362 lines
64 KiB
XML
362 lines
64 KiB
XML
<document id="DA407A8990CCFD84C20E854DA933AF98" ID-DOI="10.11646/zootaxa.4756.1.1" ID-GBIF-Dataset="4d48d52e-6505-4f62-909f-5da5c2cfb489" ID-ISSN="1175-5334" ID-Zenodo-Dep="3736895" ID-ZooBank="urn:lsid:zoobank.org:pub:9FF62616-A7FA-4331-AC51-0F534400631D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1585767638433" checkinUser="carolina" docAuthor="Kluge, Nikita J." docDate="2020" docId="039787A6FFFA840E8CFBFF7FDA20FA63" docLanguage="en" docName="zootaxa.4756.1.1.pdf.imf" docOrigin="Zootaxa 4756 (1)" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Thraulodes nigrabdominalis Kluge, 2020, sp. n." docType="treatment" docVersion="7" lastPageNumber="107" masterDocId="FFAEFFDEFF9D84648C6CFFE6DB3EFFA2" masterDocTitle="Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species" masterLastPageNumber="142" masterPageNumber="1" pageNumber="104" updateTime="1698734461607" updateUser="plazi">
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<mods:title id="33EC3202D0C881B0F06D2000F4D9CD9A">Systematic position of Thraulodes Ulmer 1920 (Ephemeroptera: Leptophlebiidae) and descriptions of new and little-known species</mods:title>
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<mods:name id="C6567C11441038BD35EFDA6524496441" type="personal">
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<mods:namePart id="819D63E70FFFDD0DD9B04C92C5DC7B3E">Kluge, Nikita J.</mods:namePart>
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<mods:title id="91E80FC1437C8F484CBD7A2F4E2F6B9F">Zootaxa</mods:title>
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<mods:part id="F0E2281946D82D114F2011861A7AB802">
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<mods:date id="1155E8CDD0B8DF5AC7A34E6FC5B18E7D">2020</mods:date>
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<mods:number id="DBF1DA0C110D8E79CD8059131D4725E3">2020-03-26</mods:number>
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<mods:detail id="F776336BBA605253E24AC5E25BA4C3AA" type="volume">
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<mods:number id="B873447EE57FA376B8AD30907CD142AC">4756</mods:number>
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<mods:detail id="F74D5BB32E1168B935D13E9C1733D53D" type="issue">
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<mods:classification id="F63DB7B80E76A0EC269096DF2966B016">journal article</mods:classification>
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<mods:identifier id="5E265EF8D45E4F04E203A7EF08E5D760" type="DOI">10.11646/zootaxa.4756.1.1</mods:identifier>
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<mods:identifier id="E56773ADA204BAFA60053A117C5FE558" type="GBIF-Dataset">4d48d52e-6505-4f62-909f-5da5c2cfb489</mods:identifier>
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<mods:identifier id="0F7B81BBDA99444B079D6FB1B9B30D83" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="496FF6496426110A438CC837677A6CEE" type="Zenodo-Dep">3736895</mods:identifier>
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<treatment id="039787A6FFFA840E8CFBFF7FDA20FA63" ID-DOI="http://doi.org/10.5281/zenodo.3811808" ID-GBIF-Taxon="163420768" ID-Zenodo-Dep="3811808" LSID="urn:lsid:plazi:treatment:039787A6FFFA840E8CFBFF7FDA20FA63" httpUri="http://treatment.plazi.org/id/039787A6FFFA840E8CFBFF7FDA20FA63" lastPageId="106" lastPageNumber="107" pageId="103" pageNumber="104">
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<subSubSection id="C324653BFFFA84038CFBFF7FD95BFF16" box="[151,613,153,180]" pageId="103" pageNumber="104" type="nomenclature">
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<paragraph id="8B8136B0FFFA84038CFBFF7FD95BFF16" blockId="103.[151,613,153,216]" box="[151,613,153,180]" pageId="103" pageNumber="104">
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<heading id="D0C981DCFFFA84038CFBFF7FD95BFF16" bold="true" box="[151,613,153,180]" fontSize="10" level="2" pageId="103" pageNumber="104" reason="2">
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<emphasis id="B94AEAA2FFFA84038CFBFF7FD95BFF16" bold="true" box="[151,613,153,180]" pageId="103" pageNumber="104">
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14.
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<taxonomicName id="4C3E4D33FFFA84038CAFFF7FD922FF16" box="[195,540,153,180]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="103" pageNumber="104" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
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<emphasis id="B94AEAA2FFFA84038CAFFF7FD922FF16" bold="true" box="[195,540,153,180]" italics="true" pageId="103" pageNumber="104">Thraulodes nigrabdominalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27957D9FFFA84038E4FFF7CD95BFF16" box="[547,613,154,180]" pageId="103" pageNumber="104" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C324653BFFFA84038CFBFF5BDA7AFF7A" box="[151,324,189,216]" pageId="103" pageNumber="104" type="description">
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<paragraph id="8B8136B0FFFA84038CFBFF5BDA7AFF7A" blockId="103.[151,613,153,216]" box="[151,324,189,216]" pageId="103" pageNumber="104">
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(
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<figureCitation id="13052A35FFFA84038CF3FF5BDA05FF7A" box="[159,315,189,216]" captionStart-0="FIGURES 450–463" captionStart-1="FIGURES 464–473" captionStart-2="FIGURES 474–478" captionStartId-0="104.[151,264,1826,1851]" captionStartId-1="106.[151,264,188,213]" captionStartId-2="107.[151,264,1689,1714]" captionTargetBox-0="[157,1429,166,1806]" captionTargetBox-2="[154,1422,186,1652]" captionTargetId-0="figure@104.[157,1429,166,1806]" captionTargetId-1="figure@105.[151,1436,181,1987]" captionTargetId-2="figure@107.[151,1436,179,1661]" captionTargetPageId-0="104" captionTargetPageId-1="105" captionTargetPageId-2="107" captionText-0="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." captionText-1="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." captionText-2="FIGURES 474–478. Thraulodes nigrabdominalis sp. n. 474, genitalia of male imago in ventral view; 475, their subimaginal exuviae; 476, hind wing; 477–478, protogonostyli and protopenis of presumably associated larva, with subimaginal gonostyli and styliger at early stage of development shown by interrupted line (474–476, holotype)." figureDoi-0="http://doi.org/10.5281/zenodo.3737022" figureDoi-1="http://doi.org/10.5281/zenodo.3737024" figureDoi-2="http://doi.org/10.5281/zenodo.3737026" httpUri-0="https://zenodo.org/record/3737022/files/figure.png" httpUri-1="https://zenodo.org/record/3737024/files/figure.png" httpUri-2="https://zenodo.org/record/3737026/files/figure.png" pageId="103" pageNumber="104">Figs 450–478</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C324653BFFFA84038CFBFF19DF48FEBB" box="[151,1142,255,281]" pageId="103" pageNumber="104" type="etymology">
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<paragraph id="8B8136B0FFFA84038CFBFF19DF48FEBB" blockId="103.[151,1437,255,2045]" box="[151,1142,255,281]" pageId="103" pageNumber="104">
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<emphasis id="B94AEAA2FFFA84038CFBFF19DA20FEBB" bold="true" box="[151,286,255,281]" pageId="103" pageNumber="104">Etymology.</emphasis>
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Allusion to uniformly dark brown, nearly black color of abdomen of imago.
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</paragraph>
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</subSubSection>
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<subSubSection id="C324653BFFFA84038CABFEC4DF0EFE24" pageId="103" pageNumber="104" type="materials_examined">
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<paragraph id="8B8136B0FFFA84038CABFEC4DF0EFE24" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
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<emphasis id="B94AEAA2FFFA84038CABFEC4DA8EFE9F" bold="true" box="[199,432,290,317]" pageId="103" pageNumber="104">Material examined.</emphasis>
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<materialsCitation id="3B563CEDFFFA84038DD9FEC5D958FEC3" ID-GBIF-Occurrence="2594509925" collectingDate="2006-02-23" collectingDateMax="2006-02-25" collectingDateMin="2006-02-23" collectionCode="XXXIII" collectorName="Aquaytia & Pampa Yurac & N. Kluge." country="Peru" latitude="-9.066667" longLatPrecision="1300" longitude="-75.95" pageId="103" pageNumber="104" specimenCount="2" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B94AEAA2FFFA84038DD9FEC5D912FE9F" bold="true" box="[437,556,291,317]" pageId="103" pageNumber="104">
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<typeStatus id="54858812FFFA84038DD9FEC5D918FE9F" box="[437,550,291,317]" pageId="103" pageNumber="104" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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S-I
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<specimenCount id="9D38FD39FFFA84038E39FEC5D956FE9F" box="[597,616,291,317]" pageId="103" pageNumber="104" type="male">♂</specimenCount>
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{
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<specimenCount id="9D38FD39FFFA84038E1AFEC5D9D8FE9F" box="[630,742,291,317]" pageId="103" pageNumber="104" type="generic">specimen</specimenCount>
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[
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<collectionCode id="ED2FAE75FFFA84038E9EFEC4D86EFE9F" box="[754,848,290,317]" pageId="103" pageNumber="104">XXXIII</collectionCode>
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] (5) 2006} PPERU, Aquaytia, Pampa Yurac (
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<geoCoordinate id="EE0A5077FFFA8403892BFEC5DEA9FE9F" box="[1351,1431,291,317]" degrees="9" direction="south" minutes="04" orientation="latitude" pageId="103" pageNumber="104" precision="925" value="-9.066667">9°04’S</geoCoordinate>
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,
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<geoCoordinate id="EE0A5077FFFA84038CFBFEA0DA3FFEC3" box="[151,257,326,353]" degrees="75" direction="west" minutes="57" orientation="longitude" pageId="103" pageNumber="104" precision="925" value="-75.95">75°57’W</geoCoordinate>
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),
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<date id="FF801070FFFA84038D79FEA0DA88FEC3" box="[277,438,326,353]" pageId="103" pageNumber="104" value="2006-02-23" valueMax="2006-02-25" valueMin="2006-02-23">
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<collectingDate id="EFC4E998FFFA84038D79FEA0DA88FEC3" box="[277,438,326,353]" pageId="103" pageNumber="104" value="2006-02-23" valueMax="2006-02-25" valueMin="2006-02-23">23–25.II.2006</collectingDate>
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</date>
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, coll. N. Kluge.
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</materialsCitation>
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<materialsCitation id="3B563CEDFFFA84038E00FEA1DF0EFE24" ID-GBIF-Occurrence="2594509902" collectingDate="2006-02-23" collectingDateMax="2006-02-25" collectingDateMin="2006-02-23" collectionCode="XXXIII" collectorName="Aquaytia & Pampa Yurac & N. Kluge." latitude="-9.066667" longLatPrecision="1300" longitude="-75.95" pageId="103" pageNumber="104" specimenCount="6" specimenCount-female="1" specimenCount-larva="4" specimenCount-male="1" typeStatus="paratype">
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<emphasis id="B94AEAA2FFFA84038E00FEA1D9DAFEC3" bold="true" box="[620,740,327,353]" pageId="103" pageNumber="104">
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<typeStatus id="54858812FFFA84038E00FEA1D9DAFEC3" box="[620,740,327,353]" pageId="103" pageNumber="104" type="paratype">Paratypes</typeStatus>
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</emphasis>
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(presumably associated): the same locality, date and collector:
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<specimenCount id="9D38FD39FFFA84038CFBFE8DDB80FE27" box="[151,190,363,389]" pageId="103" pageNumber="104" type="larva">1 L</specimenCount>
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<specimenCount id="9D38FD39FFFA84038CD3FE8DDBEDFE27" box="[191,211,363,389]" pageId="103" pageNumber="104" type="male">♂</specimenCount>
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(last instar),
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<specimenCount id="9D38FD39FFFA84038D0AFE8DDAAFFE27" box="[358,401,363,389]" pageId="103" pageNumber="104" type="larva">3 L</specimenCount>
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<specimenCount id="9D38FD39FFFA84038DFDFE8DDA9FFE27" box="[401,417,363,389]" pageId="103" pageNumber="104" type="female">♀</specimenCount>
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(penultimate instar), 1 larval exuviae (penultimate instar).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C324653BFFFA840E8CABFE68D8E7FD53" lastPageId="106" lastPageNumber="107" pageId="103" pageNumber="104" type="description">
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<paragraph id="8B8136B0FFFA84038CABFE68DA5AFE0B" blockId="103.[151,1437,255,2045]" box="[199,356,398,425]" pageId="103" pageNumber="104">
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<emphasis id="B94AEAA2FFFA84038CABFE68DA5AFE0B" bold="true" box="[199,356,398,425]" pageId="103" pageNumber="104">Descriptions.</emphasis>
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</paragraph>
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<paragraph id="8B8136B0FFFA84038CABFE55D867FD23" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
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<emphasis id="B94AEAA2FFFA84038CABFE55DA35FE6F" bold="true" box="[199,267,435,461]" italics="true" pageId="103" pageNumber="104">Larva</emphasis>
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(presumably associated). CUTICULAR COLORATION (
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<figureCitation id="13052A35FFFA84038FF5FE55DF94FE6F" box="[921,1194,434,461]" captionStart="FIGURES 450–463" captionStartId="104.[151,264,1826,1851]" captionTargetBox="[157,1429,166,1806]" captionTargetId="figure@104.[157,1429,166,1806]" captionTargetPageId="104" captionText="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="103" pageNumber="104">Figs 459–453, 456–458</figureCitation>
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). Dorsal side of head, pronotum and mesonotum brownish, with lighter and darker areas. Abdominal terga I–II or I–III either light brown, or with light brown markings; terga III–VIII either uniformly light, nearly colorless, or tergum VII bordered by reddish-brown anteriorly; terga VIII–IX contrastingly colored by intensive reddish-brown; tergum X light, nearly colorless. Femora mostly light, fore femur with subapical brown band, middle and hind femora with basal and subapical brown bands. Tibia and tarsus uniformly light brownish.
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</paragraph>
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<paragraph id="8B8136B0FFFA84038CABFD6DDA34FD4F" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
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HYPODERMAL COLORATION. Abdominal terga, besides peculiar cuticular coloration (see above) with narrow blackish lines bordering posterior margins of some terga (
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<figureCitation id="13052A35FFFA84038F0DFD49DF3AFD6B" box="[865,1028,686,713]" captionStart="FIGURES 450–463" captionStartId="104.[151,264,1826,1851]" captionTargetBox="[157,1429,166,1806]" captionTargetId="figure@104.[157,1429,166,1806]" captionTargetPageId="104" captionText="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="103" pageNumber="104">Figs 453, 459</figureCitation>
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). Tergalii gray, tracheae black (
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<figureCitation id="13052A35FFFA84038907FD49DBC0FD4F" captionStart="FIGURES 450–463" captionStartId="104.[151,264,1826,1851]" captionTargetBox="[157,1429,166,1806]" captionTargetId="figure@104.[157,1429,166,1806]" captionTargetPageId="104" captionText="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="103" pageNumber="104">Figs 454–455</figureCitation>
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).
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</paragraph>
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<paragraph id="8B8136B0FFFA84038CABFD11D90FFCDF" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
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SHAPE AND SETATION. Clypeus parallel-sided; labrum 1.3–1.4 times wider than clypeus (
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<figureCitation id="13052A35FFFA840388BCFD11DE0CFCB3" box="[1232,1330,758,785]" captionStart="FIGURES 450–463" captionStartId="104.[151,264,1826,1851]" captionTargetBox="[157,1429,166,1806]" captionTargetId="figure@104.[157,1429,166,1806]" captionTargetPageId="104" captionText="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="103" pageNumber="104">Fig. 459</figureCitation>
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). Labrum widest at midlength; initial fore margin (turned ventrally) without median emargination, shallowly concave, with all 5 denticles wide; anterior transverse setal row regular (as in
|
||
<figureCitation id="13052A35FFFA84038F32FCD9D88FFCFB" box="[862,945,831,857]" captionStart="FIGURES 86–93" captionStartId="37.[151,264,188,213]" captionTargetId="figure@36.[151,1436,342,2018]" captionTargetPageId="36" captionText="FIGURES 86–93. Thraulodes telegraphicus. 86, median longitudinal section of labrum in usual condition; 87, the same, in spread condition; 88, labrum in spread condition in dorsal view (setal rows shown as dotted lines); 89, transverse section of hind tibia; 90–93, tergalii I, III, VI and VII. Abbreviations: anter.r, anterior transverse row of setae; dent., five denticles separated by six sensilla; poster.r, posterior transverse row of setae; i.f, inner field of stout pointed setae; i-a.h, inner-anterior recurved hairs; i-a.r, inner-anterior row of stout setae; o-a.r, outer-anterior row of stout setae; o.h, outer hairs; o-p.r, outer-posterior row of stout setae; p.r, row of posterior pointed pectinate setae." figureDoi="http://doi.org/10.5281/zenodo.3736921" httpUri="https://zenodo.org/record/3736921/files/figure.png" pageId="103" pageNumber="104">Fig. 88</figureCitation>
|
||
), as wide as all 5 denticles. Maxilla with 19 pectinate setae in apical-ventral row.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABFC61D898FC67" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
|
||
Femora: Stout setae on anterior surface narrowed distally and truncated (
|
||
<figureCitation id="13052A35FFFA84038F90FC61DF61FC00" box="[1020,1119,903,930]" captionStart="FIGURES 450–463" captionStartId="104.[151,264,1826,1851]" captionTargetBox="[157,1429,166,1806]" captionTargetId="figure@104.[157,1429,166,1806]" captionTargetPageId="104" captionText="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="103" pageNumber="104">Fig. 461</figureCitation>
|
||
). Irregular row of hairs near inner margin absent on fore and middle femora, sparse on hind femur.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABFC29DF87FB90" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">Fore tibia: outer hairs form two irregular rows; inner-anterior row of recurved hairs absent; inner-anterior row of stout setae absent; inner field of stout pointed setae dense (i.e. setae longer than distances between them), consists of pointed bipectinate and smooth setae, situated irregularly (about 3–4 setae in cross section).</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABFBDDDFA1FB63" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
|
||
Hind tibia (
|
||
<figureCitation id="13052A35FFFA84038D2BFBDDDAD6FBF4" box="[327,488,1083,1110]" captionStart="FIGURES 450–463" captionStartId="104.[151,264,1826,1851]" captionTargetBox="[157,1429,166,1806]" captionTargetId="figure@104.[157,1429,166,1806]" captionTargetPageId="104" captionText="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="103" pageNumber="104">Figs 462–463</figureCitation>
|
||
): each of three rows of stout setae—outer-anterior, outer-posterior and inner-anterior ones—contains moderately long, narrowing distally stout setae with apices either blunt, or pointed; outer-anterior row, besides this, can contain long spoon-like setae; hairs located between outer-anterior and outer-posterior rows, form one irregular row (besides row of hairs posteriad of outer-posterior row of stout setae).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABFB2DDF9CFB47" blockId="103.[151,1437,255,2045]" box="[199,1186,1227,1253]" pageId="103" pageNumber="104">Claws with 2–4 denticles on rigid portion, with minute denticles on articulatory portion.</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABFB09D950FAD7" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
|
||
Tergalii (
|
||
<figureCitation id="13052A35FFFA84038D5CFB09DAEAFAAB" box="[304,468,1262,1289]" captionStart="FIGURES 450–463" captionStartId="104.[151,264,1826,1851]" captionTargetBox="[157,1429,166,1806]" captionTargetId="figure@104.[157,1429,166,1806]" captionTargetPageId="104" captionText="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="103" pageNumber="104">Figs 454–455</figureCitation>
|
||
): wide; on both lamellae main trachea mainly with branches directed toward costal margin; dorsal lamella bent backward, with costal margin more convex than anal, widest in proximal or middle part, gradually narrowed toward apex, with slender apical filament; ventral lamella widest near base, gradually narrowed toward apex, with slender apical filament.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABFA99DA6AFA43" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
|
||
Male genitalia in last larval instar (
|
||
<figureCitation id="13052A35FFFA84038E34FA99D981FA3B" box="[600,703,1407,1433]" captionStart="FIGURES 450–463" captionStartId="104.[151,264,1826,1851]" captionTargetBox="[157,1429,166,1806]" captionTargetId="figure@104.[157,1429,166,1806]" captionTargetPageId="104" captionText="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="103" pageNumber="104">Figs 460</figureCitation>
|
||
,
|
||
<figureCitation id="13052A35FFFA84038EA0FA99D80AFA3B" box="[716,820,1407,1433]" captionStart="FIGURES 474–478" captionStartId="107.[151,264,1689,1714]" captionTargetBox="[154,1422,186,1652]" captionTargetId="figure@107.[151,1436,179,1661]" captionTargetPageId="107" captionText="FIGURES 474–478. Thraulodes nigrabdominalis sp. n. 474, genitalia of male imago in ventral view; 475, their subimaginal exuviae; 476, hind wing; 477–478, protogonostyli and protopenis of presumably associated larva, with subimaginal gonostyli and styliger at early stage of development shown by interrupted line (474–476, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737026" httpUri="https://zenodo.org/record/3737026/files/figure.png" pageId="103" pageNumber="104">477–478</figureCitation>
|
||
): protogonostyli short and separated one from another by shallow concavity. Each protopenis lobe with extremely short projected gonopore-bearing process; gonopores opened caudally.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABFA0CD94EF97F" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
|
||
<emphasis id="B94AEAA2FFFA84038CABFA0CDA02F9A7" bold="true" box="[199,316,1514,1541]" italics="true" pageId="103" pageNumber="104">Subimago</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFFA84038D26FA0DD922F9A7" box="[330,540,1515,1541]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="103" pageNumber="104">Figs 464, 470, 473</figureCitation>
|
||
,
|
||
<figureCitation id="13052A35FFFA84038E4AFA0CD96AF9A7" box="[550,596,1514,1541]" captionStart="FIGURES 474–478" captionStartId="107.[151,264,1689,1714]" captionTargetBox="[154,1422,186,1652]" captionTargetId="figure@107.[151,1436,179,1661]" captionTargetPageId="107" captionText="FIGURES 474–478. Thraulodes nigrabdominalis sp. n. 474, genitalia of male imago in ventral view; 475, their subimaginal exuviae; 476, hind wing; 477–478, protogonostyli and protopenis of presumably associated larva, with subimaginal gonostyli and styliger at early stage of development shown by interrupted line (474–476, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737026" httpUri="https://zenodo.org/record/3737026/files/figure.png" pageId="103" pageNumber="104">475</figureCitation>
|
||
). CUTICULAR COLORATION. Cuticle mostly brown. Mesonotum darker brown, with darker brown antelateroparapsidal and lateroparapsidal sutures, lighter oblique stripes on posterior scutal protuberances and colorless achromozones; medioscutal and submedioscutal chromozones equally colored by brown, separated by darker brown medioscutal suture (
|
||
<figureCitation id="13052A35FFFA84038F5FF9B1D8A7F9D0" box="[819,921,1623,1650]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="103" pageNumber="104">Fig. 464</figureCitation>
|
||
). On all legs femur light brown, bordered by darker brown on outer and inner margins and apically; tibia and tarsus light brown (
|
||
<figureCitation id="13052A35FFFA84038836F99DDF81F937" box="[1114,1215,1659,1685]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="103" pageNumber="104">Fig. 470</figureCitation>
|
||
). Wings brown due to brown ring around base of each microtrichion; microtrichia blackish-brown (as in
|
||
<figureCitation id="13052A35FFFA8403880DF979DFFAF91B" box="[1121,1220,1695,1721]" captionStart="FIGURES 427–438" captionStartId="100.[151,264,1753,1778]" captionTargetBox="[151,1436,166,1733]" captionTargetId="figure@100.[151,1436,166,1733]" captionTargetPageId="100" captionText="FIGURES 427–438. Thraulodes niger sp. n. 427–428, head and thorax of male imago in lateral and dorsal view; 429, abdominal terga and sterna of male imago; 430, the same of male subimago; 431, subimaginal mesonotum; 432–433, genitalia of male imago with focus on styliger and on penis; 434, female imago; 435, its left fore wing; 436–437, its fore and middle legs; 438, subimaginal exuviae of fore wing (427–429, 432–433, 438, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737015" httpUri="https://zenodo.org/record/3737015/files/figure.png" pageId="103" pageNumber="104">Fig. 438</figureCitation>
|
||
). Abdominal terga, sterna, gonostyli and caudalii light brown.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABF901DE6FF887" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
|
||
TEXTURE. On tarsi of all legs, 1st tarsomere with microtrichia (as tibia), 2nd–5th tarsomeres coved by blunt microlepides; pointed microlepides present near apical margins of 2nd–4th tarsomeres of all legs (
|
||
<figureCitation id="13052A35FFFA8403888EF8EDDE7AF887" box="[1250,1348,1803,1829]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="103" pageNumber="104">Fig. 473</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABF8C8D95EF833" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
|
||
<emphasis id="B94AEAA2FFFA84038CABF8C8DA6EF8EB" bold="true" box="[199,336,1838,1865]" italics="true" pageId="103" pageNumber="104">Male imago</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFFA84038D31F8C9DAC5F8EB" box="[349,507,1838,1865]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="103" pageNumber="104">Figs 465–469</figureCitation>
|
||
). Coloration of body and legs nearly uniformly dark brown. Head and antennae dark brown. Dorsal eyes contiguous medially, dull orange. Thorax dorsally, laterally and ventrally equally dark brown, with pleural membranes brownish ocher.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFFA84038CABF87DDF3CF85F" blockId="103.[151,1437,255,2045]" pageId="103" pageNumber="104">
|
||
Cuticle of legs light brown; hypodermal pigmentation of femur, tibia and tarsus of all legs uniformly dark brown. In
|
||
<typeStatus id="54858812FFFA84038D65F859DA52F87B" box="[265,364,1983,2009]" pageId="103" pageNumber="104" type="holotype">holotype</typeStatus>
|
||
,relation of fore femur to fore wing length 68:260; proportions femur/tibia/tarsomeres on fore leg 68:87:3:14:11:9:8; on middle leg 63:65:3:3:3:3:7; on hind leg 77:68:3:3:3:3:7.
|
||
</paragraph>
|
||
<caption id="DF416638FFF5840C8CFBF8C4DF7DF84C" ID-DOI="http://doi.org/10.5281/zenodo.3737022" ID-Zenodo-Dep="3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="104" pageNumber="105" startId="104.[151,264,1826,1851]" targetBox="[157,1429,166,1806]" targetPageId="104">
|
||
<paragraph id="8B8136B0FFF5840C8CFBF8C4DF7DF84C" blockId="104.[151,1436,1826,2031]" pageId="104" pageNumber="105">
|
||
<emphasis id="B94AEAA2FFF5840C8CFBF8C4DA53F899" bold="true" box="[151,365,1826,1851]" pageId="104" pageNumber="105">FIGURES 450–463.</emphasis>
|
||
Larvae, presumably attributed to
|
||
<taxonomicName id="4C3E4D33FFF5840C8EAAF8C5D8D6F899" box="[710,1000,1827,1851]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="104" pageNumber="105" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF5840C8EAAF8C5D8D6F899" box="[710,1000,1827,1851]" italics="true" pageId="104" pageNumber="105">Thraulodes nigrabdominalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF5840C8F83F8C5DF16F899" bold="true" box="[1007,1064,1827,1851]" pageId="104" pageNumber="105">
|
||
<taxonomicNameLabel id="A27957D9FFF5840C8F83F8C5DF16F899" box="[1007,1064,1827,1851]" pageId="104" pageNumber="105" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF416638FFF7840E8CFBFF5AD874FEBE" ID-DOI="http://doi.org/10.5281/zenodo.3737024" ID-Zenodo-Dep="3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="106" pageNumber="107" targetBox="[151,1436,181,1987]" targetPageId="105">
|
||
<paragraph id="8B8136B0FFF7840E8CFBFF5AD874FEBE" blockId="106.[151,1443,188,285]" pageId="106" pageNumber="107">
|
||
<emphasis id="B94AEAA2FFF7840E8CFBFF5ADA53FF77" bold="true" box="[151,365,188,213]" pageId="106" pageNumber="107">FIGURES 464–473.</emphasis>
|
||
<taxonomicName id="4C3E4D33FFF7840E8D18FF5AD9A8FF76" box="[372,662,188,212]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8D18FF5AD9A8FF76" box="[372,662,188,212]" italics="true" pageId="106" pageNumber="107">Thraulodes nigrabdominalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8EF0FF5AD9E8FF76" bold="true" box="[668,726,188,212]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8EF0FF5AD9E8FF76" box="[668,726,188,212]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8136B0FFF7840E8CABFE89DE04FE70" blockId="106.[151,1437,367,1473]" pageId="106" pageNumber="107">
|
||
Fore and hind wing with longitudinal and cross veins ocher. Costal cross veins proximad of bulla either absent, of few, colorless and very thin, so that visible only under high magnification (as in
|
||
<figureCitation id="13052A35FFF7840E882FFE75DF9BFE0F" box="[1091,1189,403,429]" captionStart="FIGURES 133–142" captionStartId="45.[151,264,1898,1923]" captionTargetBox="[151,1436,166,1878]" captionTargetId="figure@45.[151,1436,166,1878]" captionTargetPageId="45" captionText="FIGURES 133–142. Thraulodes panamensis sp. n. 133, pterostigma; 134–136, male imago in lateral in ventral and dorsal view; 137–138, left half of subimaginal exuviae of mesonotum of two different specimens; 139, female imago; 140–141, genitalia in lateral and ventral view; 142, costal margin of fore wing (arrows show proximal costal cross veins) (133, 134, 136, 137, 141, 142, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3736935" httpUri="https://zenodo.org/record/3736935/files/figure.png" pageId="106" pageNumber="107">Fig. 142</figureCitation>
|
||
). Pterostigmatic cross veins moderately dense, oblique, non-branched. Hind wing with brown macula on costal brace (
|
||
<figureCitation id="13052A35FFF7840E88A6FE51DE10FE70" box="[1226,1326,439,466]" captionStart="FIGURES 474–478" captionStartId="107.[151,264,1689,1714]" captionTargetBox="[154,1422,186,1652]" captionTargetId="figure@107.[151,1436,179,1661]" captionTargetPageId="107" captionText="FIGURES 474–478. Thraulodes nigrabdominalis sp. n. 474, genitalia of male imago in ventral view; 475, their subimaginal exuviae; 476, hind wing; 477–478, protogonostyli and protopenis of presumably associated larva, with subimaginal gonostyli and styliger at early stage of development shown by interrupted line (474–476, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737026" httpUri="https://zenodo.org/record/3737026/files/figure.png" pageId="106" pageNumber="107">Fig. 476</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFF7840E8CABFE3DDA05FDBB" blockId="106.[151,1437,367,1473]" pageId="106" pageNumber="107">
|
||
Abdominal hypodermal coloration: all terga and sterna uniformly brown (
|
||
<figureCitation id="13052A35FFF7840E8F93FE3DDF5CFE54" box="[1023,1122,475,502]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="106" pageNumber="107">Fig. 466</figureCitation>
|
||
). Caudalii brown, with joinings darkened.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFF7840E8CABFDC5D880FD0B" blockId="106.[151,1437,367,1473]" pageId="106" pageNumber="107">
|
||
Genitalia (
|
||
<figureCitation id="13052A35FFF7840E8D50FDC5DAE6FD9F" box="[316,472,547,573]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="106" pageNumber="107">Figs 471–472</figureCitation>
|
||
,
|
||
<figureCitation id="13052A35FFF7840E8D8FFDC4D975FD9F" box="[483,587,546,573]" captionStart="FIGURES 474–478" captionStartId="107.[151,264,1689,1714]" captionTargetBox="[154,1422,186,1652]" captionTargetId="figure@107.[151,1436,179,1661]" captionTargetPageId="107" captionText="FIGURES 474–478. Thraulodes nigrabdominalis sp. n. 474, genitalia of male imago in ventral view; 475, their subimaginal exuviae; 476, hind wing; 477–478, protogonostyli and protopenis of presumably associated larva, with subimaginal gonostyli and styliger at early stage of development shown by interrupted line (474–476, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737026" httpUri="https://zenodo.org/record/3737026/files/figure.png" pageId="106" pageNumber="107">474–475</figureCitation>
|
||
): Styliger, gonostyli and penis uniformly brown. Dorsal extension of styliger more or less prominent. Penis lobes long, narrow and slightly divergent; each penis lobe parallel-sided, without lateral pouch, not widened apically, without ear. Telopenes in form of spear-like rolls, attached at apices of penis lobes, directed caudally-medially, stout, with groove opened dorsally-laterally.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFF7840E8CABFD54D931FD6F" blockId="106.[151,1437,367,1473]" box="[199,527,690,717]" pageId="106" pageNumber="107">
|
||
<emphasis id="B94AEAA2FFF7840E8CABFD54DAADFD6F" bold="true" box="[199,403,690,717]" italics="true" pageId="106" pageNumber="107">Female and eggs.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFF7840E8CABFD30D8E7FD53" blockId="106.[151,1437,367,1473]" box="[199,985,726,754]" pageId="106" pageNumber="107">
|
||
<emphasis id="B94AEAA2FFF7840E8CABFD30DA70FD53" bold="true" box="[199,334,726,753]" pageId="106" pageNumber="107">Dimension.</emphasis>
|
||
Fore wing length (and approximate body length)
|
||
<quantity id="4CC69B55FFF7840E8F10FD30D8EBFD53" box="[892,981,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="106" pageNumber="107" unit="mm" value="6.5">6.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C324653BFFF7840E8CABFD1CDA20FA63" pageId="106" pageNumber="107" type="discussion">
|
||
<paragraph id="8B8136B0FFF7840E8CABFD1CDEA3FB64" blockId="106.[151,1437,367,1473]" pageId="106" pageNumber="107">
|
||
<emphasis id="B94AEAA2FFF7840E8CABFD1CDAA4FCB7" bold="true" box="[199,410,762,789]" pageId="106" pageNumber="107">Stage association.</emphasis>
|
||
Winged stages were not reared from larvae, and subimaginal parts were not extracted from mature larva, so association of larva and winged stages is only presumed. This association is based on the following features.Last instar larva has tibia of hind leg tinged with blackish hypodermal pigment, that testifies about black tibia in winged stages and placing this species to the
|
||
<taxonomicName id="4C3E4D33FFF7840E8ECCFC8ED9E5FC23" box="[672,731,872,897]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="niger">
|
||
<emphasis id="B94AEAA2FFF7840E8ECCFC8ED9E5FC23" box="[672,731,872,897]" italics="true" pageId="106" pageNumber="107">niger</emphasis>
|
||
</taxonomicName>
|
||
group. In contrast to other three species of this group, the imago of
|
||
<taxonomicName id="4C3E4D33FFF7840E8CD9FC6DDAA7FC07" box="[181,409,907,933]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8CD9FC6DDAA7FC07" box="[181,409,907,933]" italics="true" pageId="106" pageNumber="107">Th. nigrabdominalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8DCCFC6DDAE1FC07" bold="true" box="[416,479,907,933]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8DCCFC6DDAE1FC07" box="[416,479,907,933]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has no hypodermal pigmentation on area of costal brace of fore wing; this coloration, when present in winged stages, is expressed on the larval protopteron as well, being developed not only in last larval instar, as well in some of the previous instars; larvae under consideration lack this pigmentation, which testifies in favor of their belonging to
|
||
<taxonomicName id="4C3E4D33FFF7840E8DAEFC11D998FBB3" box="[450,678,1015,1041]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8DAEFC11D998FBB3" box="[450,678,1015,1041]" italics="true" pageId="106" pageNumber="107">Th. nigrabdominalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8EC1FC11D9D5FBB3" bold="true" box="[685,747,1015,1041]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8EC1FC11D9D5FBB3" box="[685,747,1015,1041]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
In the examined male larva of last instar, the styliger, being at a very early stage of its development, already has a narrow-parabolic shape (
|
||
<figureCitation id="13052A35FFF7840E8F8DFBFDDF7CFB97" box="[993,1090,1051,1077]" captionStart="FIGURES 474–478" captionStartId="107.[151,264,1689,1714]" captionTargetBox="[154,1422,186,1652]" captionTargetId="figure@107.[151,1436,179,1661]" captionTargetPageId="107" captionText="FIGURES 474–478. Thraulodes nigrabdominalis sp. n. 474, genitalia of male imago in ventral view; 475, their subimaginal exuviae; 476, hind wing; 477–478, protogonostyli and protopenis of presumably associated larva, with subimaginal gonostyli and styliger at early stage of development shown by interrupted line (474–476, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737026" httpUri="https://zenodo.org/record/3737026/files/figure.png" pageId="106" pageNumber="107">Fig. 477</figureCitation>
|
||
), which is the same as in imago and subimago of
|
||
<taxonomicName id="4C3E4D33FFF7840E8D33FBD9D978FBFB" box="[351,582,1087,1113]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8D33FBD9D978FBFB" box="[351,582,1087,1113]" italics="true" pageId="106" pageNumber="107">Th. nigrabdominalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8E23FBD9D9AEFBFB" bold="true" box="[591,656,1087,1113]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8E23FBD9D9AEFBFB" box="[591,656,1087,1113]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFF7840E8ECDFBD9D87CFBFB" box="[673,834,1086,1113]" captionStart="FIGURES 474–478" captionStartId="107.[151,264,1689,1714]" captionTargetBox="[154,1422,186,1652]" captionTargetId="figure@107.[151,1436,179,1661]" captionTargetPageId="107" captionText="FIGURES 474–478. Thraulodes nigrabdominalis sp. n. 474, genitalia of male imago in ventral view; 475, their subimaginal exuviae; 476, hind wing; 477–478, protogonostyli and protopenis of presumably associated larva, with subimaginal gonostyli and styliger at early stage of development shown by interrupted line (474–476, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737026" httpUri="https://zenodo.org/record/3737026/files/figure.png" pageId="106" pageNumber="107">Figs 474–475</figureCitation>
|
||
); this differs from the sympatric species of the same group,
|
||
<taxonomicName id="4C3E4D33FFF7840E8C89FB85DA55FBDF" box="[229,363,1123,1149]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="nigripes" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8C89FB85DA55FBDF" box="[229,363,1123,1149]" italics="true" pageId="106" pageNumber="107">Th. nigripes</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8D1DFB85DA91FBDF" bold="true" box="[369,431,1123,1149]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8D1DFB85DA91FBDF" box="[369,431,1123,1149]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, whose styliger is wide-rectangular (
|
||
<figureCitation id="13052A35FFF7840E8F2AFB85D89BFBDF" box="[838,933,1123,1149]" captionStart="FIGURES 498–507" captionStartId="111.[151,264,1685,1710]" captionTargetBox="[157,1420,225,1620]" captionTargetId="figure@111.[151,1436,212,1639]" captionTargetPageId="111" captionText="FIGURES 498–507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500–501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503–506, tergalii I, III, VI and VII (498–501, 503–506, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="106" pageNumber="107">Fig. 499</figureCitation>
|
||
) and gets this shape at the same early stage of developnemt (
|
||
<figureCitation id="13052A35FFF7840E8D5BFB61DAA7FB03" box="[311,409,1158,1185]" captionStart="FIGURES 498–507" captionStartId="111.[151,264,1685,1710]" captionTargetBox="[157,1420,225,1620]" captionTargetId="figure@111.[151,1436,212,1639]" captionTargetPageId="111" captionText="FIGURES 498–507. Thraulodes nigripes sp. n. 498, larval protogonostyli and protopenis; 499, subimaginal styliger and gonostyli extracted from larva; 500–501, subimaginal penis inside larval protopenis in ventral and dorsal view; 502, larval protogonostyli with subimaginal gonostyli and styliger at earlier stage of development; 503–506, tergalii I, III, VI and VII (498–501, 503–506, holotype)." figureDoi="http://doi.org/10.5281/zenodo.3737032" httpUri="https://zenodo.org/record/3737032/files/figure.png" pageId="106" pageNumber="107">Fig. 502</figureCitation>
|
||
). In contrast to
|
||
<taxonomicName id="4C3E4D33FFF7840E8E2AFB61D993FB03" box="[582,685,1159,1185]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="niger" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8E2AFB61D993FB03" box="[582,685,1159,1185]" italics="true" pageId="106" pageNumber="107">Th. niger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8ED8FB61D9CDFB03" bold="true" box="[692,755,1159,1185]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8ED8FB61D9CDFB03" box="[692,755,1159,1185]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, larvae under consideration have no paired submedian hypodermal markings on the abdominal terga (
|
||
<figureCitation id="13052A35FFF7840E8E0CFB4DD9FEFB67" box="[608,704,1194,1221]" captionStart="FIGURES 450–463" captionStartId="104.[151,264,1826,1851]" captionTargetBox="[157,1429,166,1806]" captionTargetId="figure@104.[157,1429,166,1806]" captionTargetPageId="104" captionText="FIGURES 450–463. Larvae, presumably attributed to Thraulodes nigrabdominalis sp. n. 450–455, parts of last instar larva at the same magnification: 450, fragment of head with labrum and mandibles; 451, pronotum and mesonotum; 452, hind leg; 453, abdominal terga; 454–455, tergalii IV–V; 456–458, exuviae of young larva at the same magnification: 456–457, fore and hind legs; 458, abdominal terga and sterna; 459, young larva with brown cuticular coloration and blackish hypodermal coloration; 460, larval protogonostyli and protopenis; 461, anterior side of hind femur; 462, hind tibia with focus on outer-posterior row of stout setae; 463, the same with focus on outer-anterior and inner-anterior rows of stout setae." figureDoi="http://doi.org/10.5281/zenodo.3737022" httpUri="https://zenodo.org/record/3737022/files/figure.png" pageId="106" pageNumber="107">Fig. 459</figureCitation>
|
||
), the same as in the imago of
|
||
<taxonomicName id="4C3E4D33FFF7840E8F91FB4DDFDEFB67" box="[1021,1248,1195,1221]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8F91FB4DDFDEFB67" box="[1021,1248,1195,1221]" italics="true" pageId="106" pageNumber="107">Th. nigrabdominalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8889FB4DDE1DFB67" bold="true" box="[1253,1315,1195,1221]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8889FB4DDE1DFB67" box="[1253,1315,1195,1221]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13052A35FFF7840E8943FB4DDEAFFB64" box="[1327,1425,1195,1222]" captionStart="FIGURES 464–473" captionStartId="106.[151,264,188,213]" captionTargetId="figure@105.[151,1436,181,1987]" captionTargetPageId="105" captionText="FIGURES 464–473. Thraulodes nigrabdominalis sp. n. (holotype). 464, left half of subimaginal exuviae of mesonotum; 465, 466, male imago; 467–469, fore, middle and hind legs of male imago; 470, subimaginal exuviae of fore leg; 471, genitalia of male imago; 472, apex of penis; 473, subimaginal exuviae of tarsus." figureDoi="http://doi.org/10.5281/zenodo.3737024" httpUri="https://zenodo.org/record/3737024/files/figure.png" pageId="106" pageNumber="107">Fig. 466</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8136B0FFF7840E8CABFB28DA20FA63" blockId="106.[151,1437,367,1473]" pageId="106" pageNumber="107">
|
||
<emphasis id="B94AEAA2FFF7840E8CABFB28DA5CFB4B" bold="true" box="[199,354,1230,1257]" pageId="106" pageNumber="107">Comparison.</emphasis>
|
||
The male imago and subimago of
|
||
<taxonomicName id="4C3E4D33FFF7840E8E87FB29D8EFFB4B" box="[747,977,1231,1257]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8E87FB29D8EFFB4B" box="[747,977,1231,1257]" italics="true" pageId="106" pageNumber="107">Th. nigrabdominalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8FB4FB29DF26FB4B" bold="true" box="[984,1048,1231,1257]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8FB4FB29DF26FB4B" box="[984,1048,1231,1257]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are similar to
|
||
<taxonomicName id="4C3E4D33FFF7840E88D0FB29DE1DFB4B" box="[1212,1315,1231,1257]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="niger" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E88D0FB29DE1DFB4B" box="[1212,1315,1231,1257]" italics="true" pageId="106" pageNumber="107">Th. niger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8947FB29DE55FB4B" bold="true" box="[1323,1387,1231,1257]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8947FB29DE55FB4B" box="[1323,1387,1231,1257]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; the male imago differs from
|
||
<taxonomicName id="4C3E4D33FFF7840E8DC3FB15D92BFAAF" box="[431,533,1267,1293]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="niger" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8DC3FB15D92BFAAF" box="[431,533,1267,1293]" italics="true" pageId="106" pageNumber="107">Th. niger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8E71FB15D962FAAF" bold="true" box="[541,604,1267,1293]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8E71FB15D962FAAF" box="[541,604,1267,1293]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by an absence of coloration of costal and subcostal fields of the fore wing and by absence of paired dark markings on the abdominal terga. Both
|
||
<taxonomicName id="4C3E4D33FFF7840E8FF5FAF1DF42FA93" box="[921,1148,1303,1329]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrabdominalis" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E8FF5FAF1DF42FA93" box="[921,1148,1303,1329]" italics="true" pageId="106" pageNumber="107">Th. nigrabdominalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E88EEFAF1DFFEFA93" bold="true" box="[1154,1216,1303,1329]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E88EEFAF1DFFEFA93" box="[1154,1216,1303,1329]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C3E4D33FFF7840E889FFAF1DE67FA93" box="[1267,1369,1303,1329]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="niger" status="sp. nov.">
|
||
<emphasis id="B94AEAA2FFF7840E889FFAF1DE67FA93" box="[1267,1369,1303,1329]" italics="true" pageId="106" pageNumber="107">Th. niger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B94AEAA2FFF7840E8933FAF1DEA3FA93" bold="true" box="[1375,1437,1303,1329]" pageId="106" pageNumber="107">
|
||
<taxonomicNameLabel id="A27957D9FFF7840E8933FAF1DEA3FA93" box="[1375,1437,1303,1329]" pageId="106" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
have penis structure similar to that of
|
||
<taxonomicName id="4C3E4D33FFF7840E8E50FADDDF36FAF7" authority="Traver & Edmunds 1967" authorityName="Traver & Edmunds" authorityYear="1967" box="[572,1032,1339,1365]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="ephippiatus">
|
||
<emphasis id="B94AEAA2FFF7840E8E50FADDD9D4FAF7" box="[572,746,1339,1365]" italics="true" pageId="106" pageNumber="107">Th. ephippiatus</emphasis>
|
||
<bibRefCitation id="EFAF4B41FFF7840E8E9CFADDDF36FAF7" author="Traver, J. R. & Edmunds, G. F. Jr." box="[752,1032,1339,1365]" pageId="106" pageNumber="107" pagination="349 - 395" refId="ref72596" refString="Traver, J. R. & Edmunds, G. F. Jr. (1967) A revision of the genus Thraulodes (Ephemeroptera: Leptophlebiidae). Miscellaneous Publications of the Entomological Society of America, 5 (8), 349 - 395." type="journal article" year="1967">Traver & Edmunds 1967</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3E4D33FFF7840E887EFADDDEA2FAF7" authority="Dominguez 1987" authorityName="Dominguez" authorityYear="1987" box="[1042,1436,1339,1365]" class="Insecta" family="Leptophlebiidae" genus="Thraulus" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="cochunaensis">
|
||
<emphasis id="B94AEAA2FFF7840E887EFADDDFEBFAF7" box="[1042,1237,1339,1365]" italics="true" pageId="106" pageNumber="107">Th. cochunaensis</emphasis>
|
||
<bibRefCitation id="EFAF4B41FFF7840E88B7FADDDEA2FAF7" author="Dominguez, E." box="[1243,1436,1339,1365]" pageId="106" pageNumber="107" pagination="47 - 65" refId="ref69218" refString="Dominguez, E. (1987) El genero Thraulodes (Ephemeroptera: Leptophlebiidae) en la Republica Argentina. Acta Zoologica Lilloana, 39, 47 - 65." type="journal article" year="1987">Domínguez 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3E4D33FFF7840E8CABFAB9D9D3FADB" authority="Giordano & Dominguez 2005" authorityName="Giordano & Dominguez" authorityYear="2005" box="[199,749,1374,1401]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="basimaculatus">
|
||
<emphasis id="B94AEAA2FFF7840E8CABFAB9DAABFADB" box="[199,405,1375,1401]" italics="true" pageId="106" pageNumber="107">Th. basimaculatus</emphasis>
|
||
<bibRefCitation id="EFAF4B41FFF7840E8DF0FAB9D9D3FADB" author="Giordano, J. A. & Dominguez, E." box="[412,749,1374,1401]" pageId="106" pageNumber="107" pagination="45 - 48" refId="ref69951" refString="Giordano, J. A. & Dominguez, E. (2005) Thraulodes basimaculatus sp. n., a new species of mayfly from Bolivia: (Ephemeroptera: Leptophlebiidae: Atalophlebiinae). Zootaxa, 1040 (1), 45 - 48. https: // doi. org / 10.11646 / zootaxa. 1040.1.4" type="journal article" year="2005">Giordano & Domínguez 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
and differ from these three species by uniformly dark brown femora and tibiae of all legs. Larval cuticular coloration distinguishes this species from all other known species of
|
||
<taxonomicName id="4C3E4D33FFF7840E8CFBFA41DA29FA63" authorityName="Ulmer" authorityYear="1920" box="[151,279,1447,1473]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" kingdom="Animalia" order="Ephemeroptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94AEAA2FFF7840E8CFBFA41DA29FA63" box="[151,279,1447,1473]" italics="true" pageId="106" pageNumber="107">Thraulodes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |