165 lines
15 KiB
XML
165 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.760.24937" ID-GBIF-Dataset="173a45a1-64a2-44e8-a982-a6fbc905e816" ID-Pensoft-Pub="1313-2970-760-1" ID-ZBK="0B57270DC24C4D4DA04F15CA442E0A07" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-760-1" ModsDocOrigin="ZooKeys 760" ModsDocTitle="A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements" checkinTime="1527525277318" checkinUser="pensoft" docAuthor="Derkarabetian, Shahan, Starrett, James, Tsurusaki, Nobuo, Ubick, Darrell, Castillo, Stephanie & Hedin, Marshal" docDate="2018" docId="B3B22C897C0B5D60C441CA60697CEE52" docLanguage="en" docName="ZooKeys 760: 1-36" docOrigin="ZooKeys 760" docSource="http://dx.doi.org/10.3897/zookeys.760.24937" docTitle="CRYPTOMASTRIDAE Derkarabetian & Hedin, fam. n." docType="treatment" docUuid="A32A845F-36A7-426B-B3C1-F4E65085F356" docUuidSource="ZooBank" docVersion="6" lastPageNumber="1" masterDocId="4C02FFCBB1314B07FFAD2821FF83FFB1" masterDocTitle="A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements" masterLastPageNumber="36" masterPageNumber="1" pageNumber="1" updateTime="1643535363518" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Derkarabetian, Shahan</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Starrett, James</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tsurusaki, Nobuo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ubick, Darrell</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Castillo, Stephanie</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hedin, Marshal</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>760</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>36</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.760.24937</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.760.24937</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-760-1</mods:identifier>
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<mods:identifier type="ZBK">0B57270DC24C4D4DA04F15CA442E0A07</mods:identifier>
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<mods:identifier type="ZooBank">0B57270DC24C4D4DA04F15CA442E0A07</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="144148276" LSID="urn:lsid:zoobank.org:act:A32A845F-36A7-426B-B3C1-F4E65085F356" httpUri="http://treatment.plazi.org/id/B3B22C897C0B5D60C441CA60697CEE52" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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Family
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<taxonomicName LSID="http://zoobank.org/A32A845F-36A7-426B-B3C1-F4E65085F356" authority="Derkarabetian & Hedin" family="Cryptomastridae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Opiliones" pageId="0" pageNumber="1" rank="family">CRYPTOMASTRIDAE Derkarabetian & Hedin</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">fam. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type genus">
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<paragraph pageId="0" pageNumber="1">Type genus.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Cryptomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptomaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Cryptomaster</taxonomicName>
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Briggs, 1969
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type species">
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<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Cryptomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptomaster leviathan" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="Leviathan">Cryptomaster leviathan</taxonomicName>
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Briggs, 1969
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The
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<taxonomicName family="Cryptomastridae" lsidName="" pageId="0" pageNumber="1" rank="family">Cryptomastridae</taxonomicName>
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can be diagnosed from all other travunioids by the presence of a distal swelling on tibia II that bears enlarged setae (Figure 6A, C), a sexually dimorphic structure found only in males. Both genera are fairly distinctive.
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Cryptomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptomaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Cryptomaster</taxonomicName>
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is easily identified as the largest (>2.5 mm body length) laniatorean in the Pacific Northwest of North America (Figure 6D) and largest member of
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<taxonomicName lsidName="Travunioidea" pageId="0" pageNumber="1" rank="superfamily" superfamily="Travunioidea">Travunioidea</taxonomicName>
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, although two size forms exist (
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<bibRefCitation author="Starrett, J" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="11 - 35" title="A new monster from southwest Oregon forests: Cryptomasterbehemoth sp. n. (Opiliones, Laniatores, Travunioidea)." url="https://doi.org/10.3897/zookeys.555.6274" volume="555" year="2016">Starrett et al. 2016</bibRefCitation>
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).
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Speleomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speleomaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Speleomaster</taxonomicName>
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species are restricted to lava tubes showing extreme levels of troglomorphy with complete absence of eyes, extremely reduced pigmentation, and leg elongation (Figure 6B). Although unrelated,
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Speleomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speleomaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Speleomaster</taxonomicName>
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and
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Speleonychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speleonychia" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Speleonychia</taxonomicName>
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are both highly troglomorphic lava tube dwellers in the Pacific Northwest, found in Idaho and Washington, respectively. Aside from their disjunct geographic distribution,
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Speleomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speleomaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Speleomaster</taxonomicName>
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can be differentiated from
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Speleonychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speleonychia" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Speleonychia</taxonomicName>
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by the absence of a free ninth tergite and lateral sclerites, and by the presence of bifurcating tarsal claws of the hind legs (
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Speleonychia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speleonychia" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Speleonychia</taxonomicName>
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with a peltonychium). The cryptomastrid genera can be distinguished from the eastern North American
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<taxonomicName family="Cladonychiidae" lsidName="" pageId="0" pageNumber="1" rank="family">Cladonychiidae</taxonomicName>
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(
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Erebomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erebomaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Erebomaster</taxonomicName>
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+
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Theromaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theromaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Theromaster</taxonomicName>
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) by the spination of the pedipalpal tarsus, previously noted by
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<bibRefCitation pageId="0" pageNumber="1">Briggs (1969</bibRefCitation>
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,
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<bibRefCitation author="Briggs, TS" journalOrPublisher="Journal of Arachnology" pageId="0" pageNumber="1" pagination="205 - 214" title="Troglobitic harvestmen recently discovered in North American lava tubes (Travuniidae, Erebomastridae, Triaenonychidae: Opiliones)." url="http://www.jstor.org/stable/3705196" volume="1" year="1974">1974</bibRefCitation>
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). Cryptomastrids possess five prominent spines on the lateral margins of the pedipalpal tarsus, three on the prolateral margin and two on the retrolateral margin.
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Erebomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Erebomaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Erebomaster</taxonomicName>
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and
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Theromaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theromaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Theromaster</taxonomicName>
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possess three pairs of prominent lateral spines (in some
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Theromaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theromaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Theromaster</taxonomicName>
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, the two apical retrolateral spines are fused at the base). The
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<taxonomicName family="Cryptomastridae" lsidName="" pageId="0" pageNumber="1" rank="family">Cryptomastridae</taxonomicName>
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are unique in intestinal morphology, possessing a combination of an elongate and triangular DI (similar to
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Briggsus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Briggsus" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Briggsus</taxonomicName>
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and
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<taxonomicName class="Arachnida" family="Cladonychiidae" genus="Isolachus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isolachus" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Isolachus</taxonomicName>
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), and shorter OD2 and OD3 (similar to the
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<taxonomicName family="Paranonychidae" lsidName="" pageId="0" pageNumber="1" rank="family">Paranonychidae</taxonomicName>
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) (Suppl. material 2: Figure 2).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="included genera and species">
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<paragraph pageId="0" pageNumber="1">Included genera and species.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Cryptomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptomaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Cryptomaster</taxonomicName>
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. Described by
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<bibRefCitation pageId="0" pageNumber="1">Briggs (1969)</bibRefCitation>
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and originally included only
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Cryptomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptomaster leviathan" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="Leviathan">Cryptomaster leviathan</taxonomicName>
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Briggs, 1969 from the Coastal Range of southwestern Oregon. A second species,
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Cryptomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cryptomaster behemoth" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="Behemoth">Cryptomaster behemoth</taxonomicName>
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Starrett & Derkarabetian, 2016, was described from the west-central Cascade Range of Oregon (
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<bibRefCitation author="Starrett, J" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="11 - 35" title="A new monster from southwest Oregon forests: Cryptomasterbehemoth sp. n. (Opiliones, Laniatores, Travunioidea)." url="https://doi.org/10.3897/zookeys.555.6274" volume="555" year="2016">Starrett et al. 2016</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Speleomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speleomaster" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Speleomaster</taxonomicName>
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.
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<bibRefCitation author="Briggs, TS" journalOrPublisher="Journal of Arachnology" pageId="0" pageNumber="1" pagination="205 - 214" title="Troglobitic harvestmen recently discovered in North American lava tubes (Travuniidae, Erebomastridae, Triaenonychidae: Opiliones)." url="http://www.jstor.org/stable/3705196" volume="1" year="1974">Briggs (1974)</bibRefCitation>
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described the genus and both known species,
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Speleomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speleomaster lexi" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="Lexi">Speleomaster lexi</taxonomicName>
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Briggs, 1974 and
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<taxonomicName class="Arachnida" family="Cryptomastridae" genus="Speleomaster" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Speleomaster pecki" order="Opiliones" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="Pecki">Speleomaster pecki</taxonomicName>
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Briggs, 1974, from lava tubes of the Snake River Plain in southern Idaho.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |