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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan &amp; Machard, Patrice" docDate="2022" docId="B27B27DBA2A155E0A768173362CC9ED0" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) arabicus Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard 2022, sp. nov." docType="treatment" docUuid="20D0899F-1E9F-41EB-8247-EBA911C8DEFE" docUuidSource="ZooBank" docVersion="2" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1655206106596" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Gueorguiev, Borislav</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
<mods:affiliation>National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
<mods:nameIdentifier type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
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<mods:namePart>Wrase, David W.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Assmann, Thorsten</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
<mods:affiliation>Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Muilwijk, Jan</mods:namePart>
<mods:affiliation>Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Machard, Patrice</mods:namePart>
<mods:affiliation>29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:date>2022</mods:date>
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13. 
<taxonomicName LSID="http://zoobank.org/20D0899F-1E9F-41EB-8247-EBA911C8DEFE" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) arabicus</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
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<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3B</figureCitation>
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14E</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
</paragraph>
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<subSubSection pageId="0" pageNumber="71" type="type locality">
<paragraph pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
Saudi Arabia, 
<normalizedToken originalValue="“Hedjaz”">&quot;Hedjaz&quot;</normalizedToken>
.
</paragraph>
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<subSubSection pageId="0" pageNumber="71" type="notes on type locality">
<paragraph pageId="0" pageNumber="71">Notes on type locality.</paragraph>
<paragraph pageId="0" pageNumber="71">
Nothing is known about the exact places and circumstances of its collecting. The Hedjaz Range is a mountain range located in the Hejazi region, the western part of the country. It is very likely, as in the case of two Saudi Arabian species of 
<taxonomicName class="Insecta" family="Carabidae" genus="Acinopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acinopus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Acinopus</emphasis>
</taxonomicName>
, 
<taxonomicName genus="A." lsidName="A. brittoni" pageId="0" pageNumber="71" rank="species" species="brittoni">
<emphasis italics="true" pageId="0" pageNumber="71">A. brittoni</emphasis>
</taxonomicName>
Wrase &amp; Kataev, 2016 and 
<taxonomicName genus="A." lsidName="A. arabicus" pageId="0" pageNumber="71" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">A. arabicus</emphasis>
</taxonomicName>
Wrase &amp; Kataev, 2016 (
<bibRefCitation author="Wrase, DW" journalOrPublisher="Linzer Biologische Beitraege" pageId="0" pageNumber="71" pagination="1783 - 1806" refId="B53" refString="Wrase, DW, Kataev, BM, 2016. Four new species of genus Acinopus Dejean, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina). Linzer Biologische Beitraege 48 (2): 1783 - 1806" title="Four new species of genus Acinopus Dejean, 1821, subgenus Acinopus from southern Iran, from Sinai, and from western Saudi Arabia, and faunistic and taxonomic notes on species previously described (Coleoptera, Carabidae, Harpalini, Harpalina)." volume="48" year="2016">Wrase and Kataev 2016</bibRefCitation>
), that 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
lives in higher altitudes where habitats with enough moisture are present.
</paragraph>
<caption ID-Table-UUID="D5F1A345CA056C2DBA2597271787B1E8" httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" start="Table 3" startId="T3">
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">Table 3.</emphasis>
Morphometric data for species of the ' 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
' group of 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
.
</paragraph>
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<paragraph pageId="0" pageNumber="71">
<table pageId="0" pageNumber="71">
<tr pageId="0" pageNumber="71">
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">Species (number of samples)</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">BL/mm</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">BW/mm</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/HW</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PL</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">
PW/ 
<normalizedToken originalValue="PА">PA</normalizedToken>
</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PW/PB</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">PA/PB</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/EW</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EW/PW</th>
<th colspan="1" pageId="0" pageNumber="71" rowspan="1">EL/PL</th>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov. (3♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.20-9.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.85-3.25</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.41-1.43</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.07-1.11</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.44-1.48</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09-1.15</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.76-0.79</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.54-1.62</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.44-1.49</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.47-2.56</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov. (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.73</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">3.05</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.46</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.78</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.59</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.46</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.51</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov. (2 exx.)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">3.30-3.45</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.02-1.05</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38-1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.14-1.22</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.80-0.88</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.58-1.59</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.48</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.40-2.44</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov. (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">3.38</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.04</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.84</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.59</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.48</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.42</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
</taxonomicName>
Karsch (5♂♂, 4♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.70-9.30</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.70-3.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.28-1.41</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10-1.20</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36-1.43</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.21-1.28</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.88-0.92</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.53-1.72</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36-1.48</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.53-2.72</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName authority="brunneus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus brunneus</emphasis>
</taxonomicName>
Karsch (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.60</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.99</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.35</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.16</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.25</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.90</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.61</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.41</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.63</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
</taxonomicName>
Reitter (1♂, 6♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.60-9.00</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.70-3.20</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29-1.39</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.14-1.21</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34-1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.12-1.23</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.82-0.89</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.52-1.60</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.35-1.46</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.46-2.58</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus subsp. ferrantei" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="ferrantei">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus ferrantei</emphasis>
</taxonomicName>
Reitter (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.53</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">3.01</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.32</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.16</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.17</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.56</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.41</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.53</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName authority="insignitus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="insignitus">
<emphasis italics="true" pageId="0" pageNumber="71">P. insignitus insignitus</emphasis>
</taxonomicName>
Bedel (1♂, 4 ♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.60-8.70</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.20-3.00</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.35-1.45</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.06-1.17</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.35-1.40</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.12-1.17</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.80-0.85</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.57-1.65</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.40-1.51</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.41-2.61</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName authority="insignitus" class="Insecta" genus="P." kingdom="Animalia" lsidName="P. insignitus subsp. insignitus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="subSpecies" species="insignitus" subSpecies="insignitus">
<emphasis italics="true" pageId="0" pageNumber="71">P. insignitus insignitus</emphasis>
</taxonomicName>
Bedel (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.72</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.67</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.38</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.83</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.61</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.45</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.55</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. (2♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.30-9.00</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.90-3.20</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.30-1.32</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.34-1.38</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.09-1.11</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.79-0.83</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.62-1.63</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.45-1.47</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.65-2.72</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov. (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.65</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">3.05</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.13</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.36</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.81</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.63</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.46</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.69</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
(Reiche &amp; Saulcy) (10♂♂, 10♀♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">6.40-9.10</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.20-3.15</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.27-1.38</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.11-1.22</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.31-1.41</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10-1.20</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.81-0.90</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.52-1.62</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.29-1.48</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.42-2.60</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
(Reiche &amp; Saulcy) (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">7.82</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.74</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.33</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.16</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.35</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.16</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.56</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.37</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.49</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. (4♂♂, 1♀)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">8.70-9.90</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">3.00-3.45</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.35-1.42</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.08-1.13</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39-1.52</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.15-1.22</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.78-0.87</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.56-1.60</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39-1.46</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.41-2.60</td>
</tr>
<tr pageId="0" pageNumber="71">
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. (mean)
</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">9.28</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">3.25</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.39</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.10</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.44</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.18</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">0.82</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.59</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">1.43</td>
<td colspan="1" pageId="0" pageNumber="71" rowspan="1">2.50</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="71">
<tableNote pageId="0" pageNumber="71">For abbreviations in line 1, see &quot;Abbreviations to measurements and ratios&quot; (section &quot;Material and methods&quot;).</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="type material">
<paragraph pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♀, 'Hedjaz [w, h] // Ex-Musaeo / H.W. Bates / 1892 [w, p]' (MNHN). Paratype: 1♀, 'El Hahaz / Millinger [sic] [w, p] // El Hedjaz. / Millingen. / 1915-38.' [w, p] // 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/ 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
/ Reiche / E.B. Britton det. / 1946 [w, h&amp;p]' (NHMUK).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="materials_examined">
<paragraph pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="71">
Imprecise locality: 
<specimenCount type="female">1♀</specimenCount>
, 'Bagdad [w, h] // Ex-Musaeo / H.W. Bates / 1892' [w, p] (MNHN).
</paragraph>
<paragraph pageId="0" pageNumber="71">
TME: 
<specimenCount type="generic">3 specimens</specimenCount>
. TGE: 
<specimenCount type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="etymology">
<paragraph pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="71">The specific epithet is a Latinized adjective, based on the name of the region in which this species can be found.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="diagnosis">
<paragraph pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="71">
Among members of &quot; 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group, 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. are taxa with pronotum most constricted toward apex (PW/PA: 1.44-1.48 and 1.39-1.52, respectively; Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
). However, pronotal forms differ. Whereas the former has a subelliptic pronotum with sides to base straight or slightly convex and less constricted, the latter has a subquadrate pronotum with sides to base slightly concave and more constricted (PW/PB: 1.09-1.15 and 1.15-1.22, respectively). In addition, 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
has a darker, brown or chestnut color of integument and 5-6 setiferous punctures of anterior side of mesofemur ventrally, whereas 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. jordanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="jordanensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. jordanensis</emphasis>
</taxonomicName>
sp. nov. has a lighter, reddish-brown to rufous color and 4 setiferous punctures of anterior side of mesofemur ventrally.
</paragraph>
<paragraph pageId="0" pageNumber="71">
It is also closely related to 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov. but two species differ by a set of morphometric ratios (see 
<normalizedToken originalValue="“Diagnosis”">&quot;Diagnosis&quot;</normalizedToken>
, under 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov.).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="description">
<paragraph pageId="0" pageNumber="71">Description.</paragraph>
<paragraph pageId="0" pageNumber="71">
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of large size for 
<taxonomicName authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL: 8.20-9.40 mm; BW: 2.85-3.25 mm), with elongate, moderately to rather convex body (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3B</figureCitation>
). 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table 
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages uniformly dark reddish-brown, only terminal palpomeres slightly lighter. Integument slightly to moderately shiny, head and pronotum shinier than elytra. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Pronotum with evident microreticulation throughout, sculpticells regular isodiametric to slightly stretched. Elytra (intervals, scutellum, basal margin and lateral gutter) with distinct isodiametric sculpticells. Ventral surface with well-impressed isodiametric or slight transverse sculpticells, only epipleura, mesosternum and middle coxa with sculpticells scarcely-visible. Head impunctate, smooth or nearly smooth, with a few, very shallow wrinkles on clypeus posterior half and frontal furrows laterally. Pronotum surface mostly smooth, only basal area between foveae longitudinally wrinkled and along lateral margin with few punctures that do not reach anterior half (paratype also with several wrinkles in front of anterior transverse impression). Elytral intervals impunctate or with scattered and very shallow punctures. Abdominal ventrite 1 wrinkled medially, 2-6 smooth, neither wrinkled nor punctate. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Head</emphasis>
.
</emphasis>
More than one-third narrower than pronotum wide (PW/HW= 1.41-1.43). Eyes moderately convex. Labrum subrectangular, slightly shorter than clypeus, with anterior margin concave. Frontoclypeal suture slightly distinct in middle, indistinct at sides. Frontal furrows very shallow, subfoveolate. Paraorbital sulci straight, backward barely reaching posterior margin of eye, not reaching level of posterior supraorbital pore. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum about one time and one tenth as wide as long (PW/PL= 1.07-1.11), with widest point at medial third. Anterior transverse impression indistinct to slightly distinct, posterior transverse impression barely distinct. Sides not sinuate, smoothly convex medially and anteriorly, nearly straight posteriorly; anterior bead present laterally, lack in medial 1/8-1/10; basal bead present laterally, reduced to varying degrees or non-existent in medial half. Metepisternum as long as wide, MA/MI= 1.00. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one -and-a-half times as long as elytra wide (EL/EW= 1.54-1.62), two and a half times as long as pronotum (EL/PL= 2.47-2.56), and one and a half as wide as pronotum (EW/PW= 1.44-1.49), with widest point at beginning of third quarter. Parascutellar striole punctiform to sublinear, very shallow; striae 1-6 more impressed than striole and striae 7 and 8, moderately to indistinctly punctate; parascutellar striole short, not joining stria 1; base of stria 1 ending in parascutellar pore, striae 2-5 reaching basal bead, 6 and 7 ending little before. Interval 3 with three discal setiferous punctures on right elytron (medial one of left elytron lacking in two specimens). Umbilicate setiferous series with 15 punctures on left elytron and 16 on right elytron in holotype, with 16 punctures on each side in paratype. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Mesofemur with 4 setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with two to three long setae, one in basal third and one to two in apical half. 
<emphasis bold="true" pageId="0" pageNumber="71">
<emphasis italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Unknown. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14E</figureCitation>
). Apical gonocoxite with pointed apex and one dorsolateral ensiform seta. Spermathecal canal connected in basal third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="comparisons">
<paragraph pageId="0" pageNumber="71">Comparisons.</paragraph>
<paragraph pageId="0" pageNumber="71">
From 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
sp. nov., that inhabits areas northeast of the Al-Hedjaz region, 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
differs in: (1) dark brown color of body (vs. orange-brown color of body); (2) head narrower, compared to pronotum (PW/HW: 1.41-1.43, vs. PW/HW: 1.30-1.32); (3) pronotum with anterior angles more pointed and sides to apex more constricted (PW/PA: 1.44-1.48), vs. pronotum with anterior angles less pointed and sides to apex less constricted (PW/PA: 1.34-1.38); (4) elytra less long, compared to pronotum (EL/PL: 2.47-2.56, vs. EL/PL: 2.65-2.72); (5) meso- and metatarsomeres dorsally neither flattened nor grooved (vs. meso- and metatarsomeres dorsally partly flattened and slightly grooved).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="habitat">
<paragraph pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="71">
Like other representatives of the &quot; 
<taxonomicName baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group with known habitat preferences (
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus">
<emphasis italics="true" pageId="0" pageNumber="71">P. languidus</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunneus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunneus</emphasis>
</taxonomicName>
, 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis italics="true" pageId="0" pageNumber="71">P. brunki</emphasis>
</taxonomicName>
sp. nov.), 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov. could inhabit the wetter parts of desert and semi-desert habitats.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="distribution">
<paragraph pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="71">
The new species is known from the region of Hedjaz in Saudi Arabia, which includes the western part of the country (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
). Likely to be endemic to the Hedjaz Mountains. As far as the specimen with the locality 
<normalizedToken originalValue="“Bagdat”">&quot;Bagdat&quot;</normalizedToken>
is concerned, we believe that this inscription is either due to a mislabeling or a genuine observation. If it is the latter case, then the species probably occurs in the Iraqi areas on the border with Saudi Arabia, southwest of the city of Baghdad.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="71" type="notes">
<paragraph pageId="0" pageNumber="71">Notes.</paragraph>
<paragraph pageId="0" pageNumber="71">
The specimen from 
<normalizedToken originalValue="“Bagdad”">&quot;Bagdad&quot;</normalizedToken>
differs from the two type specimens in the larger, more quadratic and slightly wider pronotum (vs. smaller and somewhat narrow pronotum) and having less prominent anterior angles (vs. somewhat more prominent anterior angles). That is why, we have some doubt if this specimen really belongs to 
<taxonomicName class="Insecta" genus="P." kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>