" box="[696,882,2009,2033]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="paulula">
<emphasis id="DACDEAECFFEDFFA99AB15EEBFCE593CC" box="[696,803,2009,2033]" italics="true" pageId="30" pageNumber="31">C. paulula</emphasis>
Diškus
</taxonomicName>
, Tajikistan (after
<bibRefCitation id="8C284B0FFFEDFFA99C2A5EEBFB6293CC" author="Diskus, A." box="[1059,1188,2009,2033]" pageId="30" pageNumber="31" pagination="23 - 33" refId="ref47096" refString="Diskus, A. (1998) Review of the Tischeriidae (Lepidoptera) of Central Asia. Acta Zoologica Lituanica, 8 (3), 23 - 33. https: // doi. org / 10.1080 / 13921657.1998.10512245" type="journal article" year="1998">Diškus 1998</bibRefCitation>
)
</paragraph>
</caption>
<caption id="BCC66676FFECFFA8989E5E7EFA9D93C8" ID-DOI="http://doi.org/10.5281/zenodo.8261180" ID-Zenodo-Dep="8261180" httpUri="https://zenodo.org/record/8261180/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,255,1868,1892]" targetBox="[173,1421,166,1852]" targetPageId="31" targetType="figure">
<paragraph id="E80636FEFFECFFA8989E5E7EFA9D93C8" blockId="31.[151,1437,1868,2037]" pageId="31" pageNumber="32">
<emphasis id="DACDEAECFFECFFA8989E5E7EFE849358" bold="true" box="[151,322,1868,1893]" pageId="31" pageNumber="32">FIGURES 8898.</emphasis>
Valva and phallus of
<taxonomicName id="2FB94D7DFFECFFA89A0B5E7FFDB79358" authorityName="Walsingham" authorityYear="1890" box="[514,625,1869,1893]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="DACDEAECFFECFFA89A0B5E7FFDB79358" box="[514,625,1869,1893]" italics="true" pageId="31" pageNumber="32">Coptotriche</emphasis>
</taxonomicName>
. 88,
<taxonomicName id="2FB94D7DFFECFFA89A955E7FFCBC9358" authority="(De Prins)" baseAuthorityName="De Prins" baseAuthorityYear="1984" box="[668,890,1869,1893]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="berberella">
<emphasis id="DACDEAECFFECFFA89A955E7FFCD39358" box="[668,789,1869,1893]" italics="true" pageId="31" pageNumber="32">C. berberella</emphasis>
(De Prins)
</taxonomicName>
, Marocco,
<emphasis id="DACDEAECFFECFFA89BD75E7EFAD29358" bold="true" box="[990,1300,1868,1893]" pageId="31" pageNumber="32">first documentation of paratype</emphasis>
no. Diškus093 ZMUC; 89,
<taxonomicName id="2FB94D7DFFECFFA899005E43FE3393B4" authority="(Rebel)" baseAuthorityName="Rebel" box="[265,501,1905,1929]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="longiciliatella">
<emphasis id="DACDEAECFFECFFA899005E43FE6193B4" box="[265,423,1905,1929]" italics="true" pageId="31" pageNumber="32">C. longiciliatella</emphasis>
(Rebel)
</taxonomicName>
, Tenerife, Canary Is., slide no. Diškus133 ZMUC; 90,
<taxonomicName id="2FB94D7DFFECFFA89BFB5E43FAFE93B4" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis L" authorityYear="2544" box="[1010,1336,1905,1929]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="parvisacculata">
<emphasis id="DACDEAECFFECFFA89BFB5E43FB5E93B4" box="[1010,1176,1905,1929]" italics="true" pageId="31" pageNumber="32">C. parvisacculata</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<taxonomicName id="2FB94D7DFFECFFA89D365E43FE909390" authority="(Stonis et al. 2019 b)" baseAuthorityName="Stonis" baseAuthorityYear="2019" class="Magnoliopsida" family="Rosaceae" genus="Argentina" kingdom="Plantae" order="Rosales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="genus">
Argentina (
<bibRefCitation id="8C284B0FFFECFFA898975EA7FE899390" author="Stonis, J. R. &amp; Diskus, A. &amp; Remeikis, A. &amp; Katinas, L. &amp; Cumbicus Torres, N. &amp; Schuster, J. &amp; Puplesyte-Chambers, J." box="[158,335,1941,1965]" pageId="31" pageNumber="32" pagination="1 - 32" refId="ref51357" refString="Stonis, J. R., Diskus, A., Remeikis, A., Katinas, L., Cumbicus Torres, N., Schuster, J. &amp; Puplesyte-Chambers, J. (2019 b) Diagnostics of new species of Neotropical Tischeriidae (Lepidoptera), with the first record of Coptotriche Walshingham from South America. Zootaxa, 4691 (1), 1 - 32. https: // doi. org / 10.11646 / zootaxa. 4691.1.1" type="journal article" year="2019">
Stonis
<emphasis id="DACDEAECFFECFFA898D75EA7FECB9390" box="[222,269,1941,1965]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
2019b
</bibRefCitation>
)
</taxonomicName>
; 91,
<taxonomicName id="2FB94D7DFFECFFA899885EA7FDED9390" authority="Diskus" authorityName="Diskus" authorityYear="1998" box="[385,555,1941,1965]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="paulula">
<emphasis id="DACDEAECFFECFFA899885EA7FE249390" box="[385,482,1941,1965]" italics="true" pageId="31" pageNumber="32">C. paulula</emphasis>
Diškus
</taxonomicName>
, Tajikistan (
<bibRefCitation id="8C284B0FFFECFFA89A935EA7FCD59390" author="Diskus, A." box="[666,787,1941,1965]" pageId="31" pageNumber="32" pagination="23 - 33" refId="ref47096" refString="Diskus, A. (1998) Review of the Tischeriidae (Lepidoptera) of Central Asia. Acta Zoologica Lituanica, 8 (3), 23 - 33. https: // doi. org / 10.1080 / 13921657.1998.10512245" type="journal article" year="1998">Diškus 1998</bibRefCitation>
); 92,
<taxonomicName id="2FB94D7DFFECFFA89B4C5EA7FC3E9390" authority="(Braun)" baseAuthorityName="Braun" baseAuthorityYear="1972" box="[837,1016,1941,1965]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="DACDEAECFFECFFA89B4C5EA7FC6F9390" box="[837,937,1941,1965]" italics="true" pageId="31" pageNumber="32">C. confusa</emphasis>
(Braun)
</taxonomicName>
, USA (
<bibRefCitation id="8C284B0FFFECFFA89C335EA7FB2C9390" author="Stonis, J. R. &amp; Remeikis, A. &amp; Diskus, A. &amp; Davis, D. R. &amp; Solis, M. A." box="[1082,1258,1941,1965]" pageId="31" pageNumber="32" pagination="482 - 505" refId="ref52106" refString="Stonis, J. R., Remeikis, A., Diskus, A., Davis, D. R. &amp; Solis, M. A. (2020 c) American Tischeriidae (Lepidoptera) from the collection of the National Museum of Natural History, Washington D. C. Proceedings of the Entomological Society of Washington, 122 (2), 482 - 505. https: // doi. org / 10.4289 / 0013 - 8797.122.2.482" type="journal article" year="2020">
Stonis
<emphasis id="DACDEAECFFECFFA89C735EA7FB6F9390" box="[1146,1193,1941,1965]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
2020c
</bibRefCitation>
); 93,
<taxonomicName id="2FB94D7DFFECFFA89D155EA7FA509390" baseAuthorityName="De Prins" baseAuthorityYear="1984" box="[1308,1430,1941,1965]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="berberella">
<emphasis id="DACDEAECFFECFFA89D155EA7FA509390" box="[1308,1430,1941,1965]" italics="true" pageId="31" pageNumber="32">C. berberella</emphasis>
</taxonomicName>
, paratype no. Diškus089 ZMUC; 94,
<taxonomicName id="2FB94D7DFFECFFA899FC5E8BFD5393EC" box="[501,661,1977,2001]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="longiciliatella">
<emphasis id="DACDEAECFFECFFA899FC5E8BFD5393EC" box="[501,661,1977,2001]" italics="true" pageId="31" pageNumber="32">C. longiciliatella</emphasis>
</taxonomicName>
, slide no. Diškus088 ZMUC; 95,
<taxonomicName id="2FB94D7DFFECFFA89BD45E8BFAC293EC" authority="(Clemens)" baseAuthorityName="Clemens" box="[989,1284,1977,2001]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="citripenipennella">
<emphasis id="DACDEAECFFECFFA89BD45E8BFB5F93EC" box="[989,1177,1977,2001]" italics="true" pageId="31" pageNumber="32">C. citripenipennella</emphasis>
(Clemens)
</taxonomicName>
, USA, slide no. 28961 NHMUK; 96,
<taxonomicName id="2FB94D7DFFECFFA899565EEFFE0793C8" authorityName="Diskus" authorityYear="1998" box="[351,449,2013,2037]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="paulula">
<emphasis id="DACDEAECFFECFFA899565EEFFE0793C8" box="[351,449,2013,2037]" italics="true" pageId="31" pageNumber="32">C. paulula</emphasis>
</taxonomicName>
; 97,
<taxonomicName id="2FB94D7DFFECFFA899F95EEFFD2193C8" authority="Xu &amp; Dai" authorityName="Xu &amp; Dai" authorityYear="2021" box="[496,743,2013,2037]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="camptotheca">
<emphasis id="DACDEAECFFECFFA899F95EEFFD4493C8" box="[496,642,2013,2037]" italics="true" pageId="31" pageNumber="32">C. camptotheca</emphasis>
Xu &amp; Dai
</taxonomicName>
, China (
<bibRefCitation id="8C284B0FFFECFFA89B3D5EEFFC7893C8" author="Xu, J. &amp; Dai, X. &amp; Rimsaite, J. &amp; Diskus, A. &amp; Stonis, J. R." box="[820,958,2013,2037]" pageId="31" pageNumber="32" pagination="76 - 96" refId="ref53221" refString="Xu, J., Dai, X., Rimsaite, J., Diskus, A. &amp; Stonis, J. R. (2021) Discovery of the new Coptotriche species in China revealed two novel host-plant families and host-plant orders for Tischeriidae, a family of stenophagous, leafmining lepidopterans. Zootaxa, 5071 (1), 76 - 96. https: // doi. org / 10.11646 / zootaxa. 5071.1.4" type="journal article" year="2021">
Xu
<emphasis id="DACDEAECFFECFFA89B515EEFFC4193C8" box="[856,903,2013,2037]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
2021
</bibRefCitation>
); 98,
<taxonomicName id="2FB94D7DFFECFFA89BFB5EEFFA9D93C8" authority="(Stonis et al. 2019 b)" baseAuthorityName="Stonis" baseAuthorityYear="2019" box="[1010,1371,2013,2037]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="parvisacculata">
<emphasis id="DACDEAECFFECFFA89BFB5EEFFB5E93C8" box="[1010,1176,2013,2037]" italics="true" pageId="31" pageNumber="32">C. parvisacculata</emphasis>
(
<bibRefCitation id="8C284B0FFFECFFA89CAD5EEFFA9393C8" author="Stonis, J. R. &amp; Diskus, A. &amp; Remeikis, A. &amp; Katinas, L. &amp; Cumbicus Torres, N. &amp; Schuster, J. &amp; Puplesyte-Chambers, J." box="[1188,1365,2013,2037]" pageId="31" pageNumber="32" pagination="1 - 32" refId="ref51357" refString="Stonis, J. R., Diskus, A., Remeikis, A., Katinas, L., Cumbicus Torres, N., Schuster, J. &amp; Puplesyte-Chambers, J. (2019 b) Diagnostics of new species of Neotropical Tischeriidae (Lepidoptera), with the first record of Coptotriche Walshingham from South America. Zootaxa, 4691 (1), 1 - 32. https: // doi. org / 10.11646 / zootaxa. 4691.1.1" type="journal article" year="2019">
Stonis
<emphasis id="DACDEAECFFECFFA89CEC5EEFFAD393C8" box="[1253,1301,2013,2037]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
2019b
</bibRefCitation>
)
</taxonomicName>
</paragraph>
</caption>
<caption id="BCC66676FFD3FF97989E5E59FAB993D2" ID-DOI="http://doi.org/10.5281/zenodo.8261182" ID-Zenodo-Dep="8261182" httpUri="https://zenodo.org/record/8261182/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,263,1899,1923]" targetBox="[169,1406,175,1879]" targetPageId="32" targetType="figure">
<paragraph id="E80636FEFFD3FF97989E5E59FAB993D2" blockId="32.[151,1436,1899,2032]" pageId="32" pageNumber="33">
<emphasis id="DACDEAECFFD3FF97989E5E59FEA493BE" bold="true" box="[151,354,1899,1923]" pageId="32" pageNumber="33">FIGURES 99104.</emphasis>
Female genitalia of
<taxonomicName id="2FB94D7DFFD3FF979A375E59FD7093BE" authorityName="Walsingham" authorityYear="1890" box="[574,694,1899,1923]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="DACDEAECFFD3FF979A375E59FD7093BE" box="[574,694,1899,1923]" italics="true" pageId="32" pageNumber="33">Coptotriche</emphasis>
</taxonomicName>
. 99,
<taxonomicName id="2FB94D7DFFD3FF979AE75E5EFC2193BE" authority="(Frey &amp; Boll)" baseAuthorityName="Frey &amp; Boll" baseAuthorityYear="1873" box="[750,999,1899,1924]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="aenea">
<emphasis id="DACDEAECFFD3FF979AE75E5EFC8D93BE" box="[750,843,1900,1923]" italics="true" pageId="32" pageNumber="33">C. aenea</emphasis>
(Frey &amp; Boll)
</taxonomicName>
, USA,
<emphasis id="DACDEAECFFD3FF979C3E5E59FA5093B9" bold="true" box="[1079,1430,1899,1924]" pageId="32" pageNumber="33">first documentation of lectotype</emphasis>
, slide no. 28944 NHMUK; 100,
<taxonomicName id="2FB94D7DFFD3FF9799DE5EA2FD4E9395" authority="(Kasy)" baseAuthorityName="Kasy" box="[471,648,1936,1960]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="szoecsi">
<emphasis id="DACDEAECFFD3FF9799DE5EA2FDFD939A" box="[471,571,1936,1959]" italics="true" pageId="32" pageNumber="33">C. szoecsi</emphasis>
(Kasy)
</taxonomicName>
, Austria, genitalia slide no. Diškus066 ZMUC; 101,
<taxonomicName id="2FB94D7DFFD3FF979C935EA2FA5E939A" authority="(Rebel)" baseAuthorityName="Rebel" box="[1178,1432,1935,1959]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="longiciliatella">
<emphasis id="DACDEAECFFD3FF979C935EA2FA83939A" box="[1178,1349,1935,1959]" italics="true" pageId="32" pageNumber="33">C. longiciliatella</emphasis>
(Rebel)
</taxonomicName>
, Tenerife, Canary Is., slide no. Diškus068 ZMUC; 102,
<taxonomicName id="2FB94D7DFFD3FF979ABF5E86FC2193F6" authority="(Duponchel)" baseAuthorityName="Duponchel" box="[694,999,1971,1995]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="angusticolella">
<emphasis id="DACDEAECFFD3FF979ABF5E86FCA693F6" box="[694,864,1971,1995]" italics="true" pageId="32" pageNumber="33">C. angusticolella</emphasis>
(Duponchel)
</taxonomicName>
, Slovakia, slide no. Diškus079 ZMUC; 103,
<taxonomicName id="2FB94D7DFFD3FF97989E5EEAFE6A93D2" authority="(Duponchel)" baseAuthorityName="Duponchel" box="[151,428,2007,2031]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="gaunacella">
<emphasis id="DACDEAECFFD3FF97989E5EEAFEE593D2" box="[151,291,2007,2031]" italics="true" pageId="32" pageNumber="33">C. gaunacella</emphasis>
(Duponchel)
</taxonomicName>
, Poland, slide no. Diškus129 ZMUC; 104,
<taxonomicName id="2FB94D7DFFD3FF979B555EEAFBA093D2" authority="Xu &amp; Dai" authorityName="Xu &amp; Dai" authorityYear="2021" box="[860,1126,2007,2032]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="camptotheca">
<emphasis id="DACDEAECFFD3FF979B555EEAFC3C93D2" box="[860,1018,2007,2031]" italics="true" pageId="32" pageNumber="33">C. camptotheca</emphasis>
Xu &amp; Dai
</taxonomicName>
, China (
<bibRefCitation id="8C284B0FFFD3FF979CB05EE5FAB193D2" author="Stonis, J. R. &amp; Diskus, A. &amp; Remeikis, A. &amp; Katinas, L. &amp; Cumbicus Torres, N. &amp; Schuster, J. &amp; Puplesyte-Chambers, J." box="[1209,1399,2007,2031]" pageId="32" pageNumber="33" pagination="1 - 32" refId="ref51357" refString="Stonis, J. R., Diskus, A., Remeikis, A., Katinas, L., Cumbicus Torres, N., Schuster, J. &amp; Puplesyte-Chambers, J. (2019 b) Diagnostics of new species of Neotropical Tischeriidae (Lepidoptera), with the first record of Coptotriche Walshingham from South America. Zootaxa, 4691 (1), 1 - 32. https: // doi. org / 10.11646 / zootaxa. 4691.1.1" type="journal article" year="2019">
Stonis
<emphasis id="DACDEAECFFD3FF979CF65EEAFAF493D2" box="[1279,1330,2007,2031]" italics="true" pageId="32" pageNumber="33">et al.</emphasis>
2019b
</bibRefCitation>
)
</paragraph>
</caption>
<caption id="BCC66676FFD2FF96989E5E58FA4A93D3" ID-DOI="http://doi.org/10.5281/zenodo.8261184" ID-Zenodo-Dep="8261184" httpUri="https://zenodo.org/record/8261184/files/figure.png" pageId="33" pageNumber="34" startId="33.[151,257,1898,1922]" targetBox="[168,1413,180,1868]" targetPageId="33" targetType="figure">
<paragraph id="E80636FEFFD2FF96989E5E58FA4A93D3" blockId="33.[151,1437,1898,2031]" pageId="33" pageNumber="34">
<emphasis id="DACDEAECFFD2FF96989E5E58FE9993BF" bold="true" box="[151,351,1898,1922]" pageId="33" pageNumber="34">FIGURES 105113.</emphasis>
Adults of
<taxonomicName id="2FB94D7DFFD2FF9699CF5E58FDFE93BF" authorityName="Walsingham" authorityYear="1890" box="[454,568,1898,1922]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="DACDEAECFFD2FF9699CF5E58FDFE93BF" box="[454,568,1898,1922]" italics="true" pageId="33" pageNumber="34">Coptotriche</emphasis>
</taxonomicName>
. 105,
<taxonomicName id="2FB94D7DFFD2FF969A7C5E59FCA193BF" authority="(Rebel)" baseAuthorityName="Rebel" box="[629,871,1898,1922]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="longiciliatella">
<emphasis id="DACDEAECFFD2FF969A7C5E59FCD193BF" box="[629,791,1898,1922]" italics="true" pageId="33" pageNumber="34">C. longiciliatella</emphasis>
(Rebel)
</taxonomicName>
, Tenerife, Canary Is.; 106,
<taxonomicName id="2FB94D7DFFD2FF969C645E59FA5A93BF" authority="Diskus &amp; Stonis, Far" authorityName="Diskus &amp; Stonis, Far" authorityYear="2014" box="[1133,1436,1898,1922]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="DACDEAECFFD2FF969C645E59FB0D93BF" box="[1133,1227,1899,1922]" italics="true" pageId="33" pageNumber="34">C. minuta</emphasis>
Diškus &amp; Stonis, Far
</taxonomicName>
Eastern Russia, holotype (
<bibRefCitation id="8C284B0FFFD2FF9699845EBCFDF0939B" author="Stonis, J. R. &amp; Diskus, A. &amp; Rociene, A. &amp; Sruoga, V. &amp; Davis, D. R." box="[397,566,1934,1958]" pageId="33" pageNumber="34" pagination="141 - 155" refId="ref51667" refString="Stonis, J. R., Diskus, A., Rociene, A., Sruoga, V. &amp; Davis, D. R. (2014) New and little known Coptotriche and Tischeria species (Lepidoptera: Tischeriidae) from Primorskiy Kray, Russian Far East. Zootaxa, 3884 (2), 141 - 155. https: // doi. org / 10.11646 / zootaxa. 3884.2.3" type="journal article" year="2014">
Stonis
<emphasis id="DACDEAECFFD2FF9699C65EBDFE38939B" box="[463,510,1934,1958]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
2014
</bibRefCitation>
); 107,
<taxonomicName id="2FB94D7DFFD2FF969A7D5EBDFC9D939B" authority="(De Prins)" baseAuthorityName="De Prins" baseAuthorityYear="1984" box="[628,859,1934,1958]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="berberella">
<emphasis id="DACDEAECFFD2FF969A7D5EBDFD37939B" box="[628,753,1934,1958]" italics="true" pageId="33" pageNumber="34">C. berberella</emphasis>
(De Prins)
</taxonomicName>
, Marocco, paratype; 108,
<taxonomicName id="2FB94D7DFFD2FF969C465EBDFA5F939B" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis L" authorityYear="2544" box="[1103,1433,1934,1958]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="parvisacculata">
<emphasis id="DACDEAECFFD2FF969C465EBDFB3E939B" box="[1103,1272,1934,1958]" italics="true" pageId="33" pageNumber="34">C. parvisacculata</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<taxonomicName id="2FB94D7DFFD2FF96989E5E80FF3E93F7" box="[151,248,1970,1994]" class="Magnoliopsida" family="Rosaceae" genus="Argentina" kingdom="Plantae" order="Rosales" pageId="33" pageNumber="127" phylum="Tracheophyta" rank="genus">Argentina</taxonomicName>
, holotype (
<bibRefCitation id="8C284B0FFFD2FF9699695E80FDD093F7" author="Stonis, J. R. &amp; Diskus, A. &amp; Remeikis, A. &amp; Katinas, L. &amp; Cumbicus Torres, N. &amp; Schuster, J. &amp; Puplesyte-Chambers, J." box="[352,534,1970,1994]" pageId="33" pageNumber="34" pagination="1 - 32" refId="ref51357" refString="Stonis, J. R., Diskus, A., Remeikis, A., Katinas, L., Cumbicus Torres, N., Schuster, J. &amp; Puplesyte-Chambers, J. (2019 b) Diagnostics of new species of Neotropical Tischeriidae (Lepidoptera), with the first record of Coptotriche Walshingham from South America. Zootaxa, 4691 (1), 1 - 32. https: // doi. org / 10.11646 / zootaxa. 4691.1.1" type="journal article" year="2019">
Stonis
<emphasis id="DACDEAECFFD2FF9699AA5E81FE1593F7" box="[419,467,1970,1994]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
2019b
</bibRefCitation>
); 109,
<taxonomicName id="2FB94D7DFFD2FF969A5C5E81FD3093F7" box="[597,758,1970,1994]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="longiciliatella">
<emphasis id="DACDEAECFFD2FF969A5C5E81FD3093F7" box="[597,758,1970,1994]" italics="true" pageId="33" pageNumber="34">C. longiciliatella</emphasis>
</taxonomicName>
; 110,
<taxonomicName id="2FB94D7DFFD2FF969B395E81FBF093F7" authority="(Duponchel)" baseAuthorityName="Duponchel" box="[816,1078,1970,1994]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="gaunacella">
<emphasis id="DACDEAECFFD2FF969B395E81FC7393F7" box="[816,949,1970,1994]" italics="true" pageId="33" pageNumber="34">C. gaunacella</emphasis>
(Duponchel)
</taxonomicName>
, Turkmenistan; 111,
<taxonomicName id="2FB94D7DFFD2FF969CF35E81FED493D3" authority="(Duponchel)" baseAuthorityName="Duponchel" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="angusticolella">
<emphasis id="DACDEAECFFD2FF969CF35E81FA5A93F7" box="[1274,1436,1970,1994]" italics="true" pageId="33" pageNumber="34">C. angusticolella</emphasis>
(Duponchel)
</taxonomicName>
, Belgium; 112,
<taxonomicName id="2FB94D7DFFD2FF9699A05EE5FD4193D3" authority="(Gerasimov)" baseAuthorityName="Gerasimov" baseAuthorityYear="1937" box="[425,647,2006,2030]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="rosella">
<emphasis id="DACDEAECFFD2FF9699A05EE5FDC093D3" box="[425,518,2006,2030]" italics="true" pageId="33" pageNumber="34">C. rosella</emphasis>
(Gerasimov)
</taxonomicName>
, Uzbekistan; 113,
<taxonomicName id="2FB94D7DFFD2FF969B3F5EE5FBF093D3" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis NEPTI" authorityYear="2021" box="[822,1078,2006,2030]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="asiana">
<emphasis id="DACDEAECFFD2FF969B3F5EE5FC5493D3" box="[822,914,2007,2030]" italics="true" pageId="33" pageNumber="34">C. asiana</emphasis>
Diškus &amp; Stonis
</taxonomicName>
, Vietnam, paratype (
<bibRefCitation id="8C284B0FFFD2FF969CF35EE4FA4393D3" author="Xu, J. &amp; Dai, X. &amp; Rimsaite, J. &amp; Diskus, A. &amp; Stonis, J. R." box="[1274,1413,2006,2030]" pageId="33" pageNumber="34" pagination="76 - 96" refId="ref53221" refString="Xu, J., Dai, X., Rimsaite, J., Diskus, A. &amp; Stonis, J. R. (2021) Discovery of the new Coptotriche species in China revealed two novel host-plant families and host-plant orders for Tischeriidae, a family of stenophagous, leafmining lepidopterans. Zootaxa, 5071 (1), 76 - 96. https: // doi. org / 10.11646 / zootaxa. 5071.1.4" type="journal article" year="2021">
Xu
<emphasis id="DACDEAECFFD2FF969D165EE5FA8993D3" box="[1311,1359,2006,2030]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
2021
</bibRefCitation>
)
</paragraph>
</caption>
<caption id="BCC66676FFD1FF95989E5EBFFDA993F4" ID-DOI="http://doi.org/10.5281/zenodo.8261186" ID-Zenodo-Dep="8261186" httpUri="https://zenodo.org/record/8261186/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,264,1933,1957]" targetBox="[157,1425,189,1901]" targetPageId="34" targetType="figure">
<paragraph id="E80636FEFFD1FF95989E5EBFFDA993F4" blockId="34.[151,1436,1933,1994]" pageId="34" pageNumber="35">
<emphasis id="DACDEAECFFD1FF95989E5EBFFEA89398" bold="true" box="[151,366,1933,1957]" pageId="34" pageNumber="35">FIGURES 114120.</emphasis>
Adults of
<taxonomicName id="2FB94D7DFFD1FF9599D55EBFFCB59398" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2023" box="[476,883,1933,1957]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sapaensis" status="sp. nov.">
<emphasis id="DACDEAECFFD1FF9599D55EBFFD069398" box="[476,704,1933,1957]" italics="true" pageId="34" pageNumber="35">Coptotriche sapaensis</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<emphasis id="DACDEAECFFD1FF959B765EBFFC179398" bold="true" box="[895,977,1933,1957]" pageId="34" pageNumber="35">
<taxonomicNameLabel id="C1FE5797FFD1FF959B765EBFFC179398" box="[895,977,1933,1957]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, northern Vietnam. 114116, head; 117120 general view of holotype and paratypes (MfN)
</paragraph>
</caption>
<caption id="BCC66676FFD0FF94989E5EB2FE8793DD" ID-DOI="http://doi.org/10.5281/zenodo.8261188" ID-Zenodo-Dep="8261188" httpUri="https://zenodo.org/record/8261188/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,264,1920,1944]" targetBox="[158,1424,193,1889]" targetPageId="35" targetType="figure">
<paragraph id="E80636FEFFD0FF94989E5EB2FE8793DD" blockId="35.[151,1436,1920,2016]" pageId="35" pageNumber="36">
<emphasis id="DACDEAECFFD0FF94989E5EB2FEA893A5" bold="true" box="[151,366,1920,1944]" pageId="35" pageNumber="36">FIGURES 121125.</emphasis>
Male genitalia of
<taxonomicName id="2FB94D7DFFD0FF949A215EB2FC7F93A5" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2023" box="[552,953,1920,1944]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sapaensis" status="sp. nov.">
<emphasis id="DACDEAECFFD0FF949A215EB2FCCC93A5" box="[552,778,1920,1944]" italics="true" pageId="35" pageNumber="36">Coptotriche sapaensis</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<emphasis id="DACDEAECFFD0FF949BCA5EB2FBD293A5" bold="true" box="[963,1044,1920,1944]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="C1FE5797FFD0FF949BCA5EB2FBD293A5" box="[963,1044,1920,1944]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, northern Vietnam, holotype, genitalia slide no. AD1111 (MfN). 121, socii and apex of phallus; 122, uncus and apical part of phallus; 123, general view; 124, 125, spines of phallus
</paragraph>
</caption>
<caption id="BCC66676FFD7FF93989E5EB0FE5193DF" ID-DOI="http://doi.org/10.5281/zenodo.8261190" ID-Zenodo-Dep="8261190" httpUri="https://zenodo.org/record/8261190/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,264,1922,1946]" targetBox="[176,1407,198,1888]" targetPageId="36" targetType="figure">
<paragraph id="E80636FEFFD7FF93989E5EB0FE5193DF" blockId="36.[151,1436,1922,2018]" pageId="36" pageNumber="37">
<emphasis id="DACDEAECFFD7FF93989E5EB0FEA893A7" bold="true" box="[151,366,1922,1946]" pageId="36" pageNumber="37">FIGURES 126130.</emphasis>
Male genitalia of
<taxonomicName id="2FB94D7DFFD7FF939A215EB0FC7C93A7" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2023" box="[552,954,1922,1946]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sapaensis" status="sp. nov.">
<emphasis id="DACDEAECFFD7FF939A215EB0FCCD93A7" box="[552,779,1922,1946]" italics="true" pageId="36" pageNumber="37">Coptotriche sapaensis</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<emphasis id="DACDEAECFFD7FF939BCD5EB0FBD393A7" bold="true" box="[964,1045,1922,1946]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="C1FE5797FFD7FF939BCD5EB0FBD393A7" box="[964,1045,1922,1946]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, northern Vietnam, paratype, genitalia slide no. AD1106 (MfN). 126, 127, socii, tegumen and spined diaphragm; 128, valva and transtilla, ventral view; 129, valva, lateral view; 130, phallus
</paragraph>
</caption>
<caption id="BCC66676FFD6FF92989E5EB6FB2493D8" ID-DOI="http://doi.org/10.5281/zenodo.8261192" ID-Zenodo-Dep="8261192" httpUri="https://zenodo.org/record/8261192/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,264,1924,1948]" targetBox="[158,1425,188,1894]" targetPageId="37" targetType="figure">
<paragraph id="E80636FEFFD6FF92989E5EB6FB2493D8" blockId="37.[151,1436,1924,2021]" pageId="37" pageNumber="38">
<emphasis id="DACDEAECFFD6FF92989E5EB6FEAA93A0" bold="true" box="[151,364,1924,1949]" pageId="37" pageNumber="38">FIGURES 131135.</emphasis>
Female genitalia of
<taxonomicName id="2FB94D7DFFD6FF929A3E5EB7FC0593A0" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2023" box="[567,963,1925,1949]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="sapaensis" status="sp. nov.">
<emphasis id="DACDEAECFFD6FF929A3E5EB7FCDE93A1" box="[567,792,1925,1949]" italics="true" pageId="37" pageNumber="38">Coptotriche sapaensis</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<emphasis id="DACDEAECFFD6FF929BC55EB7FBDD93A0" bold="true" box="[972,1051,1925,1949]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="C1FE5797FFD6FF929BC55EB7FBDD93A0" box="[972,1051,1925,1949]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, northern Vietnam, paratype, genitalia slide no. AD1109 (MfN). 131, general view; 132, ovipositor lobes; 133, rounded accessory sac and proximal part of ductus spermathecae; 134, ovipositor lobes; 135, accessory sac and “neck” (slender proximal part) of corpus bursae
</paragraph>
</caption>
<caption id="BCC66676FFD5FF91989E5EB6FF0C93D8" ID-DOI="http://doi.org/10.5281/zenodo.8261194" ID-Zenodo-Dep="8261194" httpUri="https://zenodo.org/record/8261194/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,264,1924,1948]" targetBox="[163,1423,193,1891]" targetPageId="38" targetType="figure">
<paragraph id="E80636FEFFD5FF91989E5EB6FF0C93D8" blockId="38.[151,1436,1924,2021]" pageId="38" pageNumber="39">
<emphasis id="DACDEAECFFD5FF91989E5EB6FEAB93A0" bold="true" box="[151,365,1924,1949]" pageId="38" pageNumber="39">FIGURES 136142.</emphasis>
Bionomics of
<taxonomicName id="2FB94D7DFFD5FF919A095EB7FC4893A0" authority="Diskus &amp; Stonis" authorityName="Diskus &amp; Stonis" authorityYear="2023" box="[512,910,1925,1949]" class="Insecta" family="Tischeriidae" genus="Coptotriche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="sapaensis" status="sp. nov.">
<emphasis id="DACDEAECFFD5FF919A095EB7FD2493A1" box="[512,738,1925,1949]" italics="true" pageId="38" pageNumber="39">Coptotriche sapaensis</emphasis>
Diškus &amp; Stonis
</taxonomicName>
,
<emphasis id="DACDEAECFFD5FF919B915EB7FC2E93A0" bold="true" box="[920,1000,1925,1949]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="C1FE5797FFD5FF919B915EB7FC2E93A0" box="[920,1000,1925,1949]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
136, 137, habitat, elevation 1920 m, 15 km NW of Sapa, 22°2058”N, 103°4616”E, Lao Cai Prov., northern Vietnam; 138142, leaf mines on unidentified
<taxonomicName id="2FB94D7DFFD5FF919D035E9BFA5A93FC" box="[1290,1436,1961,1985]" class="Magnoliopsida" family="Symplocaceae" kingdom="Plantae" order="Ericales" pageId="38" pageNumber="39" phylum="Tracheophyta" rank="family">Symplocaceae</taxonomicName>
plant
</paragraph>
</caption>
<paragraph id="E80636FEFFD4FF9098CE59A5FCA69556" blockId="39.[151,1437,150,588]" pageId="39" pageNumber="40">
<emphasis id="DACDEAECFFD4FF9098CE59A5FE4F948C" box="[199,393,151,177]" italics="true" pageId="39" pageNumber="40">Female genitalia</emphasis>
(
<figureCitation id="70822A7BFFD4FF90999359A5FDFE948C" box="[410,568,150,177]" captionStart="FIGURES 131135" captionStartId="37.[151,264,1924,1948]" captionTargetBox="[158,1425,188,1894]" captionTargetId="figure-20@37.[154,1433,181,1901]" captionTargetPageId="37" captionText="FIGURES 131135. Female genitalia of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov., northern Vietnam, paratype, genitalia slide no. AD1109 (MfN). 131, general view; 132, ovipositor lobes; 133, rounded accessory sac and proximal part of ductus spermathecae; 134, ovipositor lobes; 135, accessory sac and “neck” (slender proximal part) of corpus bursae" figureDoi="http://doi.org/10.5281/zenodo.8261192" httpUri="https://zenodo.org/record/8261192/files/figure.png" pageId="39" pageNumber="40">Figs 131135</figureCitation>
) about 1445 µm long. Ovipositor lobes relatively large, with dark peg-like setae; second pair of ovipositor lobes large, with numerous relatively short setae and one very long seta on each lobe. Posterior apophyses only slightly longer than anterior apophyses; prela comprised of three pairs of stout, rod-like projections; transverse prela unusually wide. Corpus bursae gradually widening towards basal part, with numerous inconspicuous pectinations. Accessory sac globular, heavily folded. Ductus spermathecae wide proximally, slender, tube-like distally, with about 89 medium large and small coils.
</paragraph>
<paragraph id="E80636FEFFD4FF9098CE5844FA5395E6" blockId="39.[151,1437,150,588]" pageId="39" pageNumber="40">
<emphasis id="DACDEAECFFD4FF9098CE5844FE8595AD" bold="true" box="[199,323,374,400]" pageId="39" pageNumber="40">Bionomics</emphasis>
(
<figureCitation id="70822A7BFFD4FF9099595845FE2B95AC" box="[336,493,375,401]" captionStart="FIGURES 136142" captionStartId="38.[151,264,1924,1948]" captionTargetBox="[163,1423,193,1891]" captionTargetId="figure-25@38.[154,1433,181,1901]" captionTargetPageId="38" captionText="FIGURES 136142. Bionomics of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov. 136, 137, habitat, elevation 1920 m, 15 km NW of Sapa, 22°2058”N, 103°4616”E, Lao Cai Prov., northern Vietnam; 138142, leaf mines on unidentified Symplocaceae plant" figureDoi="http://doi.org/10.5281/zenodo.8261194" httpUri="https://zenodo.org/record/8261194/files/figure.png" pageId="39" pageNumber="40">Figs 136142</figureCitation>
). Host plant unidentified but is probably
<taxonomicName id="2FB94D7DFFD4FF909BA15845FB8C95AC" box="[936,1098,375,401]" class="Magnoliopsida" family="Symplocaceae" kingdom="Plantae" order="Ericales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="family">Symplocaceae</taxonomicName>
(
<figureCitation id="70822A7BFFD4FF909C5F5845FB7095AC" box="[1110,1206,375,401]" captionStart="FIGURES 136142" captionStartId="38.[151,264,1924,1948]" captionTargetBox="[163,1423,193,1891]" captionTargetId="figure-25@38.[154,1433,181,1901]" captionTargetPageId="38" captionText="FIGURES 136142. Bionomics of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov. 136, 137, habitat, elevation 1920 m, 15 km NW of Sapa, 22°2058”N, 103°4616”E, Lao Cai Prov., northern Vietnam; 138142, leaf mines on unidentified Symplocaceae plant" figureDoi="http://doi.org/10.5281/zenodo.8261194" httpUri="https://zenodo.org/record/8261194/files/figure.png" pageId="39" pageNumber="40">Fig. 138</figureCitation>
). Larvae mine leaves in February. Initially, the mine is slender with numerous lateral corridors; later, the mine widens to an elongated blotch-like mine (yellowish to pale brown, without frass). Adults occur in March. Otherwise, biology is unknown.
</paragraph>
</subSubSection>
<subSubSection id="A0A36575FFD4FF9098CE58D4FC1B961B" pageId="39" pageNumber="40" type="distribution">
<paragraph id="E80636FEFFD4FF9098CE58D4FC1B961B" blockId="39.[151,1437,150,588]" pageId="39" pageNumber="40">
<emphasis id="DACDEAECFFD4FF9098CE58D4FEA7963D" bold="true" box="[199,353,486,512]" pageId="39" pageNumber="40">Distribution.</emphasis>
This species is known from a single locality in northern
<collectingCountry id="90AE766EFFD4FF909BDD58D5FBF5963C" box="[980,1075,487,513]" name="Vietnam" pageId="39" pageNumber="40">Vietnam</collectingCountry>
near the border with
<collectingCountry id="90AE766EFFD4FF909D2958D5FAA1963C" box="[1312,1383,487,513]" name="China" pageId="39" pageNumber="40">China</collectingCountry>
: Lao Cai Province,
<quantity id="2F419B1BFFD4FF90993E5B3EFEB8961B" box="[311,382,524,551]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="39" pageNumber="40" unit="km" value="15.0">15 km</quantity>
NW Sapa, at the elevation of
<quantity id="2F419B1BFFD4FF909AC75B3EFCE5961A" box="[718,803,524,551]" metricMagnitude="3" metricUnit="m" metricValue="1.92" pageId="39" pageNumber="40" unit="m" value="1920.0">1920 m</quantity>
(
<figureCitation id="70822A7BFFD4FF909B3B5B3EFC17961B" box="[818,977,524,550]" captionStart="FIGURES 136142" captionStartId="38.[151,264,1924,1948]" captionTargetBox="[163,1423,193,1891]" captionTargetId="figure-25@38.[154,1433,181,1901]" captionTargetPageId="38" captionText="FIGURES 136142. Bionomics of Coptotriche sapaensis Diškus &amp; Stonis, sp. nov. 136, 137, habitat, elevation 1920 m, 15 km NW of Sapa, 22°2058”N, 103°4616”E, Lao Cai Prov., northern Vietnam; 138142, leaf mines on unidentified Symplocaceae plant" figureDoi="http://doi.org/10.5281/zenodo.8261194" httpUri="https://zenodo.org/record/8261194/files/figure.png" pageId="39" pageNumber="40">Figs 136, 137</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A0A36575FFD4FF9098CE5B03FB659676" box="[199,1187,561,588]" pageId="39" pageNumber="40" type="etymology">
<paragraph id="E80636FEFFD4FF9098CE5B03FB659676" blockId="39.[151,1437,150,588]" box="[199,1187,561,588]" pageId="39" pageNumber="40">
<emphasis id="DACDEAECFFD4FF9098CE5B03FE889676" bold="true" box="[199,334,561,587]" pageId="39" pageNumber="40">Etymology.</emphasis>
The species is named after Sapa (
<collectingCountry id="90AE766EFFD4FF909AC05B03FCE89676" box="[713,814,561,587]" name="Vietnam" pageId="39" pageNumber="40">Vietnam</collectingCountry>
), the locality where it was found.
</paragraph>
</subSubSection>
</treatment>
</document>