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<document id="8F0F2F8E934BA972ECD95C82E290F1F4" ID-DOI="10.11646/zootaxa.4802.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3904978" ID-ZooBank="BA145877-12CB-4C97-A18F-9E4E6F0258CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592923053381" checkinUser="plazi" docAuthor="Obando, Ranulfo González, Gironza, Nancy Carrejo, Panche, Jeferson &amp; Aldrete, Alfonso Neri García" docDate="2020" docId="039787F6D153260BFF7AFB054915FE1C" docLanguage="en" docName="zootaxa.4802.2.1.pdf" docOrigin="Zootaxa 4802 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Loneura altaquerensis Obando &amp; Gironza &amp; Panche &amp; Aldrete 2020, n. sp." docType="treatment" docVersion="4" lastPageNumber="215" masterDocId="FFAEFF8ED15F2605FFEDFFA84C2FFFCF" masterDocTitle="The genus Loneura Navás (Insecta: Psocodea: ‘ Psocoptera’: Ptiloneuridae) in Colombia, description of new species and key for identification" masterLastPageNumber="249" masterPageNumber="201" pageNumber="213" updateTime="1698776039647" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="8CEF609C6789C422C356A0CF7B5FDAB0">The genus Loneura Navás (Insecta: Psocodea: ‘ Psocoptera’: Ptiloneuridae) in Colombia, description of new species and key for identification</mods:title>
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<mods:namePart id="B4C9FF3D21D5EB0897817CCAE0023753">Obando, Ranulfo González</mods:namePart>
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<mods:namePart id="AF8616697717ADF1F9C64879406F5AE4">Gironza, Nancy Carrejo</mods:namePart>
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<mods:namePart id="8D8D04C57E255CFE521CB37AB25D3A62">Panche, Jeferson</mods:namePart>
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<mods:namePart id="55BD527CA045073DC4F06B45B755EAC6">Aldrete, Alfonso Neri García</mods:namePart>
<mods:affiliation id="D69233CA7C1B4FC5D22DDC5FA8E393C7">Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apdo. Postal 70 - 153, CdMx, MÉXICO. anga @ ib. unam. mx; https: // orcid. org / 0000 - 0001 - 7214 - 7966</mods:affiliation>
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<mods:title id="D6EDE01359C9080FD1FFA7EEF1A4C458">Zootaxa</mods:title>
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<mods:date id="1F7295D8D09C36A3839D81748130A338">2020</mods:date>
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<mods:number id="079AF11F71AEF559C918704E4053CDC1">2020-06-23</mods:number>
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<mods:number id="823EDECED1AAAC819B3FD2EC01CFF3F4">4802</mods:number>
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<treatment id="039787F6D153260BFF7AFB054915FE1C" LSID="urn:lsid:plazi:treatment:039787F6D153260BFF7AFB054915FE1C" httpUri="http://treatment.plazi.org/id/039787F6D153260BFF7AFB054915FE1C" lastPageId="14" lastPageNumber="215" pageId="12" pageNumber="213">
<subSubSection id="C324656BD1532609FF7AFB054E16FB08" box="[151,569,1197,1224]" pageId="12" pageNumber="213" type="nomenclature">
<paragraph id="8B8136E0D1532609FF7AFB054E16FB08" blockId="12.[151,569,1197,1259]" box="[151,569,1197,1224]" pageId="12" pageNumber="213">
<heading id="D0C9818CD1532609FF7AFB054E16FB08" bold="true" box="[151,569,1197,1224]" fontSize="11" level="1" pageId="12" pageNumber="213" reason="1">
<emphasis id="B94AEAF2D1532609FF7AFB054E16FB08" bold="true" box="[151,569,1197,1224]" pageId="12" pageNumber="213">
<taxonomicName id="4C3E4D63D1532609FF7AFB054D87FB07" ID-CoL="8NHS8" authority="Obando &amp; Gironza &amp; Panche &amp; Aldrete, 2020" authorityName="Obando &amp; Gironza &amp; Panche &amp; Aldrete" authorityYear="2020" box="[151,424,1197,1224]" class="Insecta" family="Ptiloneuridae" genus="Loneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="altaquerensis" status="sp. nov.">
<emphasis id="B94AEAF2D1532609FF7AFB054D87FB07" bold="true" box="[151,424,1197,1224]" italics="true" pageId="12" pageNumber="213">Loneura altaquerensis</emphasis>
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<taxonomicNameLabel id="A2795789D1532609FE5DFB054DDDFB08" box="[432,498,1197,1223]" pageId="12" pageNumber="213" rank="species">n. sp.</taxonomicNameLabel>
Male
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C324656BD1532609FF7AFB794D08FB24" box="[151,295,1233,1259]" pageId="12" pageNumber="213" type="description">
<paragraph id="8B8136E0D1532609FF7AFB794D08FB24" blockId="12.[151,569,1197,1259]" box="[151,295,1233,1259]" pageId="12" pageNumber="213">
(
<figureCitation id="13052A65D1532609FF72FB794D30FB24" box="[159,287,1233,1259]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Figs 29–33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C324656BD153260BFF7AFAB04812FED0" lastPageId="14" lastPageNumber="215" pageId="12" pageNumber="213" type="diagnosis">
<paragraph id="8B8136E0D1532609FF7AFAB04F60FAB4" blockId="12.[151,1437,1304,2015]" pageId="12" pageNumber="213">
<emphasis id="B94AEAF2D1532609FF7AFAB04D3DFAFC" bold="true" box="[151,274,1304,1331]" pageId="12" pageNumber="213">Diagnosis.</emphasis>
Mesal lateral processes of the central sclerite of the hypandrium proximally broad, with margins converging to round apices (
<figureCitation id="13052A65D1532609FE43FA954E2DFA98" box="[430,514,1341,1367]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 32</figureCitation>
). It differs from the other species in the group by the shape of the hypandrium and by the shape of the phallosome endophallic sclerites (
<figureCitation id="13052A65D1532609FD02FAC94F6DFAB4" box="[751,834,1377,1403]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 33</figureCitation>
).
</paragraph>
<paragraph id="8B8136E0D1532609FF2AFA2C4FDDF92C" blockId="12.[151,1437,1304,2015]" pageId="12" pageNumber="213">
<emphasis id="B94AEAF2D1532609FF2AFA2C4D24FA50" bold="true" box="[199,267,1412,1439]" pageId="12" pageNumber="213">Color</emphasis>
. Body light brown. Head light brown (
<figureCitation id="13052A65D1532609FD38FA2D4F04FA50" box="[725,811,1413,1439]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 31</figureCitation>
), with an ochre band from each compound eye to the epistomal sulcus and ocellar triangle; genae dark brown; postgenae light brown. Compound eyes black, ocelli hyaline, with ochre centripetal crescents. Postclypeus light brown, with posterior ochre area V–shaped; anteclypeus and labrum light brown. Maxillary palps light brown, Mx4 dark brown. Antennae light brown, flagellomeres 1–3 with apices cream. Thorax pale brown, pronotum and tergal lobes of meso- and metathorax pale brown. Legs light brown, tarsi 2–3 dark brown, fore- and middle coxae with ochre spot basally; femora with distal brown spots. Wings hyaline (
<figureCitation id="13052A65D1532609FF15F9F54DB8F9B8" box="[248,407,1629,1655]" captionStart="FIGURES 130–135" captionStartId="40.[151,264,1904,1929]" captionTargetBox="[164,1428,193,1868]" captionTargetId="figure@40.[151,1435,181,1878]" captionTargetPageId="40" captionText="FIGURES 130–135. Loneura gorgonaensis García Aldrete, González &amp; Sarria. Female. 130. Forewing. 131. Hindwing. 132. Front view of head. 133. Subgenital plate. 134. Left paraproct and epiproct. 135. Ninth sternum and left gonapophyses. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3905022" httpUri="https://zenodo.org/record/3905022/files/figure.png" pageId="12" pageNumber="213">Figs 131, 132</figureCitation>
). Forewings (
<figureCitation id="13052A65D1532609FDDDF9F54EAEF9B8" box="[560,641,1629,1655]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 29</figureCitation>
) pterostigma with proximal and distal brown bands; veins light brown, with dark brown areolae in setal insertions, a dark brown spot on vein ends at wing margin. Hindwings (
<figureCitation id="13052A65D1532609FAACF92949BBF954" box="[1345,1428,1665,1691]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 30</figureCitation>
) with a brown spot at vein ends. Abdomen light brown, clunium pale brown. Hypandrium light brown; epiproct and paraprocts cream. Phallosome brown, with endophallic sclerites dark brown.
</paragraph>
<paragraph id="8B8136E0D153260BFF2AF9454812FED0" blockId="12.[151,1437,1304,2015]" lastBlockId="14.[151,1436,153,467]" lastPageId="14" lastPageNumber="215" pageId="12" pageNumber="213">
<emphasis id="B94AEAF2D1532609FF2AF9454D4EF8C8" bold="true" box="[199,353,1773,1799]" pageId="12" pageNumber="213">Morphology.</emphasis>
Head (
<figureCitation id="13052A65D1532609FE59F9454E26F8C8" box="[436,521,1773,1799]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 31</figureCitation>
): H/MxW: 1.48; H/d: 3.85; IO/MxW: 0.75. Vertex at the same level as the upper border of the compound eyes. Outer cusp of lacinial tips broad, with four short and widely rounded denticles. Mx4/ Mx2: 1.12. Forewings (
<figureCitation id="13052A65D1532609FE49F89D4DD5F880" box="[420,506,1845,1871]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 29</figureCitation>
): L/W: 2.58. Pterostigma: lp/wp: 4.43, areola postica tall, apically rounded: al/ah: 1.36, R4+5 sinuous, M five-branched, M5 forked. Hindwings (
<figureCitation id="13052A65D1532609FC8FF8F14F97F8BC" box="[866,952,1881,1907]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 30</figureCitation>
): l/w: 2.88; M two-branched. Hypandrium (
<figureCitation id="13052A65D1532609FF4DF8D54CDAF858" box="[160,245,1917,1943]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 32</figureCitation>
): deeply cleft in the middle, posterior projections stout, blunt ended, each with a subapical macroseta on inner margin, mesal lateral processes broad, triangular. Phallosome V-shaped anteriorly, side struts slender; external parameres laminar, wide, apex rounded, bearing pores and sclerotized short teeth on the outer margins (
<figureCitation id="13052A65D1532609FAD4F86D49A1F810" box="[1337,1422,1989,2015]" captionStart="FIGURES 29–33" captionStartId="13.[151,264,1910,1935]" captionTargetBox="[167,1425,199,1873]" captionTargetId="figure@13.[151,1435,199,1885]" captionTargetPageId="13" captionText="FIGURES 29–33. Loneura altaquerensis n. sp. Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="12" pageNumber="213">Fig. 33</figureCitation>
); four pairs of endophallic sclerites, anterior pair elongate, lateral sclerites wide, with concave margins, tapered, directed posteriorly; mesal pair wide basally, mesal sclerite processes distally curved outward, distally fused to posterior sclerite, tapered and apically curved outward. Paraprocts oval, with distal setal fields; sensory fields with 22–24 trichobothria on basal rosettes. Epiproct semioval, with short and long setae.
</paragraph>
</subSubSection>
<caption id="DF416668D1522608FF7AF8DE4DFFF87D" ID-DOI="http://doi.org/10.5281/zenodo.3904992" ID-Zenodo-Dep="3904992" httpUri="https://zenodo.org/record/3904992/files/figure.png" pageId="13" pageNumber="214" startId="13.[151,264,1910,1935]" targetBox="[167,1425,199,1873]" targetPageId="13">
<paragraph id="8B8136E0D1522608FF7AF8DE4DFFF87D" blockId="13.[151,1437,1910,1971]" pageId="13" pageNumber="214">
<emphasis id="B94AEAF2D1522608FF7AF8DE4D7AF840" bold="true" box="[151,341,1910,1935]" pageId="13" pageNumber="214">FIGURES 29–33.</emphasis>
<taxonomicName id="4C3E4D63D1522608FEB1F8D04E6EF840" authority="Obando &amp; Gironza &amp; Panche &amp; Aldrete, 2020" authorityName="Obando &amp; Gironza &amp; Panche &amp; Aldrete" authorityYear="2020" box="[348,577,1911,1935]" class="Insecta" family="Ptiloneuridae" genus="Loneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Psocodea" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="altaquerensis" status="sp. nov.">
<emphasis id="B94AEAF2D1522608FEB1F8D04E6EF840" box="[348,577,1911,1935]" italics="true" pageId="13" pageNumber="214">Loneura altaquerensis</emphasis>
</taxonomicName>
<emphasis id="B94AEAF2D1522608FDAAF8DF4EAEF840" bold="true" box="[583,641,1911,1935]" pageId="13" pageNumber="214">
<taxonomicNameLabel id="A2795789D1522608FDAAF8DF4EAEF840" box="[583,641,1911,1935]" pageId="13" pageNumber="214" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Male. 29. Forewing. 30. Hindwing. 31. Front view of head. 32. Hypandrium. 33. Phallosome. Scales in mm.
</paragraph>
</caption>
<subSubSection id="C324656BD151260BFF2AFE814F64FEA8" pageId="14" pageNumber="215" type="description">
<paragraph id="8B8136E0D151260BFF2AFE814F64FEA8" blockId="14.[151,1436,153,467]" pageId="14" pageNumber="215">
<emphasis id="B94AEAF2D151260BFF2AFE814D59FE8C" bold="true" box="[199,374,297,323]" pageId="14" pageNumber="215">Measurements</emphasis>
. FW: 3400, HW: 2375, F: 775, T: 1382, t1: 625, t2: 80, t3: 120, ctt1: 21, f1: 612, f2: 525, f3: 475, Mx4: 190, IO: 390, D: 296, d: 200, IO/d: 1.95, PO: 0.68.
</paragraph>
</subSubSection>
<subSubSection id="C324656BD151260BFF2AFED84809FE60" pageId="14" pageNumber="215" type="materials_examined">
<paragraph id="8B8136E0D151260BFF2AFED84809FE60" blockId="14.[151,1436,153,467]" pageId="14" pageNumber="215">
<emphasis id="B94AEAF2D151260BFF2AFED84DB8FE44" bold="true" box="[199,407,368,395]" pageId="14" pageNumber="215">Material studied.</emphasis>
<materialsCitation id="3B563CBDD151260BFE4DFED94809FE60" collectorName="M. Gonzalez." country="Colombia" elevation="1386" latitude="1.3" location="Altaquer. Rio Nambi Natural Reserve" longLatPrecision="21" longitude="-78.08389" pageId="14" pageNumber="215" specimenCount="1" stateProvince="Narino" typeStatus="holotype">
<typeStatus id="54858842D151260BFE4DFED94E27FE44" box="[416,520,369,395]" pageId="14" pageNumber="215" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F3297670D151260BFDB4FED94EDDFE44" box="[601,754,369,395]" name="Colombia" pageId="14" pageNumber="215">COLOMBIA</collectingCountry>
.
<collectingRegion id="49FAF802D151260BFD12FED94F7FFE44" box="[767,848,369,395]" country="Colombia" name="Narino" pageId="14" pageNumber="215">Nariño</collectingRegion>
.
<location id="8EE1603BD151260BFCB6FED94921FE44" LSID="urn:lsid:plazi:treatment:039787F6D153260BFF7AFB054915FE1C:8EE1603BD151260BFCB6FED94921FE44" box="[859,1294,368,395]" country="Colombia" latitude="1.3" longLatPrecision="21" longitude="-78.08389" name="Altaquer. Rio Nambi Natural Reserve" pageId="14" pageNumber="215" stateProvince="Narino">Altaquer. Rio Ñambí Natural Reserve</location>
,
<geoCoordinate id="EE0A5027D151260BFAF1FED949B6FE44" box="[1308,1433,369,395]" degrees="1" direction="north" minutes="18" orientation="latitude" pageId="14" pageNumber="215" precision="15" seconds="00" value="1.3">1º18’00”N</geoCoordinate>
:
<geoCoordinate id="EE0A5027D151260BFF7AFE3C4D27FE60" box="[151,264,404,431]" degrees="78" direction="west" minutes="5" orientation="longitude" pageId="14" pageNumber="215" precision="15" seconds="2" value="-78.08389">78º5’2”W</geoCoordinate>
,
<quantity id="4CC69B05D151260BFEFEFE3D4D44FE60" box="[275,363,405,431]" metricMagnitude="3" metricUnit="m" metricValue="1.386" pageId="14" pageNumber="215" unit="m" value="1386.0">
<elevation id="0013D1D3D151260BFEFEFE3D4D44FE60" box="[275,363,405,431]" metricMagnitude="3" metricUnit="m" metricValue="1.386" pageId="14" pageNumber="215" unit="m" value="1386.0">1386 m</elevation>
</quantity>
.
<collectingMethod id="527F4EF7D151260BFE9BFE3D4E07FE60" box="[374,552,405,431]" pageId="14" pageNumber="215">Black light trap</collectingMethod>
.
<collectorName id="26CB5336D151260BFDDFFE3D4EE4FE60" box="[562,715,405,431]" pageId="14" pageNumber="215">M. González.</collectorName>
MUSENUV slide code 29311.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C324656BD151260BFF2AFE114915FE1C" box="[199,1338,441,467]" pageId="14" pageNumber="215" type="etymology">
<paragraph id="8B8136E0D151260BFF2AFE114915FE1C" blockId="14.[151,1436,153,467]" box="[199,1338,441,467]" pageId="14" pageNumber="215">
<emphasis id="B94AEAF2D151260BFF2AFE114D61FE1C" bold="true" box="[199,334,441,467]" pageId="14" pageNumber="215">Etymology.</emphasis>
The specific name refers to Altaquer,
<collectingRegion id="49FAF802D151260BFD17FE114FF9FE1C" box="[762,982,441,467]" country="Colombia" name="Narino" pageId="14" pageNumber="215">Nariño Department</collectingRegion>
, where the
<typeStatus id="54858842D151260BFBB8FE114898FE1C" box="[1109,1207,441,467]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
was found.
</paragraph>
</subSubSection>
</treatment>
</document>
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