treatments-xml/data/63/43/87/634387D1A358FFBA173BFA9CFE9E380C.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.6280443" ID-GBIF-Dataset="647046fe-0ee5-401d-b4f8-aea0355fde7f" ID-Zenodo-Dep="6280443" checkinTime="1471630429166" checkinUser="plazi" docAuthor="Piotr Naskrecki" docDate="2000" docId="634387D1A358FFBA173BFA9CFE9E380C" docLanguage="en" docName="Naskrecki_KCR_2000.pdf.imf" docOrigin="Philadelphia, PA: The Orthopterists Society at the Academy of Natural Sciences of Philadelphia," docStyle="DocumentStyle{}" docTitle="Eriolus" docType="treatment" docVersion="8" lastPageId="116" lastPageNumber="113" masterDocId="9F7AFFA9A32AFFCE167AFFC3FFF13C6E" masterDocTitle="Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato)." pageId="114" pageNumber="111" updateTime="1645797358202" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato).</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Piotr Naskrecki</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:originInfo>
<mods:dateIssued>2000</mods:dateIssued>
<mods:publisher>The Orthopterists Society at the Academy of Natural Sciences of Philadelphia,</mods:publisher>
<mods:place>
<mods:placeTerm>Philadelphia, PA</mods:placeTerm>
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<mods:classification>book</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.270035</mods:identifier>
<mods:identifier type="GBIF-Dataset">647046fe-0ee5-401d-b4f8-aea0355fde7f</mods:identifier>
<mods:identifier type="Zenodo-Dep">270035</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6280443" ID-Zenodo-Dep="6280443" LSID="urn:lsid:plazi:treatment:634387D1A358FFBA173BFA9CFE9E380C" httpUri="http://treatment.plazi.org/id/634387D1A358FFBA173BFA9CFE9E380C" lastPageId="116" lastPageNumber="113" pageId="114" pageNumber="111">
<paragraph blockId="114.[321,570,1375,1397]" box="[321,570,1375,1397]" pageId="114" pageNumber="111">
<heading box="[321,570,1375,1397]" centered="true" fontSize="9" level="14" pageId="114" pageNumber="111" reason="4">
<taxonomicName box="[321,429,1375,1396]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="114" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[321,429,1375,1396]" italics="true" pageId="114" pageNumber="111">ERIOLUS</emphasis>
</taxonomicName>
<collectingRegion box="[435,513,1375,1397]" country="Venezuela" name="Bolivar" pageId="114" pageNumber="111">Bolivar</collectingRegion>
, 1888
</heading>
</paragraph>
<paragraph blockId="114.[151,694,1432,1540]" pageId="114" pageNumber="111">
1888 
<collectingRegion box="[205,283,1432,1454]" country="Venezuela" name="Bolivar" pageId="114" pageNumber="111">Bolivar</collectingRegion>
, Mem. Soc. Zool. 
<collectingCountry box="[469,540,1432,1453]" name="France" pageId="114" pageNumber="111">France</collectingCountry>
1: 150; 
<typeStatus box="[618,665,1432,1453]" pageId="114" pageNumber="111">type</typeStatus>
species: 
<taxonomicName box="[296,463,1460,1481]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="114" pageNumber="111" phylum="Arthropoda" rank="species" species="caraibeus">
<emphasis box="[296,463,1460,1481]" italics="true" pageId="114" pageNumber="111">Eriolus caraibeus</emphasis>
</taxonomicName>
<collectingRegion box="[469,547,1460,1482]" country="Venezuela" name="Bolivar" pageId="114" pageNumber="111">Bolivar</collectingRegion>
, 1888
</paragraph>
<paragraph blockId="114.[151,694,1432,1540]" pageId="114" pageNumber="111">
1999 Naskrecki and Otte, Illustr. Cat. Orthop. I (
<collectionCode box="[657,694,1489,1510]" pageId="114" pageNumber="111">CD</collectionCode>
<collectionCode box="[209,268,1518,1540]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="114" pageNumber="111">ROM</collectionCode>
) &gt;&gt;full references
</paragraph>
<subSubSection pageId="114" pageNumber="111" type="diagnosis">
<paragraph blockId="114.[151,740,1574,1739]" box="[151,262,1574,1596]" pageId="114" pageNumber="111">
<heading bold="true" box="[151,262,1574,1596]" fontSize="9" level="13" pageId="114" pageNumber="111" reason="6">
<emphasis bold="true" box="[151,262,1574,1596]" pageId="114" pageNumber="111">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="114.[151,740,1574,1739]" lastBlockId="114.[771,1360,207,372]" pageId="114" pageNumber="111">
Body small as for the subfamily, slender; both sexes macropterous (
<figureCitation box="[308,387,1631,1653]" captionStart="FIG. 30" captionStartId="117.[174,218,1684,1706]" captionTargetBox="[174,1368,204,1607]" captionTargetId="figure@117.[174,1369,204,1607]" captionTargetPageId="117" captionText="FIG. 30. Eriolus aculeus. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. epiphallus, posterior and lateral views, F. titillators." httpUri="https://zenodo.org/record/270066/files/figure.png" pageId="114" pageNumber="111">Figs. 30</figureCitation>
A, 31A, 37C). Head with fastigium of vertex 0.8-1.5 times as long as diameter of eye, sharply conical or blunt. Head narrow, frons flat or weakly convex; genae without lateral carinae. Prosternum unarmed or armed with two spines. Male cerci and concealed genitalia extremely variable, titillators always well developed; male paraprocts modified into long, finger-like processes; 10th tergite in both sexes often with long, narrow lobes on posterior margin; ovipositor moderately to strongly curved upwards, pointed apically.
</paragraph>
</subSubSection>
<subSubSection lastPageId="115" lastPageNumber="112" pageId="114" pageNumber="111" type="description">
<paragraph blockId="114.[771,1361,406,1742]" box="[771,1222,406,429]" pageId="114" pageNumber="111">
<emphasis bold="true" box="[771,900,406,428]" pageId="114" pageNumber="111">Description</emphasis>
(male except where specified)
</paragraph>
<paragraph blockId="114.[771,1361,406,1742]" pageId="114" pageNumber="111">
<emphasis box="[794,875,435,456]" italics="true" pageId="114" pageNumber="111">Head.—</emphasis>
Fastigium of vertex small as for the subfamily, 0.8-1.5 times as long as diameter of eye (
<figureCitation box="[1195,1278,464,486]" captionStart="FIG. 30" captionStartId="117.[174,218,1684,1706]" captionTargetBox="[174,1368,204,1607]" captionTargetId="figure@117.[174,1369,204,1607]" captionTargetPageId="117" captionText="FIG. 30. Eriolus aculeus. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. epiphallus, posterior and lateral views, F. titillators." httpUri="https://zenodo.org/record/270066/files/figure.png" pageId="114" pageNumber="111">Figs. 30</figureCitation>
C, 31C); fastigium sharply conical, with lamelliform, ventral keel, or blunt apically and rounded underneath; dorsal surface of fastigium smooth or with weak longitudinal furrow; lateral ocelli well developed; fastigium of vertex continuous with fastigium of frons. Antennal sockets separated by distance equal to about 0.4 diameter of eye. Eyes globose, moderately projecting; frons flat or weakly convex; genae with lateral carinae. Labrum and mandibles symmetrical to weakly asymmetrical (right mandible smaller) (
<figureCitation box="[930,1003,749,771]" captionStart="FIG. 30" captionStartId="117.[174,218,1684,1706]" captionTargetBox="[174,1368,204,1607]" captionTargetId="figure@117.[174,1369,204,1607]" captionTargetPageId="117" captionText="FIG. 30. Eriolus aculeus. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. epiphallus, posterior and lateral views, F. titillators." httpUri="https://zenodo.org/record/270066/files/figure.png" pageId="114" pageNumber="111">Fig. 30</figureCitation>
B).
</paragraph>
<paragraph blockId="114.[771,1361,406,1742]" pageId="114" pageNumber="111">
<emphasis box="[794,1023,778,799]" italics="true" pageId="114" pageNumber="111">Thorax and wings.—</emphasis>
Dorsal surface of pronotum smooth, flat; anterior margin of pronotum straight, posterior margin straight or weakly convex; lateral lobes with posterior angle narrowly rounded, with well developed humeral sinus. Thoracic auditory spiracle large, elliptical, completely hidden under lateral lobes of pronotum; posterior margin of spiracle with finger-like projection. Prosternum unarmed or armed with two, thin, widely separated spines, meso- and metasternum unarmed, without well developed basisternal lobes.
</paragraph>
<paragraph blockId="114.[771,1361,406,1742]" pageId="114" pageNumber="111">
Wings in both sexes fully developed, surpassing apices of hind 
<taxonomicName box="[893,971,1092,1114]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="114" pageNumber="111" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
. Stridulatory apparatus of male well developed; stridulatory file straight or weakly sinusoidal (
<figureCitation box="[816,894,1148,1171]" captionStart="FIG. 50" captionStartId="148.[151,195,1684,1706]" captionTargetBox="[204,1306,204,1665]" captionTargetId="figure@148.[204,1307,204,1665]" captionTargetPageId="148" captionText="FIG. 50. Stridulatory files of Costa Rican species of Conocephalinae s. l. A. Erioloides longinoi, C. Erioloides sikesi, C. Erioloides acutidentis, D. Eriolus duplidentis, E. Eriolus aculeus, F. Eriolus penicillus, G. Subria scutellaris, H. Subria crassicerca." httpUri="https://zenodo.org/record/270086/files/figure.png" pageId="114" pageNumber="111">Figs. 50</figureCitation>
D-F), teeth relatively thick and narrow to very thin and wide; stridulatory area of left wing without secondary venation, rectangular or nearly triangular; mirror of right wing rectangular to nearly triangular, with very well developed veinlet next to AA1. Posterior margin of front wing straight or weakly convex; apex of front wing narrowly rounded.
</paragraph>
<paragraph blockId="114.[771,1361,406,1742]" pageId="114" pageNumber="111">
<emphasis box="[794,868,1349,1370]" italics="true" pageId="114" pageNumber="111">Legs.—</emphasis>
Fore coxa with an elongate, sometimes laterally flattened, forward projecting spine dorsally; middle and hind coxa without spine; all trochanters unarmed. All 
<taxonomicName box="[812,887,1434,1456]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="114" pageNumber="111" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
unarmed dorsally but with small spines on anterior ventral margins, posterior ventral margins unarmed; genicular lobes of all 
<taxonomicName box="[1092,1167,1491,1513]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="114" pageNumber="111" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
armed with long spines, sometimes posterior lobes of fore and/or mid 
<taxonomicName box="[771,846,1548,1570]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="114" pageNumber="111" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
unarmed. Front and mid tibia unarmed dorsally, their ventral margins with short, immovable spines; tympanum on fore tibia bilaterally closed, tympanal slits facing forward, tympanal area moderately swollen; hind tibia armed on all four dorsal and ventral margins; apex of tibia with two pairs of ventral and one pair of dorsal movable spurs.
</paragraph>
<caption ID-Table-UUID="BF95664FA359FFBD16D4FF0CFC933C8A" box="[174,866,207,229]" httpUri="http://table.plazi.org/id/BF95664FA359FFBD16D4FF0CFC933C8A" pageId="115" pageNumber="112" targetBox="[174,1376,265,978]" targetIsTable="true" targetPageId="115">
<paragraph blockId="115.[174,1376,207,431]" box="[174,866,207,229]" pageId="115" pageNumber="112">
<emphasis bold="true" box="[174,289,207,228]" pageId="115" pageNumber="112">TABLE 22.</emphasis>
Key for identification of Costa Rican species of 
<taxonomicName box="[795,866,207,228]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="genus">Eriolus</taxonomicName>
</paragraph>
</caption>
<paragraph blockId="115.[174,1376,207,431]" pageId="115" pageNumber="112">
<table box="[174,1376,265,978]" gridcols="5" gridrows="4" pageId="115" pageNumber="112">
<tr box="[174,1376,265,431]" gridrow="0" pageId="115" pageNumber="112">
<th box="[174,287,265,431]" gridcol="0" gridrow="0" pageId="115" pageNumber="112">
Species 
<taxonomicName box="[174,287,323,344]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="acutipennis">
<emphasis box="[174,287,323,344]" italics="true" pageId="115" pageNumber="112">acutipennis</emphasis>
</taxonomicName>
</th>
<th box="[310,557,265,431]" gridcol="1" gridrow="0" pageId="115" pageNumber="112">Male cercus straight, with small sub-basal tooth internally</th>
<th box="[589,867,265,431]" gridcol="2" gridrow="0" pageId="115" pageNumber="112">
10th tergite (both sexes) with two small, triangular lobes (
<figureCitation box="[658,727,351,373]" captionStart="FIG. 31" captionStartId="118.[151,195,1684,1706]" captionTargetBox="[174,1329,204,1624]" captionTargetId="figure@118.[174,1329,204,1625]" captionTargetPageId="118" captionText="FIG. 31. Eriolus duplidentis. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. titillators." httpUri="https://zenodo.org/record/270067/files/figure.png" pageId="115" pageNumber="112">Fig. 31</figureCitation>
E)
</th>
<th box="[884,1062,265,431]" gridcol="3" gridrow="0" pageId="115" pageNumber="112">
Ovipositor sickle-shaped, as long as half of hind femur (
<figureCitation box="[892,962,409,431]" captionStart="FIG. 31" captionStartId="118.[151,195,1684,1706]" captionTargetBox="[174,1329,204,1624]" captionTargetId="figure@118.[174,1329,204,1625]" captionTargetPageId="118" captionText="FIG. 31. Eriolus duplidentis. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. titillators." httpUri="https://zenodo.org/record/270067/files/figure.png" pageId="115" pageNumber="112">Fig. 31</figureCitation>
F)
</th>
<th box="[1081,1376,265,431]" gridcol="4" gridrow="0" pageId="115" pageNumber="112">Male subgenital plate with styli located on widely separated, narrow lobes</th>
</tr>
<tr box="[174,1376,524,632]" gridrow="1" pageId="115" pageNumber="112">
<th box="[174,287,524,632]" gridcol="0" gridrow="1" pageId="115" pageNumber="112">
<taxonomicName box="[174,247,524,545]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="aculeus">
<emphasis box="[174,247,524,545]" italics="true" pageId="115" pageNumber="112">aculeus</emphasis>
</taxonomicName>
</th>
<td box="[310,557,524,632]" gridcol="1" gridrow="1" pageId="115" pageNumber="112">
<emphasis italics="true" pageId="115" pageNumber="112">with large, oblique, apical arm</emphasis>
</td>
<td box="[589,867,524,632]" gridcol="2" gridrow="1" pageId="115" pageNumber="112">
<emphasis italics="true" pageId="115" pageNumber="112">
with two long, thin, parallel lobes (
<figureCitation box="[744,816,553,575]" captionStart="FIG. 29" captionStartId="112.[151,195,1655,1677]" captionTargetBox="[204,1333,204,1625]" captionTargetId="figure@112.[204,1333,204,1625]" captionTargetPageId="112" captionText="FIG. 29. Species of Pluviasilva. A, C, F-J. P. levis: A. male - habitus, C. ovipositor, F. male cerci, lateral view, G. male cerci, dorsal view, H. titillators, I. male head and pronotum, dorsal view, J. male face; B, D-E. P. mexicana: B. male - habitus, D. male cerci, lateral view, E. male cerci, dorsal view." httpUri="https://zenodo.org/record/270065/files/figure.png" pageId="115" pageNumber="112">Fig. 29</figureCitation>
D)
</emphasis>
</td>
<td box="[884,1062,524,632]" gridcol="3" gridrow="1" pageId="115" pageNumber="112">
<emphasis italics="true" pageId="115" pageNumber="112">
saber-shaped, as long as 0.7 of hind femur (
<figureCitation box="[892,963,610,632]" captionStart="FIG. 29" captionStartId="112.[151,195,1655,1677]" captionTargetBox="[204,1333,204,1625]" captionTargetId="figure@112.[204,1333,204,1625]" captionTargetPageId="112" captionText="FIG. 29. Species of Pluviasilva. A, C, F-J. P. levis: A. male - habitus, C. ovipositor, F. male cerci, lateral view, G. male cerci, dorsal view, H. titillators, I. male head and pronotum, dorsal view, J. male face; B, D-E. P. mexicana: B. male - habitus, D. male cerci, lateral view, E. male cerci, dorsal view." httpUri="https://zenodo.org/record/270065/files/figure.png" pageId="115" pageNumber="112">Fig. 29</figureCitation>
A)
</emphasis>
</td>
<td box="[1081,1376,524,632]" gridcol="4" gridrow="1" pageId="115" pageNumber="112">
<emphasis italics="true" pageId="115" pageNumber="112">
<heading box="[1081,1351,524,546]" centered="true" fontSize="9" level="14" pageId="115" pageNumber="112" reason="4">with small apical incision</heading>
and long styli
</emphasis>
</td>
</tr>
<tr box="[174,1376,697,805]" gridrow="2" pageId="115" pageNumber="112">
<th box="[174,287,697,805]" gridcol="0" gridrow="2" pageId="115" pageNumber="112">
<taxonomicName box="[174,268,697,718]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="penicillus">
<emphasis box="[174,268,697,718]" italics="true" pageId="115" pageNumber="112">penicillus</emphasis>
</taxonomicName>
</th>
<td box="[310,557,697,805]" gridcol="1" gridrow="2" pageId="115" pageNumber="112">
<emphasis italics="true" pageId="115" pageNumber="112">with thin, finger-like subapical spine</emphasis>
</td>
<td box="[589,867,697,805]" gridcol="2" gridrow="2" pageId="115" pageNumber="112">
<emphasis italics="true" pageId="115" pageNumber="112">
with two long, thin, convergent lobes (
<figureCitation box="[597,667,754,776]" captionStart="FIG. 31" captionStartId="118.[151,195,1684,1706]" captionTargetBox="[174,1329,204,1624]" captionTargetId="figure@118.[174,1329,204,1625]" captionTargetPageId="118" captionText="FIG. 31. Eriolus duplidentis. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. titillators." httpUri="https://zenodo.org/record/270067/files/figure.png" pageId="115" pageNumber="112">Fig. 31</figureCitation>
A)
</emphasis>
</td>
<td box="[884,1062,697,805]" gridcol="3" gridrow="2" pageId="115" pageNumber="112">
<emphasis italics="true" pageId="115" pageNumber="112">
saber-shaped, as long as 0.8-0.9 of hind femur (
<figureCitation box="[892,962,783,805]" captionStart="FIG. 31" captionStartId="118.[151,195,1684,1706]" captionTargetBox="[174,1329,204,1624]" captionTargetId="figure@118.[174,1329,204,1625]" captionTargetPageId="118" captionText="FIG. 31. Eriolus duplidentis. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. titillators." httpUri="https://zenodo.org/record/270067/files/figure.png" pageId="115" pageNumber="112">Fig. 31</figureCitation>
B)
</emphasis>
</td>
<td box="[1081,1376,697,805]" gridcol="4" gridrow="2" pageId="115" pageNumber="112">
<emphasis italics="true" pageId="115" pageNumber="112">
<heading box="[1081,1351,697,719]" centered="true" fontSize="9" level="14" pageId="115" pageNumber="112" reason="4">with small apical incision</heading>
and long styli
</emphasis>
</td>
</tr>
<tr box="[174,1376,869,978]" gridrow="3" pageId="115" pageNumber="112">
<th box="[174,287,869,978]" gridcol="0" gridrow="3" pageId="115" pageNumber="112">
<heading box="[174,283,869,890]" centered="true" fontSize="9" level="14" pageId="115" pageNumber="112" reason="4">
<taxonomicName box="[174,283,869,890]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="duplidentis">
<emphasis box="[174,283,869,890]" italics="true" pageId="115" pageNumber="112">duplidentis</emphasis>
</taxonomicName>
</heading>
</th>
<td box="[310,557,869,978]" gridcol="1" gridrow="3" pageId="115" pageNumber="112">conical, with small, thick basal spine and large protuberance midlength</td>
<td box="[589,867,869,978]" gridcol="2" gridrow="3" pageId="115" pageNumber="112">
with shallow, widely triangular incision (
<figureCitation box="[597,668,927,949]" captionStart="FIG. 30" captionStartId="117.[174,218,1684,1706]" captionTargetBox="[174,1368,204,1607]" captionTargetId="figure@117.[174,1369,204,1607]" captionTargetPageId="117" captionText="FIG. 30. Eriolus aculeus. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. epiphallus, posterior and lateral views, F. titillators." httpUri="https://zenodo.org/record/270066/files/figure.png" pageId="115" pageNumber="112">Fig. 30</figureCitation>
D)
</td>
<td box="[884,1062,869,978]" gridcol="3" gridrow="3" pageId="115" pageNumber="112">
sickle-shaped, as long as half of hind femur (
<figureCitation box="[892,963,956,978]" captionStart="FIG. 30" captionStartId="117.[174,218,1684,1706]" captionTargetBox="[174,1368,204,1607]" captionTargetId="figure@117.[174,1369,204,1607]" captionTargetPageId="117" captionText="FIG. 30. Eriolus aculeus. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. epiphallus, posterior and lateral views, F. titillators." httpUri="https://zenodo.org/record/270066/files/figure.png" pageId="115" pageNumber="112">Fig. 30</figureCitation>
A)
</td>
<td box="[1081,1376,869,978]" gridcol="4" gridrow="3" pageId="115" pageNumber="112">without styli and with small, triangular apical incision</td>
</tr>
</table>
</paragraph>
<paragraph blockId="115.[174,763,1152,1731]" pageId="115" pageNumber="112">
<emphasis box="[196,317,1152,1173]" italics="true" pageId="115" pageNumber="112">Abdomen.—</emphasis>
Dorsal surface of abdominal terga smooth, unmodified. Male 10th tergite often with hind margin bearing two, widely separated, narrow lobes (
<figureCitation box="[659,731,1210,1233]" captionStart="FIG. 30" captionStartId="117.[174,218,1684,1706]" captionTargetBox="[174,1368,204,1607]" captionTargetId="figure@117.[174,1369,204,1607]" captionTargetPageId="117" captionText="FIG. 30. Eriolus aculeus. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. epiphallus, posterior and lateral views, F. titillators." httpUri="https://zenodo.org/record/270066/files/figure.png" pageId="115" pageNumber="112">Fig. 30</figureCitation>
D); female 10th tergite similar to that of conspecific males, often with a pair of long, narrow, lobes; supraanal plate in both sexes small, broadly rounded apically; male cercus variable but always with one or two inner, blunt spines; spines basal, subapical or apical; paraprocts modified into long, finger-like processes (
<figureCitation box="[552,633,1386,1408]" captionStart="FIG. 30" captionStartId="117.[174,218,1684,1706]" captionTargetBox="[174,1368,204,1607]" captionTargetId="figure@117.[174,1369,204,1607]" captionTargetPageId="117" captionText="FIG. 30. Eriolus aculeus. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. epiphallus, posterior and lateral views, F. titillators." httpUri="https://zenodo.org/record/270066/files/figure.png" pageId="115" pageNumber="112">Figs. 30</figureCitation>
D, 31D, 32A, 32E); female cercus, simple, narrowly conical. Subgenital plate of male with a very shallow apical emargination; with or without styli; female subgenital plate approximately triangular, with shallow apical incision and sometimes with small lobe in the middle of apical incision. Male internal genitalia extremely variable but always with well developed titillators (
<figureCitation box="[461,544,1592,1614]" captionStart="FIG. 30" captionStartId="117.[174,218,1684,1706]" captionTargetBox="[174,1368,204,1607]" captionTargetId="figure@117.[174,1369,204,1607]" captionTargetPageId="117" captionText="FIG. 30. Eriolus aculeus. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. epiphallus, posterior and lateral views, F. titillators." httpUri="https://zenodo.org/record/270066/files/figure.png" pageId="115" pageNumber="112">Figs. 30</figureCitation>
E-F, 31E, 32D, 32G). Ovipositor narrow, moderately to strongly upcurved, its margins usually parallel; both dorsal and ventral margins of ovipositor smooth, its apex pointed; ovipositor always shorter than hind femur (
<figureCitation box="[437,517,1709,1731]" captionStart="FIG. 30" captionStartId="117.[174,218,1684,1706]" captionTargetBox="[174,1368,204,1607]" captionTargetId="figure@117.[174,1369,204,1607]" captionTargetPageId="117" captionText="FIG. 30. Eriolus aculeus. A. female - habitus, B. female face, C. female head and pronotum, dorsal view, D. male cerci, dorsal and posterior views, E. epiphallus, posterior and lateral views, F. titillators." httpUri="https://zenodo.org/record/270066/files/figure.png" pageId="115" pageNumber="112">Figs. 30</figureCitation>
A, 31A, 32B, 32F).
</paragraph>
<paragraph blockId="115.[794,1384,1152,1731]" pageId="115" pageNumber="112">
<emphasis box="[817,950,1152,1173]" italics="true" pageId="115" pageNumber="112">Coloration.—</emphasis>
General coloration pale green (
<figureCitation box="[1282,1354,1152,1174]" captionStart="FIG. 37" captionStartId="135.[174,218,1655,1677]" captionTargetBox="[227,1329,204,1640]" captionTargetId="figure@135.[227,1330,204,1640]" captionTargetPageId="135" captionText="FIG. 37. Costa Rican species of Conocephalinae s. l. A. Erioloides longipennis - female ovipositing, B. Erioloides longinoi - male, C. Eriolus penicillus - female, D. Conocephalus cinereus - male, macropterous form, E. Neoconocephalus triops - male, brown form, F. Sphyrometopa atlantica - male, inset - nymph." httpUri="https://zenodo.org/record/270073/files/figure.png" pageId="115" pageNumber="112">Fig. 37</figureCitation>
C); face green, sometimes clypeus and labrum brown; pronotum sometimes with two thin, light stripes along dorsal edges, and darker, brown or reddish markings in metazona; tegmina sometimes with irregularly distributes small, black dots.
</paragraph>
</subSubSection>
<subSubSection lastPageId="116" lastPageNumber="113" pageId="115" pageNumber="112" type="discussion">
<paragraph blockId="115.[794,1384,1152,1731]" lastBlockId="116.[151,740,807,1122]" lastPageId="116" lastPageNumber="113" pageId="115" pageNumber="112">
<emphasis box="[817,930,1328,1349]" italics="true" pageId="115" pageNumber="112">Remarks.—</emphasis>
The genus 
<taxonomicName box="[1049,1120,1328,1349]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis box="[1049,1120,1328,1349]" italics="true" pageId="115" pageNumber="112">Eriolus</emphasis>
</taxonomicName>
was originally described to include only one species, 
<taxonomicName box="[1103,1297,1357,1378]" class="Insecta" family="Tettigoniidae" genus="Erioloides" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="caraibeus">
<emphasis box="[1103,1297,1357,1378]" italics="true" pageId="115" pageNumber="112">Erioloides caraibeus</emphasis>
</taxonomicName>
<collectingRegion box="[1306,1383,1357,1379]" country="Venezuela" name="Bolivar" pageId="115" pageNumber="112">Bolivar</collectingRegion>
from 
<collectingCountry box="[853,913,1386,1408]" name="Cuba" pageId="115" pageNumber="112">Cuba</collectingCountry>
. Subsequent authors included in this genus numerous additional, often unrelated species. 
<bibRefCitation author="Hebard, M." journalOrPublisher="Trans. Amer. Ent .. Soc" pageId="115" pageNumber="112" pagination="79 - 156" part="53" refString="Hebard, M. 1927. Studies in the Tettigoniidae of Panama. Trans. Amer. Ent .. Soc. 53: 79 - 156." title="Studies in the Tettigoniidae of Panama" type="journal article" year="1927">Hebard (1927)</bibRefCitation>
moved 4 of these species (
<taxonomicName authority="Pictet et Saussure" authorityName="Pictet et Saussure" class="Insecta" family="Tettigoniidae" genus="Erioloides" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="consobrinus">
<emphasis box="[1145,1290,1445,1467]" italics="true" pageId="115" pageNumber="112">E. consobrinus</emphasis>
Pictet et Saussure
</taxonomicName>
, 
<taxonomicName authority="Redtenbacher" authorityName="Redtenbacher" box="[899,1129,1474,1496]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="frater">
<emphasis box="[899,978,1475,1496]" italics="true" pageId="115" pageNumber="112">E. frater</emphasis>
Redtenbacher
</taxonomicName>
, 
<taxonomicName authority="Saussure" authorityName="Saussure" box="[1139,1377,1475,1496]" class="Insecta" family="Tettigoniidae" genus="Erioloides" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="mexicanum">
<emphasis box="[1139,1277,1475,1496]" italics="true" pageId="115" pageNumber="112">E. mexicanum</emphasis>
Saussure
</taxonomicName>
, 
<taxonomicName authority="Redtenbacher" authorityName="Redtenbacher" box="[794,1055,1504,1526]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="spiniger">
<emphasis box="[794,900,1505,1526]" italics="true" pageId="115" pageNumber="112">E. spiniger</emphasis>
Redtenbacher
</taxonomicName>
) to a new genus, 
<taxonomicName box="[1242,1338,1504,1525]" class="Insecta" family="Tettigoniidae" genus="Erioloides" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis box="[1242,1338,1504,1525]" italics="true" pageId="115" pageNumber="112">Erioloides</emphasis>
</taxonomicName>
. He also indicated the possibility that some other species 
<emphasis italics="true" pageId="115" pageNumber="112">
(
<taxonomicName authority="Redtenbacher" authorityName="Redtenbacher" class="Insecta" family="Tettigoniidae" genus="Erioloides" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="variety" species="jamaicensis" variety="longipennis">
E. jamaicensis Bruner, 
<emphasis box="[1011,1160,1562,1584]" italics="true" pageId="115" pageNumber="112">E. longipennis</emphasis>
Redtenbacher
</taxonomicName>
</emphasis>
<emphasis box="[1321,1329,1563,1584]" italics="true" pageId="115" pageNumber="112">)</emphasis>
, unknown to him, may also belong to his new genus. 
<taxonomicName box="[794,1009,1621,1642]" class="Insecta" family="Tettigoniidae" genus="Erioloides" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis box="[794,1009,1621,1642]" italics="true" pageId="115" pageNumber="112">Erioloides longipennis</emphasis>
</taxonomicName>
is formally transferred to 
<taxonomicName box="[1287,1383,1621,1642]" class="Insecta" family="Tettigoniidae" genus="Erioloides" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis box="[1287,1383,1621,1642]" italics="true" pageId="115" pageNumber="112">Erioloides</emphasis>
</taxonomicName>
in this work, while 
<taxonomicName box="[1010,1151,1651,1672]" class="Insecta" family="Tettigoniidae" genus="Erioloides" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="species" species="jamaicensis">
<emphasis box="[1010,1151,1651,1672]" italics="true" pageId="115" pageNumber="112">E. jamaicensis</emphasis>
</taxonomicName>
will be treated in an upcoming revision of the genus 
<taxonomicName box="[1153,1250,1680,1701]" class="Insecta" family="Tettigoniidae" genus="Erioloides" kingdom="Animalia" order="Orthoptera" pageId="115" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis box="[1153,1250,1680,1701]" italics="true" pageId="115" pageNumber="112">Erioloides</emphasis>
</taxonomicName>
and related genera (Naskrecki and Cohn, in prep.). Having exam- ined the 
<typeStatus box="[247,341,807,829]" pageId="116" pageNumber="113">holotype</typeStatus>
of 
<taxonomicName authority="Karny, I" authorityName="Karny, I" box="[378,647,807,829]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="116" pageNumber="113" phylum="Arthropoda" rank="species" species="nigrifrons">
<emphasis box="[378,554,807,828]" italics="true" pageId="116" pageNumber="113">Eriolus nigrifrons</emphasis>
Karny, I
</taxonomicName>
came to the conclusion that it is more closely related to the genus 
<taxonomicName box="[151,218,866,887]" class="Insecta" family="Tettigoniidae" genus="Vestria" kingdom="Animalia" order="Orthoptera" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis box="[151,218,866,887]" italics="true" pageId="116" pageNumber="113">Vestria</emphasis>
</taxonomicName>
than 
<taxonomicName box="[282,352,866,887]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis box="[282,352,866,887]" italics="true" pageId="116" pageNumber="113">Eriolus</emphasis>
</taxonomicName>
, and is here formally transferred to the former.
</paragraph>
<caption ID-Table-UUID="BF95664FA35EFFBA16EDFF0CFD123D6C" httpUri="http://table.plazi.org/id/BF95664FA35EFFBA16EDFF0CFD123D6C" pageId="116" pageNumber="113" targetBox="[151,1344,293,765]" targetIsTable="true" targetPageId="116">
<caption box="[151,702,207,229]" pageId="116" pageNumber="113">
<paragraph blockId="116.[151,739,207,258]" box="[151,702,207,229]" pageId="116" pageNumber="113">
<emphasis bold="true" box="[151,267,207,228]" pageId="116" pageNumber="113">TABLE 23.</emphasis>
Body measurements of species of 
<taxonomicName box="[631,702,207,228]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">Eriolus</taxonomicName>
</paragraph>
</caption>
<paragraph blockId="116.[151,739,207,258]" box="[151,739,236,258]" pageId="116" pageNumber="113">(all measurements are lengths in mm: range, mean±SD)</paragraph>
</caption>
<paragraph pageId="116" pageNumber="113">
<table box="[151,1344,293,765]" gridcols="6" gridrows="5" pageId="116" pageNumber="113">
<tr box="[151,1344,293,315]" gridrow="0" pageId="116" pageNumber="113">
<th box="[151,264,293,315]" gridcol="0" gridrow="0" pageId="116" pageNumber="113">Species</th>
<th box="[287,476,293,315]" gridcol="1" gridrow="0" pageId="116" pageNumber="113">Body with wings</th>
<th box="[529,718,293,315]" gridcol="2" gridrow="0" pageId="116" pageNumber="113">Tegmen</th>
<th box="[756,909,293,315]" gridcol="3" gridrow="0" pageId="116" pageNumber="113">Pronotum</th>
<th box="[944,1134,293,315]" gridcol="4" gridrow="0" pageId="116" pageNumber="113">Hind femur</th>
<th box="[1156,1344,293,315]" gridcol="5" gridrow="0" pageId="116" pageNumber="113">Ovipositor</th>
</tr>
<tr box="[151,1344,351,428]" gridrow="1" pageId="116" pageNumber="113">
<th box="[151,264,351,428]" gridcol="0" gridrow="1" pageId="116" pageNumber="113">
<taxonomicName box="[151,264,351,372]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="116" pageNumber="113" phylum="Arthropoda" rank="species" species="acutipennis">
<emphasis box="[151,264,351,372]" italics="true" pageId="116" pageNumber="113">acutipennis</emphasis>
</taxonomicName>
male female
</th>
<td box="[287,476,351,428]" gridcol="1" gridrow="1" pageId="116" pageNumber="113">22.6 27.9</td>
<td box="[529,718,351,428]" gridcol="2" gridrow="1" pageId="116" pageNumber="113">15.5 21.5</td>
<td box="[756,909,351,428]" gridcol="3" gridrow="1" pageId="116" pageNumber="113">5.0 4.8</td>
<td box="[944,1134,351,428]" gridcol="4" gridrow="1" pageId="116" pageNumber="113">12.1 12.1</td>
<td box="[1156,1344,351,428]" gridcol="5" gridrow="1" pageId="116" pageNumber="113">na 6.4</td>
</tr>
<tr box="[151,1344,462,540]" gridrow="2" pageId="116" pageNumber="113">
<th box="[151,264,462,540]" gridcol="0" gridrow="2" pageId="116" pageNumber="113">
<taxonomicName box="[151,224,462,483]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="116" pageNumber="113" phylum="Arthropoda" rank="species" species="aculeus">
<emphasis box="[151,224,462,483]" italics="true" pageId="116" pageNumber="113">aculeus</emphasis>
</taxonomicName>
male female
</th>
<td box="[287,476,462,540]" gridcol="1" gridrow="2" pageId="116" pageNumber="113">27.4-31.4, 30±1.4 31.8-36.2, 34±1.9</td>
<td box="[529,718,462,540]" gridcol="2" gridrow="2" pageId="116" pageNumber="113">21.2-24.3, 23.2±1.1 25-28.2, 26.6±1.3</td>
<td box="[756,909,462,540]" gridcol="3" gridrow="2" pageId="116" pageNumber="113">4.5-4.9, 4.7±0.2 4.8-5.3, 5±0.2</td>
<td box="[944,1134,462,540]" gridcol="4" gridrow="2" pageId="116" pageNumber="113">11.4-13.5, 12.5±0.7 13.1-13.6, 13.4±0.2</td>
<td box="[1156,1344,462,540]" gridcol="5" gridrow="2" pageId="116" pageNumber="113">na 8.4-10.1, 9.3±0.7</td>
</tr>
<tr box="[151,1344,574,652]" gridrow="3" pageId="116" pageNumber="113">
<th box="[151,264,574,652]" gridcol="0" gridrow="3" pageId="116" pageNumber="113">
<taxonomicName box="[151,245,574,595]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="116" pageNumber="113" phylum="Arthropoda" rank="species" species="penicillus">
<emphasis box="[151,245,574,595]" italics="true" pageId="116" pageNumber="113">penicillus</emphasis>
</taxonomicName>
male female
</th>
<td box="[287,476,574,652]" gridcol="1" gridrow="3" pageId="116" pageNumber="113">29.6-31.9, 30.8±1.6 32.8-36.4, 34.9±1.3</td>
<td box="[529,718,574,652]" gridcol="2" gridrow="3" pageId="116" pageNumber="113">24.5-26.4, 25.5±1.3 26.1-30, 28.2±1.3</td>
<td box="[756,909,574,652]" gridcol="3" gridrow="3" pageId="116" pageNumber="113">4.6-4.7, 4.7±0.1 4.7-5.1, 4.9±0.1</td>
<td box="[944,1134,574,652]" gridcol="4" gridrow="3" pageId="116" pageNumber="113">11.7-12.8, 12.3±0.8 12.5-13.9, 13.3±0.5</td>
<td box="[1156,1344,574,652]" gridcol="5" gridrow="3" pageId="116" pageNumber="113">na 10.5-12.1, 11.1±0.6</td>
</tr>
<tr box="[151,1344,686,765]" gridrow="4" pageId="116" pageNumber="113">
<th box="[151,264,686,765]" gridcol="0" gridrow="4" pageId="116" pageNumber="113">
<heading box="[151,260,686,707]" centered="true" fontSize="9" level="14" pageId="116" pageNumber="113" reason="4">
<taxonomicName box="[151,260,686,707]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="116" pageNumber="113" phylum="Arthropoda" rank="species" species="duplidentis">
<emphasis box="[151,260,686,707]" italics="true" pageId="116" pageNumber="113">duplidentis</emphasis>
</taxonomicName>
</heading>
male female
</th>
<td box="[287,476,686,765]" gridcol="1" gridrow="4" pageId="116" pageNumber="113">32-34.3, 32.7±0.9 33-36.3, 34.4±1.2</td>
<td box="[529,718,686,765]" gridcol="2" gridrow="4" pageId="116" pageNumber="113">25-27.8, 26.5±1.1 26-29.4, 27.7±1.2</td>
<td box="[756,909,686,765]" gridcol="3" gridrow="4" pageId="116" pageNumber="113">5-5.7, 5.2±0.3 4.8-5.4, 5.1±0.2</td>
<td box="[944,1134,686,765]" gridcol="4" gridrow="4" pageId="116" pageNumber="113">12.7-13.6, 13.2±0.4 13.1-14.9, 13.8±0.7</td>
<td box="[1156,1344,686,765]" gridcol="5" gridrow="4" pageId="116" pageNumber="113">na 6.4-7.3, 6.9±0.3</td>
</tr>
</table>
</paragraph>
<paragraph blockId="116.[151,740,807,1122]" pageId="116" pageNumber="113">
In 
<collectingCountry box="[205,322,924,946]" name="Costa Rica" pageId="116" pageNumber="113">Costa Rica</collectingCountry>
, the genus 
<taxonomicName box="[450,521,924,945]" class="Insecta" family="Tettigoniidae" genus="Eriolus" kingdom="Animalia" order="Orthoptera" pageId="116" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis box="[450,521,924,945]" italics="true" pageId="116" pageNumber="113">Eriolus</emphasis>
</taxonomicName>
is represented by 4 species. Both males and females can be easily identified to species based on their unique genitalic characters. Nothing is known about the biology and acoustic behavior these species. They sometimes come to light and most of them were collected by canopy fogging in primary, lowland wet forests.
</paragraph>
</subSubSection>
</treatment>
</document>