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<document id="A28535F75E4407486B0F5890922BC079" ID-DOI="10.3897/zookeys.1214.122439" ID-ZooBank="8B42B493-BD13-4D4C-9161-55978636D055" ID-publisher-id="122439" URI-arpha="463DF256-3712-5219-BB26-3A5BD76AE6E1" URI-zoobank="http://zoobank.org/8B42B493-BD13-4D4C-9161-55978636D055" article-type="research-article" checkinTime="1728496588403" checkinUser="pensoft" docAuthor="Lan, Chang-Ting, Wu, Li, Luo, Tao, Liu, Ye-Wei, Zhou, Jia-Jun, Yu, Jing, Zhao, Xin-Rui, Xiao, Ning &amp; Zhou, Jiang" docDate="2024" docId="E615223BF2B05FB696BD9D895AE03E23" docLanguage="en" docName="ZooKeys 1214: 237-264" docOrigin="ZooKeys 1214" docSource="https://zookeys.pensoft.net/article/122439/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Triplophysa yaluwang Lan, Liu, Zhou &amp; Zhou 2024, sp. nov." docType="treatment" docUuid="DE306B1B-F770-4E79-9B9E-400CEC202266" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="264" masterDocId="463DF25637125219BB263A5BD76AE6E1" masterDocTitle="Two new hypogean species of the genus Triplophysa (Osteichthyes, Cypriniformes, Nemacheilidae) from Guizhou Province, Southwest China, with underestimated diversity" masterLastPageNumber="264" masterPageNumber="237" pageNumber="237" updateTime="1728496979665" updateUser="ExternalLinkService">
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<mods:title id="EF13D4AE731519AEAF99080ABD36ACF2">Two new hypogean species of the genus Triplophysa (Osteichthyes, Cypriniformes, Nemacheilidae) from Guizhou Province, Southwest China, with underestimated diversity</mods:title>
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<mods:namePart id="C76C2B64916CC371A7781D6EB5252274">Lan, Chang-Ting</mods:namePart>
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<mods:namePart id="7B1DC7A032AD4C3CBDDF9BDC06D8674E">Wu, Li</mods:namePart>
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<mods:namePart id="1F75EC09989F5FDB59E4A213E8D5C59F">Luo, Tao</mods:namePart>
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<mods:affiliation id="FF3F3E0040BE257DA65412B42E3DB340">School of Life Sciences, Guizhou Normal University, Guiyang 550025, Guizhou, China</mods:affiliation>
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<mods:namePart id="048654877F20C714C6F981826D5DFEBA">Liu, Ye-Wei</mods:namePart>
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<mods:affiliation id="33432BC70373832F98903E3E931C62D1">School of Karst Science, Guizhou Normal University, Guiyang 550001, Guizhou, China</mods:affiliation>
</mods:name>
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<mods:namePart id="CAD0EC6C318961A60BCA0C5697560C99">Zhou, Jia-Jun</mods:namePart>
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<mods:affiliation id="95442ADB0C09F0594FA59FEFAC71283C">Guangxi Key Laboratory for Forest Ecology and Conservation, College of Forestry, Guangxi University, Nanning 530004, Guangxi, China &amp; Zhejiang Forest Resource Monitoring Center, Hangzhou 310020, Zhejiang, China</mods:affiliation>
</mods:name>
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<mods:namePart id="52BA99749E806DDD274F7A13FD63CE59">Yu, Jing</mods:namePart>
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<mods:namePart id="4B54EEBD66A47B311D5C3F5CB2267D0A">Zhao, Xin-Rui</mods:namePart>
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<mods:namePart id="2B5B36AB292F8EFC4DBA37F9E49FDFD7">Xiao, Ning</mods:namePart>
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<mods:affiliation id="59AFFE3CA5790967A509F8F6AF13AC68">Zhejiang Forestry Survey Planning and Design Company Limited, Hangzhou 310020, Zhejiang, China</mods:affiliation>
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<mods:namePart id="9F8182C991694243D0C2C202EBB7767C">Zhou, Jiang</mods:namePart>
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<mods:date id="812930A17A5061068B53A3BA1C8C0461">2024</mods:date>
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<treatment id="E615223BF2B05FB696BD9D895AE03E23" ID-arpha="E615223B-F2B0-5FB6-96BD-9D895AE03E23" ID-zoobank="https://zoobank.org/DE306B1B-F770-4E79-9B9E-400CEC202266" LSID="urn:lsid:zoobank.org:act:DE306B1B-F770-4E79-9B9E-400CEC202266" LSID-ZBK="urn:lsid:zoobank.org:act:DE306B1B-F770-4E79-9B9E-400CEC202266" httpUri="http://treatment.plazi.org/id/E615223BF2B05FB696BD9D895AE03E23">
<subSubSection id="FA4FB6660A38E939DF606AB516A24686" type="nomenclature">
<paragraph id="3860902C075AA794AD8FA5451A8E26FC">
<taxonomicName id="3B42DBC4CB4FC8D05D8B70BE296AB295" authority="Lan, Liu, Zhou &amp; Zhou" authorityName="Lan, Liu, Zhou &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="AF3C2B6C2214E8835519E7737036F4E9" italics="true">Triplophysa yaluwang</emphasis>
Lan, Liu, Zhou &amp; Zhou
</taxonomicName>
<taxonomicNameLabel id="C67FC3C3816AFAF018AC45ACC0874232" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="99048F066762BB45D46BCFFEF9C00466" type="description">
<paragraph id="F7EDC34417381C45A5C523BAB5CC9358">
<figureCitation id="B96B8682129EEB5CED336FA51D9F74AE" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological characteristics of holotype GZNU 20240118001 of Triplophysa yaluwang sp. nov. in preservative (10 % formalin) A dorsal view B ventral view C lateral view D dorsal view of head E lateral view of head, and F ventral view of head." figureDoi="10.3897/zookeys.1214.122439.figure6" httpUri="https://binary.pensoft.net/fig/1150826">Figs 6</figureCitation>
,
<figureCitation id="E9AFBF6CFD9D36A90888DDCD8776C371" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ecological photographs of Triplophysa yaluwang sp. nov. and closely related species in life A Triplophysa yaluwang sp. nov. (paratype, GZNU 20240118002) B Triplophysa yaluwang sp. nov. (paratype, GZNU 20240118005), and C T. guizhouensis." figureDoi="10.3897/zookeys.1214.122439.figure7" httpUri="https://binary.pensoft.net/fig/1150827">7</figureCitation>
,
<tableCitation id="5C06B4F694B1E4A7F09CA8D56D84EC31" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Morphological comparison of Triplophysa ziyunensis sp. nov. (TZ), Triplophysa yaluwang sp. nov. (TY), T. wudangensis (TW), T. rosa (TR), T. qingzhenensis (TQ), and T. guizhouensis (TG). All units in mm. P - values are at the 95 % significance level." httpUri="http://table.plazi.org/id/0E8A3204548C4BF47AC722F1629B7376" tableUuid="0E8A3204548C4BF47AC722F1629B7376">Table 5</tableCitation>
<xref id="92ADEA4777EE1E6558DFEE4883FE7BE2" ref-type="supplementary-material" rid="S1">, Suppl. material 1</xref>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E3ZFK" type="type material">
<paragraph id="DD188A0B8A5943C1391CB25106190AA4">
<heading id="8C0F3AA4354584F574B6A9B97B4EE60E" reason="title">Type material.</heading>
</paragraph>
<paragraph id="CA2DDC68C92CC9C3723D70214587DE4F">
<materialsCitation id="31B1187F162521C12014E4EB912FD0EC" collectingDate="2024-01-18" collectionCode="GZNU" collectorName="Jia-Jun Zhou" country="China" county="Ziyun County" elevation="1276" latitude="25.899088" location="Xinzhai Village" longLatPrecision="1" longitude="106.07921" municipality="Maoying Town" specimenCode="GZNU 20240118001" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<emphasis id="900F0E80F4609201D93731B155B201E7" bold="true">
<emphasis id="33DAB64F83252DBF7C417C1DF31D13EC" bold="true" italics="true">
<typeStatus id="9D8C3B16DDF7B8C7BEBF994D3D4236B7" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCode id="72AE1A75BE8EF51E7B79BABA8AA34582" collectionCode="GZNU">GZNU 20240118001</specimenCode>
(Fig.
<figureCitation id="2B3182AAA1CD7449DA3B9B70A121989E" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological characteristics of holotype GZNU 20240118001 of Triplophysa yaluwang sp. nov. in preservative (10 % formalin) A dorsal view B ventral view C lateral view D dorsal view of head E lateral view of head, and F ventral view of head." figureDoi="10.3897/zookeys.1214.122439.figure6" httpUri="https://binary.pensoft.net/fig/1150826">6</figureCitation>
),
<quantity id="C4BB953CFACCE34BB2050AB4BBFD9635" metricMagnitude="-2" metricUnit="m" metricValue="8.76" unit="mm" value="87.6">87.6 mm</quantity>
total length (
<abbrev id="ABBRID0EL1FK" xlink_title="total length">TL</abbrev>
),
<quantity id="67980DD9A5FC7BB775667A10719B68FF" metricMagnitude="-2" metricUnit="m" metricValue="7.39" unit="mm" value="73.9">73.9 mm</quantity>
standard length (SL), collected by
<collectorName id="1A58B93D1C0F1B5262E18DAF8FEDAC70">Jia-Jun Zhou</collectorName>
on
<date id="805D113C9142BDE1AE9F308F1DABD6D0" value="2024-01-18">
<collectingDate id="10ECF4869E562B5FB6C872924EE6103C" value="2024-01-18">18 January 2024</collectingDate>
</date>
, in
<location id="8B1E9645B7E3F699629A004EE7CD5352" LSID="urn:lsid:plazi:treatment:E615223BF2B05FB696BD9D895AE03E23:8B1E9645B7E3F699629A004EE7CD5352" country="China" county="Ziyun County" latitude="25.899088" longLatPrecision="1" longitude="106.07921" municipality="Maoying Town" name="Xinzhai Village" stateProvince="Guizhou">Xinzhai Village</location>
,
<collectingMunicipality id="74CD6579FF34F6C21115FE1CA2F4949B">Maoying Town</collectingMunicipality>
,
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,
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,
<collectingCountry id="CC51DB251F7F38E685E3DFF17D6EB6C6" name="China">China</collectingCountry>
(
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<geoCoordinate id="DC8A0CF227192739C85A8DD5B439ABAB" degrees="25.89908752" direction="north" orientation="latitude" precision="1" value="25.899088">25.89908752 ° N</geoCoordinate>
,
<geoCoordinate id="D2AFA46526A5F9129CDDEE279C59358A" degrees="106.07921141" direction="east" orientation="longitude" precision="1" value="106.07921">106.07921141 ° E</geoCoordinate>
</named-content>
,
<elevation id="2163E01FEC51B9A0816753D3C8732A11" metricMagnitude="3" metricUnit="m" metricValue="1.276" unit="m" value="1276.0">
<quantity id="C418114ADCD197610FBBC929681B509E" metricMagnitude="3" metricUnit="m" metricValue="1.276" unit="m" value="1276.0">1276 m</quantity>
a. s. l.
</elevation>
; Fig.
<figureCitation id="5E65AA1B9D182B75EAC8B1A43D790771" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sample collection localities and distributions of the two new species and 39 hypogean species of the genus Triplophysa in southern China. The base maps are from the Standard Map Service website (http://bzdt.ch.mnr.gov.cn/index.html)." figureDoi="10.3897/zookeys.1214.122439.figure1" httpUri="https://binary.pensoft.net/fig/1150821">1</figureCitation>
).
</materialsCitation>
</paragraph>
<caption id="3E54B31AA11A4B6A5A7D3BE3E44B2E87" ID-DOI="10.3897/zookeys.1214.122439.figure6" ID-arpha="1C637152-54BC-5A5D-85B6-C0C1B5C146F6" httpUri="https://binary.pensoft.net/fig/1150826" startId="F6">
<paragraph id="ACAE64C29BCB4E894D63339C9BDC7460">
<label id="5C5C035F1AA4F29B309B72CF2CC0CA45">Figure 6.</label>
</paragraph>
<paragraph id="F66C4625D099E38560E4A8EFBF0FC887">
Morphological characteristics of holotype GZNU 20240118001 of
<taxonomicName id="06FB1C496119F7A90D0CE9EF3E6AC1C9" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="96A0E2791366D7AAE4694EE5A420B696" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="51D75D929A7A6E7A720F5677E469A1E1" rank="species">sp. nov.</taxonomicNameLabel>
in preservative (10 % formalin)
<emphasis id="34FF08D33C0E0BD6BBA7E6A09820DA57" bold="true">A</emphasis>
dorsal view
<emphasis id="32E6C0983A8F0779D97E326A4F680E52" bold="true">B</emphasis>
ventral view
<emphasis id="22DA3FAB93238F5C0E3BF05A562FB6A8" bold="true">C</emphasis>
lateral view
<emphasis id="B718BE87CA2CFBC79373DEFADE0A0781" bold="true">D</emphasis>
dorsal view of head
<emphasis id="3A36BCF08996EC43211393BABBDB93A6" bold="true">E</emphasis>
lateral view of head, and
<emphasis id="C378D02A34EC9ECF6258285A629EF64A" bold="true">F</emphasis>
ventral view of head.
</paragraph>
</caption>
<paragraph id="0F34713C1DDE6C94187185B34681EAF2">
<emphasis id="A906AE93F9F9FEB19DD58E0393E9F70D" bold="true">
<emphasis id="D23E3020EA75BAFCAB6C9096C2F99F8C" bold="true" italics="true">
<typeStatus id="B4CEF123BCD4BBF60A0082EDA0FC454D" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount id="48D07EE9A179BB19F8C8E40534F044FC" count="4" type="generic">Four specimens</specimenCount>
from the same locality as the
<typeStatus id="0849526A6FB3A2606413F6A1507E24C9" type="holotype">holotype</typeStatus>
:
<collectionCode id="34B2E1A20F9E9FEE809E7A14ADB52B0C">GZNU</collectionCode>
20240118002118005,
<quantity id="3CAEE600167956F759BFC8AAB80C011F" metricMagnitude="-2" metricUnit="m" metricValue="6.9" metricValueMax="8.39" metricValueMin="5.41" unit="mm" value="69.0" valueMax="83.9" valueMin="54.1">54.183.9 mm</quantity>
SL, collected by Jia-Jun Zhou and Ye-Wei Liu on
<date id="923098369FCB5B62146B6A5D26B7C726" value="2023-09-27">27 September 2023</date>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EG3FK" type="diagnosis">
<paragraph id="7C99EC5603F8081D72EE18133C5A7041">
<heading id="80AF0D5A8521200E92CB6CB0F0078472" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="F53092F9554F29FF12D1E3D209E7C191">
<taxonomicName id="E59ACF8CBB61D080E689CF3A2AFBC465" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="6FDB83625BF0487FE0969E3A2371C485" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="68241757D992614EFD4F91E1F911DAE5" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from other hypogean species of the genus
<taxonomicName id="C72B3DF111A5BF6D84FED4317D9C1CF2" authorityName="Rendahl" authorityYear="1933" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="genus">
<emphasis id="471ECF1EB1060CB0A1E196AB1A43229F" italics="true">Triplophysa</emphasis>
</taxonomicName>
by the following characteristics in combination: (1) body naked, scaleless, with irregular pale dark brownish brown markings, except ventral; (2) eyes reduced, diameter 4.66.1 % of head length; (3) pelvic-fin tip reaching anus; (4) tip of pectoral fin not reaching to pelvic fin origin; (5) anterior and posterior nostrils closely set, with the anterior nostril elongated to a barbel-like tip; (6) tip of outrostral barbel extending backward, not reaching to anterior margin of eye; (7) lateral line complete; (8) posterior chamber of air bladder degenerated; and (9) dorsal-fin rays iii- 7, pectoral-fin rays i- 9, pelvic-fin rays i- 5, anal-fin rays i- 5, and 14 branched caudal-fin rays.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E53FK" type="description">
<paragraph id="3122E2515406EAE8ADD69ADF3D6C0521">
<heading id="67A782F9ED7F90268A09682F35BF3C59" reason="title">Description.</heading>
</paragraph>
<paragraph id="68D598DB1F58D6CA3BCD5C023BA541D6">
Morphological data of
<taxonomicName id="EFCDE20C6B6A1C1C541275B26A585580" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="89590045723B78B0767DAD3E104DBE47" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FA6B3BB2B9915D2F244A8019C8EA88EF" rank="species">sp. nov.</taxonomicNameLabel>
specimens are provided in Table
<tableCitation id="903BC4BC3EB1F517BE1ACCB1F4EA734F" captionStart="Table 5" captionStartId="T5" captionText="Table 5. Morphological comparison of Triplophysa ziyunensis sp. nov. (TZ), Triplophysa yaluwang sp. nov. (TY), T. wudangensis (TW), T. rosa (TR), T. qingzhenensis (TQ), and T. guizhouensis (TG). All units in mm. P - values are at the 95 % significance level." httpUri="http://table.plazi.org/id/0E8A3204548C4BF47AC722F1629B7376" tableUuid="0E8A3204548C4BF47AC722F1629B7376">5</tableCitation>
and Suppl. material
<xref id="A51D89420BC0F2F4125F74F52712B532" ref-type="supplementary-material" rid="S1">1</xref>
. Body elongated and cylindrical, posterior portion gradually compressed from dorsal fin to caudal-fin base, with deepest body depth anterior to dorsal-fin origin, deepest body depth 1216 % of SL. Dorsal profile slightly convex from snout to dorsal-fin insertion, then straight from posterior portion of dorsal-fin origin to caudal-fin base. Ventral profile flat. Head short, length 2627 % of SL, slightly depressed and flattened, width slightly greater than depth (HW / HD = 1.11.3). Snout slightly pointed, and snout length 4352 % of HL. Mouth inferior and curved, mouth corner situated below anterior nostril, upper and lower lips smooth, lower lip with V-shaped median notch. Three pairs of barbels are present: inner rostral barbel long, length 1627 % of HL, backward extending to corner of mouth; out rostral barbel long, length 3944 % of HL, backward extending to beyond anterior margin of eyes. Maxillary barbel not extending to posterior margin of operculum, length 2236 % of HL. Anterior and posterior nostrils closely set, length
<quantity id="60A7E9235F70F46CEC927FF0C95F1B67" metricMagnitude="-4" metricUnit="m" metricValue="6.3" metricValueMax="8.2" metricValueMin="4.4" unit="mm" value="0.63" valueMax="0.82" valueMin="0.44">0.440.82 mm</quantity>
. Anterior nostril tube long, with an elongated short barbel-like tip, tip of posterior nostril extending backwards not reaching to anterior margin of eye. Eyes reduced, with diameter 56 % of HL. Gill opening small, gill rakers not developed, nine inner gill rakers on first gill arch (
<emphasis id="B36233AD329B5C3ED75909438CC776B0" italics="true">n</emphasis>
= 1).
</paragraph>
<paragraph id="DCE928F01439EE743AD6BA3DD3DBC5E5">
Dorsal-fin rays iii- 7, pectoral-fin rays i- 9, pelvic-fin rays i- 56, anal-fin rays i- 5, 14 branched caudal-fin rays. Dorsal fin short, length 1922 % of SL, distal margin emarginated, origin anterior to pelvic-fin insertion and situated slightly posterior to the midpoint between snout tip and caudal-fin base, first branched ray longest, shorter than head length, tip of dorsal fin vertical to anus. Pectoral fin moderately developed, length 1925 % of SL, tip of pectoral fin extending backward almost to the midpoint between origin of pectoral and pelvic fin origins, not reaching to pelvic fin origin. Pelvic fin length 1617 % of SL, vertically aligned with second branched ray of dorsal fin, tips of pelvic fin reaching to anus. Anal fin length 1618 % of SL, distal margin truncated, origin close to anus, tips of anal fin not reaching caudal-fin base, distance between tips of anal fin and anus 2.2 × the eye diameter. Caudal fin forked, upper lobe slightly longer than lower lobe, tips pointed, caudal peduncle length ~
<quantity id="AA7D49B4F5965F9EFFD910944A386850" metricMagnitude="-2" metricUnit="m" metricValue="1.2" unit="mm" value="12.0">12 mm</quantity>
, caudal peduncle depth ~
<quantity id="01B137392706118CB2AC328AF1396D9D" metricMagnitude="-3" metricUnit="m" metricValue="4.8" unit="mm" value="4.8">4.8 mm</quantity>
, with weak adipose crests along both dorsal and ventral sides. Total vertebrae: 40 (
<emphasis id="70A191791298AF07FAB61A664311F630" italics="true">n</emphasis>
= 1).
</paragraph>
<paragraph id="09DB5E0AA58E1F4C3130C65E4D8AE11B">Cephalic lateral line system developed. Lateral line complete, exceeding tip of pectoral fin and reaching base of caudal fin. Two chambers of air bladder, anterior chamber dumbbell-shaped and membranous, open on both sides, slightly closed posteriorly; posterior chamber degenerated, slightly filling the body cavity, connected with anterior chamber by a long, slender tube.</paragraph>
</subSubSection>
<subSubSection id="SECID0E54FK" type="Coloration">
<paragraph id="C6539B6DE08F3A10BD1B2E3F0D741920">
<heading id="23191C76F2A998BFE05FA9B971AE139F" reason="title">Coloration.</heading>
</paragraph>
<paragraph id="32931F737D6EB3EFF326C8DCDC06EBCE">
In cave water, living fish were semi-translucent with a pale pink body with irregular dark brownish brown patches on the Entire body (Fig.
<figureCitation id="73FB71B96A803AD6E913A1CE45D37400" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ecological photographs of Triplophysa yaluwang sp. nov. and closely related species in life A Triplophysa yaluwang sp. nov. (paratype, GZNU 20240118002) B Triplophysa yaluwang sp. nov. (paratype, GZNU 20240118005), and C T. guizhouensis." figureDoi="10.3897/zookeys.1214.122439.figure7" httpUri="https://binary.pensoft.net/fig/1150827">7</figureCitation>
). After fixation in 10 % formalin, the body color was white, and the dark brown color lightened (Fig.
<figureCitation id="EDDFF90C278DB27FA682A1A6DF8DA97C" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Morphological characteristics of holotype GZNU 20240118001 of Triplophysa yaluwang sp. nov. in preservative (10 % formalin) A dorsal view B ventral view C lateral view D dorsal view of head E lateral view of head, and F ventral view of head." figureDoi="10.3897/zookeys.1214.122439.figure6" httpUri="https://binary.pensoft.net/fig/1150826">6</figureCitation>
).
</paragraph>
<caption id="B9679A48B2B5FCB13457EA6D178AEBBA" ID-DOI="10.3897/zookeys.1214.122439.figure7" ID-arpha="08B1CEB9-73F0-5B10-BCFE-9A76D6A0FE14" httpUri="https://binary.pensoft.net/fig/1150827" startId="F7">
<paragraph id="FB02DC403EDB245077FBBF607EF91825">
<label id="823F68CFC047A7058E22BBFAC09C2909">Figure 7.</label>
</paragraph>
<paragraph id="996E699494F9A2602DB8F470858AE47E">
Ecological photographs of
<taxonomicName id="83A81949D78941390563D89271161AF6" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="2A7EFF4918AB14FCC8012923CE90AD88" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="18AAC3F3D218A6FE99F337397AFAF8D2" rank="species">sp. nov.</taxonomicNameLabel>
and closely related species in life
<emphasis id="C5F56C6ABAFB034FA2CEFFCAD84BB08B" bold="true">A</emphasis>
<taxonomicName id="8BF1356B3EB6F7D73380859164EE476E" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="4FF2F908A561B9916DC5A977174071ED" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F97ABE1540C1855FCD20EED1659C41D7" rank="species">sp. nov.</taxonomicNameLabel>
(paratype, GZNU 20240118002)
<emphasis id="2CD8EFB378645E14166091678EBE95EE" bold="true">B</emphasis>
<taxonomicName id="C068CEC6C21E07B17EF4355C695A3A82" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="411D651AD9F6BC81F050B1CFD0978DF7" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0B7C5DD378A3025E72A8FEBAAC620F51" rank="species">sp. nov.</taxonomicNameLabel>
(paratype, GZNU 20240118005), and
<emphasis id="9E4410F6E460DD3C8BF67C7A6861A4AC" bold="true">C</emphasis>
<taxonomicName id="D1D30D774F084FD47DCE02A7E489E148" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="guizhouensis">
<emphasis id="33777C24EF5A67CB2F47E6121B96DCBC" italics="true">T. guizhouensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ELAGK" type="variation">
<paragraph id="DA9C8FB80E6BB94C70500E550C4DDECF">
<heading id="92404A0EF3B48CD3FBEE49066ACF0DB8" reason="title">Variations.</heading>
</paragraph>
<paragraph id="AB9AFF2BF51DBD8003E68F29A20A278A">
Among the
<specimenCount id="9E481259C7801AB0ACE77558C674E6C8" count="5" type="generic">five specimens</specimenCount>
collected,
<collectionCode id="8E4EC1CDD1C62CE0BB372DD1A8D11126">GZNU</collectionCode>
20240118002118004 are essentially identical to the
<typeStatus id="15E66093D1B3FAD0FB4254BC8AE69035" type="holotype">holotype</typeStatus>
in fin characteristics and body coloration.
<collectionCode id="A2E43A3B319525CA7E46B4E62949B6B9">GZNU</collectionCode>
20240118005 differs from the
<typeStatus id="20F68076355B28D7A0D4FBDF7133A5DC" type="holotype">holotype</typeStatus>
by the absence of body pigmentation and the absence of the eye (Fig.
<figureCitation id="682BE2ED46E70CBAFC879460D90E01C7" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ecological photographs of Triplophysa yaluwang sp. nov. and closely related species in life A Triplophysa yaluwang sp. nov. (paratype, GZNU 20240118002) B Triplophysa yaluwang sp. nov. (paratype, GZNU 20240118005), and C T. guizhouensis." figureDoi="10.3897/zookeys.1214.122439.figure7" httpUri="https://binary.pensoft.net/fig/1150827">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EVAGK" type="description">
<paragraph id="1B763CDFEA6E89CF3DF00240384879C0">
<heading id="99844E03A0249823A5E6EA3D2FCB579D" reason="title">Secondary sex characteristics.</heading>
</paragraph>
<paragraph id="BC32D0569B0B83227222B4C01D0442B1">
Secondary sex characteristics were not observed in the specimens of
<taxonomicName id="48D71D3D5FCDB493163DD6F13C804CE9" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="6FD73F1DB2A93518060F48BE30D5D4F8" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E472E307D7FD16B1EE0A72E24E2FD094" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EGBGK" type="comparisons">
<paragraph id="CA35A04013B32A639FADB654C1E7F22D">
<heading id="FF86F6EB6D40A6B61B1FE60FEDBFCA1E" reason="title">Comparisons.</heading>
</paragraph>
<paragraph id="390094E2E675B07300397C8DA3622936">
Detailed morphological comparative data of
<taxonomicName id="9BA14F7D29CF360FA41416DDD82E9137" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="DD6FE841C83A1239C0974E72B8524F09" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="752614F0F70C19AD0FA660274FA7D5A5" rank="species">sp. nov.</taxonomicNameLabel>
with
<taxonomicName id="D159425B50DFB14B527CCCAE87C47E36" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="ziyunensis" status="sp. nov.">
<emphasis id="1B8F969585522FED9CB3A0CFAEF85B94" italics="true">Triplophysa ziyunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="578FBB3E2BCF65F0876B1E257FAD2A75" rank="species">sp. nov.</taxonomicNameLabel>
and the 39 hypogean species of
<taxonomicName id="847113D10AE57E9CF875FC056AF6AA85" authorityName="Rendahl" authorityYear="1933" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="genus">
<emphasis id="AE288A48493850FA73F6F37033B90CDD" italics="true">Triplophysa</emphasis>
</taxonomicName>
are given in Table
<tableCitation id="70412CC55C5F7C045A735E5F77DFDE8E" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of the diagnostic features of the two new species described here with those selected for the 39 recognized hypogean species of the genus Triplophysa. Modified from Wu et al. (2018 b), Liu et al. (2022), and Luo et al. (2023). “ ” indicates uncertainty." httpUri="http://table.plazi.org/id/A655E81F294C66F97A3FA26414C2D66D" tableUuid="A655E81F294C66F97A3FA26414C2D66D">2</tableCitation>
.
<taxonomicName id="83A9FE294DE2E40BC1354365A2C73864" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="7AA37BBB5DDF683F743B238D443D7E30" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B40E8DAFFBF7072465C48AC3D42FD90D" rank="species">sp. nov.</taxonomicNameLabel>
is genetically close to
<taxonomicName id="DE388E1C3367617A93F35DCDC3BA3C6E" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="guizhouensis">
<emphasis id="7AC956D852CC4578A45404C7C159A18E" italics="true">T. guizhouensis</emphasis>
</taxonomicName>
,
<taxonomicName id="8CB2A67F2A8879EA1DC304D221E8A1D0" authorityName="Ren, Yang &amp; Chen" authorityYear="2012" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="longliensis">
<emphasis id="CA4BEF34C37DE32817490A138C17A940" italics="true">T. longliensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="A081FDF378AC97C538D3A3F5BB8F1BC6" authorityName="Chen &amp; Peng" authorityYear="2019" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="sanduensis">
<emphasis id="2C9AD00B9C4511BCB2536B8E46A25050" italics="true">T. sanduensis</emphasis>
</taxonomicName>
, but it can be distinguished in combination with morphological characteristics.
</paragraph>
<paragraph id="27568AAAAC3675B3B8F176DE317122B8">
<taxonomicName id="147DE6D8F0384633AA89A7F447BA06DA" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="8804B0959016CAC6CB943F2BEB3CA631" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A38D7F70EFCE712710ACE8E727342406" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="D625E23BF5F40573CBC742B40E942BD6" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="ziyunensis" status="sp. nov.">
<emphasis id="C4306B19878D09889594142271EB97F9" italics="true">Triplophysa ziyunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9405019248A893B457BD68A067249BF0" rank="species">sp. nov.</taxonomicNameLabel>
by having dorsal fin distal margin being emarginated (vs truncated), total vertebrae 40 (vs 39), seven branched dorsal fin rays (vs 8), nine branched pectoral fin rays (vs 10), and 14 branched caudal fin rays (vs 16).
</paragraph>
<paragraph id="3EDAA54FFF0ADC6DD9EC1923FB347608">
<taxonomicName id="F4FD32D3E20D3F07832F62CBF44A6BA0" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="05137904500AA5B56D183A1D0C3F6C14" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7ABD5808F817F579158F3476A6F8D8AE" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from
<taxonomicName id="23B49A40F698AEAA6096CA93B84F6221" authorityName="Ren, Yang &amp; Chen" authorityYear="2012" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="longliensis">
<emphasis id="56FD54DAD2B241B138127BD59893F524" italics="true">T. longliensis</emphasis>
</taxonomicName>
by having eyes reduced, small diameter 4.66.1 % of HL (vs normal, diameter 9.511.5 % of HL), interorbital width, 24.326.0 % of HL (vs 31.437.5 of HL), total vertebrae 40 (vs 42), degenerated posterior chamber of air bladder (vs developed), seven branched dorsal-fin rays (vs 8), nine branched pectoral-fin rays (vs 10), and 14 branched caudal-fin rays (vs 1516).
</paragraph>
<paragraph id="97B3FD22562243C7B3C3F995BB1DEAC3">
<taxonomicName id="0A8F34760E96F94B0868815B7A4D8E86" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="BF25B997661DD23DEBAC37D9FF4F07AF" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3C876C68CC55C2F67A7C0D9782255AE7" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from
<taxonomicName id="00077BAD38881CB0FBE264D9E5255B5E" authorityName="Chen &amp; Peng" authorityYear="2019" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="sanduensis">
<emphasis id="9F50DA88C59364F2E057DAF7A15C4DCB" italics="true">T. sanduensis</emphasis>
</taxonomicName>
by having eyes reduced, small diameter 4.66.1 % of HL (vs normal, diameter 11.915.4 % of HL), interorbital width, 24.326.0 % of HL (vs 31.240.2 of HL), total vertebrae 40 (vs 41), dorsal-fin rays, iii, 7 (vs ii, 89), three unbranched anal-fin rays (vs 1), 14 branched caudal-fin rays (vs 1718), and tip of pelvic fin reaching anus (vs not reaching anus).
</paragraph>
<paragraph id="0AF6280044B4D960225363A75D2293C6">
<taxonomicName id="CB8BC0126530F3473D83C5F1F1EE0855" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="DD35C8D527BFFFDC840C6FCDF9F09860" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C6C6F03B635D21F85CE8888517F80C02" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4F0567BA5FB498EC5A957356A8F73525" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="guizhouensis">
<emphasis id="9C94AFBB75C69F06EEF38D33FCF89A54" italics="true">T. guizhouensis</emphasis>
</taxonomicName>
by having eyes reduced, diameter 4.66.1 % of HL (vs normal, diameter 9.412.1 % of HL), dorsal fin distal margin being emarginated (vs truncated), body scaleless (vs body covered by sparse scales), degenerated posterior chamber of air bladder (vs developed), seven branched dorsal fin rays (vs 8), five branched anal-fin rays (vs 6), and tip of pelvic fin reaching anus (vs not reaching anus).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZGGK" type="ecology">
<paragraph id="B4B2232C07DABF56010DE56442D61032">
<heading id="2D978ACA6BB5134B9F90704909670248" reason="title">Ecology and distribution.</heading>
</paragraph>
<paragraph id="BB92414AB56CF604399B505197AE7CEE">
The new species
<taxonomicName id="B0F27BDE0FD98BD15D25A1BD8283EAB7" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="66168B30B54F1B02401FA2584EFEAEA9" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="01C36124C8D3FC6CF3559C24793F032C" rank="species">sp. nov.</taxonomicNameLabel>
was found in one cave far from the village of Xinzhai Village, Maoying Town, Ziyun County,
<collectingRegion id="698163F052851EF15873775AD857579C" country="China" name="Guizhou">Guizhou Province</collectingRegion>
,
<collectingCountry id="B3B949270438B8E45DD62E9A65801D9E" name="China">China</collectingCountry>
(Fig.
<figureCitation id="CC89D7B52A05E0274E4AA9E5E0593451" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sample collection localities and distributions of the two new species and 39 hypogean species of the genus Triplophysa in southern China. The base maps are from the Standard Map Service website (http://bzdt.ch.mnr.gov.cn/index.html)." figureDoi="10.3897/zookeys.1214.122439.figure1" httpUri="https://binary.pensoft.net/fig/1150821">1</figureCitation>
), in a water system where the underground river is a tributary of the Hongshui River. The cave habitat is a vertical shaft with an entrance located halfway up the mountainside. The underground river is approximately
<quantity id="191BCE4B5836884227E24C14F4D37FFE" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</quantity>
deep from the entrance, and the accessible portion is around
<quantity id="DA4D63BB398A3C6D1473FFEF52405695" metricMagnitude="2" metricUnit="m" metricValue="2.0" unit="m" value="200.0">200 m</quantity>
long,
<quantity id="2964B1B11CB4A5D0A1B1BFA56BCA0FCE" metricMagnitude="0" metricUnit="m" metricValue="3.0" unit="m" value="3.0">3 m</quantity>
wide, and
<quantity id="180EBCC00343EA6D434C8A90F7EDDE5D" metricMagnitude="0" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" unit="m" value="1.5" valueMax="2.0" valueMin="1.0">12 m</quantity>
deep. In this cave, the new species is sympatric with
<taxonomicName id="239AD76581ECBDE0EE5E2D8235CECC83" baseAuthorityName="Pellegrin" baseAuthorityYear="1931" family="Cyprinidae" genus="Sinocyclocheilus" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="multipunctatus">
<emphasis id="BB74423FE71193B7513B31713AEA2832" italics="true">Sinocyclocheilus multipunctatus</emphasis>
</taxonomicName>
and some unnamed spiders.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZHGK" type="remarks">
<paragraph id="0E83258168EBF2D389394BA31348C6F0">
<heading id="F36DA1A123D9455EC069B8084488DA8C" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="B0453CD935F97D0CCBCC4191F792C2CE">
The new species,
<taxonomicName id="5CB6ABE25B4567E938789281F07A5B5D" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang" status="sp. nov.">
<emphasis id="CDB00FAAAFCA7BEAFD9D0D59FC5537B2" italics="true">Triplophysa yaluwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB66EBC1A766B581B325A6C9077C4BF6" rank="species">sp. nov.</taxonomicNameLabel>
, inhabits the underground rivers of the
<typeStatus id="83D0E2F2108F12F1E03095E11B4B01FC">type</typeStatus>
locality. Eyes are present and reduced, and with irregular dark brownish brown patches on the head and body. Therefore, this species can be considered as a stygophile fish within the hypogean group of the genus
<taxonomicName id="D7BFBEB9874A50269596FC82C4F9AFEB" authorityName="Rendahl" authorityYear="1933" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="genus">
<emphasis id="F863413A95FC77592E07F86AC9887798" italics="true">Triplophysa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERIGK" type="etymology">
<paragraph id="64BC4564213A8ACBD394F25270828B5C">
<heading id="F355E5C0D9008E6FEEF1E1C35E29B3EE" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="B4D33ECD55D94F0B7C15101571964B9B">
The specific epithet
<taxonomicName id="5DEA355F89D1665BB161F8EACD30A883" authorityName="Lan &amp; Wu &amp; Luo &amp; Liu &amp; Zhou &amp; Yu &amp; Zhao &amp; Xiao &amp; Zhou" authorityYear="2024" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" phylum="Chordata" rank="species" species="yaluwang">
<emphasis id="4C1508CB834158963C21A94F3244483A" italics="true">yaluwang</emphasis>
</taxonomicName>
comes from King Yalu, a hero to the Miao people of Ziyun County,
<collectingRegion id="0A04E4A679EEA0765FD8373FD47DC29C" country="China" name="Guizhou">Guizhou Province</collectingRegion>
,
<collectingCountry id="E61D48BE887ADC0BB38FA49FB40D6897" name="China">China</collectingCountry>
, where the
<typeStatus id="D2CCCE95F6D917DC0FDEFE10C3BE176E">type</typeStatus>
locality is found. He was the 18
<superScript id="AEBA0A3984CAAF5C0EC0301E1C6D0009">th</superScript>
generation leader of the Miao ancestors in western
<collectingCountry id="9A962655ADFF2F24F2AB514A4A270DBA" name="China">China</collectingCountry>
and led the Miao people through many trials and tribulations. He eventually carved out a suitable land for his people to live in near the
<typeStatus id="0C6EA56CF38241950EF637739E6AF5F5">type</typeStatus>
locality. His deeds have been preserved in the form of a song, which has been organized into the first full-length heroic epic of the Hmong, King Yalu. We propose the common English name “ King Yalu high-plateau loach ” and the Chinese name “ Yà Lǔ Wáng Gāo Yuán Qīu (亚鲁王高原鳅) ”.
</paragraph>
</subSubSection>
</treatment>
</document>