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<document id="DEA1F7492ED1E07D47AAB302B6E8AFFD" ID-CLB-Dataset="25544" ID-DOI="10.11646/zootaxa.4647.1.31" ID-GBIF-Dataset="2a05f92b-5125-41c8-ba0d-df4afd5c8e3d" ID-ISSN="1175-5326" ID-Zenodo-Dep="3353347" ID-ZooBank="D00376D7-BE82-45C9-ACE6-A8789E3F9B9B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564251352740" checkinUser="plazi" docAuthor="Mašán, Peter &amp; Babaeian, Esmaeil" docDate="2019" docId="63478799FFD3FFDF04F5EA91FE58FA27" docLanguage="en" docName="zootaxa.4647.1.31.pdf" docOrigin="Zootaxa 4647 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cosmolaelaps ekaterinae Mašán &amp; Babaeian 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="501" masterDocId="9F7EFFE1FFD2FFD90462EE3CFF91FFC0" masterDocTitle="A new myrmecophilous mite species of the genus Cosmolaelaps Berlese, 1903 (Acari, Mesostigmata, Laelapidae) from Central Europe (Slovakia)" masterLastPageNumber="505" masterPageNumber="495" pageNumber="496" updateTime="1698747683202" updateUser="ExternalLinkService">
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<mods:title id="E4D9B69E0F55A551FEB00F8A5569286C">A new myrmecophilous mite species of the genus Cosmolaelaps Berlese, 1903 (Acari, Mesostigmata, Laelapidae) from Central Europe (Slovakia)</mods:title>
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<mods:name id="A84A24F094738AA5A1E2BE84709086F9" type="personal">
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<mods:namePart id="7F4B67F9876FE4CAFE907D5258139B89">Mašán, Peter</mods:namePart>
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<mods:namePart id="A67BC673077A9ABA359705139CA94729">Babaeian, Esmaeil</mods:namePart>
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<mods:title id="CF21F1424F61AE82980BAD543098C013">Zootaxa</mods:title>
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<mods:date id="4E189D2A10BF6B819136DD6532789EC9">2019</mods:date>
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<mods:number id="09D681A45E5CBB4E9A19C45D1B420B8D">2019-07-26</mods:number>
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<treatment id="63478799FFD3FFDF04F5EA91FE58FA27" ID-GBIF-Taxon="159398417" LSID="urn:lsid:plazi:treatment:63478799FFD3FFDF04F5EA91FE58FA27" httpUri="http://treatment.plazi.org/id/63478799FFD3FFDF04F5EA91FE58FA27" lastPageId="6" lastPageNumber="501" pageId="1" pageNumber="496">
<subSubSection id="A3F46504FFD3FFD804F5EA91FE88FB2B" pageId="1" pageNumber="496" type="nomenclature">
<paragraph id="EB51368FFFD3FFD804F5EA91FDBBFB07" blockId="1.[151,554,1197,1259]" box="[151,554,1197,1224]" pageId="1" pageNumber="496">
<heading id="B01981E3FFD3FFD804F5EA91FDBBFB07" bold="true" box="[151,554,1197,1224]" fontSize="11" level="1" pageId="1" pageNumber="496" reason="1">
<emphasis id="D99AEA9DFFD3FFD804F5EA91FDBBFB07" bold="true" box="[151,554,1197,1224]" pageId="1" pageNumber="496">
<taxonomicName id="2CEE4D0CFFD3FFD804F5EA91FE54FB08" authority="Mašán &amp; Babaeian, 2019" authorityName="Mašán &amp; Babaeian" authorityYear="2019" box="[151,453,1197,1224]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="496" phylum="Arthropoda" rank="species" species="ekaterinae" status="sp. nov.">
<emphasis id="D99AEA9DFFD3FFD804F5EA91FE54FB08" bold="true" box="[151,453,1197,1224]" italics="true" pageId="1" pageNumber="496">Cosmolaelaps ekaterinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C2A957E6FFD3FFD805AFEA91FDBBFB07" box="[461,554,1197,1223]" pageId="1" pageNumber="496" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="EB51368FFFD3FFD804F5EAEDFE88FB2B" blockId="1.[151,554,1197,1259]" box="[151,281,1233,1259]" pageId="1" pageNumber="496">
(
<figureCitation id="73D52A0AFFD3FFD804FDEAEDFE81FB2B" box="[159,272,1233,1259]" captionStart-0="FIGURES 17" captionStart-1="FIGURES 811" captionStart-2="FIGURES 1218" captionStartId-0="2.[151,275,1862,1889]" captionStartId-1="3.[151,275,1735,1762]" captionStartId-2="5.[151,275,1888,1915]" captionTargetBox-0="[151,1435,183,1836]" captionTargetBox-1="[162,1431,195,1696]" captionTargetBox-2="[151,1436,179,1862]" captionTargetId-0="figure@2.[151,1436,179,1836]" captionTargetId-1="figure@3.[151,1436,179,1709]" captionTargetId-2="figure@5.[151,1436,179,1862]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." captionText-1="FIGURES 811. Cosmolaelaps ekaterinae sp. nov., female, legs IIV. 8. Leg I; 9. Leg II; 10. Leg III; 11. Leg IV." captionText-2="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi-0="http://doi.org/10.5281/zenodo.3416846" figureDoi-1="http://doi.org/10.5281/zenodo.3416848" figureDoi-2="http://doi.org/10.5281/zenodo.3416850" httpUri-0="https://zenodo.org/record/3416846/files/figure.png" httpUri-1="https://zenodo.org/record/3416848/files/figure.png" httpUri-2="https://zenodo.org/record/3416850/files/figure.png" pageId="1" pageNumber="496">Figs 118</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A3F46504FFD3FFDF04F5EB26FE58FA27" lastPageId="6" lastPageNumber="501" pageId="1" pageNumber="496" type="materials_examined">
<paragraph id="EB51368FFFD3FFD804F5EB26FE32FAF3" blockId="1.[151,1437,1305,1512]" box="[151,419,1305,1331]" pageId="1" pageNumber="496">
<emphasis id="D99AEA9DFFD3FFD804F5EB26FE32FAF3" box="[151,419,1305,1331]" italics="true" pageId="1" pageNumber="496">Type material examined</emphasis>
</paragraph>
<paragraph id="EB51368FFFD3FFD804F5EB01FC7EFA27" blockId="1.[151,1437,1305,1512]" pageId="1" pageNumber="496">
<materialsCitation id="5B863CD2FFD3FFD804F5EB01FB9AFA5F" ID-GBIF-Occurrence="2413160320" collectingDate="2018-10-24" country="Slovakia" elevation="190" location="Chlaba Village" municipality="Burda Mountains" pageId="1" pageNumber="496" specimenCount="2" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="3455882DFFD3FFD804F5EB01FF6EFA97" box="[151,255,1341,1367]" pageId="1" pageNumber="496" type="holotype">Holotype</typeStatus>
<specimenCount id="FDE8FD06FFD3FFD8056AEB01FEC8FA97" box="[264,345,1341,1367]" pageId="1" pageNumber="496" type="female">female</specimenCount>
: SW
<collectingCountry id="93F9761FFFD3FFD805F4EB01FE6AFA97" box="[406,507,1341,1367]" name="Slovakia" pageId="1" pageNumber="496">Slovakia</collectingCountry>
,
<collectingMunicipality id="0B35ACF5FFD3FFD80665EB01FD5EFA97" box="[519,719,1341,1367]" pageId="1" pageNumber="496">Burda Mountains</collectingMunicipality>
,
<location id="EE316054FFD3FFD806B9EB01FC1BFA97" LSID="urn:lsid:plazi:treatment:63478799FFD3FFDF04F5EA91FE58FA27:EE316054FFD3FFD806B9EB01FC1BFA97" box="[731,906,1341,1367]" country="Slovakia" municipality="Burda Mountains" name="Chlaba Village" pageId="1" pageNumber="496">Chľaba Village</location>
,
<location id="EE316054FFD3FFD807F4EB01FBEFFA97" LSID="urn:lsid:plazi:treatment:63478799FFD3FFDF04F5EA91FE58FA27:EE316054FFD3FFD807F4EB01FBEFFA97" box="[918,1150,1341,1367]" country="Slovakia" municipality="Burda Mountains" name="Kovacov Settlement" pageId="1" pageNumber="496">Kováčov Settlement</location>
, edge of broadleaved de- ciduous forest (
<emphasis id="D99AEA9DFFD3FFD80529EB5DFDD0FABB" box="[331,577,1377,1403]" italics="true" pageId="1" pageNumber="496">
<location id="EE316054FFD3FFD80529EB5DFDD0FABB" LSID="urn:lsid:plazi:treatment:63478799FFD3FFDF04F5EA91FE58FA27:EE316054FFD3FFD80529EB5DFDD0FABB" box="[331,577,1377,1403]" country="Slovakia" municipality="Burda Mountains" name="Carpineto-Quercetum" pageId="1" pageNumber="496">Carpineto-Quercetum</location>
</emphasis>
) and meadow, subterraneous nest of
<taxonomicName id="2CEE4D0CFFD3FFD80793EB5EFB3CFABB" baseAuthorityName="Nylander" baseAuthorityYear="1846" box="[1009,1197,1377,1403]" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="496" phylum="Arthropoda" rank="species" species="umbratus">
<emphasis id="D99AEA9DFFD3FFD80793EB5EFB3CFABB" box="[1009,1197,1377,1403]" italics="true" pageId="1" pageNumber="496">
<location id="EE316054FFD3FFD80793EB5EFBABFABB" LSID="urn:lsid:plazi:treatment:63478799FFD3FFDF04F5EA91FE58FA27:EE316054FFD3FFD80793EB5EFBABFABB" box="[1009,1082,1378,1403]" country="Slovakia" municipality="Burda Mountains" name="Lasius" pageId="1" pageNumber="496">Lasius</location>
umbratus
</emphasis>
</taxonomicName>
colonized by
<taxonomicName id="2CEE4D0CFFD3FFD80131EB5EFE83FA5F" baseAuthorityName="Latreille" baseAuthorityYear="1798" class="Insecta" family="Formicidae" genus="Lasius" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="496" phylum="Arthropoda" rank="species" species="fuliginosus">
<emphasis id="D99AEA9DFFD3FFD80131EB5EFE83FA5F" italics="true" pageId="1" pageNumber="496">
<location id="EE316054FFD3FFD80131EB5EFA0DFABB" LSID="urn:lsid:plazi:treatment:63478799FFD3FFDF04F5EA91FE58FA27:EE316054FFD3FFD80131EB5EFA0DFABB" box="[1363,1436,1378,1403]" country="Slovakia" municipality="Burda Mountains" name="Lasius" pageId="1" pageNumber="496">Lasius</location>
fuliginosus
</emphasis>
</taxonomicName>
in rotting tree stump (
<taxonomicName id="2CEE4D0CFFD3FFD80670EBB9FDE1FA5F" box="[530,624,1413,1439]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="496" phylum="Tracheophyta" rank="genus">
<emphasis id="D99AEA9DFFD3FFD80670EBB9FDE1FA5F" box="[530,624,1413,1439]" italics="true" pageId="1" pageNumber="496">
<location id="EE316054FFD3FFD80670EBB9FDE1FA5F" LSID="urn:lsid:plazi:treatment:63478799FFD3FFDF04F5EA91FE58FA27:EE316054FFD3FFD80670EBB9FDE1FA5F" box="[530,624,1413,1439]" country="Slovakia" municipality="Burda Mountains" name="Quercus" pageId="1" pageNumber="496">Quercus</location>
</emphasis>
</taxonomicName>
<specimenCount id="FDE8FD06FFD3FFD8061AEBB9FD0BFA5F" box="[632,666,1413,1439]" pageId="1" pageNumber="496" type="generic">sp.</specimenCount>
),
<quantity id="2C169B6AFFD3FFD806D2EBB9FD69FA5F" box="[688,760,1413,1439]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="496" unit="m" value="190.0">
<elevation id="60C3D1BCFFD3FFD806D2EBB9FD69FA5F" box="[688,760,1413,1439]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="496" unit="m" value="190.0">190 m</elevation>
</quantity>
a.s.l.,
<date id="9F50104FFFD3FFD80721EBB9FB9AFA5F" box="[835,1035,1413,1439]" pageId="1" pageNumber="496" value="2018-10-24">
<collectingDate id="8F14E9A7FFD3FFD80721EBB9FB9AFA5F" box="[835,1035,1413,1439]" pageId="1" pageNumber="496" value="2018-10-24">October 24, 2018</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="5B863CD2FFD3FFD8007AEBB9FDE4FA03" ID-GBIF-Occurrence="2413160273" collectingDate="2018-10-24" country="Slovakia" elevation="190" location="Chlaba Village" municipality="Burda Mountains" pageId="1" pageNumber="496" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="3455882DFFD3FFD8007AEBB9FB12FA5F" box="[1048,1155,1413,1439]" pageId="1" pageNumber="496" type="paratype">paratypes</typeStatus>
(eight
<specimenCount id="FDE8FD06FFD3FFD800B7EBB9FABDFA5F" box="[1237,1324,1413,1439]" pageId="1" pageNumber="496" type="female">females</specimenCount>
and three
<specimenCount id="FDE8FD06FFD3FFD804F5EB95FF4FFA04" box="[151,222,1449,1476]" pageId="1" pageNumber="496" type="male">males</specimenCount>
): with the same data as for holotype
</materialsCitation>
.
<materialsCitation id="5B863CD2FFD3FFD8061DEB95FC7EFA27" ID-GBIF-Occurrence="2413160506" collectionCode="IZSAS, JAZM" pageId="1" pageNumber="496" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="holotype">
The
<typeStatus id="3455882DFFD3FFD806D3EB95FC82FA03" box="[689,787,1449,1475]" pageId="1" pageNumber="496" type="holotype">holotype</typeStatus>
and seven
<typeStatus id="3455882DFFD3FFD807F3EB95FC6DFA03" box="[913,1020,1449,1475]" pageId="1" pageNumber="496" type="paratype">paratypes</typeStatus>
(five
<specimenCount id="FDE8FD06FFD3FFD8005DEB95FB07FA03" box="[1087,1174,1449,1475]" pageId="1" pageNumber="496" type="female">females</specimenCount>
and two
<specimenCount id="FDE8FD06FFD3FFD8009CEB95FAD0FA04" box="[1278,1345,1449,1476]" pageId="1" pageNumber="496" type="male">males</specimenCount>
) are deposited in
<collectionCode id="8DFFAE4AFFD3FFD8056EEBF1FECCFA27" box="[268,349,1485,1511]" pageId="1" pageNumber="496">IZSAS</collectionCode>
; four
<typeStatus id="3455882DFFD3FFD805FCEBF1FD98FA27" box="[414,521,1485,1511]" pageId="1" pageNumber="496" type="paratype">paratypes</typeStatus>
(three
<specimenCount id="FDE8FD06FFD3FFD8063BEBF1FD21FA27" box="[601,688,1485,1511]" pageId="1" pageNumber="496" type="female">females</specimenCount>
and one
<specimenCount id="FDE8FD06FFD3FFD80774EBF1FCDFFA28" box="[790,846,1485,1512]" pageId="1" pageNumber="496" type="male">male</specimenCount>
) are in
<collectionCode id="8DFFAE4AFFD3FFD807C0EBF1FC7EFA27" box="[930,1007,1485,1511]" pageId="1" pageNumber="496">JAZM</collectionCode>
</materialsCitation>
</paragraph>
<paragraph id="EB51368FFFD3FFD8078DEBF1FC62FA27" blockId="1.[151,1437,1305,1512]" box="[1007,1011,1485,1511]" pageId="1" pageNumber="496">.</paragraph>
<paragraph id="EB51368FFFD3FFD804F5E806FEFBF993" blockId="1.[151,1437,1593,1979]" box="[151,362,1593,1619]" pageId="1" pageNumber="496">
<emphasis id="D99AEA9DFFD3FFD804F5E806FE98F993" box="[151,265,1594,1619]" italics="true" pageId="1" pageNumber="496">Diagnosis</emphasis>
(
<emphasis id="D99AEA9DFFD3FFD8057BE805FEF0F993" box="[281,353,1593,1619]" italics="true" pageId="1" pageNumber="496">Adults</emphasis>
)
</paragraph>
<paragraph id="EB51368FFFD3FFD804F5E861FB18F87B" blockId="1.[151,1437,1593,1979]" pageId="1" pageNumber="496">
Idiosoma small and less than 400 μm in length. Dorsal shield surface essentially smooth, having 39 pairs of setae plus 23 unpaired setae (Jx) on posteromedian area. Dorsal shield setae (except j1, z1 and J5) with well-developed basal protuberance, narrow, markedly attenuate distally and conspicuously elongate so their tips reaching beyond the insertions of the two following setae. In female, presternal platelets free, weakly sclerotized; sternal shield smooth on whole surface, almost unsclerotized in front portion so its anterior margin not distinctly defined; epigynal shield tongue-shaped, normally with one Λ-like and two or three V-like transverse sculptural lines in its posterior portion; setae ad1 on femora I and
<collectionCode id="8DFFAE4AFFD3FFD8067EE909FDBEF88F" box="[540,559,1845,1871]" pageId="1" pageNumber="496">II</collectionCode>
and ad2 on femur
<collectionCode id="8DFFAE4AFFD3FFD80763E909FC8FF88F" box="[769,798,1845,1871]" pageId="1" pageNumber="496">IV</collectionCode>
thickened; pd2 on femur
<collectionCode id="8DFFAE4AFFD3FFD8005FE909FBC1F88F" box="[1085,1104,1845,1871]" pageId="1" pageNumber="496">II</collectionCode>
and ad1 on femora
<collectionCode id="8DFFAE4AFFD3FFD8014FE909FAD8F88F" box="[1325,1353,1845,1871]" pageId="1" pageNumber="496">III</collectionCode>
and
<collectionCode id="8DFFAE4AFFD3FFD8011DE909FA0DF88F" box="[1407,1436,1845,1871]" pageId="1" pageNumber="496">IV</collectionCode>
thickened and spatulate. In male, holoventral shield present; cheliceral spermatodactyl well developed, about twice as long as fixed digit; femur
<collectionCode id="8DFFAE4AFFD3FFD805B4E941FE78F857" box="[470,489,1917,1943]" pageId="1" pageNumber="496">II</collectionCode>
and telotarsus
<collectionCode id="8DFFAE4AFFD3FFD806F2E941FD3CF857" box="[656,685,1917,1943]" pageId="1" pageNumber="496">IV</collectionCode>
bearing a ventral spur. Peritrematal shields terminally slightly widened and truncate. Anterior margin of epistome convex, regularly and densely denticulate.
</paragraph>
<caption id="BF916607FFD0FFDB04F5E97AFCE6F868" ID-DOI="http://doi.org/10.5281/zenodo.3416846" ID-Zenodo-Dep="3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="2" pageNumber="497" startId="2.[151,275,1862,1889]" targetBox="[151,1435,183,1836]" targetPageId="2">
<paragraph id="EB51368FFFD0FFDB04F5E97AFCE6F868" blockId="2.[151,1436,1862,1961]" pageId="2" pageNumber="497">
<emphasis id="D99AEA9DFFD0FFDB04F5E97AFEDDF8A0" bold="true" box="[151,332,1862,1889]" pageId="2" pageNumber="497">FIGURES 17.</emphasis>
<taxonomicName id="2CEE4D0CFFD0FFDB0531E97AFDFFF8A0" authority="Mašán &amp; Babaeian, 2019" authorityName="Mašán &amp; Babaeian" authorityYear="2019" box="[339,622,1862,1888]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="497" phylum="Arthropoda" rank="species" species="ekaterinae" status="sp. nov.">
<emphasis id="D99AEA9DFFD0FFDB0531E97AFDFFF8A0" box="[339,622,1862,1888]" italics="true" pageId="2" pageNumber="497">Cosmolaelaps ekaterinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C2A957E6FFD0FFDB0614E97BFD5EF8A1" box="[630,719,1863,1889]" pageId="2" pageNumber="497" rank="species">sp. nov.</taxonomicNameLabel>
, female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view.
</paragraph>
</caption>
<caption id="BF916607FFD1FFDA04F5E8FBFA1EF922" ID-DOI="http://doi.org/10.5281/zenodo.3416848" ID-Zenodo-Dep="3416848" box="[151,1423,1735,1762]" httpUri="https://zenodo.org/record/3416848/files/figure.png" pageId="3" pageNumber="498" startId="3.[151,275,1735,1762]" targetBox="[162,1431,195,1696]" targetPageId="3">
<paragraph id="EB51368FFFD1FFDA04F5E8FBFA1EF922" blockId="3.[151,1423,1735,1762]" box="[151,1423,1735,1762]" pageId="3" pageNumber="498">
<emphasis id="D99AEA9DFFD1FFDA04F5E8FBFEC9F921" bold="true" box="[151,344,1735,1762]" pageId="3" pageNumber="498">FIGURES 811.</emphasis>
<taxonomicName id="2CEE4D0CFFD1FFDA053DE8F4FDEAF922" authority="Mašán &amp; Babaeian, 2019" authorityName="Mašán &amp; Babaeian" authorityYear="2019" box="[351,635,1736,1762]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="498" phylum="Arthropoda" rank="species" species="ekaterinae" status="sp. nov.">
<emphasis id="D99AEA9DFFD1FFDA053DE8F4FDEAF922" box="[351,635,1736,1762]" italics="true" pageId="3" pageNumber="498">Cosmolaelaps ekaterinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C2A957E6FFD1FFDA06E0E8F4FD4AF922" box="[642,731,1736,1762]" pageId="3" pageNumber="498" rank="species">sp. nov.</taxonomicNameLabel>
, female, legs IIV. 8. Leg I; 9. Leg
<collectionCode id="8DFFAE4AFFD1FFDA000BE8F4FBEEF922" box="[1129,1151,1736,1762]" pageId="3" pageNumber="498">II</collectionCode>
; 10. Leg
<collectionCode id="8DFFAE4AFFD1FFDA0085E8F4FA97F922" box="[1255,1286,1736,1762]" pageId="3" pageNumber="498">III</collectionCode>
; 11. Leg
<collectionCode id="8DFFAE4AFFD1FFDA010FE8F4FA1AF922" box="[1389,1419,1736,1762]" pageId="3" pageNumber="498">IV</collectionCode>
.
</paragraph>
</caption>
<paragraph id="EB51368FFFD1FFDA04F5E908FEEDF88E" blockId="3.[151,1436,1844,2014]" box="[151,380,1844,1870]" pageId="3" pageNumber="498">
<emphasis id="D99AEA9DFFD1FFDA04F5E908FE8AF88D" box="[151,283,1844,1869]" italics="true" pageId="3" pageNumber="498">Description</emphasis>
(
<emphasis id="D99AEA9DFFD1FFDA054EE908FEE5F88E" box="[300,372,1844,1870]" italics="true" pageId="3" pageNumber="498">Adults</emphasis>
)
</paragraph>
<paragraph id="EB51368FFFD1FFDD04F5E964FE93FE83" blockId="3.[151,1436,1844,2014]" lastBlockId="4.[151,1437,153,1871]" lastPageId="4" lastPageNumber="499" pageId="3" pageNumber="498">
<emphasis id="D99AEA9DFFD1FFDA04F5E964FF64F8B2" bold="true" box="[151,245,1880,1906]" pageId="3" pageNumber="498">Female.</emphasis>
<emphasis id="D99AEA9DFFD1FFDA049CE964FE2DF8B2" box="[254,444,1880,1906]" italics="true" pageId="3" pageNumber="498">Dorsal idiosoma</emphasis>
(
<figureCitation id="73D52A0AFFD1FFDA05AFE964FD84F8B2" box="[461,533,1880,1906]" captionStart="FIGURES 17" captionStartId="2.[151,275,1862,1889]" captionTargetBox="[151,1435,183,1836]" captionTargetId="figure@2.[151,1436,179,1836]" captionText="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="3" pageNumber="498">Figs 1</figureCitation>
,
<figureCitation id="73D52A0AFFD1FFDA0647E964FDD5F8B2" box="[549,580,1880,1906]" captionStart="FIGURES 1218" captionStartId="5.[151,275,1888,1915]" captionTargetBox="[151,1436,179,1862]" captionTargetId="figure@5.[151,1436,179,1862]" captionText="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="3" pageNumber="498">12</figureCitation>
). Idiosoma oblong, ovoid, 375395 μm long and 230245 μm wide (n=5). Dorsal shield entire, almost completely covering dorsum, smooth except for a few lineae close to anterolateral margins, 370390 μm long, and 225240 μm wide nearly at level of setae z5, bearing 39 pairs of setae (j1j6, z1z6, s1s6, r2r5 on podonotal region; J1J5, Z1Z5, S1S5, Px2, Px3 and two or rarely three unpaired Jx setae on opisthonotal region). Almost all dorsal shield setae similar in length, modified, scimitar-shaped, having small basal protuberance and attenuate and elongate distal section (
<figureCitation id="73D52A0AFFD6FFDD0726EE81FC18FF17" box="[836,905,189,215]" captionStart="FIGURES 17" captionStartId="2.[151,275,1862,1889]" captionTargetBox="[151,1435,183,1836]" captionTargetId="figure@2.[151,1436,179,1836]" captionText="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="4" pageNumber="499">Fig. 2</figureCitation>
); only vertical (j1), paravertical (z1) and clunal setae (Z5) unmodified, simple, smooth, needle-like and markedly shorter than other setae on the shield; z1 shortest (2125 μm), j1 (3137 μm) similar to or slightly shorter than Z5 (3343 μm). Most dorsal shield setae 90105 μm in length.
</paragraph>
<paragraph id="EB51368FFFD6FFDD04A5EF71FAF5FBBF" blockId="4.[151,1437,153,1871]" pageId="4" pageNumber="499">
<emphasis id="D99AEA9DFFD6FFDD04A5EF71FE19FEA7" box="[199,392,333,359]" italics="true" pageId="4" pageNumber="499">Ventral idiosoma</emphasis>
(
<figureCitation id="73D52A0AFFD6FFDD05FBEF71FE4EFEA7" box="[409,479,333,359]" captionStart="FIGURES 17" captionStartId="2.[151,275,1862,1889]" captionTargetBox="[151,1435,183,1836]" captionTargetId="figure@2.[151,1436,179,1836]" captionText="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="4" pageNumber="499">Fig. 3</figureCitation>
). Tritosternum with long columnar base and two long and distinctly pilose laciniae. Presternal region and anterior part of sternal shield weakly sclerotized and defined, with a pair of small and indistinctive presternal platelets. Sternal shield subquadrate, 105114 μm long, 8593 μm wide at level of setae st2, 7983 μm wide at level of constriction between coxae
<collectionCode id="8DFFAE4AFFD6FFDD06C4EF85FD2AFE13" box="[678,699,441,467]" pageId="4" pageNumber="499">II</collectionCode>
, and smooth throughout except for a few lineae on lateral margins; lateral margins well sclerotized, posterior margin usually straight, sometimes slightly convex or concave; shield with two pairs of distinct lyrifissures (
<date id="9F50104FFFD6FFDD0628EC3DFDE1FDDB" box="[586,624,513,539]" pageId="4" pageNumber="499">iv1</date>
,
<date id="9F50104FFFD6FFDD061EEC3DFD30FDDC" box="[636,673,513,540]" pageId="4" pageNumber="499">iv2</date>
) and three pairs of subequal setae (st1st
<date id="9F50104FFFD6FFDD001AEC3DFB43FDDB" box="[1144,1234,513,539]" pageId="4" pageNumber="499" value="1929-03-24">3 2429</date>
μm). Metasternal setae (st
<date id="9F50104FFFD6FFDD0490EC19FEDEFDFF" box="[242,335,548,575]" pageId="4" pageNumber="499" value="1928-04-25">4 2528</date>
μm) and associated lyrifissures (
<date id="9F50104FFFD6FFDD06A1EC19FD78FD80" box="[707,745,549,576]" pageId="4" pageNumber="499">iv3</date>
) placed on soft integument,
<date id="9F50104FFFD6FFDD0049EC19FBDEFDFF" box="[1067,1103,549,575]" pageId="4" pageNumber="499">iv3</date>
rarely on posterior margin of sternal shield (on one side only). Epigynal shield oblong, well elongate, slightly wider in anterior portion, tongueshaped, 185195 μm long, 6776 μm wide at level of genital setae (st
<date id="9F50104FFFD6FFDD07C1EC50FC6FFD47" box="[931,1022,620,647]" pageId="4" pageNumber="499" value="1926-05-24">5 2426</date>
μm), hyaline anteriorly (hyaline part approximately reaching level of
<date id="9F50104FFFD6FFDD0665ECADFDBFFD6C" box="[519,558,657,684]" pageId="4" pageNumber="499">iv2</date>
), regularly rounded posteriorly, bearing one pair of setae (but asymmetrically duplicate in illustrated specimen) and with typical pattern of transverse lines as in
<figureCitation id="73D52A0AFFD6FFDD004BEC89FBFFFD0F" box="[1065,1134,693,719]" captionStart="FIGURES 17" captionStartId="2.[151,275,1862,1889]" captionTargetBox="[151,1435,183,1836]" captionTargetId="figure@2.[151,1436,179,1836]" captionText="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="4" pageNumber="499">Figs 4</figureCitation>
and
<figureCitation id="73D52A0AFFD6FFDD00C1EC89FB53FD0F" box="[1187,1218,693,719]" captionStart="FIGURES 1218" captionStartId="5.[151,275,1888,1915]" captionTargetBox="[151,1436,179,1862]" captionTargetId="figure@5.[151,1436,179,1862]" captionText="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="4" pageNumber="499">14</figureCitation>
; genital lyrifissures (
<date id="9F50104FFFD6FFDD04C2ECE4FF54FD33" box="[160,197,728,755]" pageId="4" pageNumber="499">iv5</date>
) situated on soft integument behind st5. Peritremes normal, with anterior ends reaching slightly beyond dorsal setae s1; peritrematal shields narrow, with short post-stigmatic sections terminally slightly widened and truncate. Soft integument between setae ZV1 and oval metapodal platelets with two pairs of small, narrow, irregular sclerites; anterior pair transversely oriented. A slot-like adgenital sclerite close to each lateral margin of epigynal shield present (one or both of them often partly or completely fused to the shield). Anal shield subtriangular, widely rounded anteriorly, similar in length and width (5862 μm long and wide), smooth or with short transverse lines in anterior part, bearing three circum-anal setae, and a pair of marginal gland pores (gv3) more or less aligned with posterior margin of anal opening; postanal seta slightly thickened, moderately longer than adanals (pa 2832 μm, ad 1720 μm); anus with submedial position on the shield. Soft lateral and opisthogastrict integument bearing 1719 pairs of setae. Opisthogastric setae JV1, JV2, JV3, ZV1 and ZV2 simple, needle-like and shorter when compared with other scimitar-shaped setae on opisthogaster; setae in JV and ZV rows becoming progressively longer posteriorly (JV
<date id="9F50104FFFD6FFDD01F1EA7DFF4CFBBF" pageId="4" pageNumber="499" value="1928-01-24">1 2428</date>
μm, JV
<date id="9F50104FFFD6FFDD0555EA59FE00FBBF" box="[311,401,1125,1151]" pageId="4" pageNumber="499" value="1933-02-26">2 2633</date>
μm, JV3 3944 μm, JV4 5563 μm; ZV
<date id="9F50104FFFD6FFDD073CEA59FC25FBBF" box="[862,948,1125,1151]" pageId="4" pageNumber="499" value="1926-01-23">1 2326</date>
μm, ZV2 3438 μm, ZV3 5159 μm).
</paragraph>
<paragraph id="EB51368FFFD6FFDD04A5EAB5FF4BFB2B" blockId="4.[151,1437,153,1871]" pageId="4" pageNumber="499">
<emphasis id="D99AEA9DFFD6FFDD04A5EAB5FE5EFB63" box="[199,463,1161,1187]" italics="true" pageId="4" pageNumber="499">Sperm induction system</emphasis>
(
<figureCitation id="73D52A0AFFD6FFDD05BFEAB5FDB0FB63" box="[477,545,1161,1187]" captionStart="FIGURES 17" captionStartId="2.[151,275,1862,1889]" captionTargetBox="[151,1435,183,1836]" captionTargetId="figure@2.[151,1436,179,1836]" captionText="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="4" pageNumber="499">Fig. 3</figureCitation>
). All parts including infundibula, tubuli and sacculus foemineus well sclerotized and distinct. Tubuli long, moderately widened at solenostomes, each with a poculiform distal end entering the sacculus.
</paragraph>
<paragraph id="EB51368FFFD6FFDD04A5EAC9FAE0FA27" blockId="4.[151,1437,153,1871]" pageId="4" pageNumber="499">
<emphasis id="D99AEA9DFFD6FFDD04A5EAC9FE44FACF" box="[199,469,1269,1295]" italics="true" pageId="4" pageNumber="499">Gnathosomal structures</emphasis>
(
<figureCitation id="73D52A0AFFD6FFDD0587EAC9FDDAFACF" box="[485,587,1268,1295]" captionStart="FIGURES 17" captionStartId="2.[151,275,1862,1889]" captionTargetBox="[151,1435,183,1836]" captionTargetId="figure@2.[151,1436,179,1836]" captionText="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="4" pageNumber="499">Figs 57</figureCitation>
,
<figureCitation id="73D52A0AFFD6FFDD063BEAC8FDE9FACF" box="[601,632,1268,1295]" captionStart="FIGURES 1218" captionStartId="5.[151,275,1888,1915]" captionTargetBox="[151,1436,179,1862]" captionTargetId="figure@5.[151,1436,179,1862]" captionText="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="4" pageNumber="499">15</figureCitation>
). Epistome regularly convex, with anterior margin bearing denticles of similar sizes (
<figureCitation id="73D52A0AFFD6FFDD0553EB25FEE7FAF3" box="[305,374,1304,1331]" captionStart="FIGURES 17" captionStartId="2.[151,275,1862,1889]" captionTargetBox="[151,1435,183,1836]" captionTargetId="figure@2.[151,1436,179,1836]" captionText="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="4" pageNumber="499">Fig. 5</figureCitation>
). Deutosternal furrow with six transverse lines of 818 denticles each; corniculi horn-like, parallel to each other, not reaching middle of palp femur; internal malae with median projections contiguous and with lightly serrate margins; laterally, four to six setiform tines on each side, some tines subdivided (
<figureCitation id="73D52A0AFFD6FFDD00D9EB5DFB6EFABC" box="[1211,1279,1377,1404]" captionStart="FIGURES 17" captionStartId="2.[151,275,1862,1889]" captionTargetBox="[151,1435,183,1836]" captionTargetId="figure@2.[151,1436,179,1836]" captionText="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="4" pageNumber="499">Fig. 6</figureCitation>
). Palp apotele 2-tined. Cheliceral digits of similar lengths; movable digit 2934 μm long, with two closely set teeth in addition to distal hook; fixed digit with five or six small teeth in addition to apical hook and minute setiform
<emphasis id="D99AEA9DFFD6FFDD0166EB95FA05FA03" box="[1284,1428,1449,1475]" italics="true" pageId="4" pageNumber="499">pilus dentilis</emphasis>
; arthrodial process as a coronet-like fringe; dorsal cheliceral seta and antiaxial lyrifissure not discerned (
<figureCitation id="73D52A0AFFD6FFDD017CEBF1FAF2FA27" box="[1310,1379,1485,1511]" captionStart="FIGURES 17" captionStartId="2.[151,275,1862,1889]" captionTargetBox="[151,1435,183,1836]" captionTargetId="figure@2.[151,1436,179,1836]" captionText="FIGURES 17. Cosmolaelaps ekaterinae sp. nov., female. 1. Idiosoma, dorsal view; 2. Medial dorsal seta, variant forms, lateral view; 3. Idiosoma, ventral view; 4. Epigynal shield, with typical ornamentation pattern; 5. Epistome; 6. Gnathosoma with palp, ventral view; 7. Chelicera, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3416846" httpUri="https://zenodo.org/record/3416846/files/figure.png" pageId="4" pageNumber="499">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="EB51368FFFD6FFDD04A5EBCEFF53F88F" blockId="4.[151,1437,153,1871]" pageId="4" pageNumber="499">
<emphasis id="D99AEA9DFFD6FFDD04A5EBCEFF6DF9CB" box="[199,252,1522,1547]" italics="true" pageId="4" pageNumber="499">Legs</emphasis>
(
<figureCitation id="73D52A0AFFD6FFDD0568EBCDFEEDF9CB" box="[266,380,1521,1547]" captionStart="FIGURES 811" captionStartId="3.[151,275,1735,1762]" captionTargetBox="[162,1431,195,1696]" captionTargetId="figure@3.[151,1436,179,1709]" captionText="FIGURES 811. Cosmolaelaps ekaterinae sp. nov., female, legs IIV. 8. Leg I; 9. Leg II; 10. Leg III; 11. Leg IV." figureDoi="http://doi.org/10.5281/zenodo.3416848" httpUri="https://zenodo.org/record/3416848/files/figure.png" pageId="4" pageNumber="499">Figs 811</figureCitation>
). All legs with well-developed pretarsus and ambulacral apparatus (including pulvillus and two claws), shorter than idiosoma: legs I 300325 μm, legs
<collectionCode id="8DFFAE4AFFD6FFDD0765E829FC8BF9EF" box="[775,794,1557,1583]" pageId="4" pageNumber="499">II</collectionCode>
225250 μm, legs
<collectionCode id="8DFFAE4AFFD6FFDD0790E829FB9FF9EF" box="[1010,1038,1557,1583]" pageId="4" pageNumber="499">III</collectionCode>
205220 μm, and legs
<collectionCode id="8DFFAE4AFFD6FFDD0174E829FAA2F9EF" box="[1302,1331,1557,1583]" pageId="4" pageNumber="499">IV</collectionCode>
270290 μm long. Leg segments not spurred ventrally, with normal chaetotactic pattern: leg I—coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-1, femur 2-2/1, 3/3-2, genu 2-3/2, 3/1-2, tibia 2-3/2, 3/1-2; leg II—coxa 0-0/1, 0/1-0, trochanter 1-0/1, 0/2-1, femur 2-3/1, 2/2-1, genu 2-3/1, 2/1-2, tibia 2-2/1, 2/1-2; leg III—coxa 0-0/1, 0/1-0, trochanter 1-1/1, 0/2-0, femur 1-2/1, 1/0-1, genu 2-2/1, 2/1-1, tibia 2-1/1, 2/1-1; leg IV—coxa 0-0/1, 0/0-0, trochanter 1-1/1, 0/2-0, femur 1-2/1, 1/0-1, genu 2-2/1, 3/0-1, tibia 2-1/1, 3/1-2; tarsi IIIV—18 setae each. Leg setae mostly uniform in shape, smooth and needle-like; setae ad1 on femora I and
<collectionCode id="8DFFAE4AFFD6FFDD06BBE8D1FD7DF8C7" box="[729,748,1773,1799]" pageId="4" pageNumber="499">II</collectionCode>
and ad2 on femur
<collectionCode id="8DFFAE4AFFD6FFDD07A1E8D1FC71F8C7" box="[963,992,1773,1799]" pageId="4" pageNumber="499">IV</collectionCode>
moderately thickened; pd2 on femur
<collectionCode id="8DFFAE4AFFD6FFDD01E8E8D1FA0CF8C7" box="[1418,1437,1773,1799]" pageId="4" pageNumber="499">II</collectionCode>
and ad1 on femora
<collectionCode id="8DFFAE4AFFD6FFDD050DE92DFE13F8EB" box="[367,386,1809,1835]" pageId="4" pageNumber="499">II</collectionCode>
and
<collectionCode id="8DFFAE4AFFD6FFDD05DAE92DFE45F8EB" box="[440,468,1809,1835]" pageId="4" pageNumber="499">III</collectionCode>
modified, thickened, widened toward their apex and spatulate (see detail in
<figureCitation id="73D52A0AFFD6FFDD014AE92DFF25F88F" captionStart="FIGURES 811" captionStartId="3.[151,275,1735,1762]" captionTargetBox="[162,1431,195,1696]" captionTargetId="figure@3.[151,1436,179,1709]" captionText="FIGURES 811. Cosmolaelaps ekaterinae sp. nov., female, legs IIV. 8. Leg I; 9. Leg II; 10. Leg III; 11. Leg IV." figureDoi="http://doi.org/10.5281/zenodo.3416848" httpUri="https://zenodo.org/record/3416848/files/figure.png" pageId="4" pageNumber="499">Figs 9 and 11</figureCitation>
).
</paragraph>
<caption id="BF916607FFD7FFDC04F5E95CFDA0F826" ID-DOI="http://doi.org/10.5281/zenodo.3416850" ID-Zenodo-Dep="3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="5" pageNumber="500" startId="5.[151,275,1888,1915]" targetBox="[151,1436,179,1862]" targetPageId="5">
<paragraph id="EB51368FFFD7FFDC04F5E95CFDA0F826" blockId="5.[151,1436,1888,2023]" pageId="5" pageNumber="500">
<emphasis id="D99AEA9DFFD7FFDC04F5E95CFEFBF8BA" bold="true" box="[151,362,1888,1915]" pageId="5" pageNumber="500">FIGURES 1218.</emphasis>
<taxonomicName id="2CEE4D0CFFD7FFDC0516E95DFD03F8BB" authority="Mašán &amp; Babaeian, 2019" authorityName="Mašán &amp; Babaeian" authorityYear="2019" box="[372,658,1889,1915]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="500" phylum="Arthropoda" rank="species" species="ekaterinae" status="sp. nov.">
<emphasis id="D99AEA9DFFD7FFDC0516E95DFD03F8BB" box="[372,658,1889,1915]" italics="true" pageId="5" pageNumber="500">Cosmolaelaps ekaterinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C2A957E6FFD7FFDC06FEE95DFD64F8BB" box="[668,757,1889,1915]" pageId="5" pageNumber="500" rank="species">
sp.
<date id="9F50104FFFD7FFDC06A7E95DFD64F8BB" box="[709,757,1889,1915]" pageId="5" pageNumber="500">nov.</date>
</taxonomicNameLabel>
12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg
<collectionCode id="8DFFAE4AFFD7FFDC0119E9B9FA09F85F" box="[1403,1432,1925,1951]" pageId="5" pageNumber="500">IV</collectionCode>
, male, lateroventral view; 17. Tibia and genu of leg
<collectionCode id="8DFFAE4AFFD7FFDC06B6E995FD60F803" box="[724,753,1961,1987]" pageId="5" pageNumber="500">IV</collectionCode>
, male, ventrolateral view; 18. Femur and trochanter of leg
<collectionCode id="8DFFAE4AFFD7FFDC01E1E995FA09F803" box="[1411,1432,1961,1987]" pageId="5" pageNumber="500">II</collectionCode>
, male, ventrolateral view. Not scaled.
</paragraph>
</caption>
<paragraph id="EB51368FFFD4FFDF04A5EEA5FDEDFDA3" blockId="6.[151,1437,152,611]" pageId="6" pageNumber="501">
<emphasis id="D99AEA9DFFD4FFDF04A5EEA5FE95FF73" bold="true" box="[199,260,153,179]" pageId="6" pageNumber="501">Male</emphasis>
(
<figureCitation id="73D52A0AFFD4FFDF0575EEA5FE5AFF73" box="[279,459,152,179]" captionStart="FIGURES 1218" captionStartId="5.[151,275,1888,1915]" captionTargetBox="[151,1436,179,1862]" captionTargetId="figure@5.[151,1436,179,1862]" captionText="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="6" pageNumber="501">Figs 13, 1518</figureCitation>
). Idiosoma 355370 μm long (n=3). Dorsal and ventral setal lengths proportionally shorter than in female. Holoventral shield 285295 μm long and 125130 μm wide (behind coxae
<collectionCode id="8DFFAE4AFFD4FFDF016AEE81FAB8FF17" box="[1288,1321,189,215]" pageId="6" pageNumber="501">IV</collectionCode>
), with ten pairs of setae (st1st5, JV1JV3, ZV1 and ZV2) in addition to three circumanal setae; sternogenital surface smooth but posterior region of the shield with distinct reticula represented mostly by transverse lines (
<figureCitation id="73D52A0AFFD4FFDF00AEEF39FAB0FEDF" box="[1228,1313,261,287]" captionStart="FIGURES 1218" captionStartId="5.[151,275,1888,1915]" captionTargetBox="[151,1436,179,1862]" captionTargetId="figure@5.[151,1436,179,1862]" captionText="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="6" pageNumber="501">Fig. 13</figureCitation>
). Posterior ends of peritrematal shields free, not fused to holoventral shield behind the coxae
<collectionCode id="8DFFAE4AFFD4FFDF0055EF15FBC4FE83" box="[1079,1109,297,323]" pageId="6" pageNumber="501">IV</collectionCode>
. Soft integument with 1214 pairs of setae. Cheliceral digits of similar lengths, with indistinguishable dentation; spermatodactyl well developed, elongated, slim, progressively narrowed toward apex, with straight proximal section directed anteriorly, a small ventromedial projection pointed backward, moderate submedial curvature, distal section directed upward, and club-like tip (
<figureCitation id="73D52A0AFFD4FFDF04A7EF85FE8BFE13" box="[197,282,440,467]" captionStart="FIGURES 1218" captionStartId="5.[151,275,1888,1915]" captionTargetBox="[151,1436,179,1862]" captionTargetId="figure@5.[151,1436,179,1862]" captionText="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="6" pageNumber="501">Fig. 15</figureCitation>
); movable digit with spermatodactyl 5558 μm in length, approximately twice longer than fixed digit (2427 μm). Legs similar to those in female, except for femur
<collectionCode id="8DFFAE4AFFD4FFDF0731EFE1FCF7FE37" box="[851,870,477,503]" pageId="6" pageNumber="501">II</collectionCode>
with a spur-like ventral projection associated with anteroventral seta (
<figureCitation id="73D52A0AFFD4FFDF050FEC3DFE2EFDDB" box="[365,447,513,539]" captionStart="FIGURES 1218" captionStartId="5.[151,275,1888,1915]" captionTargetBox="[151,1436,179,1862]" captionTargetId="figure@5.[151,1436,179,1862]" captionText="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="6" pageNumber="501">Fig. 18</figureCitation>
), ventral base of telotarsus
<collectionCode id="8DFFAE4AFFD4FFDF0691EC3DFC81FDDB" box="[755,784,513,539]" pageId="6" pageNumber="501">IV</collectionCode>
with small pointed or blunt projection (
<figureCitation id="73D52A0AFFD4FFDF00ADEC3DFAB0FDDC" box="[1231,1313,513,540]" captionStart="FIGURES 1218" captionStartId="5.[151,275,1888,1915]" captionTargetBox="[151,1436,179,1862]" captionTargetId="figure@5.[151,1436,179,1862]" captionText="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="6" pageNumber="501">Fig. 16</figureCitation>
), and genu
<collectionCode id="8DFFAE4AFFD4FFDF04F5EC19FF25FDFF" box="[151,180,549,575]" pageId="6" pageNumber="501">IV</collectionCode>
with markedly thickened and shortened anteroventral seta having its apex truncate or bluntly rounded (
<figureCitation id="73D52A0AFFD4FFDF0159EC19FA1EFDFF" box="[1339,1423,549,575]" captionStart="FIGURES 1218" captionStartId="5.[151,275,1888,1915]" captionTargetBox="[151,1436,179,1862]" captionTargetId="figure@5.[151,1436,179,1862]" captionText="FIGURES 1218. Cosmolaelaps ekaterinae sp. nov. 12. Idiosoma, female, dorsal view; 13. Holoventral shield, male; 14. Sternal and epigynal shield, female; 15. Chelicera, male, lateroventral view; 16. Tarsus and tibia of leg IV, male, lateroventral view; 17. Tibia and genu of leg IV, male, ventrolateral view; 18. Femur and trochanter of leg II, male, ventrolateral view. Not scaled." figureDoi="http://doi.org/10.5281/zenodo.3416850" httpUri="https://zenodo.org/record/3416850/files/figure.png" pageId="6" pageNumber="501">Fig. 17</figureCitation>
). Other characters similar to those of female.
</paragraph>
<paragraph id="EB51368FFFD4FFDF04F5EC89FE9EFD0F" blockId="6.[151,1437,693,791]" box="[151,271,693,719]" pageId="6" pageNumber="501">
<emphasis id="D99AEA9DFFD4FFDF04F5EC89FE9EFD0F" box="[151,271,693,719]" italics="true" pageId="6" pageNumber="501">Etymology</emphasis>
</paragraph>
<paragraph id="EB51368FFFD4FFDF04F5ECE5FB54FCD7" blockId="6.[151,1437,693,791]" pageId="6" pageNumber="501">This species is named in honour of our colleague, acarologist Katya Sidorchuk (Paleontological Institute, Russian Academy of Sciences, Moscow), who has passed away recently, as a dedication to her memory.</paragraph>
<paragraph id="EB51368FFFD4FFDF04F5ED56FEC2FC43" blockId="6.[151,1437,874,1511]" box="[151,339,874,899]" pageId="6" pageNumber="501">
<emphasis id="D99AEA9DFFD4FFDF04F5ED56FEC2FC43" box="[151,339,874,899]" italics="true" pageId="6" pageNumber="501">Taxonomic notes</emphasis>
</paragraph>
<paragraph id="EB51368FFFD4FFDF04F5EDB1FE58FA27" blockId="6.[151,1437,874,1511]" pageId="6" pageNumber="501">
The dorsal shield chaetotaxy of
<taxonomicName id="2CEE4D0CFFD4FFDF059DEDB1FC8AFC67" authority="Mašán &amp; Babaeian, 2019" authorityName="Mašán &amp; Babaeian" authorityYear="2019" box="[511,795,909,935]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="ekaterinae" status="sp. nov.">
<emphasis id="D99AEA9DFFD4FFDF059DEDB1FC8AFC67" box="[511,795,909,935]" italics="true" pageId="6" pageNumber="501">Cosmolaelaps ekaterinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C2A957E6FFD4FFDF0741EDB1FCEDFC67" box="[803,892,909,935]" pageId="6" pageNumber="501" rank="species">sp. nov.</taxonomicNameLabel>
, with remarkably elongate and distally attenuate setae, is unique and quite unlike any other species in the genus. Other congeners have much broader and shorter dorsal setae, and usually with their tips reaching prior to or only slightly beyond the insertions of the following setae. Nevertheless, the new species is most similar to
<taxonomicName id="2CEE4D0CFFD4FFDF06CCEDC5FB0BFBD3" authority="Ma &amp; Lin, 2009" authorityName="Ma &amp; Lin" authorityYear="2009" box="[686,1178,1017,1043]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF06CCEDC5FC4FFBD3" box="[686,990,1017,1043]" italics="true" pageId="6" pageNumber="501">Cosmolaelaps leptochaetes</emphasis>
<bibRefCitation id="8F7F4B7EFFD4FFDF0786EDC5FB0BFBD3" author="Ma, L. M. &amp; Lin, J. Z." box="[996,1178,1017,1043]" pageId="6" pageNumber="501" pagination="27 - 37" refId="ref6765" refString="Ma, L. M. &amp; Lin, J. Z. (2009) On five new species of the genera Hypoaspis and Cosmolaelaps, with supplementary descriptions of Cosmolaelaps xiajiangensis and Haemolaelaps cordatus (Acari: Mesostigmata: Laelapidae). Acta Arachnologica Sinica, 18, 27 - 37." type="journal article" year="2009">Ma &amp; Lin, 2009</bibRefCitation>
</taxonomicName>
, described from
<collectingCountry id="93F9761FFFD4FFDF0130EDC5FA09FBD3" box="[1362,1432,1017,1043]" name="China" pageId="6" pageNumber="501">China</collectingCountry>
, and reported from
<collectingCountry id="93F9761FFFD4FFDF0509EA21FE09FBF7" box="[363,408,1053,1079]" name="Iran" pageId="6" pageNumber="501">Iran</collectingCountry>
(
<bibRefCitation id="8F7F4B7EFFD4FFDF05C5EA21FD34FBF7" author="Ramroodi, S. &amp; Hajizadeh, J. &amp; Joharchi, O." box="[423,677,1053,1079]" pageId="6" pageNumber="501" refId="ref7076" refString="Ramroodi, S., Hajizadeh, J. &amp; Joharchi, O. (2014) Faunistic survey on laelapid mites (Acari: Mesostigmata) of Guilan province with new records for Iran. Poster (P 4 - 23). Abstract book of the XIV. International Congress of Acarology, Kyoto, Japan, 2014, 153." type="journal volume" year="2014">
Ramroodi
<emphasis id="D99AEA9DFFD4FFDF0642EA22FDC8FBF7" box="[544,601,1053,1079]" italics="true" pageId="6" pageNumber="501">et al.</emphasis>
, 2014
</bibRefCitation>
). The new species can be easily distinguished from
<taxonomicName id="2CEE4D0CFFD4FFDF008DEA21FA0DFBF7" authorityName="Ma &amp; Lin" authorityYear="2009" box="[1263,1436,1053,1079]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF008DEA21FA0DFBF7" box="[1263,1436,1053,1079]" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
by the following characters: idiosoma 275295 μm long (
<quantity id="2C169B6AFFD4FFDF0779EA7CFC0AFB9B" box="[795,923,1088,1115]" metricMagnitude="1" metricUnit="m" metricValue="1.56845" metricValueMax="1.6382999999999999" metricValueMin="1.4986" pageId="6" pageNumber="501" unit="in" value="617.5" valueMax="645.0" valueMin="590.0">590645 in</quantity>
<taxonomicName id="2CEE4D0CFFD4FFDF07C0EA7DFBDEFB9B" authorityName="Ma &amp; Lin" authorityYear="2009" box="[930,1103,1089,1115]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF07C0EA7DFBDEFB9B" box="[930,1103,1089,1115]" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
); dorsal setae longer, with tip reaching beyond the insertions of two following setae in a row (setae only reaching beyond the insertion of the following seta in
<taxonomicName id="2CEE4D0CFFD4FFDF055FEAB5FE7AFB63" authorityName="Ma &amp; Lin" authorityYear="2009" box="[317,491,1161,1187]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF055FEAB5FE7AFB63" box="[317,491,1161,1187]" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
); anterior margin of sternal shield and small presternal platelets weakly sclerotized, poorly defined (the entire sternal shield and relatively large presternal platelets well sclerotized and defined in
<taxonomicName id="2CEE4D0CFFD4FFDF01E1EA91FEB2FB2B" authorityName="Ma &amp; Lin" authorityYear="2009" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF01E1EA91FEB2FB2B" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
); peritremes relatively shorter, with anterior ends reaching slightly beyond the dorsal setae s1 (peritremes longer, reaching close to the setae z
<quantity id="2C169B6AFFD4FFDF061AEAC9FD30FACF" box="[632,673,1269,1295]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="501" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="2CEE4D0CFFD4FFDF06CAEAC9FCC7FACF" authorityName="Ma &amp; Lin" authorityYear="2009" box="[680,854,1269,1295]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF06CAEAC9FCC7FACF" box="[680,854,1269,1295]" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
); setae of JV and ZV rows becoming progressively longer posteriorly (all opisthogastric setae of similar length in
<taxonomicName id="2CEE4D0CFFD4FFDF0736EB25FB90FAF3" authorityName="Ma &amp; Lin" authorityYear="2009" box="[852,1025,1305,1331]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF0736EB25FB90FAF3" box="[852,1025,1305,1331]" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
); spermatodactyl about twice longer than fixed digit of chelicera (only slightly longer than fixed digit in
<taxonomicName id="2CEE4D0CFFD4FFDF07FFEB01FBDAFA97" authorityName="Ma &amp; Lin" authorityYear="2009" box="[925,1099,1341,1367]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF07FFEB01FBDAFA97" box="[925,1099,1341,1367]" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
); male femur
<collectionCode id="8DFFAE4AFFD4FFDF008CEB01FA90FA97" box="[1262,1281,1341,1367]" pageId="6" pageNumber="501">II</collectionCode>
with a strong ventral spur associated with short and thin anteroventral seta (apparently with no ventral spur on femur
<collectionCode id="8DFFAE4AFFD4FFDF0125EB5DFACBFABB" box="[1351,1370,1377,1403]" pageId="6" pageNumber="501">II</collectionCode>
in
<taxonomicName id="2CEE4D0CFFD4FFDF01E1EB5DFEB2FA5F" authorityName="Ma &amp; Lin" authorityYear="2009" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF01E1EB5DFEB2FA5F" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
, whereas anteroventral seta long and markedly thickened); female tarsus
<collectionCode id="8DFFAE4AFFD4FFDF0001EBB9FBE7FA5F" box="[1123,1142,1413,1439]" pageId="6" pageNumber="501">II</collectionCode>
relatively short and wide, stout (apparently narrower and longer in shape in
<taxonomicName id="2CEE4D0CFFD4FFDF06ADEB95FCEDFA03" authorityName="Ma &amp; Lin" authorityYear="2009" box="[719,892,1449,1475]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF06ADEB95FCEDFA03" box="[719,892,1449,1475]" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
); male tarsus
<collectionCode id="8DFFAE4AFFD4FFDF007CEB95FBAAFA03" box="[1054,1083,1449,1475]" pageId="6" pageNumber="501">IV</collectionCode>
with a small ventral projection (absent in
<taxonomicName id="2CEE4D0CFFD4FFDF056EEBF1FE29FA27" authorityName="Ma &amp; Lin" authorityYear="2009" box="[268,440,1485,1511]" class="Arachnida" family="Laelapidae" genus="Cosmolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="501" phylum="Arthropoda" rank="species" species="leptochaetes">
<emphasis id="D99AEA9DFFD4FFDF056EEBF1FE29FA27" box="[268,440,1485,1511]" italics="true" pageId="6" pageNumber="501">C. leptochaetes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>