275 lines
37 KiB
XML
275 lines
37 KiB
XML
<document id="FFD648A898D6F0AE235052E016437834" ID-DOI="10.11646/zootaxa.2901.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5284936" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629727970278" checkinUser="felipe" docAuthor="Yanes, Yurena, Santana, Jesús, Artiles, Miguel, Deniz, Francisco, Martín, Javier, Alonso, María R. & Ibáñez, Miguel" docDate="2011" docId="8146737DFFE9FFD6FF838D4BA0F3E0E0" docLanguage="en" docName="zt02901p051.pdf" docOrigin="Zootaxa 2901 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2901.1.3" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Napaeus grohi Alonso, Ibanez & Santana 2011, n. sp." docType="treatment" docVersion="4" lastPageNumber="49" masterDocId="7D7F0B05FFE4FFD8FF148B76A25FE52B" masterDocTitle="Five new Napaeus species (Gastropoda: Pulmonata: Enidae) from Gran Canaria and El Hierro (Canary Islands)" masterLastPageNumber="51" masterPageNumber="35" pageNumber="48" updateTime="1699102865397" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="F0B34BBA8BAB6272D328FB889DE63E70" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="6FD4039C04342D2A81AC5A7E9DDBFA43">
|
||
<mods:title id="348463351230B7ECC11CB04546049576">Five new Napaeus species (Gastropoda: Pulmonata: Enidae) from Gran Canaria and El Hierro (Canary Islands)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="BF797713DC7FA3C8C16F2FAF12B26832" type="personal">
|
||
<mods:role id="2CA4658D9E05559D8C0E8A07B36FA1B0">
|
||
<mods:roleTerm id="34D6BFA3F9170FBADC954F582B319A2B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="CE5B65DD9085C552AC2C1828B0AE8755">Yanes, Yurena</mods:namePart>
|
||
<mods:affiliation id="F2BAAAB3DE984228D080F85132CF40B3">Instituto Andaluz de Ciencias de la Tierra (CSIC-Universidad de Granada), Camino del Jueves s / n, 18100, Armilla, Granada, Spain. E-mail: yurenayanes @ ugr. es</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="CF434A2B33A478F182654CD82A793D9F" type="personal">
|
||
<mods:role id="0D0E44D09F08CB4D3BAA90E49020BADA">
|
||
<mods:roleTerm id="ECA0AA2646A9E65D1EAF883660EE01E1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A85EE5295A5A5F5405C1B7C11C682078">Santana, Jesús</mods:namePart>
|
||
<mods:affiliation id="F0672B9B61B1938D6B46D74E08D3FB20">Las Palmas de Gran Canaria, Gran Canaria, Canary Islands, Spain. E-mail: CASIOPEA _ 69 @ terra. es; franciscodeniz @ yahoo. es Arinaga, Gran Canaria, Canary Islands, Spain. E-mail: miguel _ artiles @ yahoo. es</mods:affiliation>
|
||
<mods:nameIdentifier id="59A247231C396FFA9745A142D9D28962" type="email">_69@terra.es</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="48FC0D788D0CABFBE1DB27F389F2372E" type="personal">
|
||
<mods:role id="3B3183201435B3F76591A220D52DDAAE">
|
||
<mods:roleTerm id="489DD285A974F8422FBE7283C7856A51">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6BE9F760653D9525D85A0887404BA2F5">Artiles, Miguel</mods:namePart>
|
||
<mods:affiliation id="618F3CBA860B51146DF21A931FBA1A64">Las Palmas de Gran Canaria, Gran Canaria, Canary Islands, Spain. E-mail: CASIOPEA _ 69 @ terra. es; franciscodeniz @ yahoo. es Arinaga, Gran Canaria, Canary Islands, Spain. E-mail: miguel _ artiles @ yahoo. es</mods:affiliation>
|
||
<mods:nameIdentifier id="ED94720B0969BE72CDA1A48F11E8CE9A" type="email">_69@terra.es</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="F3EE0A35C137E663D23E07748F471E81" type="personal">
|
||
<mods:role id="17712D5B87224290411FA11BB82B3D18">
|
||
<mods:roleTerm id="823D24DBD14DF3DCFE2B35A426159C33">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="12046EEB7499146FDC188E17AABA804F">Deniz, Francisco</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="61A93599D216BB9A50FFC22339FDEB21" type="personal">
|
||
<mods:role id="1109DB30068DCA7172B6BD1FE9614ED9">
|
||
<mods:roleTerm id="3AF6CC4BF41EE025E19EB0A8049C0848">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="40A6CE0D276EE571B3C05188B452725F">Martín, Javier</mods:namePart>
|
||
<mods:affiliation id="F12F8D47AAE076B50570EDBD993381B3">El Coromoto, La Laguna, Tenerife, Canary Islands, Spain. E-mail: javimarba @ hotmail. com</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="9ACD498FD3CF14808FDAA4CD0AEC3AFA" type="personal">
|
||
<mods:role id="C2FF1586F6C56AB1892B4DBD5FA903B9">
|
||
<mods:roleTerm id="19D04BCEB92B2E6B1D2EE05EBE29C08C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BBBF867C67EF0B7D009F89B98193F0BF">Alonso, María R.</mods:namePart>
|
||
<mods:affiliation id="FD2E9CF45404DA0AFC88EDD9238F7B29">Departamento de Biologia Animal, Universidad de La Laguna, E- 38206 La Laguna, Tenerife, Islas Canarias, Spain.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="C93FEA8CB6E2DD84D2677B3CF656AC7C" type="personal">
|
||
<mods:role id="1590665F0A4B1E4B84EF1E9FB5BCFB7B">
|
||
<mods:roleTerm id="0B3BE3210FFE0787E4A117FA1B6183C5">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4CFD23F2FF2B53053AA62BD62F8CCC30">Ibáñez, Miguel</mods:namePart>
|
||
<mods:affiliation id="3188516D6EC8F5350F85EE5C057159D8">Departamento de Biologia Animal, Universidad de La Laguna, E- 38206 La Laguna, Tenerife, Islas Canarias, Spain.</mods:affiliation>
|
||
<mods:nameIdentifier id="E03EC0829D182B9156AE6FC2EEC5C566" type="email">mibanez@ull.es</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource id="FDF5EACBA60B0803BB0123F3F7F4842F">text</mods:typeOfResource>
|
||
<mods:relatedItem id="FE78EE6D1340D42063B17994AB635B25" type="host">
|
||
<mods:titleInfo id="3EDEE9C4F1D36F444C3E36D1AF797DD4">
|
||
<mods:title id="C3FC0D5043465ADF35645C8BD074AFCE">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="D7774CBC302B8090482302C56C345B3F">
|
||
<mods:date id="C54B2DE4DC56DDD5BF635B850385739F">2011</mods:date>
|
||
<mods:detail id="AAC6533C1DC529654FBFCB75E99927AE" type="pubDate">
|
||
<mods:number id="9F4904FCA08D368C9079D1F1A7ABDA2C">2011-06-01</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="7E532AE7336453105BB34C66B00C9508" type="volume">
|
||
<mods:number id="41A6DEAB11B11C7BF7617B2069142EE7">2901</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="298CFD11012251D9DB8D0FE26DA1A747" type="issue">
|
||
<mods:number id="E883E28E0243F73615F779011E38D1A5">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="A6E2CDF4248BB8769E108DC142EE445B" unit="page">
|
||
<mods:start id="FA3D3A34C7DFD95B67E3BB3E8B98A429">35</mods:start>
|
||
<mods:end id="DB8FDE93368628BCA778D31A30FDD19B">51</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="3E79EAB00BF65D5281AFA7BCD70E31B8">
|
||
<mods:url id="2B1A823B4E19FA9B5AB0B3D6154447B8">https://biotaxa.org/Zootaxa/article/view/zootaxa.2901.1.3</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="2358724C77D55CBD0D5E2580B7BC19A7">journal article</mods:classification>
|
||
<mods:identifier id="2B8B98A7F4FFD2599A7BC726E0B7662F" type="DOI">10.11646/zootaxa.2901.1.3</mods:identifier>
|
||
<mods:identifier id="05ED7B65B907BBC049656AFBAF8085F7" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="0F501DCEAB0A7F10C790AFDEF6F0EF46" type="Zenodo-Dep">5284936</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="8146737DFFE9FFD6FF838D4BA0F3E0E0" ID-DOI="http://doi.org/10.5281/zenodo.5292372" ID-Zenodo-Dep="5292372" LSID="urn:lsid:plazi:treatment:8146737DFFE9FFD6FF838D4BA0F3E0E0" httpUri="http://treatment.plazi.org/id/8146737DFFE9FFD6FF838D4BA0F3E0E0" lastPageId="14" lastPageNumber="49" pageId="13" pageNumber="48">
|
||
<subSubSection id="41F591E0FFE9FFD5FF838D4BA087E37C" box="[151,728,1597,1623]" pageId="13" pageNumber="48" type="nomenclature">
|
||
<paragraph id="0950C26BFFE9FFD5FF838D4BA087E37C" blockId="13.[151,728,1597,1656]" box="[151,728,1597,1623]" pageId="13" pageNumber="48">
|
||
<heading id="52187507FFE9FFD5FF838D4BA087E37C" bold="true" box="[151,728,1597,1623]" fontSize="11" level="1" pageId="13" pageNumber="48" reason="1">
|
||
<emphasis id="3B9B1E79FFE9FFD5FF838D4BA087E37C" bold="true" box="[151,728,1597,1623]" pageId="13" pageNumber="48">
|
||
<taxonomicName id="CEEFB9E8FFE9FFD5FF838D4BA0D3E37C" ID-CoL="45NPV" authority="Alonso, Ibanez & Santana" authorityName="Alonso, Ibanez & Santana" authorityYear="2011" box="[151,652,1597,1623]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="48" phylum="Mollusca" rank="species" species="grohi" status="sp. nov.">
|
||
<emphasis id="3B9B1E79FFE9FFD5FF838D4BA31CE37C" bold="true" box="[151,323,1597,1623]" italics="true" pageId="13" pageNumber="48">Napaeus grohi</emphasis>
|
||
Alonso, Ibáñez & Santana
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="20A8A302FFE9FFD5FD8D8D4BA087E37C" box="[665,728,1597,1623]" pageId="13" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41F591E0FFE9FFD5FF838D16A396E353" box="[151,457,1632,1656]" pageId="13" pageNumber="48" type="description">
|
||
<paragraph id="0950C26BFFE9FFD5FF838D16A396E353" blockId="13.[151,728,1597,1656]" box="[151,457,1632,1656]" pageId="13" pageNumber="48">
|
||
<figureCitation id="91D4DEEEFFE9FFD5FF838D16A2BCE353" box="[151,227,1632,1656]" captionStart="FIGURE 1" captionStartId="3.[151,248,1648,1671]" captionTargetBox="[156,1430,207,1614]" captionTargetId="figure-40@3.[151,1436,195,1624]" captionTargetPageId="3" captionText="FIGURE 1. Geographical distribution of the new species." figureDoi="http://doi.org/10.5281/zenodo.5284942" httpUri="https://zenodo.org/record/5284942/files/figure.png" pageId="13" pageNumber="48">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="91D4DEEEFFE9FFD5FFE58D16A34CE353" box="[241,275,1632,1656]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="13" pageNumber="48">3 J</figureCitation>
|
||
,
|
||
<figureCitation id="91D4DEEEFFE9FFD5FE0B8D17A333E353" box="[287,364,1632,1656]" captionStart="FIGURE 7" captionStartId="9.[151,248,1358,1381]" captionTargetBox="[171,1416,198,1320]" captionTargetId="figure-301@9.[151,1436,193,1337]" captionTargetPageId="9" captionText="FIGURE 7. Scatter plots of some shell measurements for the new species and the most similar species. AS, aperture surface area (plane view); BS, body whorl surface area (plane view); FS, first whorls surface area (plane view); PS, penultimate whorl surface area (plane view); SB, shell breadth; SH, shell height; SS, shell surface area (plane view). The dimensions are in mm (lengths) or mm2 (surface areas)." figureDoi="http://doi.org/10.5281/zenodo.5284962" httpUri="https://zenodo.org/record/5284962/files/figure.png" pageId="13" pageNumber="48">7 A–C</figureCitation>
|
||
,
|
||
<figureCitation id="91D4DEEEFFE9FFD5FE638D16A3FEE353" box="[375,417,1632,1656]" captionStart="FIGURE 9" captionStartId="12.[151,248,1756,1779]" captionTargetBox="[151,1436,916,1735]" captionTargetId="figure-423@12.[151,1436,916,1735]" captionTargetPageId="12" captionText="FIGURE 9. Details of shell ornamentation of holotypes (penultimate whorl). A. Napaeus arinagaensis n. sp. B. N. validoi n. sp. C. N. grohi n. sp." figureDoi="http://doi.org/10.5281/zenodo.5284970" httpUri="https://zenodo.org/record/5284970/files/figure.png" pageId="13" pageNumber="48">9 C</figureCitation>
|
||
,
|
||
<figureCitation id="91D4DEEEFFE9FFD5FEB98D16A396E353" box="[429,457,1632,1656]" captionStart="FIGURE 10" captionStartId="13.[151,248,1075,1098]" captionTargetBox="[151,1436,414,1054]" captionTargetId="figure-450@13.[151,1436,414,1054]" captionTargetPageId="13" captionText="FIGURE 10. Napaeus grohi n. sp., holotype. A. Genital system. B. Detail of the flagellum. C. Internal anatomy of penis. A4–A5, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix (although this appears be bent it is in fact only wrinkled); e, epiphallus; f, flagellum; go, genital orifice; o, free oviduct; par, penial appendix retractor; p, penis; pr, penis retractor; sph, sphincter between epiphallus and penis; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5284974" httpUri="https://zenodo.org/record/5284974/files/figure.png" pageId="13" pageNumber="48">10</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41F591E0FFE9FFD5FF838DDEA133E207" pageId="13" pageNumber="48" type="materials_examined">
|
||
<paragraph id="0950C26BFFE9FFD5FF838DDEA677E3EB" blockId="13.[151,1437,1703,2019]" box="[151,1064,1703,1728]" pageId="13" pageNumber="48">
|
||
<emphasis id="3B9B1E79FFE9FFD5FF838DDEA36FE3EB" bold="true" box="[151,304,1703,1728]" inLineHeading="true" pageId="13" pageNumber="48" reason="1">Type locality.</emphasis>
|
||
Pista del Derrabado (El Hierro;
|
||
<collectionCode id="6FFE5AAEFFE9FFD5FD8D8DDFA085E3EB" box="[665,730,1705,1728]" pageId="13" pageNumber="48">UTM</collectionCode>
|
||
: 28RDR
|
||
<quantity id="CE176F8EFFE9FFD5FC2D8DDEA19EE3EB" box="[825,961,1704,1728]" metricMagnitude="3" metricUnit="m" metricValue="6.18285" pageId="13" pageNumber="48" unit="m" value="6182.85">6182, 850 m</quantity>
|
||
altitude).
|
||
</paragraph>
|
||
<paragraph id="0950C26BFFE9FFD5FFD18DBAA133E207" blockId="13.[151,1437,1703,2019]" pageId="13" pageNumber="48">
|
||
<materialsCitation id="B987C836FFE9FFD5FFD18DBAA3FDE223" collectingDate="1993-01-05" collectionCode="TFMC" collectorName="Leg. M. Ibanez" pageId="13" pageNumber="48" specimenCode="MT 0419" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="3B9B1E79FFE9FFD5FFD18DBAA366E3CF" bold="true" box="[197,313,1740,1764]" inLineHeading="true" pageId="13" pageNumber="48" reason="1">
|
||
<typeStatus id="D6547CC9FFE9FFD5FFD18DBAA36BE3CF" box="[197,308,1740,1764]" pageId="13" pageNumber="48" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="91D4DEEEFFE9FFD5FE5E8DBAA3FDE3C8" box="[330,418,1740,1764]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="13" pageNumber="48">Fig. 3 J</figureCitation>
|
||
; ethanol-preserved specimen):
|
||
<collectionCode id="6FFE5AAEFFE9FFD5FC108DBAA10FE3CF" box="[772,848,1740,1764]" pageId="13" pageNumber="48">TFMC</collectionCode>
|
||
(
|
||
<specimenCode id="59496A10FFE9FFD5FC768DBAA18EE3CF" box="[866,977,1740,1764]" collectionCode="TFMC" pageId="13" pageNumber="48">MT 0419</specimenCode>
|
||
, shell and dissected soft body).
|
||
<collectorName id="A41AA7BDFFE9FFD5FA578DBAA2BDE223" pageId="13" pageNumber="48">Leg. M. Ibáñez</collectorName>
|
||
,
|
||
<date id="7D51E4ABFFE9FFD5FFFA8D86A3FDE223" box="[238,418,1776,1801]" pageId="13" pageNumber="48" value="1993-01-05">
|
||
<collectingDate id="6D151D43FFE9FFD5FFFA8D86A3FDE223" box="[238,418,1776,1801]" pageId="13" pageNumber="48" value="1993-01-05">5 January 1993</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="B987C836FFE9FFD5FEA48D86A137E207" collectionCode="JSGC" location="Ermita de La Caridad" municipality="Lomo Blanco" pageId="13" pageNumber="48" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="3B9B1E79FFE9FFD5FEA48D86A077E223" bold="true" box="[432,552,1776,1800]" pageId="13" pageNumber="48">
|
||
<typeStatus id="D6547CC9FFE9FFD5FEA48D86A077E223" box="[432,552,1776,1800]" pageId="13" pageNumber="48" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
. 2 shells (
|
||
<collectionCode id="6FFE5AAEFFE9FFD5FDB58D86A0BCE223" box="[673,739,1776,1800]" pageId="13" pageNumber="48">JSGC</collectionCode>
|
||
) collected in 2006 from the
|
||
<location id="0C3094B0FFE9FFD5FB378D86A742E223" LSID="urn:lsid:plazi:treatment:8146737DFFE9FFD6FF838D4BA0F3E0E0:0C3094B0FFE9FFD5FB378D86A742E223" box="[1059,1309,1776,1800]" municipality="Lomo Blanco" name="Ermita de La Caridad" pageId="13" pageNumber="48">Ermita de La Caridad</location>
|
||
and
|
||
<collectingMunicipality id="E9345811FFE9FFD5FA4D8D87A2B8E207" pageId="13" pageNumber="48">Lomo Blanco</collectingMunicipality>
|
||
(northwest of the island); deposited in the
|
||
<collectionCode id="6FFE5AAEFFE9FFD5FDA08C62A0ABE207" box="[692,756,1812,1836]" pageId="13" pageNumber="48">JSGC</collectionCode>
|
||
collection
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41F591E0FFE9FFD5FFD18C4EA344E25F" pageId="13" pageNumber="48" type="etymology">
|
||
<paragraph id="0950C26BFFE9FFD5FFD18C4EA344E25F" blockId="13.[151,1437,1703,2019]" pageId="13" pageNumber="48">
|
||
<emphasis id="3B9B1E79FFE9FFD5FFD18C4EA314E27B" bold="true" box="[197,331,1848,1872]" inLineHeading="true" pageId="13" pageNumber="48" reason="1">Etymology.</emphasis>
|
||
The specific name is dedicated to our friend Klaus Groh, who has spent several decades working in malacology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41F591E0FFE9FFD5FFD18CF4A156E2C8" pageId="13" pageNumber="48" type="distribution">
|
||
<paragraph id="0950C26BFFE9FFD5FFD18CF4A156E2C8" blockId="13.[151,1437,1703,2019]" pageId="13" pageNumber="48">
|
||
<emphasis id="3B9B1E79FFE9FFD5FFD18CF4A016E2B0" box="[197,585,1922,1947]" inLineHeading="true" pageId="13" pageNumber="48" reason="1">
|
||
<emphasis id="3B9B1E79FFE9FFD5FFD18CF4A3B7E2B0" bold="true" box="[197,488,1922,1947]" pageId="13" pageNumber="48">Distribution and habitat</emphasis>
|
||
(
|
||
<figureCitation id="91D4DEEEFFE9FFD5FEE38CF5A067E2B0" box="[503,568,1923,1947]" captionStart="FIGURE 1" captionStartId="3.[151,248,1648,1671]" captionTargetBox="[156,1430,207,1614]" captionTargetId="figure-40@3.[151,1436,195,1624]" captionTargetPageId="3" captionText="FIGURE 1. Geographical distribution of the new species." figureDoi="http://doi.org/10.5281/zenodo.5284942" httpUri="https://zenodo.org/record/5284942/files/figure.png" pageId="13" pageNumber="48">Fig. 1</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Endemic to El Hierro, occupying an area of about
|
||
<quantity id="CE176F8EFFE9FFD5FB918CF5A694E2B0" box="[1157,1227,1923,1947]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="48" unit="km" value="10.0">10 km</quantity>
|
||
<superScript id="FE9A6F23FFE9FFD5FBD88CF6A68BE2A5" attach="left" box="[1228,1236,1920,1934]" fontSize="6" pageId="13" pageNumber="48">2</superScript>
|
||
of the humid laurel forest on the El Golfo northern slopes, within an altitudinal range between 850 and
|
||
<quantity id="CE176F8EFFE9FFD5FB638CD1A68EE295" box="[1143,1233,1959,1983]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="48" unit="m" value="1200.0">1200 m</quantity>
|
||
. This species is a ground-dweller, associated with the humid laurel forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41F591E0FFEAFFD6FFD18BE1A0C4E414" pageId="14" pageNumber="49" type="diagnosis">
|
||
<paragraph id="0950C26BFFEAFFD6FFD18BE1A0C4E414" blockId="14.[151,1437,151,1483]" pageId="14" pageNumber="49">
|
||
<emphasis id="3B9B1E79FFEAFFD6FFD18BE1A363E59B" bold="true" box="[197,316,151,176]" inLineHeading="true" pageId="14" pageNumber="49" reason="1">Diagnosis.</emphasis>
|
||
Shell obese, conic-ovate, with weakly convex whorls increasing regularly in size and shallow but distinct suture. Body whorl large. Aperture with five-sided appearance. Penis globose, without penial papilla but with sphincter between epiphallus and penis. Penial cavity with eight longitudinal, wrinkled folds, four of these thickened and alternating with the other four. Penial retractor muscle inserting on distal end of penis. Bursa copulatrix very large, pear shaped, without diverticulum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="41F591E0FFEAFFD6FFD18A3DA0F3E0E0" pageId="14" pageNumber="49" type="description">
|
||
<paragraph id="0950C26BFFEAFFD6FFD18A3DA387E7E7" blockId="14.[151,1437,151,1483]" pageId="14" pageNumber="49">
|
||
<emphasis id="3B9B1E79FFEAFFD6FFD18A3DA30CE44F" bold="true" box="[197,339,331,356]" inLineHeading="true" pageId="14" pageNumber="49" reason="1">Description.</emphasis>
|
||
Body brown, covered with blackish-brown spots, denser in cephalic area. Shell (
|
||
<figureCitation id="91D4DEEEFFEAFFD6FBCB8A3AA769E44F" box="[1247,1334,332,356]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="14" pageNumber="49">Fig. 3 J</figureCitation>
|
||
) dextral, obese (SB/SH index), conic-ovate, with whorls increasing regularly in size, 6½–7 weakly convex whorls. Body whorl large (BH/SH index), comprising almost ¾ of shell surface area (BS/SS index). Protoconch smooth, shiny, with 1½ whorls. Aperture long and wide (AH/SH and AB/SB indices). Angle (
|
||
<figureCitation id="91D4DEEEFFEAFFD6FCFD8AC1A672E4E4" box="[1001,1069,439,463]" captionStart="FIGURE 2" captionStartId="4.[151,248,908,931]" captionTargetBox="[158,1428,208,867]" captionTargetId="figure-435@4.[151,1436,193,887]" captionTargetPageId="4" captionText="FIGURE 2. Drawings of the shell of the holotype of Napaeus arinagaensis n. sp., showing the placement of the measurements obtained (in º, 2 mm or mm). α, upper palatal angle (i.e. the angle between the columella and the upper palatal margin of the aperture); AB, aperture breadth; AH, aperture height; AP, aperture perimeter; AS, aperture surface area (plane view); BH, body whorl height (at columella level); BP, body whorl perimeter; BS, body whorl surface area (plane view); FB, first whorls breadth; FH, first whorls height; FP, first whorls perimeter; FS, first whorls surface area (plane view); PB, penultimate whorl breadth; PH, penultimate whorl height; PP, penultimate whorl perimeter; PS, penultimate whorl surface area (plane view); SB, shell breadth; SH, shell height; SP, shell perimeter; SS, shell surface area (plane view); T, distance between columella and periphery of penultimate whorl above aperture; U, highest distance between columella and palatal edge of aperture." figureDoi="http://doi.org/10.5281/zenodo.5284944" httpUri="https://zenodo.org/record/5284944/files/figure.png" pageId="14" pageNumber="49">Fig. 2</figureCitation>
|
||
) between columella and start of upper palatal margin about 120º; upper palatal margin transitioning into central part of palatal margin with a rounded, blunt angle (ca. 50º), so that the central part is almost parallel with the columella; (angle between them
|
||
<emphasis id="3B9B1E79FFEAFFD6FA6E8977A7C3E733" box="[1402,1436,513,536]" italics="true" pageId="14" pageNumber="49">ca.</emphasis>
|
||
170º); junction of palatal and columellar edges also slightly angular, resulting in a five-sided appearance of the aperture. Peristome discontinuous, expanded as moderate whitish lip, more developed in lower part of palatal edge and reflected on columellar edge, partly covering umbilical slit.
|
||
<typeStatus id="D6547CC9FFEAFFD6FC65891AA186E7AF" box="[881,985,620,644]" pageId="14" pageNumber="49" type="holotype">Holotype</typeStatus>
|
||
with callosity between peristome edges and small nodule at junction of parietal and palatal margins. Aperture extending laterally beyond penultimate whorl by about 28% (
|
||
<emphasis id="3B9B1E79FFEAFFD6FE5789C3A302E7E7" box="[323,349,693,716]" italics="true" pageId="14" pageNumber="49">ca</emphasis>
|
||
0.95 mm).
|
||
</paragraph>
|
||
<paragraph id="0950C26BFFEAFFD6FFD189A1A199E63F" blockId="14.[151,1437,151,1483]" pageId="14" pageNumber="49">
|
||
Shell colour pale-brown, with first whorls darker than rest. Shell ornamentation almost smooth, characterised by numerous weak, radial oblique ribs (
|
||
<figureCitation id="91D4DEEEFFEAFFD6FD36898AA022E63F" box="[546,637,764,788]" captionStart="FIGURE 9" captionStartId="12.[151,248,1756,1779]" captionTargetBox="[151,1436,916,1735]" captionTargetId="figure-423@12.[151,1436,916,1735]" captionTargetPageId="12" captionText="FIGURE 9. Details of shell ornamentation of holotypes (penultimate whorl). A. Napaeus arinagaensis n. sp. B. N. validoi n. sp. C. N. grohi n. sp." figureDoi="http://doi.org/10.5281/zenodo.5284970" httpUri="https://zenodo.org/record/5284970/files/figure.png" pageId="14" pageNumber="49">Fig. 9 C</figureCitation>
|
||
). Shell with gloss despite ribs.
|
||
</paragraph>
|
||
<paragraph id="0950C26BFFEAFFD6FFD18869A02CE1E4" blockId="14.[151,1437,151,1483]" pageId="14" pageNumber="49">
|
||
Genital system (
|
||
<figureCitation id="91D4DEEEFFEAFFD6FE6F8869A392E61C" box="[379,461,799,823]" captionStart="FIGURE 10" captionStartId="13.[151,248,1075,1098]" captionTargetBox="[151,1436,414,1054]" captionTargetId="figure-450@13.[151,1436,414,1054]" captionTargetPageId="13" captionText="FIGURE 10. Napaeus grohi n. sp., holotype. A. Genital system. B. Detail of the flagellum. C. Internal anatomy of penis. A4–A5, parts of the penial appendix; ag, albumen gland; bc, bursa copulatrix (although this appears be bent it is in fact only wrinkled); e, epiphallus; f, flagellum; go, genital orifice; o, free oviduct; par, penial appendix retractor; p, penis; pr, penis retractor; sph, sphincter between epiphallus and penis; v, vagina; vd, vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5284974" httpUri="https://zenodo.org/record/5284974/files/figure.png" pageId="14" pageNumber="49">Fig. 10</figureCitation>
|
||
; only
|
||
<specimenCount id="1FE909E2FFEAFFD6FD068856A0F5E61C" box="[530,682,799,823]" pageId="14" pageNumber="49" type="generic">one specimen</specimenCount>
|
||
dissected, the
|
||
<typeStatus id="D6547CC9FFEAFFD6FC5B8869A1E9E61C" box="[847,950,799,823]" pageId="14" pageNumber="49" type="holotype">holotype</typeStatus>
|
||
). Atrium very short. Penis globose, without sections, about one-third the length of epiphallus, without penial papilla but with sphincter between epiphallus and penis. Penial cavity with eight longitudinal, wrinkled folds, four of them thickened and alternating with the other four which are narrower. Penial retractor muscle inserting on distal end of penis. Epiphallus tubular, not subdivided into portions, opening distally into penis. Flagellum short. Vas deferens opens laterally into proximal end of epiphallus. Penial appendix arising from distal portion of penis near atrium, slightly shorter than penis and epiphallus combined. Part A
|
||
<subScript id="956BC02EFFEAFFD6FE3A8F71A369E13E" attach="left" box="[302,310,1031,1045]" fontSize="6" pageId="14" pageNumber="49">1</subScript>
|
||
of penial appendix short, tubular, clearly differentiated from globular part A
|
||
<subScript id="956BC02EFFEAFFD6FBB88F71A6EBE13E" attach="left" box="[1196,1204,1031,1045]" fontSize="6" pageId="14" pageNumber="49">2</subScript>
|
||
. Short appendicular retractor muscle inserting laterally in middle zone of part A
|
||
<subScript id="956BC02EFFEAFFD6FC298F58A11AE117" attach="left" box="[829,837,1070,1084]" fontSize="6" pageId="14" pageNumber="49">1</subScript>
|
||
, joined to short penial retractor on diaphragm walls. Part A
|
||
<subScript id="956BC02EFFEAFFD6FFF48F23A2B7E148" attach="left" box="[224,232,1109,1123]" fontSize="6" pageId="14" pageNumber="49">3</subScript>
|
||
slender, shorter than penis. Part A
|
||
<subScript id="956BC02EFFEAFFD6FD7A8F23A029E148" attach="left" box="[622,630,1109,1123]" fontSize="6" pageId="14" pageNumber="49">4</subScript>
|
||
shorter and more slender than A
|
||
<subScript id="956BC02EFFEAFFD6FCFD8F23A1AEE148" attach="left" box="[1001,1009,1109,1123]" fontSize="6" pageId="14" pageNumber="49">3</subScript>
|
||
, passing gradually into expanded and longer A
|
||
<subScript id="956BC02EFFEAFFD6FFEF8F0DA35CE1A2" attach="left" box="[251,259,1147,1161]" fontSize="6" pageId="14" pageNumber="49">5</subScript>
|
||
. Free oviduct three to four times longer than vagina. Vagina firmly attached to body tegument by short connective fibres. Bursa copulatrix complex without diverticulum, vesicle pear shaped, very large, ca. twice as long as wide; bursa duct tubular and very thick.
|
||
</paragraph>
|
||
<paragraph id="0950C26BFFEAFFD6FFD18FACA288E074" blockId="14.[151,1437,151,1483]" pageId="14" pageNumber="49">
|
||
<emphasis id="3B9B1E79FFEAFFD6FFD18FACA041E1D8" bold="true" box="[197,542,1242,1267]" inLineHeading="true" pageId="14" pageNumber="49" reason="1">Comparison with other taxa.</emphasis>
|
||
Shell morphology:
|
||
<taxonomicName id="CEEFB9E8FFEAFFD6FDE38FAAA1C8E1D8" authorityName="Alonso, Ibanez & Santana" authorityYear="2011" box="[759,919,1243,1267]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="grohi">
|
||
<emphasis id="3B9B1E79FFEAFFD6FDE38FAAA1C8E1D8" box="[759,919,1243,1267]" italics="true" pageId="14" pageNumber="49">Napaeus grohi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFEAFFD6FCB38FADA1A2E1D8" box="[935,1021,1243,1267]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="14" pageNumber="49">Fig. 3 J</figureCitation>
|
||
) is comparable in form and size with
|
||
<taxonomicName id="CEEFB9E8FFEAFFD6FF838E76A2B2E03C" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[151,237,1280,1303]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="josei">
|
||
<emphasis id="3B9B1E79FFEAFFD6FF838E76A2B2E03C" box="[151,237,1280,1303]" italics="true" pageId="14" pageNumber="49">N. josei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFEAFFD6FFE88F89A303E03C" box="[252,348,1279,1303]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="14" pageNumber="49">Fig. 3 A</figureCitation>
|
||
) and
|
||
<taxonomicName id="CEEFB9E8FFEAFFD6FE8E8E76A041E03C" baseAuthorityName="Webb & Berthelot" baseAuthorityYear="1833" box="[410,542,1279,1303]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="3B9B1E79FFEAFFD6FE8E8E76A041E03C" box="[410,542,1279,1303]" italics="true" pageId="14" pageNumber="49">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="91D4DEEEFFEAFFD6FD3A8F89A0D2E03C" box="[558,653,1279,1303]" captionStart="FIGURE 3" captionStartId="5.[151,250,1849,1872]" captionTargetBox="[176,1412,193,1835]" captionTargetId="figure-155@5.[176,1412,193,1835]" captionTargetPageId="5" captionText="FIGURE 3. Shells of: A. Napaeus josei n. sp., holotype. B. N. nanodes, from Barranco del Agua, Tenerife (AIT). C. N. badiosus, from Barranco de Valle Seco, Tenerife. D. N. venegueraensis n. sp., holotype. E. N. chrysaloides, from Tamadaba, Gran Canaria (AIT). F. N. validoi n. sp., holotype. G. N. obesatus, from Barranco del Pagador, Gran Canaria (AIT). H. N. bajamarensis, holotype (from Yanes et al. 2009). J. N. grohi n. sp., holotype. K. N. osoriensis, from Osorio, Gran Canaria (AIT). L. N. myosotis, from Montaña de Guía, Gran Canaria (AIT). M. N. exilis, from Altos de Gáldar, Gran Canaria (AIT). N. N. arinagaensis n. sp., holotype. O. N. moquinianus, from Brezal del Palmital, Gran Canaria (AIT). P. N. isletae, holotype." figureDoi="http://doi.org/10.5281/zenodo.5284948" httpUri="https://zenodo.org/record/5284948/files/figure.png" pageId="14" pageNumber="49">Fig. 3 C</figureCitation>
|
||
), its whorls being flatter and the ornamentation weaker than in both of those species. In
|
||
<taxonomicName id="CEEFB9E8FFEAFFD6FE5B8E52A3FBE010" authorityName="Yanes & Santana & Artiles & Deniz & Martín & Alonso & Ibáñez" authorityYear="2011" box="[335,420,1316,1339]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="josei">
|
||
<emphasis id="3B9B1E79FFEAFFD6FE5B8E52A3FBE010" box="[335,420,1316,1339]" italics="true" pageId="14" pageNumber="49">N. josei</emphasis>
|
||
</taxonomicName>
|
||
the shell is broader and in
|
||
<taxonomicName id="CEEFB9E8FFEAFFD6FDDA8E52A10DE010" baseAuthorityName="Webb & Berthelot" baseAuthorityYear="1833" box="[718,850,1315,1339]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="badiosus">
|
||
<emphasis id="3B9B1E79FFEAFFD6FDDA8E52A10DE010" box="[718,850,1315,1339]" italics="true" pageId="14" pageNumber="49">N. badiosus</emphasis>
|
||
</taxonomicName>
|
||
it is more slender (excluding the aperture) than in
|
||
<taxonomicName id="CEEFB9E8FFEAFFD6FA978E52A28EE074" authorityName="Alonso, Ibanez & Santana" authorityYear="2011" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="grohi">
|
||
<emphasis id="3B9B1E79FFEAFFD6FA978E52A28EE074" italics="true" pageId="14" pageNumber="49">N. grohi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="0950C26BFFEAFFD6FFD18E1DA0F3E0E0" blockId="14.[151,1437,151,1483]" pageId="14" pageNumber="49">
|
||
Genital anatomy:
|
||
<taxonomicName id="CEEFB9E8FFEAFFD6FE928E1AA380E0A8" authorityName="Alonso, Ibanez & Santana" authorityYear="2011" box="[390,479,1387,1411]" class="Gastropoda" family="Enidae" genus="Napaeus" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="49" phylum="Mollusca" rank="species" species="grohi">
|
||
<emphasis id="3B9B1E79FFEAFFD6FE928E1AA380E0A8" box="[390,479,1387,1411]" italics="true" pageId="14" pageNumber="49">N. grohi</emphasis>
|
||
</taxonomicName>
|
||
differs from all species of the genus for which the anatomy is known by the insertion of the penial retractor muscle on the distal end of the penis, near the genital orifice and by the very large, pear shaped vesicle of the bursa copulatrix complex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |