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<mods:title id="ACA9AF92DE08E63802F5D521C8642725">Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution</mods:title>
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<heading id="D0C981F2B9A0FF11C1E36D0DAC919C22" bold="true" box="[151,746,1342,1368]" fontSize="11" level="1" pageId="212" pageNumber="213" reason="1">
<emphasis id="B94AEA8CB9A0FF11C1E36D0DAC919C22" bold="true" box="[151,746,1342,1368]" pageId="212" pageNumber="213">
Subtribe
<taxonomicName id="4C3E4D1DB9A0FF11C07F6D0DAC919C22" ID-CoL="8VRCG" authority="Newton &amp; Thayer, 1992" authorityName="Newton &amp; Thayer" authorityYear="1992" box="[267,746,1342,1368]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="subTribe" subTribe="Clavilispinina">
Clavilispinina
<bibRefCitation id="EFAF4B6FB9A0FF11C0B56D0DAC919C22" author="Newton, A. F. &amp; Thayer, M. K." box="[449,746,1342,1368]" pageId="212" pageNumber="213" pagination="1 - 92" refId="ref220616" refString="Newton, A. F. &amp; Thayer, M. K. (1992) Current classification and family-group names in Staphyliniformia (Coleoptera). Fieldiana Zoology, new series, 67, 1 - 92." type="journal article" year="1992">Newton &amp; Thayer, 1992</bibRefCitation>
</taxonomicName>
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<paragraph id="8B81369EB9A0FF11C1E36DBEACCD9CDC" blockId="212.[151,725,1421,1486]" box="[151,694,1421,1446]" pageId="212" pageNumber="213">
<emphasis id="B94AEA8CB9A0FF11C1E36DBEAEAD9CDC" bold="true" box="[151,214,1421,1446]" pageId="212" pageNumber="213">0737.</emphasis>
<taxonomicName id="4C3E4D1DB9A0FF11C1A96DBEAC539CDC" authority="Fagel, 1955 b: 78" authorityName="Fagel" authorityPageNumber="78" authorityYear="1955" box="[221,552,1421,1446]" class="Insecta" family="Staphylinidae" genus="Allotrochus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A0FF11C1A96DBEAF1F9CDC" bold="true" box="[221,356,1421,1446]" italics="true" pageId="212" pageNumber="213">Allotrochus</emphasis>
<bibRefCitation id="EFAF4B6FB9A0FF11C01F6DBEAC539CDC" author="Fagel, G." box="[363,552,1421,1446]" pageId="212" pageNumber="213" pagination="1946 - 1949" refId="ref190927" refString="Fagel, G. (1955 b) Osoriinae (Coleoptera Polyphaga) Fam. Staphylinidae. In: Exploration du Parc National de l'Upemba, Mission G. F. de Witte, en collaboration avec W. Adam, A Janssens, L. Van Meel et R. Verheyen (1946 - 1949). Fasc. 39. Institut des Parcs Nationaux du Congo Belge, Bruxelles. 134 pp." type="book chapter" year="1955">Fagel, 1955b: 78</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13052A1BB9A0FF11C3286DBEACD69CDC" box="[604,685,1421,1446]" captionStart="FIGURES 8899" captionStartId="328.[151,264,1862,1885]" captionTargetBox="[156,1436,198,1832]" captionTargetId="figure@328.[151,1436,193,1840]" captionTargetPageId="328" captionText="FIGURES 8899. Habitus; Piestinae (8891); Osoriinae (9299); 88, after Pérez et al. 2017; 89, Peru: department of Madre de Dios (MUSM); 90, Peru: department of Cuzco (MUSM); 91, after Brunke et al. 2011; 92, after Sharp 1887a; 93, after Sharp 1887a; 94, after Irmler 1991; 95, after Irmler 2001b; 96, after Bierig 1931; 97, after Sharp 1887a; 98, after Caron et al. 2012; 99, after Bruch 1942; slightly modified. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3257248/files/figure.png" pageId="212" pageNumber="213">Fig. 92</figureCitation>
)
</paragraph>
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<paragraph id="8B81369EB9A0FF11C1E36D85ACAE9CB7" blockId="212.[151,725,1421,1486]" box="[151,725,1460,1486]" pageId="212" pageNumber="213">
<typeStatus id="5485883CB9A0FF11C1E36D85AEAA9CB4" box="[151,209,1462,1486]" pageId="212" pageNumber="213">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A0FF11C0436D87ACAE9CB7" authority="Fauvel, 1898" authorityName="Fauvel" authorityYear="1898" box="[311,725,1460,1485]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="curticollis">
<emphasis id="B94AEA8CB9A0FF11C0436D87AC479CB7" box="[311,572,1460,1485]" italics="true" pageId="212" pageNumber="213">Holotrochus curticollis</emphasis>
Fauvel, 1898
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB9A0FF11C1E36E30AD6A9F66" blockId="212.[151,1436,1539,1876]" box="[151,785,1539,1564]" pageId="212" pageNumber="213">5 species are included in this genus from Latin America.</paragraph>
<paragraph id="8B81369EB9A0FF11C1E36E19AB119F39" blockId="212.[151,1436,1539,1876]" box="[151,1386,1578,1603]" pageId="212" pageNumber="213">
Characters/Description:
<bibRefCitation id="EFAF4B6FB9A0FF11C0DC6E19AD549F38" author="Navarrete-Heredia, J. L. &amp; Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[424,815,1578,1603]" pageId="212" pageNumber="213" refId="ref220178" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Universidad de Guadalajara y CONABIO, Mexico. xii + 401 pp." type="book" year="2002">
Navarrete-Heredia
<emphasis id="B94AEA8CB9A0FF11C30B6E18ACCC9F39" box="[639,695,1578,1603]" italics="true" pageId="212" pageNumber="213">et al.</emphasis>
2002: 213
</bibRefCitation>
(characters in key);
<bibRefCitation id="EFAF4B6FB9A0FF11C5616E18AAAC9F38" author="Newton, A. F." box="[1045,1239,1578,1602]" pageId="212" pageNumber="213" pagination="1 - 52" refId="ref220402" refString="Newton, A. F. (2017) Nomenclatural and taxonomic changes in Staphyliniformia (Coleoptera). Insecta Mundi, 0595, 1 - 52." type="book chapter" year="2017">Newton 2017: 23</bibRefCitation>
(characters).
</paragraph>
<paragraph id="8B81369EB9A0FF11C1E36E62AFF09FEA" blockId="212.[151,1436,1539,1876]" pageId="212" pageNumber="213">
Key:
<bibRefCitation id="EFAF4B6FB9A0FF11C1A06E63AFE69F13" author="Irmler, U." box="[212,413,1616,1641]" pageId="212" pageNumber="213" pagination="247 - 250" refId="ref201901" refString="Irmler, U. (2000 a) The neotropical species of the genus Allotrochus Fagel, 1955. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie, 70, 247 - 250." type="journal article" year="2000">Irmler 2000a: 247</bibRefCitation>
(Neotropical species);
<bibRefCitation id="EFAF4B6FB9A0FF11C3D46E63ADB09F13" author="Naomi, S. I. &amp; Irmler, U." box="[672,971,1616,1641]" pageId="212" pageNumber="213" pagination="105 - 110" refId="ref219758" refString="Naomi, S. I. &amp; Irmler, U. (2012) First record of the genus Allotrochus Fagel, 1955 (Coleoptera: Staphylinidae: Osoriinae) from the Oriental region, with description of the new species A. takahashii from Ishigaki Is., Nansei Islands, Japan. Biogeography, 14, 105 - 110." type="journal article" year="2012">Naomi &amp; Irmler 2012: 106</bibRefCitation>
(species of the genus);
<bibRefCitation id="EFAF4B6FB9A0FF11C5A66E63ABE79F13" author="Irmler, U." box="[1234,1436,1616,1641]" pageId="212" pageNumber="213" pagination="145 - 172" refId="ref203218" refString="Irmler, U. (2015 b) Osoriinae of Cuba with description of new species and an identification key (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 55 (1), 145 - 172." type="journal article" year="2015">Irmler 2015b: 158</bibRefCitation>
(Cuban species).
</paragraph>
<paragraph id="8B81369EB9A0FF11C1E36EADAF5E9FCD" blockId="212.[151,1436,1539,1876]" box="[151,293,1694,1719]" pageId="212" pageNumber="213">Distribution.</paragraph>
<paragraph id="8B81369EB9A0FF11C1DE6EF4AFFB9FA4" blockId="212.[151,1436,1539,1876]" box="[170,384,1733,1758]" pageId="212" pageNumber="213">
West Indies:
<collectingCountry id="F329760EB9A0FF11C0496EF6AF079FA4" box="[317,380,1733,1758]" name="Cuba" pageId="212" pageNumber="213">Cuba</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9A0FF11C1DE6EDFAC2C9E7F" blockId="212.[151,1436,1539,1876]" box="[170,599,1772,1797]" pageId="212" pageNumber="213">
Central America:
<collectingCountry id="F329760EB9A0FF11C0066EDFAFB19E7F" box="[370,458,1772,1797]" name="Mexico" pageId="212" pageNumber="213">Mexico</collectingCountry>
,
<collectingCountry id="F329760EB9A0FF11C0A26EDFAC289E7F" box="[470,595,1772,1797]" name="Guatemala" pageId="212" pageNumber="213">Guatemala</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9A0FF11C1DE6F27ACA59E57" blockId="212.[151,1436,1539,1876]" box="[170,734,1812,1837]" pageId="212" pageNumber="213">
South America:
<collectingCountry id="F329760EB9A0FF11C0156F27AFA19E57" box="[353,474,1812,1837]" name="Venezuela" pageId="212" pageNumber="213">Venezuela</collectingCountry>
,
<collectingCountry id="F329760EB9A0FF11C0926F27AC3D9E57" box="[486,582,1812,1837]" name="Ecuador" pageId="212" pageNumber="213">Ecuador</collectingCountry>
,
<collectingCountry id="F329760EB9A0FF11C3256F26ACFC9E56" box="[593,647,1813,1836]" name="Peru" pageId="212" pageNumber="213">Peru</collectingCountry>
,
<collectingCountry id="F329760EB9A0FF11C3E66F27ACA19E57" box="[658,730,1812,1837]" name="Brazil" pageId="212" pageNumber="213">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9A0FF11C1DE6F08ADCD9E2E" blockId="212.[151,1436,1539,1876]" box="[170,950,1851,1876]" pageId="212" pageNumber="213">
Other regions: Ethiopian,
<collectingRegion id="49FAF87CB9A0FF11C0B86F08AC509E2E" box="[460,555,1851,1876]" country="Republic of the Congo" name="Orientale" pageId="212" pageNumber="213">Oriental</collectingRegion>
, Australian, Oceanic, Madagascan.
</paragraph>
<paragraph id="8B81369EB9A0FF11C1E36FBBAC809EDB" blockId="212.[151,923,1928,2031]" box="[151,763,1928,1953]" pageId="212" pageNumber="213">
<emphasis id="B94AEA8CB9A0FF11C1E36FBBAEAD9EDB" bold="true" box="[151,214,1928,1953]" pageId="212" pageNumber="213">0738.</emphasis>
<taxonomicName id="4C3E4D1DB9A0FF11C1A96FBBAC159EDB" authority="Bernhauer, 1926: 255" authorityName="Bernhauer" authorityPageNumber="255" authorityYear="1926" box="[221,622,1928,1953]" class="Insecta" family="Staphylinidae" genus="Clavilispinus" kingdom="Animalia" order="Coleoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A0FF11C1A96FBBAF0F9EDB" bold="true" box="[221,372,1928,1953]" italics="true" pageId="212" pageNumber="213">Clavilispinus</emphasis>
Bernhauer, 1926: 255
</taxonomicName>
(
<figureCitation id="13052A1BB9A0FF11C3D66FBBAC889EDB" box="[674,755,1928,1953]" captionStart="FIGURES 8899" captionStartId="328.[151,264,1862,1885]" captionTargetBox="[156,1436,198,1832]" captionTargetId="figure@328.[151,1436,193,1840]" captionTargetPageId="328" captionText="FIGURES 8899. Habitus; Piestinae (8891); Osoriinae (9299); 88, after Pérez et al. 2017; 89, Peru: department of Madre de Dios (MUSM); 90, Peru: department of Cuzco (MUSM); 91, after Brunke et al. 2011; 92, after Sharp 1887a; 93, after Sharp 1887a; 94, after Irmler 1991; 95, after Irmler 2001b; 96, after Bierig 1931; 97, after Sharp 1887a; 98, after Caron et al. 2012; 99, after Bruch 1942; slightly modified. Scale bars: 1 mm." httpUri="https://zenodo.org/record/3257248/files/figure.png" pageId="212" pageNumber="213">Fig. 94</figureCitation>
)
</paragraph>
<paragraph id="8B81369EB9A0FF11C1E36F83AD629EB2" blockId="212.[151,923,1928,2031]" box="[151,793,1967,1992]" pageId="212" pageNumber="213">
<typeStatus id="5485883CB9A0FF11C1E36F83AEAA9EB2" box="[151,209,1968,1992]" pageId="212" pageNumber="213">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A0FF11C0436F9CAD629EB2" authority="Bernhauer, 1926" authorityName="Bernhauer" authorityYear="1926" box="[311,793,1967,1992]" class="Insecta" family="Staphylinidae" genus="Paralispinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="species" species="siargaoanus">
<emphasis id="B94AEA8CB9A0FF11C0436F9CAC2C9EB2" box="[311,599,1967,1992]" italics="true" pageId="212" pageNumber="213">Paralispinus siargaoanus</emphasis>
Bernhauer, 1926
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9A0FF11C1BB6FE4ADE09E95" box="[207,923,2006,2031]" pageId="212" pageNumber="213" type="reference_group">
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<treatmentCitationGroup id="AB2E11B0B9A0FF11C1BB6FE4ADE09E95" box="[207,923,2006,2031]" pageId="212" pageNumber="213">
<taxonomicName id="4C3E4D1DB9A0FF11C1BB6FE4AF899E95" ID-CoL="8XJK4" authority="Fauvel, 1864: 56" authorityName="Fauvel" authorityPageNumber="56" authorityYear="1864" box="[207,498,2006,2031]" class="Insecta" family="Staphylinidae" genus="Ancaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="212" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A0FF11C1BB6FE4AF559E94" box="[207,302,2007,2030]" italics="true" pageId="212" pageNumber="213">Ancaeus</emphasis>
<treatmentCitation id="0A9F108FB9A0FF11C0406FE5AF899E95" author="Fauvel, A." box="[308,498,2006,2031]" page="56" pageId="212" pageNumber="213" year="1864">
<bibRefCitation id="EFAF4B6FB9A0FF11C0406FE5AF899E95" author="Fauvel, A." box="[308,498,2006,2031]" pageId="212" pageNumber="213" pagination="3 - 62" refId="ref191389" refString="Fauvel, A. (1864) Etudes sur les Staphylinides de l'Amerique centrale, principalement du Mexique. Notices Entomologiques, 2, 3 - 62, 66, pl. 1." type="journal article" year="1864">Fauvel, 1864: 56</bibRefCitation>
</treatmentCitation>
</taxonomicName>
[synonymized by
<treatmentCitation id="0A9F108FB9A0FF11C3B56FE4ADEF9E95" author="Cameron, M." box="[705,916,2007,2031]" page="42" pageId="212" pageNumber="213" year="1930">
<bibRefCitation id="EFAF4B6FB9A0FF11C3B56FE4ADEF9E95" author="Cameron, M." box="[705,916,2007,2031]" pageId="212" pageNumber="213" pagination="1 - 3" refId="ref179466" refString="Cameron, M. (1930) Coleoptera. Staphylinidae. Vol. I. In: Stephenson, J. (Ed.), The Fauna of British India including Ceylon and Burma. Taylor &amp; Francis, London. xvii + 471 pp., pls. 1 - 3." type="book chapter" year="1930">Cameron 1930: 42</bibRefCitation>
</treatmentCitation>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9A1FF10C07368ACAD3D99CD" box="[263,838,158,183]" pageId="213" pageNumber="214" type="materials_examined">
<paragraph id="8B81369EB9A1FF10C07368ACAD3D99CD" blockId="213.[207,1077,158,339]" box="[263,838,158,183]" pageId="213" pageNumber="214">
<typeStatus id="5485883CB9A1FF10C07368ACAF3A99CD" box="[263,321,159,183]" pageId="213" pageNumber="214">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A1FF10C0D368ACAD3D99CD" authority="Fauvel, 1864" authorityName="Fauvel" authorityYear="1864" box="[423,838,158,183]" class="Insecta" family="Staphylinidae" genus="Ancaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="species" species="megacephalus">
<emphasis id="B94AEA8CB9A1FF10C0D368ACACD699CD" box="[423,685,158,183]" italics="true" pageId="213" pageNumber="214">Ancaeus megacephalus</emphasis>
<bibRefCitation id="EFAF4B6FB9A1FF10C3C068ADAD3D99CD" author="Fauvel, A." box="[692,838,158,183]" pageId="213" pageNumber="214" pagination="3 - 62" refId="ref191389" refString="Fauvel, A. (1864) Etudes sur les Staphylinides de l'Amerique centrale, principalement du Mexique. Notices Entomologiques, 2, 3 - 62, 66, pl. 1." type="journal article" year="1864">Fauvel, 1864</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9A1FF10C1BB68F6AA5499A4" box="[207,1071,197,222]" pageId="213" pageNumber="214" type="reference_group">
<paragraph id="8B81369EB9A1FF10C1BB68F6AA5499A4" blockId="213.[207,1077,158,339]" box="[207,1071,197,222]" pageId="213" pageNumber="214">
<treatmentCitationGroup id="AB2E11B0B9A1FF10C1BB68F6AA5499A4" box="[207,1071,197,222]" pageId="213" pageNumber="214">
<taxonomicName id="4C3E4D1DB9A1FF10C1BB68F6AC1399A4" ID-CoL="8XN4S" authority="Bernhauer, 1937: 579" authorityName="Bernhauer" authorityPageNumber="579" authorityYear="1937" box="[207,616,197,222]" class="Insecta" family="Staphylinidae" genus="Neolispinodes" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A1FF10C1BB68F6AF1599A4" box="[207,366,197,222]" italics="true" pageId="213" pageNumber="214">Neolispinodes</emphasis>
<treatmentCitation id="0A9F108FB9A1FF10C00068F6AC1399A4" author="Bernhauer, M." box="[372,616,197,222]" page="579" pageId="213" pageNumber="214" year="1937">
<bibRefCitation id="EFAF4B6FB9A1FF10C00068F6AC1399A4" author="Bernhauer, M." box="[372,616,197,222]" pageId="213" pageNumber="214" pagination="577 - 619" refId="ref173457" refString="Bernhauer, M. (1937) Neuheiten der ostafrikanischen Staphylinidenfauna (45. Beitrag zur afrikanischen Fauna). In: Festschrift zum 60. Geburtstage von Professor Dr. Embrik Strand. Vol. 2. Izdevnieciba &quot; Latvija &quot;, Riga, pp. 577 - 619." type="book chapter" year="1937">Bernhauer, 1937: 579</bibRefCitation>
</treatmentCitation>
</taxonomicName>
[synonymized by
<treatmentCitation id="0A9F108FB9A1FF10C24368F5AA5299A4" author="Herman, L. H." box="[823,1065,197,222]" page="1232" pageId="213" pageNumber="214" year="2001">
<bibRefCitation id="EFAF4B6FB9A1FF10C24368F5AA5299A4" author="Herman, L. H." box="[823,1065,197,222]" pageId="213" pageNumber="214" pagination="1067 - 1806" refId="ref199974" refString="Herman, L. H. (2001 d) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. III. Oxyteline Group. Bulletin of the American Museum of Natural History, 265, 1067 - 1806." type="journal article" year="2001">Herman 2001d: 1232</bibRefCitation>
</treatmentCitation>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9A1FF10C07368DEAD3D987F" box="[263,838,236,261]" pageId="213" pageNumber="214" type="materials_examined">
<paragraph id="8B81369EB9A1FF10C07368DEAD3D987F" blockId="213.[207,1077,158,339]" box="[263,838,236,261]" pageId="213" pageNumber="214">
<typeStatus id="5485883CB9A1FF10C07368DEAF3A987F" box="[263,321,237,261]" pageId="213" pageNumber="214">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A1FF10C0D368DEAD3D987F" authority="Fauvel, 1864" authorityName="Fauvel" authorityYear="1864" box="[423,838,236,261]" class="Insecta" family="Staphylinidae" genus="Ancaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="species" species="megacephalus">
<emphasis id="B94AEA8CB9A1FF10C0D368DEACD6987F" box="[423,685,236,261]" italics="true" pageId="213" pageNumber="214">Ancaeus megacephalus</emphasis>
<bibRefCitation id="EFAF4B6FB9A1FF10C3C068DFAD3D987F" author="Fauvel, A." box="[692,838,236,261]" pageId="213" pageNumber="214" pagination="3 - 62" refId="ref191389" refString="Fauvel, A. (1864) Etudes sur les Staphylinides de l'Amerique centrale, principalement du Mexique. Notices Entomologiques, 2, 3 - 62, 66, pl. 1." type="journal article" year="1864">Fauvel, 1864</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9A1FF10C1BB6920AA4E9851" box="[207,1077,275,300]" pageId="213" pageNumber="214" type="reference_group">
<paragraph id="8B81369EB9A1FF10C1BB6920AA4E9851" blockId="213.[207,1077,158,339]" box="[207,1077,275,300]" pageId="213" pageNumber="214">
<treatmentCitationGroup id="AB2E11B0B9A1FF10C1BB6920AA4E9851" box="[207,1077,275,300]" pageId="213" pageNumber="214">
<taxonomicName id="4C3E4D1DB9A1FF10C1BB6920AC2C9856" ID-CoL="8XNLY" authority="Bernhauer, 1921 a: 67" authorityName="Bernhauer" authorityPageNumber="67" authorityYear="1921" box="[207,599,275,300]" class="Insecta" family="Staphylinidae" genus="Paralispinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A1FF10C1BB6920AF259856" box="[207,350,275,300]" italics="true" pageId="213" pageNumber="214">Paralispinus</emphasis>
<treatmentCitation id="0A9F108FB9A1FF10C0116920AC2C9856" author="Bernhauer, M." box="[357,599,275,300]" page="67" pageId="213" pageNumber="214" year="1921">
<bibRefCitation id="EFAF4B6FB9A1FF10C0116920AC2C9856" author="Bernhauer, M." box="[357,599,275,300]" pageId="213" pageNumber="214" pagination="65 - 77" refId="ref171995" refString="Bernhauer, M. (1921 a) Zur Staphylinidenfauna von Sud-Amerika. (24. Beitrag.). Deutsche Entomologische Zeitschrift, 1921 (1), 65 - 77." type="journal article" year="1921">Bernhauer, 1921a: 67</bibRefCitation>
</treatmentCitation>
</taxonomicName>
[synonymized by
<treatmentCitation id="0A9F108FB9A1FF10C2526920AA559851" author="Blackwelder, R. E." box="[806,1070,275,300]" page="291" pageId="213" pageNumber="214" year="1952">
<bibRefCitation id="EFAF4B6FB9A1FF10C2526920AA559851" author="Blackwelder, R. E." box="[806,1070,275,300]" pageId="213" pageNumber="214" refId="ref175744" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. Bulletin of the United States National Museum, 200, iv + 483 pp." type="book" year="1952">Blackwelder 1952: 291</bibRefCitation>
</treatmentCitation>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3246515B9A1FF10C0736908AF949DD7" pageId="213" pageNumber="214" type="materials_examined">
<paragraph id="8B81369EB9A1FF10C0736908AD3D9829" blockId="213.[207,1077,158,339]" box="[263,838,314,339]" pageId="213" pageNumber="214">
<typeStatus id="5485883CB9A1FF10C0736908AF3A9829" box="[263,321,315,339]" pageId="213" pageNumber="214">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A1FF10C0D36908AD3D9829" authority="Fauvel, 1864" authorityName="Fauvel" authorityYear="1864" box="[423,838,314,339]" class="Insecta" family="Staphylinidae" genus="Ancaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="species" species="megacephalus">
<emphasis id="B94AEA8CB9A1FF10C0D36908ACD69829" box="[423,685,314,339]" italics="true" pageId="213" pageNumber="214">Ancaeus megacephalus</emphasis>
<bibRefCitation id="EFAF4B6FB9A1FF10C3C06909AD3D9829" author="Fauvel, A." box="[692,838,314,339]" pageId="213" pageNumber="214" pagination="3 - 62" refId="ref191389" refString="Fauvel, A. (1864) Etudes sur les Staphylinides de l'Amerique centrale, principalement du Mexique. Notices Entomologiques, 2, 3 - 62, 66, pl. 1." type="journal article" year="1864">Fauvel, 1864</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB9A1FF10C1E369BBAD6498DB" blockId="213.[151,1436,392,963]" box="[151,799,392,417]" pageId="213" pageNumber="214">14 species are included in this genus from Latin America.</paragraph>
<paragraph id="8B81369EB9A1FF10C1E3699CACA29B6C" blockId="213.[151,1436,392,963]" pageId="213" pageNumber="214">
Characters/Description:
<bibRefCitation id="EFAF4B6FB9A1FF10C0C7699CACCE98B2" author="Bernhauer, M." box="[435,693,431,456]" pageId="213" pageNumber="214" pagination="65 - 77" refId="ref171995" refString="Bernhauer, M. (1921 a) Zur Staphylinidenfauna von Sud-Amerika. (24. Beitrag.). Deutsche Entomologische Zeitschrift, 1921 (1), 65 - 77." type="journal article" year="1921">Bernhauer 1921a: 67</bibRefCitation>
(characters);
<bibRefCitation id="EFAF4B6FB9A1FF10C212699CAAF998BD" author="Blackwelder, R. E." box="[870,1154,431,456]" pageId="213" pageNumber="214" refId="ref175651" refString="Blackwelder, R. E. (1943) Monograph of the West-Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, viii + 658 pp. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book" year="1943">Blackwelder 1943: 120</bibRefCitation>
(characters in key as
<taxonomicName id="4C3E4D1DB9A1FF10C1BB69E5AF259895" authorityName="Bernhauer" authorityYear="1921" box="[207,350,470,495]" class="Insecta" family="Staphylinidae" genus="Paralispinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A1FF10C1BB69E5AF259895" box="[207,350,470,495]" italics="true" pageId="213" pageNumber="214">Paralispinus</emphasis>
</taxonomicName>
), 156 (characters as
<taxonomicName id="4C3E4D1DB9A1FF10C32B69E5AC959895" authorityName="Bernhauer" authorityYear="1921" box="[607,750,470,495]" class="Insecta" family="Staphylinidae" genus="Paralispinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A1FF10C32B69E5AC959895" box="[607,750,470,495]" italics="true" pageId="213" pageNumber="214">Paralispinus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFAF4B6FB9A1FF10C27869E5AAD09895" author="Navarrete-Heredia, J. L. &amp; Newton, A. F. &amp; Thayer, M. K. &amp; Ashe, J. S. &amp; Chandler, D. S." box="[780,1195,470,495]" pageId="213" pageNumber="214" refId="ref220178" refString="Navarrete-Heredia, J. L., Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2002) Guia ilustrada para los generos de Staphylinidae (Coleoptera) de Mexico. Universidad de Guadalajara y CONABIO, Mexico. xii + 401 pp." type="book" year="2002">
Navarrete-Heredia
<emphasis id="B94AEA8CB9A1FF10C29D69E4AA539895" box="[1001,1064,470,495]" italics="true" pageId="213" pageNumber="214">et al.</emphasis>
2002: 213
</bibRefCitation>
(characters in key);
<bibRefCitation id="EFAF4B6FB9A1FF10C1BB69CDAF819B6C" author="Newton, A. F. &amp; Peck, S. B." box="[207,506,509,534]" pageId="213" pageNumber="214" pagination="131 - 158" refId="ref220547" refString="Newton, A. F. &amp; Peck, S. B. (2006) Family Staphylinidae, the rove beetles. In: Peck, S. B. (Ed.), The beetles of the Galapagos Islands, Ecuador: evolution, ecology, and diversity (Insecta: Coleoptera). NRC Research Press, Ottawa, pp. 131 - 158." type="book chapter" year="2006">Newton &amp; Peck 2006: 142</bibRefCitation>
(characters in key).
</paragraph>
<paragraph id="8B81369EB9A1FF10C1E36A16ABF79B1E" blockId="213.[151,1436,392,963]" pageId="213" pageNumber="214">
Key:
<bibRefCitation id="EFAF4B6FB9A1FF10C1A16A17AFA59B47" author="Blackwelder, R. E." box="[213,478,548,573]" pageId="213" pageNumber="214" refId="ref175651" refString="Blackwelder, R. E. (1943) Monograph of the West-Indian beetles of the family Staphylinidae. Bulletin of the United States National Museum, 182, viii + 658 pp. https: // doi. org / 10.5479 / si. 03629236.182. i" type="book" year="1943">Blackwelder 1943: 157</bibRefCitation>
(species of the West Indies as
<taxonomicName id="4C3E4D1DB9A1FF10C2356A17ADAB9B47" authorityName="Bernhauer" authorityYear="1921" box="[833,976,548,573]" class="Insecta" family="Staphylinidae" genus="Paralispinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A1FF10C2356A17ADAB9B47" box="[833,976,548,573]" italics="true" pageId="213" pageNumber="214">Paralispinus</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFAF4B6FB9A1FF10C29D6A17AAE79B47" author="Irmler, U." box="[1001,1180,548,573]" pageId="213" pageNumber="214" pagination="85 - 91" refId="ref201770" refString="Irmler, U. (1991) Neue Arten der Gattung Clavilispinus Blackwelder (Col., Staphylinidae) aus der Neotropis. Entomologische Blatter fur Biologie und Systematik der Kafer, 87 (1 - 2), 85 - 91." type="journal article" year="1991">Irmler 1991: 85</bibRefCitation>
(Neotropical species);
<bibRefCitation id="EFAF4B6FB9A1FF10C1BB6A78AFE09B1E" author="Irmler, U." box="[207,411,587,612]" pageId="213" pageNumber="214" pagination="349 - 360" refId="ref202109" refString="Irmler, U. (2003 b) Distribution, redescription and description of new species of the Clavilispinus exiguus group (Coleoptera: Staphylinidae, Osoriinae) in the Neotropical region. Amazoniana, 17 (3 - 4), 349 - 360." type="journal article" year="2003">Irmler 2003b: 350</bibRefCitation>
(Neotropical species of
<taxonomicName id="4C3E4D1DB9A1FF10C3D96A78ADE69B19" baseAuthorityName="Erichson" baseAuthorityYear="1840" box="[685,925,587,612]" class="Insecta" family="Staphylinidae" genus="Clavilispinus" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="B94AEA8CB9A1FF10C3D96A78ADE69B19" box="[685,925,587,612]" italics="true" pageId="213" pageNumber="214">Clavilispinus exiguus</emphasis>
</taxonomicName>
group);
<bibRefCitation id="EFAF4B6FB9A1FF10C2896A78AAB29B1E" author="Irmler, U." box="[1021,1225,587,612]" pageId="213" pageNumber="214" pagination="145 - 172" refId="ref203218" refString="Irmler, U. (2015 b) Osoriinae of Cuba with description of new species and an identification key (Coleoptera: Staphylinidae). Acta Entomologica Musei Nationalis Pragae, 55 (1), 145 - 172." type="journal article" year="2015">Irmler 2015b: 158</bibRefCitation>
(Cuban species).
</paragraph>
<paragraph id="8B81369EB9A1FF10C1E36A41AF5E9BF0" blockId="213.[151,1436,392,963]" box="[151,293,626,651]" pageId="213" pageNumber="214">Distribution.</paragraph>
<paragraph id="8B81369EB9A1FF10C1DE6AA9ADFE9BA3" blockId="213.[151,1436,392,963]" pageId="213" pageNumber="214">
West Indies:
<collectingCountry id="F329760EB9A1FF10C03D6AAAAFFC9BC8" box="[329,391,665,690]" name="Cuba" pageId="213" pageNumber="214">Cuba</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C0EC6AAAAF8E9BC8" box="[408,501,665,690]" name="Jamaica" pageId="213" pageNumber="214">Jamaica</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C3726AAAAC8C9BC8" box="[518,759,665,690]" name="Dominican Republic" pageId="213" pageNumber="214">Dominican Republic</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C27C6AA9ADED9BC8" box="[776,918,665,690]" name="Puerto Rico" pageId="213" pageNumber="214">Puerto Rico</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C2D36AAAADB09BCB" box="[935,971,665,689]" name="United States of America" pageId="213" pageNumber="214">US</collectingCountry>
Virgin Isl.,
<collectingCountry id="F329760EB9A1FF10C5106AAAAA959BC8" box="[1124,1262,665,690]" name="Guadeloupe" pageId="213" pageNumber="214">Guadeloupe</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C58B6AAAAB169BC8" box="[1279,1389,665,690]" name="Saint Lucia" pageId="213" pageNumber="214">St. Lucia</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C40A6AAAAFB59BA3" name="Saint Vincent and the Grenadines" pageId="213" pageNumber="214">St. Vincent &amp; Grenadines</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C0AD6AF3AC409BA3" box="[473,571,704,729]" name="Grenada" pageId="213" pageNumber="214">Grenada</collectingCountry>
, Trinidad-Tobago,
<collectingCountry id="F329760EB9A1FF10C2606AF3ADFA9BA3" box="[788,897,704,729]" name="Barbados" pageId="213" pageNumber="214">Barbados</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9A1FF10C1DE6AD4AAEC9B85" blockId="213.[151,1436,392,963]" box="[170,1175,743,768]" pageId="213" pageNumber="214">
Central America:
<collectingCountry id="F329760EB9A1FF10C0066AD4AFB19A7A" box="[370,458,743,768]" name="Mexico" pageId="213" pageNumber="214">Mexico</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C0A26AD4AC299A7A" box="[470,594,743,768]" name="Guatemala" pageId="213" pageNumber="214">Guatemala</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C3296AD4ACDC9A7A" box="[605,679,743,768]" name="Belize" pageId="213" pageNumber="214">Belize</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C3C76AD4AD589A7A" box="[691,803,743,768]" name="Honduras" pageId="213" pageNumber="214">Honduras</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C25A6AD4ADDF9A7A" box="[814,932,743,768]" name="Nicaragua" pageId="213" pageNumber="214">Nicaragua</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C2DB6AD4AA579A7A" box="[943,1068,743,768]" name="Costa Rica" pageId="213" pageNumber="214">Costa Rica</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C5436ADBAAE89B85" box="[1079,1171,744,767]" name="Panama" pageId="213" pageNumber="214">Panama</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9A1FF10C1DE6B3EAF889A34" blockId="213.[151,1436,392,963]" pageId="213" pageNumber="214">
South America:
<collectingCountry id="F329760EB9A1FF10C0056B3EAF999A5C" box="[369,482,781,806]" name="Colombia" pageId="213" pageNumber="214">Colombia</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C0816B3EAC159A5C" box="[501,622,781,806]" name="Venezuela" pageId="213" pageNumber="214">Venezuela</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C3F56B3DACA09A5C" box="[641,731,782,806]" name="Guyana" pageId="213" pageNumber="214">Guyana</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C3996B3EAD209A5C" box="[749,859,781,806]" name="Suriname" pageId="213" pageNumber="214">Suriname</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C21A6B3EAA649A5C" box="[878,1055,781,806]" name="French Guiana" pageId="213" pageNumber="214">French Guiana</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C5456B3EAAEA9A5C" box="[1073,1169,781,806]" name="Ecuador" pageId="213" pageNumber="214">Ecuador</collectingCountry>
,
<collectingRegion id="49FAF87CB9A1FF10C5D06B3EAB619A5C" box="[1188,1306,781,806]" country="Ecuador" name="Galapagos" pageId="213" pageNumber="214">Galapagos</collectingRegion>
Isl.,
<collectingCountry id="F329760EB9A1FF10C4166B3CABE39A5C" box="[1378,1432,783,806]" name="Peru" pageId="213" pageNumber="214">Peru</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C1BB6B07AF5F9A37" box="[207,292,820,845]" name="Bolivia" pageId="213" pageNumber="214">Bolivia</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C0446B07AF0C9A37" box="[304,375,820,845]" name="Brazil" pageId="213" pageNumber="214">Brazil</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C0F66B05AF959A34" box="[386,494,822,846]" name="Paraguay" pageId="213" pageNumber="214">Paraguay</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9A1FF10C1DE6B6FAAD79A0F" blockId="213.[151,1436,392,963]" box="[170,1196,860,885]" pageId="213" pageNumber="214">
Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan,
<collectingRegion id="49FAF87CB9A1FF10C2286B6FADC19A0F" box="[860,954,860,885]" country="Republic of the Congo" name="Orientale" pageId="213" pageNumber="214">Oriental</collectingRegion>
, Australian, Oceanic.
</paragraph>
<paragraph id="8B81369EB9A1FF10C1E36BB0AC219AE1" blockId="213.[151,1436,392,963]" box="[151,602,899,924]" pageId="213" pageNumber="214">
<emphasis id="B94AEA8CB9A1FF10C1E36BB0AEAD9AE6" bold="true" box="[151,214,899,924]" pageId="213" pageNumber="214">0739.</emphasis>
<taxonomicName id="4C3E4D1DB9A1FF10C1A96BB0AC219AE1" authority="Irmler, 2016 a: 109" authorityName="Irmler" authorityPageNumber="109" authorityYear="2016" box="[221,602,899,924]" class="Insecta" family="Staphylinidae" genus="Geotrochopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B94AEA8CB9A1FF10C1A96BB0AFF99AE6" bold="true" box="[221,386,899,924]" italics="true" pageId="213" pageNumber="214">Geotrochopsis</emphasis>
<bibRefCitation id="EFAF4B6FB9A1FF10C0FD6BB0AC219AE1" author="Irmler, U." box="[393,602,899,924]" pageId="213" pageNumber="214" pagination="109 - 116" refId="ref203322" refString="Irmler, U. (2016 a) A new genus and new species of neotropical Thoracophorini (Coleoptera: Staphylinidae: Osoriinae). Bonn zoological Bulletin, 64 (2), 109 - 116." type="journal article" year="2016">Irmler, 2016a: 109</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB9A1FF10C1E36B98AC9C9AB9" blockId="213.[151,1436,392,963]" box="[151,743,938,963]" pageId="213" pageNumber="214">
<typeStatus id="5485883CB9A1FF10C1E36B98AEAA9AB9" box="[151,209,939,963]" pageId="213" pageNumber="214">Type</typeStatus>
species:
<taxonomicName id="4C3E4D1DB9A1FF10C0436B99AC9C9AB9" authority="Irmler, 2016" authorityName="Irmler" authorityYear="2016" box="[311,743,938,963]" class="Insecta" family="Staphylinidae" genus="Geotrochopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="B94AEA8CB9A1FF10C0436B99AC2F9AB9" box="[311,596,938,963]" italics="true" pageId="213" pageNumber="214">Geotrochopsis pubescens</emphasis>
Irmler, 2016
</taxonomicName>
</paragraph>
<paragraph id="8B81369EB9A1FF10C1E36BCBADA09D6B" blockId="213.[151,987,1016,1197]" box="[151,987,1016,1041]" pageId="213" pageNumber="214">3 species are known from and are restricted to Central and South America.</paragraph>
<paragraph id="8B81369EB9A1FF10C1E36C13AC1F9D4D" blockId="213.[151,987,1016,1197]" box="[151,612,1054,1080]" pageId="213" pageNumber="214">
Key:
<bibRefCitation id="EFAF4B6FB9A1FF10C1A06C2DAFDD9D4D" author="Irmler, U." box="[212,422,1054,1079]" pageId="213" pageNumber="214" pagination="109 - 116" refId="ref203322" refString="Irmler, U. (2016 a) A new genus and new species of neotropical Thoracophorini (Coleoptera: Staphylinidae: Osoriinae). Bonn zoological Bulletin, 64 (2), 109 - 116." type="journal article" year="2016">Irmler, 2016a: 114</bibRefCitation>
(key to species).
</paragraph>
<paragraph id="8B81369EB9A1FF10C1E36C76AF5E9D24" blockId="213.[151,987,1016,1197]" box="[151,293,1093,1118]" pageId="213" pageNumber="214">Distribution.</paragraph>
<paragraph id="8B81369EB9A1FF10C1DE6C5FAC239DFF" blockId="213.[151,987,1016,1197]" box="[170,600,1132,1157]" pageId="213" pageNumber="214">
Central America:
<collectingCountry id="F329760EB9A1FF10C0066C5FAFB19DFF" box="[370,458,1132,1157]" name="Mexico" pageId="213" pageNumber="214">Mexico</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C0A26C5EAC2F9DFF" box="[470,596,1132,1157]" name="Costa Rica" pageId="213" pageNumber="214">Costa Rica</collectingCountry>
.
</paragraph>
<paragraph id="8B81369EB9A1FF10C1DE6CA7AF949DD7" blockId="213.[151,987,1016,1197]" box="[170,495,1172,1197]" pageId="213" pageNumber="214">
South America:
<collectingCountry id="F329760EB9A1FF10C0156CA6AFEC9DD6" box="[353,407,1173,1196]" name="Peru" pageId="213" pageNumber="214">Peru</collectingCountry>
,
<collectingCountry id="F329760EB9A1FF10C0D76CA7AF909DD7" box="[419,491,1172,1197]" name="Brazil" pageId="213" pageNumber="214">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>