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<document id="4F643FE23922C8CDB94A41CFC5F6C27A" ID-DOI="10.11646/zootaxa.3741.4.2" ID-GBIF-Dataset="26b2bc4d-8855-43a3-b7ef-1541ed82c8ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="247812" ID-ZooBank="6D837E20-C644-432D-9AAC-8F30D3B9854A" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460704005188" checkinUser="plazi" docAuthor="Behan-Pelletier, Valerie M. &amp; Walter, David E." docDate="2013" docId="443887FA660AFFB1FF6FFF67FC874A87" docLanguage="en" docName="zt03741p489.pdf" docOrigin="Zootaxa 3741 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tectoribates campestris Behan-Pelletier &amp; Walter, 2013, sp. nov." docType="treatment" docVersion="8" lastPageNumber="481" masterDocId="B801FF82661EFFA7FFF8FFFFFFD54B72" masterDocTitle="Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida)" masterLastPageNumber="489" masterPageNumber="459" pageNumber="479" updateTime="1698356775934" updateUser="plazi">
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<mods:title id="42791DC8804DD5B06262C246BA4E12E4">Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida)</mods:title>
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<mods:name id="DB40051EF52F6CE7F292D4ADA5E1D5F7" type="personal">
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<mods:namePart id="4B3A9EE23609A05D1D4DAF6136DB85B9">Behan-Pelletier, Valerie M.</mods:namePart>
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<mods:namePart id="648D04EF076A8C2B4C2AEB3052A99F28">Walter, David E.</mods:namePart>
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<mods:title id="9D8C0B979F78B95FC690EC724B28C60B">Zootaxa</mods:title>
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<mods:date id="84F8921270983B917B7001C6497B82BB">2013</mods:date>
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<treatment id="443887FA660AFFB1FF6FFF67FC874A87" ID-DOI="http://doi.org/10.5281/zenodo.5671485" ID-GBIF-Taxon="119583775" ID-Zenodo-Dep="5671485" LSID="urn:lsid:plazi:treatment:443887FA660AFFB1FF6FFF67FC874A87" httpUri="http://treatment.plazi.org/id/443887FA660AFFB1FF6FFF67FC874A87" lastPageId="22" lastPageNumber="481" pageId="20" pageNumber="479">
<subSubSection id="848B6567660AFFB3FF6FFF67FEEF4BA1" pageId="20" pageNumber="479" type="nomenclature">
<paragraph id="CC2E36EC660AFFB3FF6FFF67FDC04BC0" blockId="20.[151,533,152,211]" box="[151,533,152,178]" pageId="20" pageNumber="479">
<heading id="97668180660AFFB3FF6FFF67FDC04BC0" bold="true" box="[151,533,152,178]" fontSize="11" level="1" pageId="20" pageNumber="479" reason="1">
<emphasis id="FEE5EAFE660AFFB3FF6FFF67FDC04BC0" bold="true" box="[151,533,152,178]" pageId="20" pageNumber="479">
<taxonomicName id="0B914D6F660AFFB3FF6FFF67FE654BC0" box="[151,432,152,178]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="479" phylum="Arthropoda" rank="species" species="campestris" status="sp. nov.">
<emphasis id="FEE5EAFE660AFFB3FF6FFF67FE654BC0" bold="true" box="[151,432,152,178]" italics="true" pageId="20" pageNumber="479">Tectoribates campestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D65785660AFFB3FE4FFF67FDC04BC0" box="[439,533,152,178]" pageId="20" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="CC2E36EC660AFFB3FF6FFF45FEEF4BA1" blockId="20.[151,533,152,211]" box="[151,314,186,211]" pageId="20" pageNumber="479">
(
<figureCitation id="54AA2A69660AFFB3FF67FF45FEDC4BA1" box="[159,265,186,211]" captionStart="FIGURES 11 A F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="20" pageNumber="479">Figs. 11F</figureCitation>
,
<figureCitation id="54AA2A69660AFFB3FEECFF44FEE44BA1" box="[276,305,187,211]" captionStart="FIGURE 12" captionStartId="21.[151,250,1967,1989]" captionTargetBox="[232,1336,196,1910]" captionTargetId="figure@21.[224,1346,193,1915]" captionTargetPageId="21" captionText="FIGURE 12. Tectoribates campestris sp. nov., adult female, A, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247823/files/figure.png" pageId="20" pageNumber="479">12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="848B6567660AFFB3FF6FFEFDFC234A8A" pageId="20" pageNumber="479" type="materials_examined">
<paragraph id="CC2E36EC660AFFB3FF6FFEFDFC234A8A" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF6FFEFDFEAF4A69" bold="true" box="[151,378,258,283]" pageId="20" pageNumber="479">Material examined</emphasis>
.
<typeStatus id="132A884E660AFFB3FE70FEFDFE264A69" box="[392,499,258,283]" pageId="20" pageNumber="479" type="holotype">Holotype</typeStatus>
, adult female,
<collectingCountry id="B486767C660AFFB3FD6BFEFDFD184A69" box="[659,717,258,283]" name="United States of America" pageId="20" pageNumber="479">USA</collectingCountry>
, Kansas, Konza LTER, US-UK grassland Project, Site B4, 4 yr. burn cycle,
<date id="B82F102C660AFFB3FEE2FED5FE5E4A30" box="[282,395,298,322]" pageId="20" pageNumber="479" value="1999-05-18">18.v.1999</date>
, 39°
<geoCoordinate id="A9A5502B660AFFB3FE40FED5FE284A30" box="[440,509,298,322]" direction="north" orientation="latitude" pageId="20" pageNumber="479" precision="555" value="4.78">4.78N</geoCoordinate>
96°
<geoCoordinate id="A9A5502B660AFFB3FDDFFED6FD5B4A30" box="[551,654,297,322]" direction="west" orientation="longitude" pageId="20" pageNumber="479" precision="55" value="-35.847">35.847W</geoCoordinate>
(V. Behan-Pelletier) from rocky slope with
<emphasis id="FEE5EAFE660AFFB3FB83FED5FB084A33" box="[1147,1245,298,321]" italics="true" pageId="20" pageNumber="479">Baptesia</emphasis>
, deposited in the CNC,
<typeStatus id="132A884E660AFFB3FF19FEB1FEC44A14" box="[225,273,334,358]" pageId="20" pageNumber="479">type</typeStatus>
number 24150.
<typeStatus id="132A884E660AFFB3FE37FEB1FDEA4A14" box="[463,575,334,358]" pageId="20" pageNumber="479" type="paratype">Paratypes</typeStatus>
, 8 with same data as
<typeStatus id="132A884E660AFFB3FCC2FEB2FC4A4A14" box="[826,927,333,358]" pageId="20" pageNumber="479" type="holotype">holotype</typeStatus>
; with same data as
<typeStatus id="132A884E660AFFB3FB7BFEB2FB3D4A14" box="[1155,1256,333,358]" pageId="20" pageNumber="479" type="holotype">holotype</typeStatus>
, except, 4 from upper litter and soil in depression; 3 from seep in depression; Konza LTER, Gallery Forest along King Creek, 39°04N 96°35W,
<date id="B82F102C660AFFB3FEACFE67FE174AC2" box="[340,450,408,432]" pageId="20" pageNumber="479" value="1999-05-19">19.v.1999</date>
(V. Behan-Pelletier &amp; M.
<collectingCountry id="B486767C660AFFB3FD13FE67FCEA4AC3" box="[747,831,408,433]" name="Virgin Islands" pageId="20" pageNumber="479">St.John</collectingCountry>
) 2 from litter and soil under riparian grasses;
<collectingCountry id="B486767C660AFFB3FAB9FE67FA4D4AC3" box="[1345,1432,408,433]" name="Canada" pageId="20" pageNumber="479">Canada</collectingCountry>
, Ontario, Almonte, Alvar burnt Lands,
<date id="B82F102C660AFFB3FDBFFE43FD6A4AA7" box="[583,703,444,469]" pageId="20" pageNumber="479" value="1990-06-26">26.vi.1990</date>
(V. Behan-Pelletier &amp; B. Eamer) 4 from
<emphasis id="FEE5EAFE660AFFB3FB76FE42FA494AA7" box="[1166,1436,444,469]" italics="true" pageId="20" pageNumber="479">Penstamon, Campanula</emphasis>
under Juniper.
<typeStatus id="132A884E660AFFB3FEC4FE1EFE7C4A8B" box="[316,425,481,505]" pageId="20" pageNumber="479" type="paratype">Paratypes</typeStatus>
deposited in the CNC, PMAE.IZ, USNM and RNC.
</paragraph>
</subSubSection>
<subSubSection id="848B6567660AFFB3FF3FFDFBFAA1484D" pageId="20" pageNumber="479" type="diagnosis">
<paragraph id="CC2E36EC660AFFB3FF3FFDFBFAA1484D" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FFDFBFEEF496F" bold="true" box="[199,314,516,541]" pageId="20" pageNumber="479">Diagnosis</emphasis>
. Adult. Total length 280300. Integument with micropuncta between genital and anal plates subequal in size, though more spaced than micropuncta on rest of body. Rostral margin dentate, with medial small U-shaped indentation. Notogaster anteriorly with distinct hexagonal pattern. Octotaxic system expressed as saccules about
<quantity id="0B699B09660AFFB3FEBBFD8FFEBA49FB" box="[323,367,624,649]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="479" unit="in" value="5.0">5 in</quantity>
diameter. Notogastral setae slightly roughened, 1321, with seta
<emphasis id="FEE5EAFE660AFFB3FBADFD8EFBB449FA" box="[1109,1121,625,648]" italics="true" pageId="20" pageNumber="479">c</emphasis>
longest; mutual distance of
<emphasis id="FEE5EAFE660AFFB3FF6FFD6BFF0549C3" box="[151,208,660,689]" italics="true" pageId="20" pageNumber="479">h1h1</emphasis>
about 15, of
<emphasis id="FEE5EAFE660AFFB3FE9DFD6AFE4A49C3" box="[357,415,661,689]" italics="true" pageId="20" pageNumber="479">p1p1</emphasis>
about 30. Lamella 7578 long, of which cusp 4347; about 16 wide at level of insertion of seta
<emphasis id="FEE5EAFE660AFFB3FF32FD44FF0B49A6" box="[202,222,699,724]" italics="true" pageId="20" pageNumber="479">le</emphasis>
. Lamellar cleft extending posteriorly about 14 from region of attachment of lamella to prodorsum; distance between lamellae in this region (width of cleft) about 2. Lamellar cusp with short tapered medial and lateral dentes,
<date id="B82F102C660AFFB3FF6FFCFCFE8E4869" box="[151,347,771,796]" pageId="20" pageNumber="479" value="1914-11-16" valueMax="1914-11-10" valueMin="1914-11-16">1116 and 1014</date>
long, respectively. Lamellar seta arising anteriorly on cusp, about 41 long. Bothridial seta clubshaped, 4957. Tutorium 6873 long, of which cusp 2126. Custodium about 12 long. Epimeral seta
<emphasis id="FEE5EAFE660AFFB3FAFDFCD8FAF5484D" box="[1285,1312,807,831]" italics="true" pageId="20" pageNumber="479">1c</emphasis>
2630.
</paragraph>
</subSubSection>
<subSubSection id="848B6567660AFFB3FF3FFCB4FE144CEC" pageId="20" pageNumber="479" type="description">
<paragraph id="CC2E36EC660AFFB3FF3FFCB4FB7148F5" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FFCB4FE8C4816" bold="true" box="[199,345,843,868]" pageId="20" pageNumber="479">Description.</emphasis>
Adults.
<emphasis id="FEE5EAFE660AFFB3FE40FCB3FD904811" box="[440,581,844,867]" italics="true" pageId="20" pageNumber="479">Dimensions:</emphasis>
Total length: females (
<emphasis id="FEE5EAFE660AFFB3FCB1FCB3FC824811" box="[841,855,844,867]" italics="true" pageId="20" pageNumber="479">n</emphasis>
= 7) 298 (290300); males (
<emphasis id="FEE5EAFE660AFFB3FB62FCB3FB7D4811" box="[1178,1192,844,867]" italics="true" pageId="20" pageNumber="479">n</emphasis>
= 7) 292 (range 290 300). Notogastral width: females (
<emphasis id="FEE5EAFE660AFFB3FDE0FC8FFDF348F5" box="[536,550,880,903]" italics="true" pageId="20" pageNumber="479">n</emphasis>
= 6) 191 (185200); males (
<emphasis id="FEE5EAFE660AFFB3FC90FC8FFCA348F5" box="[872,886,880,903]" italics="true" pageId="20" pageNumber="479">n</emphasis>
= 6) 186 (range 175190).
</paragraph>
<paragraph id="CC2E36EC660AFFB3FF3FFC6BFD124F65" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FFC6BFE9B48D9" box="[199,334,916,939]" italics="true" pageId="20" pageNumber="479">Integument:</emphasis>
Prodorsum, notogaster, pedotecta, anal plates, genital plates, venter and mentum punctate; puncta between genital and anal plates subequal in size, more widely spaced than puncta elsewhere on body. Lamellae, pedotectum, mentum, coxisternal region with fine irregular striae; tutorial cusp with distinct striae. Rostrum without distinct striae close to medial indentation.
</paragraph>
<paragraph id="CC2E36EC660AFFB3FF3FFBDCFB724E29" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FFBDCFE984F4E" box="[199,333,1059,1084]" italics="true" pageId="20" pageNumber="479">Prodorsum:</emphasis>
Rostrum with distinct, convex medial crest proximally, rostral margin dentate, with medial small U-shaped indentation. Rostrum without deep groove ventral to seta
<emphasis id="FEE5EAFE660AFFB3FC6AFBB7FC7C4F2D" box="[914,937,1096,1119]" italics="true" pageId="20" pageNumber="479">ro</emphasis>
. Lamella 7578 long, of which cusp 4347; about 16 wide at level of insertion of seta
<emphasis id="FEE5EAFE660AFFB3FD8AFB94FD534FF6" box="[626,646,1131,1156]" italics="true" pageId="20" pageNumber="479">le</emphasis>
. Lamellar cusps parallel and closely adjacent medially. Lamellar cleft extending posteriorly about 14 from region of attachment of lamella to prodorsum; distance between lamellae in this region (width of cleft) about 2. Lamellar cusp with two short, tapered dentes; medial dens longer than lateral dens,
<date id="B82F102C660AFFB3FF23FB29FE774F9D" box="[219,418,1238,1263]" pageId="20" pageNumber="479" value="1914-11-16" valueMax="1914-11-10" valueMin="1914-11-16">1116 and 1014</date>
long, respectively. Seta
<emphasis id="FEE5EAFE660AFFB3FD46FB27FD004F9D" box="[702,725,1240,1263]" italics="true" pageId="20" pageNumber="479">ro</emphasis>
4454 long, strongly barbed along length, acuminate, curved anteromedially. Lamellar seta arising anteriorly on cusp, about 41 long, heavily barbed, acuminate distally. Seta
<emphasis id="FEE5EAFE660AFFB3FA7EFB03FA494E61" box="[1414,1436,1276,1299]" italics="true" pageId="20" pageNumber="479">in</emphasis>
about 72 long, barbed (subequally to
<emphasis id="FEE5EAFE660AFFB3FDCEFAE0FD9F4E4A" box="[566,586,1311,1336]" italics="true" pageId="20" pageNumber="479">le</emphasis>
), acuminate; mutual distance about 2333. Bothridial seta club-shaped, 49 57, head directed slightly anteromedially (
<figureCitation id="54AA2A69660AFFB3FD88FABCFD164E29" box="[624,707,1347,1372]" captionStart="FIGURE 12" captionStartId="21.[151,250,1967,1989]" captionTargetBox="[232,1336,196,1910]" captionTargetId="figure@21.[224,1346,193,1915]" captionTargetPageId="21" captionText="FIGURE 12. Tectoribates campestris sp. nov., adult female, A, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247823/files/figure.png" pageId="20" pageNumber="479">Fig. 12</figureCitation>
). Seta
<emphasis id="FEE5EAFE660AFFB3FCF7FABBFCFD4E29" box="[783,808,1348,1371]" italics="true" pageId="20" pageNumber="479">ex</emphasis>
1623. Dorsophragmata separate.
</paragraph>
<paragraph id="CC2E36EC660AFFB3FF3FFA98FDCB4EBA" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FFA98FDC54EF2" box="[199,528,1383,1408]" italics="true" pageId="20" pageNumber="479">Lateral region of podosoma:</emphasis>
Tutorium 6072 long, of which cusp 2126, extending anterior of insertion of rostral seta. Custodium about 12 long. Porose area Al about
<quantity id="0B699B09660AFFB3FCAEFA74FC514ED6" box="[854,900,1419,1444]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="20" pageNumber="479" unit="in" value="8.0">8 in</quantity>
diameter (difficult to see because of proximal position close to base of tutorium).
</paragraph>
<paragraph id="CC2E36EC660AFFB3FF3FFA2BFF2A4D28" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FFA2BFE994E99" box="[199,332,1492,1515]" italics="true" pageId="20" pageNumber="479">Notogaster:</emphasis>
Longer than wide: 1.2:1. Anteriorly with distinct hexagonal pattern (
<figureCitation id="54AA2A69660AFFB3FB83FA2CFB3D4E99" box="[1147,1256,1491,1516]" captionStart="FIGURES 11 A F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="20" pageNumber="479">Figs. 11F</figureCitation>
,
<figureCitation id="54AA2A69660AFFB3FB00FA2CFAC24E99" box="[1272,1303,1491,1515]" captionStart="FIGURE 12" captionStartId="21.[151,250,1967,1989]" captionTargetBox="[232,1336,196,1910]" captionTargetId="figure@21.[224,1346,193,1915]" captionTargetPageId="21" captionText="FIGURE 12. Tectoribates campestris sp. nov., adult female, A, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247823/files/figure.png" pageId="20" pageNumber="479">12</figureCitation>
). Octotaxic system expressed as saccules about
<quantity id="0B699B09660AFFB3FDCCFA08FDB74D62" box="[564,610,1527,1552]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="20" pageNumber="479" unit="in" value="5.0">5 in</quantity>
diameter (
<figureCitation id="54AA2A69660AFFB3FD26FA08FCEA4D7D" box="[734,831,1527,1552]" captionStart="FIGURES 11 A F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="20" pageNumber="479">Fig. 11F</figureCitation>
, insert). Notogastral setae slightly roughened, 1021 long, with seta
<emphasis id="FEE5EAFE660AFFB3FEBAF9E3FE9B4D41" box="[322,334,1564,1587]" italics="true" pageId="20" pageNumber="479">c</emphasis>
longest;
<emphasis id="FEE5EAFE660AFFB3FE4DF9E4FE044D46" box="[437,465,1563,1588]" italics="true" pageId="20" pageNumber="479">lm</emphasis>
posteromediad Sa and
<emphasis id="FEE5EAFE660AFFB3FD2EF9E4FD394D46" box="[726,748,1563,1588]" italics="true" pageId="20" pageNumber="479">lp</emphasis>
anteromediad S1; mutual distance of
<emphasis id="FEE5EAFE660AFFB3FB6CF9E4FB1B4D4A" box="[1172,1230,1563,1592]" italics="true" pageId="20" pageNumber="479">h1h1</emphasis>
about 15, of
<emphasis id="FEE5EAFE660AFFB3FA9BF9E3FA494D4A" box="[1379,1436,1564,1592]" italics="true" pageId="20" pageNumber="479">p1p1</emphasis>
about 30.
</paragraph>
<paragraph id="CC2E36EC660AFFB3FF3FF99AFF364D98" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FF99AFEAC4D0C" box="[199,377,1637,1662]" italics="true" pageId="20" pageNumber="479">Ventral Region:</emphasis>
Epimeral setae mostly 715 long, weakly barbed, with
<emphasis id="FEE5EAFE660AFFB3FC06F999FBCC4D0C" box="[1022,1049,1638,1662]" italics="true" pageId="20" pageNumber="479">1c</emphasis>
more heavily barbed, and thicker than other epimeral setae, 2630. Genital, aggenital, anal and adanal setae weakly barbed; genital setae about 12, other setae about 6. Genital setae 2+4 or 3+3. Lyrifissure
<emphasis id="FEE5EAFE660AFFB3FCE3F952FCEA4DB4" box="[795,831,1709,1734]" italics="true" pageId="20" pageNumber="479">iad</emphasis>
anterolateral of anal plate. Postanal porose area about
<date id="B82F102C660AFFB3FF6FF92DFF364D98" box="[151,227,1745,1770]" pageId="20" pageNumber="479" value="2011-10-05">11 x 5.</date>
</paragraph>
<paragraph id="CC2E36EC660AFFB3FF3FF90AFE644C43" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<taxonomicName id="0B914D6F660AFFB3FF3FF90AFE164C7C" authority="Axillary" authorityName="Axillary" box="[199,451,1781,1806]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="20" pageNumber="479" phylum="Nematoda" rank="genus">
<emphasis id="FEE5EAFE660AFFB3FF3FF90AFE8A4C7C" box="[199,351,1781,1806]" italics="true" pageId="20" pageNumber="479">Gnathosoma:</emphasis>
Axillary
</taxonomicName>
saccule at base of palp
<date id="B82F102C660AFFB3FD28F90AFD264C7C" box="[720,755,1781,1806]" pageId="20" pageNumber="479">6 x</date>
2. Mentum with small
<taxonomicName id="0B914D6F660AFFB3FC02F908FBE74C7C" box="[1018,1074,1783,1806]" class="Arachnida" family="Tegoribatidae" genus="Oribata" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="479" phylum="Arthropoda" rank="species" species="tecta">tecta</taxonomicName>
, anterolateral of seta
<emphasis id="FEE5EAFE660AFFB3FADAF90AFAE54C7C" box="[1314,1328,1781,1806]" italics="true" pageId="20" pageNumber="479">h</emphasis>
. Relative lengths of setae:
<emphasis id="FEE5EAFE660AFFB3FEACF8E6FE7F4C40" box="[340,426,1817,1842]" italics="true" pageId="20" pageNumber="479">m&gt;a&gt;h</emphasis>
.
</paragraph>
<paragraph id="CC2E36EC660AFFB3FF3FF8C0FCA44C0B" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FF8C0FED34C24" box="[199,262,1855,1878]" italics="true" pageId="20" pageNumber="479">Legs:</emphasis>
Setae
<emphasis id="FEE5EAFE660AFFB3FEB7F8C2FEB34C24" box="[335,358,1853,1878]" italics="true" pageId="20" pageNumber="479">l”</emphasis>
of genu and tibia I more barbed than other setae on these segments, that of genu I spinose; about 13 and 17, respectively. Small knob lateral of solenidia on tibia I.
</paragraph>
<paragraph id="CC2E36EC660AFFB3FF3FF878FE144CEC" blockId="20.[151,1436,258,2022]" box="[199,449,1925,1950]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FF878FE904CEC" box="[199,325,1927,1950]" italics="true" pageId="20" pageNumber="479">Immatures.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="848B6567660AFFB3FF3FF856FD4C4C94" pageId="20" pageNumber="479" type="etymology">
<paragraph id="CC2E36EC660AFFB3FF3FF856FD4C4C94" blockId="20.[151,1436,258,2022]" pageId="20" pageNumber="479">
<emphasis id="FEE5EAFE660AFFB3FF3FF856FE934CB0" bold="true" box="[199,326,1961,1986]" pageId="20" pageNumber="479">Etymology</emphasis>
. The specific epithet
<taxonomicName id="0B914D6F660AFFB3FDC3F854FD614CB0" box="[571,692,1963,1986]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="20" pageNumber="479" phylum="Arthropoda" rank="species" species="campestris">
<emphasis id="FEE5EAFE660AFFB3FDC3F854FD614CB0" box="[571,692,1963,1986]" italics="true" pageId="20" pageNumber="479">campestris</emphasis>
</taxonomicName>
comes from the Latin meaning associated with fields and refers to the distribution of this species in grassland.
</paragraph>
</subSubSection>
<caption id="98EE6664660BFFB2FF6FF850FDC74C91" httpUri="https://zenodo.org/record/247823/files/figure.png" pageId="21" pageNumber="480" targetBox="[232,1336,196,1910]" targetPageId="21">
<paragraph id="CC2E36EC660BFFB2FF6FF850FDC74C91" blockId="21.[151,1435,1967,2019]" pageId="21" pageNumber="480">
<emphasis id="FEE5EAFE660BFFB2FF6FF850FEF04CB7" bold="true" box="[151,293,1967,1989]" pageId="21" pageNumber="480">FIGURE 12.</emphasis>
<taxonomicName id="0B914D6F660BFFB2FEC9F850FDF34CB7" box="[305,550,1967,1989]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="21" pageNumber="480" phylum="Arthropoda" rank="species" species="campestris" status="sp. nov.">
<emphasis id="FEE5EAFE660BFFB2FEC9F850FDF34CB7" box="[305,550,1967,1989]" italics="true" pageId="21" pageNumber="480">Tectoribates campestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D65785660BFFB2FDCAF850FD5D4CB7" box="[562,648,1967,1989]" pageId="21" pageNumber="480" rank="species">
<emphasis id="FEE5EAFE660BFFB2FDCAF850FD5D4CB7" bold="true" box="[562,648,1967,1989]" pageId="21" pageNumber="480">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult female, A, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm.
</paragraph>
</caption>
<subSubSection id="848B65676608FFB1FF3FFF68FC874A87" pageId="22" pageNumber="481" type="discussion">
<paragraph id="CC2E36EC6608FFB1FF3FFF68FE6C4A32" blockId="22.[151,1436,151,501]" pageId="22" pageNumber="481">
<emphasis id="FEE5EAFE6608FFB1FF3FFF68FEE14BC2" bold="true" box="[199,308,151,176]" pageId="22" pageNumber="481">Remarks</emphasis>
. This species is most similar to the
<typeStatus id="132A884E6608FFB1FD05FF67FCF84BC2" box="[765,813,152,176]" pageId="22" pageNumber="481">type</typeStatus>
species
<taxonomicName id="0B914D6F6608FFB1FC61FF68FB4D4BC2" box="[921,1176,151,176]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="FEE5EAFE6608FFB1FC61FF68FB4D4BC2" box="[921,1176,151,176]" italics="true" pageId="22" pageNumber="481">Tectoribates proximus</emphasis>
</taxonomicName>
with which it shares notogastral saccules and a dentate rostral margin. It differs in having: medial and lateral dentes on the lamellar cusps subequal in length or with the medial dens slightly longer (medial shorter than lateral in
<taxonomicName id="0B914D6F6608FFB1FB1AFF1EFAB24B8A" box="[1250,1383,225,248]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="FEE5EAFE6608FFB1FB1AFF1EFAB24B8A" box="[1250,1383,225,248]" italics="true" pageId="22" pageNumber="481">T. proximus</emphasis>
</taxonomicName>
); an epimeral setation of 3-1-2-3 (3-1-3-
<quantity id="0B699B096608FFB1FDD5FEFBFD8E4A6F" box="[557,603,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="22" pageNumber="481" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="0B914D6F6608FFB1FD9CFEFAFD3D4A6E" box="[612,744,261,284]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="FEE5EAFE6608FFB1FD9CFEFAFD3D4A6E" box="[612,744,261,284]" italics="true" pageId="22" pageNumber="481">T. proximus</emphasis>
</taxonomicName>
according to Bernini 1973); a shorter body length, 280300 (
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<taxonomicName id="0B914D6F6608FFB1FEDEFED6FE7D4A32" box="[294,424,297,320]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="FEE5EAFE6608FFB1FEDEFED6FE7D4A32" box="[294,424,297,320]" italics="true" pageId="22" pageNumber="481">T. proximus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="CC2E36EC6608FFB1FF3FFEB3FC874A87" blockId="22.[151,1436,151,501]" pageId="22" pageNumber="481">
Populations from the Konza Tallgrass prairie site collected in
<date id="B82F102C6608FFB1FC86FEB2FC244A16" box="[894,1009,332,357]" pageId="22" pageNumber="481" value="1999-05">May 1999</date>
had a sex ratio of 1:1. The only study of population dynamics and reproduction in
<taxonomicName id="0B914D6F6608FFB1FD74FE90FCC04AFA" box="[652,789,367,392]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE6608FFB1FD74FE90FCC04AFA" box="[652,789,367,392]" italics="true" pageId="22" pageNumber="481">Tectoribates</emphasis>
</taxonomicName>
is that of Smelyansky (2006) who studied characteristics of reproduction in field populations of
<taxonomicName id="0B914D6F6608FFB1FDB2FE6AFD6E4ADE" box="[586,699,405,428]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="22" pageNumber="481" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="FEE5EAFE6608FFB1FDB2FE6AFD6E4ADE" box="[586,699,405,428]" italics="true" pageId="22" pageNumber="481">T. ornatus</emphasis>
</taxonomicName>
from the Trans-Volga region of the Russian steppe during June to August. He found that gravid females decreased from over 80% of female specimens collected to 0% during the 3 months, while sex ratio changed from 21% male to 62% male.
</paragraph>
</subSubSection>
</treatment>
</document>