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<document ID-DOI="http://doi.org/10.5281/zenodo.6280456" ID-GBIF-Dataset="647046fe-0ee5-401d-b4f8-aea0355fde7f" ID-GBIF-Taxon="124407419" ID-Zenodo-Dep="6280456" checkinTime="1471630429166" checkinUser="plazi" docAuthor="Piotr Naskrecki" docDate="2000" docId="634387D1A356FFB31579FD12FBB638EE" docLanguage="en" docName="Naskrecki_KCR_2000.pdf.imf" docOrigin="Philadelphia, PA: The Orthopterists Society at the Academy of Natural Sciences of Philadelphia," docStyle="DocumentStyle{}" docTitle="Subria sylvestris Naskrecki and Morris, sp. n." docType="treatment" docVersion="8" lastPageId="125" lastPageNumber="122" masterDocId="9F7AFFA9A32AFFCE167AFFC3FFF13C6E" masterDocTitle="Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato)." pageId="124" pageNumber="121" updateTime="1645797358202" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Katydids of Costa Rica / Vol. 1, Systematics and bioacoustics of the cone-head katydids (Orthoptera: Tettigoniidae: Conocephalinae sensu lato).</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Piotr Naskrecki</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:originInfo>
<mods:dateIssued>2000</mods:dateIssued>
<mods:publisher>The Orthopterists Society at the Academy of Natural Sciences of Philadelphia,</mods:publisher>
<mods:place>
<mods:placeTerm>Philadelphia, PA</mods:placeTerm>
</mods:place>
</mods:originInfo>
<mods:classification>book</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.270035</mods:identifier>
<mods:identifier type="GBIF-Dataset">647046fe-0ee5-401d-b4f8-aea0355fde7f</mods:identifier>
<mods:identifier type="Zenodo-Dep">270035</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6280456" ID-GBIF-Taxon="124407419" ID-Zenodo-Dep="6280456" LSID="urn:lsid:plazi:treatment:634387D1A356FFB31579FD12FBB638EE" httpUri="http://treatment.plazi.org/id/634387D1A356FFB31579FD12FBB638EE" lastPageId="125" lastPageNumber="122" pageId="124" pageNumber="121">
<subSubSection box="[771,1258,721,743]" pageId="124" pageNumber="121" type="nomenclature">
<paragraph blockId="124.[771,1258,721,799]" box="[771,1258,721,743]" pageId="124" pageNumber="121">
<taxonomicName authority="Naskrecki and Morris" authorityName="Naskrecki and Morris" box="[771,1192,721,743]" class="Insecta" family="Tettigoniidae" genus="Subria" kingdom="Animalia" order="Orthoptera" pageId="124" pageNumber="121" phylum="Arthropoda" rank="species" species="sylvestris" status="sp. n.">
<emphasis bold="true" box="[771,952,721,743]" italics="true" pageId="124" pageNumber="121">Subria sylvestris</emphasis>
Naskrecki and Morris
</taxonomicName>
,
<taxonomicNameLabel box="[1201,1258,722,743]" pageId="124" pageNumber="121" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="124" pageNumber="121" type="description">
<paragraph blockId="124.[771,1258,721,799]" box="[771,1085,749,771]" pageId="124" pageNumber="121">
Common name: Forest
<taxonomicName box="[1019,1085,749,771]" class="Insecta" family="Tettigoniidae" genus="Subria" kingdom="Animalia" order="Orthoptera" pageId="124" pageNumber="162" phylum="Arthropoda" rank="genus">subria</taxonomicName>
</paragraph>
<paragraph blockId="124.[771,1258,721,799]" box="[771,1243,777,799]" pageId="124" pageNumber="121">
<figureCitation box="[771,851,777,799]" captionStart="FIG. 33" captionStartId="123.[174,218,1626,1648]" captionTargetBox="[180,1374,189,1618]" captionTargetId="figure@123.[180,1374,189,1621]" captionTargetPageId="123" captionText="FIG. 33. Subria spp. A. S. sylvestris - male, habitus, B. S. sylvestris - male head and pronotum, dorsal view, C. S. scutellaris - ditto, D. S. crassicerca - ditto, E. S. scutellaris - male head and pronotum, lateral view, F. S. crassicerca - ditto, G. S. scutellaris - apex of right titillator, H. S. crassicerca - ditto, I. S. sylvestris - titillators, J. S. sylvestris - ovipositor, K. S. sylvestris - female subgenital plate, L. S. nitida - ditto." httpUri="https://zenodo.org/record/270069/files/figure.png" pageId="124" pageNumber="121">Figs. 33</figureCitation>
A-B, 33I-K, 36H, 38D, 58C-D,
<figureCitation box="[1164,1243,777,799]" captionStart="MAPS 13 - 18" captionStartId="159.[174,246,1632,1654]" captionTargetBox="[253,1276,174,1610]" captionTargetId="figure@159.[253,1276,174,1610]" captionTargetPageId="159" captionText="MAPS 13 - 18. Distribution of species of Conocephalinae s. l. in Costa Rica. 13. Sphyrometopa femorata, S. atlantica, Eppia truncatipennis, 14. Erioloides longipennis, E. macrocephalus, E. longinoi, E. brevipennis, 15. Erioloides, consobrinus, E. acutidentis, E. latiscobinus, E. sikesi, 16. Eriolus aculeus, E. acutipennis, E. duplidentis, E. penicillus, 17. Subria sylvestris, S. crassicerca, S. scutellaris, 18. Podacanthophorus alas, P. nelciae, P. vargasi, P. maylinae." httpUri="https://zenodo.org/record/270097/files/figure.png" pageId="124" pageNumber="121">Map 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="124" pageNumber="121" type="materials_examined">
<paragraph blockId="124.[771,1334,833,912]" box="[771,1289,833,855]" pageId="124" pageNumber="121">
<typeStatus box="[771,823,834,855]" pageId="124" pageNumber="121">Type</typeStatus>
locality:
<collectingCountry box="[920,1035,833,855]" name="Costa Rica" pageId="124" pageNumber="121">Costa Rica</collectingCountry>
:
<collectingRegion box="[1043,1129,833,855]" country="Costa Rica" name="Heredia" pageId="124" pageNumber="121">Heredia</collectingRegion>
Prov., La Selva
</paragraph>
<paragraph blockId="124.[771,1334,833,912]" pageId="124" pageNumber="121">
Biological Station;
<typeStatus box="[1026,1073,861,882]" pageId="124" pageNumber="121">type</typeStatus>
depository: Academy of Natural Sciences, Philadelphia
<typeStatus box="[1177,1271,890,912]" pageId="124" pageNumber="121">holotype</typeStatus>
male
</paragraph>
</subSubSection>
<subSubSection lastPageId="125" lastPageNumber="122" pageId="124" pageNumber="121" type="description">
<paragraph blockId="124.[771,1361,949,1440]" pageId="124" pageNumber="121">
<emphasis box="[794,1043,949,970]" italics="true" pageId="124" pageNumber="121">Diagnostic description.—</emphasis>
General characteristics as described above. Male pronotum short, barely covering basal half of stridulatory apparatus (
<figureCitation box="[1162,1235,1007,1029]" captionStart="FIG. 33" captionStartId="123.[174,218,1626,1648]" captionTargetBox="[180,1374,189,1618]" captionTargetId="figure@123.[180,1374,189,1621]" captionTargetPageId="123" captionText="FIG. 33. Subria spp. A. S. sylvestris - male, habitus, B. S. sylvestris - male head and pronotum, dorsal view, C. S. scutellaris - ditto, D. S. crassicerca - ditto, E. S. scutellaris - male head and pronotum, lateral view, F. S. crassicerca - ditto, G. S. scutellaris - apex of right titillator, H. S. crassicerca - ditto, I. S. sylvestris - titillators, J. S. sylvestris - ovipositor, K. S. sylvestris - female subgenital plate, L. S. nitida - ditto." httpUri="https://zenodo.org/record/270069/files/figure.png" pageId="124" pageNumber="121">Fig. 33</figureCitation>
B). Tegmina surpassing apices of hind
<taxonomicName box="[1055,1130,1037,1059]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="124" pageNumber="121" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
by about 1/3 of their length (
<figureCitation box="[852,923,1066,1088]" captionStart="FIG. 33" captionStartId="123.[174,218,1626,1648]" captionTargetBox="[180,1374,189,1618]" captionTargetId="figure@123.[180,1374,189,1621]" captionTargetPageId="123" captionText="FIG. 33. Subria spp. A. S. sylvestris - male, habitus, B. S. sylvestris - male head and pronotum, dorsal view, C. S. scutellaris - ditto, D. S. crassicerca - ditto, E. S. scutellaris - male head and pronotum, lateral view, F. S. crassicerca - ditto, G. S. scutellaris - apex of right titillator, H. S. crassicerca - ditto, I. S. sylvestris - titillators, J. S. sylvestris - ovipositor, K. S. sylvestris - female subgenital plate, L. S. nitida - ditto." httpUri="https://zenodo.org/record/270069/files/figure.png" pageId="124" pageNumber="121">Fig. 33</figureCitation>
A, 36H) (in one female from La Selva Biological Station tegmina are abnormally long, surpassing the apices of the hind
<taxonomicName box="[992,1066,1125,1147]" class="Insecta" family="Tettigoniidae" genus="Sphyrometopa" kingdom="Animalia" order="Orthoptera" pageId="124" pageNumber="121" phylum="Arthropoda" rank="species" species="femora">femora</taxonomicName>
by more than half the length of the tegmen). Stridulatory file of male weakly curved,
<quantity box="[771,854,1183,1205]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="124" pageNumber="121" unit="mm" value="2.1">2.1 mm</quantity>
long, with 170-179 (n=3) wide and thin teeth, maximum width of file 142 μm (
<figureCitation box="[1125,1198,1213,1235]" captionStart="FIG. 51" captionStartId="149.[174,218,1684,1706]" captionTargetBox="[212,1337,204,1665]" captionTargetId="figure@149.[212,1338,204,1667]" captionTargetPageId="149" captionText="FIG. 51. Stridulatory files of Costa Rican species of Conocephalinae s. l. A. Subria sylvestris, C. Podacanthophorus maylinae, C. Podacanthophorus alas, D. Podacanthophorus vargasi, E. Pluviasilva levis, F. Pluviasilva mexicana." httpUri="https://zenodo.org/record/270087/files/figure.png" pageId="124" pageNumber="121">Fig. 51</figureCitation>
A); teeth of file uniform in thickness, evenly spaced and smoothly narrowing towards proximal end. Male cercus weakly incurved, with incised apex, and large, blunt, ventral tooth subapically (
<figureCitation box="[962,1032,1330,1352]" captionStart="FIG. 38" captionStartId="136.[151,195,1655,1677]" captionTargetBox="[219,1298,204,1639]" captionTargetId="figure@136.[219,1299,204,1640]" captionTargetPageId="136" captionText="FIG. 38. Eggs and cerci of Costa Rican species of Conocephalinae s. l. A-C. eggs, with enlarged fragments of the chorion: A. Copiphora rhinoceros, B. Lirometopum coronatum, C. Erioloides longipennis, hatched egg, D-F. male right cerci of Subria spp., dorsal views: D. S. sylvestris, E. S. scutellaris, F. S. crassicerca." httpUri="https://zenodo.org/record/270074/files/figure.png" pageId="124" pageNumber="121">Fig. 38</figureCitation>
D). Ovipositor distinctly curved upwards and slightly widened midlength, slightly longer than half of hind femur (ratio ovipositor/hind femur 0.65- 0.74) (
<figureCitation box="[835,906,1418,1440]" captionStart="FIG. 33" captionStartId="123.[174,218,1626,1648]" captionTargetBox="[180,1374,189,1618]" captionTargetId="figure@123.[180,1374,189,1621]" captionTargetPageId="123" captionText="FIG. 33. Subria spp. A. S. sylvestris - male, habitus, B. S. sylvestris - male head and pronotum, dorsal view, C. S. scutellaris - ditto, D. S. crassicerca - ditto, E. S. scutellaris - male head and pronotum, lateral view, F. S. crassicerca - ditto, G. S. scutellaris - apex of right titillator, H. S. crassicerca - ditto, I. S. sylvestris - titillators, J. S. sylvestris - ovipositor, K. S. sylvestris - female subgenital plate, L. S. nitida - ditto." httpUri="https://zenodo.org/record/270069/files/figure.png" pageId="124" pageNumber="121">Fig. 33</figureCitation>
J).
</paragraph>
<paragraph blockId="124.[771,1360,1477,1733]" lastBlockId="125.[174,763,206,424]" lastPageId="125" lastPageNumber="122" pageId="124" pageNumber="121">
<emphasis box="[794,935,1477,1498]" italics="true" pageId="124" pageNumber="121">Coloration —</emphasis>
Unlike other species of the genus,
<taxonomicName class="Insecta" family="Tettigoniidae" genus="Subria" kingdom="Animalia" order="Orthoptera" pageId="124" pageNumber="121" phylum="Arthropoda" rank="species" species="sylvestris">
<emphasis italics="true" pageId="124" pageNumber="121">S. sylvestris</emphasis>
</taxonomicName>
exhibits a remarkable color dimorphism, having both green and brown (
<figureCitation box="[1070,1143,1535,1557]" captionStart="FIG. 36" captionStartId="134.[151,196,1655,1677]" captionTargetBox="[239,1270,204,1639]" captionTargetId="figure@134.[239,1270,204,1640]" captionTargetPageId="134" captionText="FIG. 36. Costa Rican species of Conocephalinae s. l. A. Conocephalus angustifrons - male, B. Caulopsis cuspidata - female, C. Acantheremus colwelli - male, D. Vestria punctata - male and female, in copula, E. Copiphora hastata - male, F. Podacanthophorus vargasi - female, G. Copiphora rhinoceros - male eating a snail, H. Subria sylvestris - male, brown form." httpUri="https://zenodo.org/record/270072/files/figure.png" pageId="124" pageNumber="121">Fig. 36</figureCitation>
H) color morphs occurring within the same population. The material examined included 35% of brown individuals and 65% of green ones (including olive green ones). There seem to be no correlation between the color and season or geographical distribution. Most individuals of the species lack a wide, dark band on the pronotum, typical of the remaining two species of the genus. Instead, the sides of pronotum have two, thin, white or light brown stripes. Occasionally the brown band, or intermediate stages are present. Tegmina often have several irregularly distributed, diffused dark spots. The outer pagina of the hind femur may or may not have a distinct reticulate pattern (the pattern is present in most brown individuals but is only rarely distinct in green ones).
</paragraph>
<paragraph blockId="125.[196,468,458,480]" box="[196,468,458,480]" pageId="125" pageNumber="122">
<emphasis box="[196,368,459,480]" italics="true" pageId="125" pageNumber="122">Measurements.—</emphasis>
<tableCitation box="[375,464,458,480]" captionStart="TABLE 25" captionStartId="124.[151,231,207,228]" captionTargetBox="[151,1323,293,661]" captionText="TABLE 25. Body measurements of Costa Rican species of Subria (all measurements are lengths in mm: range, mean ± SD)" httpUri="http://table.plazi.org/id/BF95664FA356FFB216EDFF0CFD123D6C" pageId="125" pageNumber="122" tableUuid="BF95664FA356FFB216EDFF0CFD123D6C">Table 25</tableCitation>
.
</paragraph>
<paragraph blockId="125.[174,762,514,928]" pageId="125" pageNumber="122">
<emphasis box="[196,346,515,536]" italics="true" pageId="125" pageNumber="122">Bioacoustics.—</emphasis>
The call of
<taxonomicName box="[465,581,514,536]" class="Insecta" family="Tettigoniidae" genus="Subria" kingdom="Animalia" order="Orthoptera" pageId="125" pageNumber="122" phylum="Arthropoda" rank="species" species="sylvestris">
<emphasis box="[465,581,514,536]" italics="true" pageId="125" pageNumber="122">S. sylvestris</emphasis>
</taxonomicName>
is low duty, with sound production accounting for only about 14% of the calling cycle. It consists of short (147-175 ms) phrases produced at intervals of 0.85 -
<geoCoordinate box="[490,539,598,619]" direction="south" orientation="latitude" pageId="125" pageNumber="122" precision="5555" value="-4.0">4.0 S</geoCoordinate>
, each phrase consists of two short (38 - 78 ms) pulses (
<figureCitation box="[532,614,626,648]" captionStart="FIG. 58" captionStartId="156.[151,195,1655,1677]" captionTargetBox="[234,1272,204,1607]" captionTargetId="figure@156.[234,1272,204,1607]" captionTargetPageId="156" captionText="FIG. 58. Oscillograms of male airborne calls of Costa Rican species of Conocephalinae s. l. A. Vestria puncta, B. ditto, 0.5 s fragment, C. Subria sylvestris, D. ditto, 0.5 s fragment, E. Subria scutellaris, F. ditto, 0.5 s fragment, G. Podacanthophorus alas, 0.5 s fragment." httpUri="https://zenodo.org/record/270094/files/figure.png" pageId="125" pageNumber="122">Figs. 58</figureCitation>
C-D) (all data apply to songs recorded at temperature 26½C). Typical of most
<taxonomicName box="[233,375,682,704]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="125" pageNumber="122" phylum="Arthropoda" rank="family">Tettigoniidae</taxonomicName>
, it is a low Q call, with most energy allocated between 6 and 12 kHz in the audible range, and above 20 kHz (the upper frequency range of the call is unknown because call components above 22 kHz were not recorded due to technical constraints of the equipment used). Males start calling at dusk and continue probably all night. Calls are undetectable in field conditions, even on relatively quiet nights. In laboratory conditions the calls are audible from a distance of about
<quantity box="[653,733,906,928]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="125" pageNumber="122" unit="m" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection pageId="125" pageNumber="122" type="biology_ecology">
<paragraph blockId="125.[174,763,962,1376]" pageId="125" pageNumber="122">
<emphasis box="[196,476,962,984]" italics="true" pageId="125" pageNumber="122">
Remarks —
<taxonomicName box="[315,476,962,983]" class="Insecta" family="Tettigoniidae" genus="Subria" kingdom="Animalia" order="Orthoptera" pageId="125" pageNumber="122" phylum="Arthropoda" rank="species" species="sylvestris">Subria sylvestris</taxonomicName>
</emphasis>
is one of the most common species of
<taxonomicName box="[288,435,990,1012]" class="Insecta" family="Tettigoniidae" kingdom="Animalia" order="Orthoptera" pageId="125" pageNumber="122" phylum="Arthropoda" rank="tribe" tribe="Copiphorinae">Copiphorinae</taxonomicName>
in the lowland rain forests of
<collectingCountry box="[174,292,1018,1040]" name="Costa Rica" pageId="125" pageNumber="122">Costa Rica</collectingCountry>
, occurring in both primary and secondary forests at elevations ranging from sea level to about
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="125" pageNumber="122" unit="m" value="700.0">700 m</quantity>
(
<figureCitation box="[208,290,1074,1096]" captionStart="MAPS 13 - 18" captionStartId="159.[174,246,1632,1654]" captionTargetBox="[253,1276,174,1610]" captionTargetId="figure@159.[253,1276,174,1610]" captionTargetPageId="159" captionText="MAPS 13 - 18. Distribution of species of Conocephalinae s. l. in Costa Rica. 13. Sphyrometopa femorata, S. atlantica, Eppia truncatipennis, 14. Erioloides longipennis, E. macrocephalus, E. longinoi, E. brevipennis, 15. Erioloides, consobrinus, E. acutidentis, E. latiscobinus, E. sikesi, 16. Eriolus aculeus, E. acutipennis, E. duplidentis, E. penicillus, 17. Subria sylvestris, S. crassicerca, S. scutellaris, 18. Podacanthophorus alas, P. nelciae, P. vargasi, P. maylinae." httpUri="https://zenodo.org/record/270097/files/figure.png" pageId="125" pageNumber="122">Map 17</figureCitation>
). It can be found on broad leafed understory vegetation as well as in the canopy. In the field individuals of this species have been seen eating generative parts of plants (e.g. flowers of
<taxonomicName box="[435,502,1159,1180]" class="Liliopsida" family="Costaceae" genus="Costus" kingdom="Plantae" order="Zingiberales" pageId="125" pageNumber="122" phylum="Tracheophyta" rank="genus">
<emphasis box="[435,502,1159,1180]" italics="true" pageId="125" pageNumber="122">Costus</emphasis>
</taxonomicName>
sp., fallen fruits), decaying leaves, tree bark (
<taxonomicName box="[491,763,1186,1207]" class="Magnoliopsida" family="Fabaceae" genus="Pentaclethra" kingdom="Plantae" order="Fabales" pageId="125" pageNumber="122" phylum="Tracheophyta" rank="species" species="macroloba">
<emphasis box="[491,763,1186,1207]" italics="true" pageId="125" pageNumber="122">Pentaclethra macroloba</emphasis>
</taxonomicName>
(
<taxonomicName box="[181,316,1214,1236]" class="Magnoliopsida" family="Mimosaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="125" pageNumber="122" phylum="Magnoliophyta" rank="family">Mimosaceae</taxonomicName>
)), and occasionally insects. In laboratory conditions they can thrive for months on a diet consisting of lettuce, carrot, and apples.
</paragraph>
</subSubSection>
<subSubSection pageId="125" pageNumber="122" type="discussion">
<paragraph blockId="125.[174,763,962,1376]" pageId="125" pageNumber="122">
This new species is very closely related to
<taxonomicName box="[622,701,1298,1320]" class="Insecta" family="Tettigoniidae" genus="Subria" kingdom="Animalia" order="Orthoptera" pageId="125" pageNumber="122" phylum="Arthropoda" rank="species" species="nitida">
<emphasis box="[622,701,1298,1320]" italics="true" pageId="125" pageNumber="122">S. nitida</emphasis>
</taxonomicName>
, from which it differs in the shape of male cerci and female subgenital plate (
<figureCitation box="[357,428,1354,1376]" captionStart="FIG. 33" captionStartId="123.[174,218,1626,1648]" captionTargetBox="[180,1374,189,1618]" captionTargetId="figure@123.[180,1374,189,1621]" captionTargetPageId="123" captionText="FIG. 33. Subria spp. A. S. sylvestris - male, habitus, B. S. sylvestris - male head and pronotum, dorsal view, C. S. scutellaris - ditto, D. S. crassicerca - ditto, E. S. scutellaris - male head and pronotum, lateral view, F. S. crassicerca - ditto, G. S. scutellaris - apex of right titillator, H. S. crassicerca - ditto, I. S. sylvestris - titillators, J. S. sylvestris - ovipositor, K. S. sylvestris - female subgenital plate, L. S. nitida - ditto." httpUri="https://zenodo.org/record/270069/files/figure.png" pageId="125" pageNumber="122">Fig. 33</figureCitation>
L).
</paragraph>
</subSubSection>
<subSubSection pageId="125" pageNumber="122" type="etymology">
<paragraph blockId="125.[174,762,1410,1460]" pageId="125" pageNumber="122">
<emphasis box="[196,331,1410,1431]" italics="true" pageId="125" pageNumber="122">Etymology.—</emphasis>
The specific epithet simply reflects the preferred habitat of the new species.
</paragraph>
</subSubSection>
<subSubSection pageId="125" pageNumber="122" type="materials_examined">
<paragraph blockId="125.[174,762,1495,1742]" lastBlockId="125.[794,1384,206,1152]" pageId="125" pageNumber="122">
<emphasis box="[196,373,1495,1516]" italics="true" pageId="125" pageNumber="122">Material examined</emphasis>
(
<typeStatus box="[391,484,1495,1517]" pageId="125" pageNumber="122">holotype</typeStatus>
and
<typeStatus box="[533,637,1496,1517]" pageId="125" pageNumber="122">paratypes</typeStatus>
).
<emphasis box="[647,673,1496,1517]" italics="true" pageId="125" pageNumber="122"></emphasis>
<emphasis bold="true" pageId="125" pageNumber="122">
<collectingCountry name="Costa Rica" pageId="125" pageNumber="122">COSTA RICA</collectingCountry>
:
</emphasis>
Las Ánimas,
<date box="[378,512,1524,1546]" pageId="125" pageNumber="122" value="1930-04-18">18 April 1930</date>
(coll. C.H. Lankester) -
<specimenCount pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[259,327,1552,1574]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="125" pageNumber="122">ANSP</collectionCode>
);
<emphasis bold="true" box="[346,498,1551,1573]" pageId="125" pageNumber="122">
<collectingRegion box="[346,437,1551,1573]" country="Costa Rica" name="Alajuela" pageId="125" pageNumber="122">Alajuela</collectingRegion>
Prov.
</emphasis>
, Sector San Ramón, P.N.
<collectingRegion box="[174,297,1580,1601]" country="Costa Rica" name="Guanacaste" pageId="125" pageNumber="122">Guanacaste</collectingRegion>
, elev.
<quantity box="[363,429,1580,1602]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="125" pageNumber="122" unit="m" value="620.0">620 m</quantity>
,
<date box="[442,762,1579,1602]" pageId="125" pageNumber="122" value="1994-08-15" valueMax="1994-09-02" valueMin="1994-08-15">15 August - 2 September 1994</date>
(coll. H. García) -
<specimenCount box="[369,469,1608,1630]" pageId="125" pageNumber="122" type="female">2 females</specimenCount>
(
<collectionCode box="[485,550,1608,1630]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
);
<emphasis bold="true" box="[570,756,1607,1629]" pageId="125" pageNumber="122">
<collectingRegion box="[570,694,1607,1629]" country="Costa Rica" name="Guanacaste" pageId="125" pageNumber="122">Guanacaste</collectingRegion>
Prov.
</emphasis>
, Estac. Pitilla,
<quantity box="[316,369,1636,1658]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="125" pageNumber="122" unit="km" value="9.0">9 km</quantity>
S. Santa Cecilia, elev.
<quantity box="[606,672,1636,1657]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="125" pageNumber="122" unit="m" value="700.0">700 m</quantity>
,
<date pageId="125" pageNumber="122" value="1993-06-15">15 June 1993</date>
(coll. C. Moraga) -
<specimenCount box="[420,518,1664,1686]" pageId="125" pageNumber="122" type="female">9 females</specimenCount>
(
<collectionCode box="[530,595,1664,1686]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); Estac. Pitilla,
<quantity metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="125" pageNumber="122" unit="km" value="9.0">9 km</quantity>
S. Santa Cecilia, P.N.
<collectingRegion box="[435,557,1692,1713]" country="Costa Rica" name="Guanacaste" pageId="125" pageNumber="122">Guanacaste</collectingRegion>
, elev.
<quantity box="[621,686,1692,1713]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="125" pageNumber="122" unit="m" value="700.0">700 m</quantity>
,
<date pageId="125" pageNumber="122" value="1993-06-03">3 June 1993</date>
(coll. C. Moraga) -
<specimenCount box="[425,514,1720,1742]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[527,592,1720,1742]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); Estac. Pitilla,
<quantity box="[750,830,206,1741]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="125" pageNumber="122" unit="km" value="9.0">9 km</quantity>
St. Cecilia, elev.
<quantity box="[1029,1100,206,228]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="125" pageNumber="122" unit="m" value="700.0">700 m</quantity>
,
<date box="[1114,1254,206,228]" pageId="125" pageNumber="122" value="1988-07-15">15 July 1988</date>
(coll. GNP Biodiversity Survey) -
<specimenCount box="[1016,1082,234,256]" pageId="125" pageNumber="122" type="male">1 male</specimenCount>
(
<collectionCode box="[1092,1156,234,256]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); same locality,
<date pageId="125" pageNumber="122" value="1989-05-15">15 May 1989</date>
(coll. GNP Biodiversity Survey) -
<specimenCount box="[1175,1240,262,284]" pageId="125" pageNumber="122" type="male">1 male</specimenCount>
(
<collectionCode box="[1251,1315,262,284]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); same locality,
<date box="[878,1035,290,312]" pageId="125" pageNumber="122" value="1994-08-15">15 August 1994</date>
(coll. C. Moraga) -
<specimenCount box="[1228,1295,290,312]" pageId="125" pageNumber="122" type="male">1 male</specimenCount>
(
<collectionCode box="[1307,1372,290,312]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); Estac. Pitilla,
<quantity box="[930,981,318,340]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="125" pageNumber="122" unit="km" value="9.0">9 km</quantity>
S Santa Cecilia, elev.
<quantity box="[1197,1260,318,340]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="125" pageNumber="122" unit="m" value="700.0">700 m</quantity>
,
<date pageId="125" pageNumber="122" value="1989-09-15">15 September 1989</date>
(coll. C. Moraga and P. Rios) -
<specimenCount box="[1205,1293,346,368]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[1306,1371,346,368]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); Estac. Pitilla,
<quantity box="[937,991,374,396]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="125" pageNumber="122" unit="km" value="9.0">9 km</quantity>
S Santa Cecilia, elev.
<quantity box="[1224,1291,374,396]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="125" pageNumber="122" unit="m" value="700.0">700 m</quantity>
,
<date pageId="125" pageNumber="122" value="1991-05-15">15 May 1991</date>
(coll. P. Rios, C. Moraga and R. Blanco) -
<specimenCount box="[1292,1382,402,424]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[801,869,430,452]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
);
<emphasis bold="true" box="[890,1043,430,452]" pageId="125" pageNumber="122">
<collectingRegion box="[890,979,430,452]" country="Costa Rica" name="Heredia" pageId="125" pageNumber="122">Heredia</collectingRegion>
Prov.
</emphasis>
, Estac. Magsasay, P.N. Braulio Carillo, elev.
<quantity box="[931,996,459,480]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="125" pageNumber="122" unit="m" value="200.0">200 m</quantity>
,
<date box="[1006,1138,458,480]" pageId="125" pageNumber="122" value="1990-06-15">15 June 1990</date>
(coll. E. Alcazar) -
<specimenCount pageId="125" pageNumber="122" type="female">4 females</specimenCount>
(
<collectionCode box="[870,935,486,508]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); Puerto Viejo, La Selva Biological Station, elev.
<quantity box="[846,953,514,536]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="125" pageNumber="122" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50 - 150 m</quantity>
,
<geoCoordinate box="[963,1057,514,536]" direction="north" orientation="latitude" pageId="125" pageNumber="122" precision="925" value="10.433333">10° 26' N</geoCoordinate>
,
<geoCoordinate box="[1067,1152,515,536]" direction="west" orientation="longitude" pageId="125" pageNumber="122" precision="925" value="-84.01667">84° 1' W</geoCoordinate>
,
<date box="[1162,1382,514,536]" pageId="125" pageNumber="122" value="1994-02-17" valueMax="1994-02-22" valueMin="1994-02-17">17 - 22 February 1994</date>
(coll. P. Naskrecki) -
<specimenCount box="[1005,1073,542,564]" pageId="125" pageNumber="122" type="male">1 male</specimenCount>
(
<collectionCode box="[1085,1120,542,563]" pageId="125" pageNumber="122">PN</collectionCode>
collection); same locality,
<date box="[794,941,570,592]" pageId="125" pageNumber="122" value="1994-03-15">15 March 1994</date>
(coll. P. Naskrecki) -
<specimenCount box="[1153,1232,570,592]" pageId="125" pageNumber="122" type="male">2 males</specimenCount>
,
<specimenCount box="[1240,1336,570,592]" pageId="125" pageNumber="122" type="female">3 females</specimenCount>
(
<collectionCode box="[1348,1383,571,592]" pageId="125" pageNumber="122">PN</collectionCode>
collection); same locality,
<date box="[1047,1193,598,620]" pageId="125" pageNumber="122" value="1994-04-01" valueMax="1994-04-04" valueMin="1994-04-01">1 - 4 April 1994</date>
(coll. P. Naskrecki) -
<specimenCount box="[811,893,626,648]" pageId="125" pageNumber="122" type="male">6 males</specimenCount>
,
<specimenCount box="[906,998,626,648]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
, 1 nymph (incl.
<typeStatus box="[1175,1267,626,648]" pageId="125" pageNumber="122">holotype</typeStatus>
,
<typeStatus box="[1290,1375,626,648]" pageId="125" pageNumber="122">allotype</typeStatus>
) (
<collectionCode box="[801,872,654,676]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="125" pageNumber="122">ANSP</collectionCode>
); same locality,
<date box="[1051,1293,654,676]" pageId="125" pageNumber="122" value="1995-12-07" valueMax="1995-12-10" valueMin="1995-12-07">7 - 10 December 1995</date>
(coll. P. Naskrecki) -
<specimenCount box="[921,997,682,704]" pageId="125" pageNumber="122" type="male">2 males</specimenCount>
(
<collectionCode box="[1008,1042,683,704]" pageId="125" pageNumber="122">PN</collectionCode>
collection); same locality,
<date pageId="125" pageNumber="122" value="1998-04-13" valueMax="1998-05-10" valueMin="1998-04-13">13 April - 10 May 1998</date>
(coll. P. Naskrecki) -
<specimenCount box="[1163,1235,710,732]" pageId="125" pageNumber="122" type="male">1 male</specimenCount>
,
<specimenCount box="[1245,1334,710,732]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[1348,1383,710,731]" pageId="125" pageNumber="122">PN</collectionCode>
collection);
<emphasis bold="true" box="[912,1041,738,760]" pageId="125" pageNumber="122">
<collectingRegion box="[912,983,738,760]" country="Costa Rica" name="Limon" pageId="125" pageNumber="122">Limón</collectingRegion>
Prov.
</emphasis>
, Cuatro Esquinas, Tortuguero N. P.,
<date box="[824,1019,766,788]" pageId="125" pageNumber="122" value="1989-09-15">15 September 1989</date>
(coll. J. Solano) -
<specimenCount box="[1204,1293,766,788]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[1306,1371,766,788]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); same locality, elev.
<quantity box="[989,1053,795,816]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="125" pageNumber="122" unit="m" value="100.0">100 m</quantity>
,
<date box="[1062,1224,794,816]" pageId="125" pageNumber="122" value="1990-01-15">15 January 1990</date>
(coll. J. Solano) -
<specimenCount box="[811,901,822,844]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[917,982,822,844]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); Estac. Hitoy-Cerere Res. Biol. Hitoy Cerere, Río Cerere, elev.
<quantity box="[1068,1136,851,872]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="125" pageNumber="122" unit="m" value="100.0">100 m</quantity>
,
<date box="[1150,1288,850,872]" pageId="125" pageNumber="122" value="1991-05-15">15 May 1991</date>
(coll. A. Moreno) -
<specimenCount box="[908,979,878,900]" pageId="125" pageNumber="122" type="male">1 male</specimenCount>
(
<collectionCode box="[994,1059,878,900]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); same locality, elev.
<quantity box="[1280,1347,878,900]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="125" pageNumber="122" unit="m" value="100.0">100 m</quantity>
,
<date pageId="125" pageNumber="122" value="1992-06-20">20 June 1992</date>
(coll. F.A. Quesada) -
<specimenCount box="[1107,1191,906,928]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[1202,1266,906,928]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="125" pageNumber="122">INBio</collectionCode>
); La Emilia, near Guápiles, elev.
<quantity box="[1009,1093,934,956]" metricMagnitude="2" metricUnit="m" metricValue="3.048" pageId="125" pageNumber="122" unit="m" value="304.8">304.8 m</quantity>
,
<date box="[1105,1268,934,955]" pageId="125" pageNumber="122" value="1923-08-24">24 August 1923</date>
-
<specimenCount box="[1292,1382,934,956]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[801,869,963,985]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="125" pageNumber="122">ANSP</collectionCode>
); La Lola (
<quantity box="[984,1053,962,984]" metricMagnitude="2" metricUnit="m" metricValue="8.04672" pageId="125" pageNumber="122" unit="mi" value="0.5">0.5 mi</quantity>
. W
<collectingRegion box="[1097,1259,962,984]" country="Peru" name="Madre de Dios" pageId="125" pageNumber="122">Madre de Dios</collectingRegion>
),
<date pageId="125" pageNumber="122" value="1961-10-02">2 October 1961</date>
(coll. Hubbell, Cantrall, Cohn) -
<specimenCount box="[1179,1276,990,1012]" pageId="125" pageNumber="122" type="female">4 females</specimenCount>
(
<collectionCode box="[1289,1372,990,1012]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="125" pageNumber="122">UMMZ</collectionCode>
);
<emphasis bold="true" box="[794,970,1018,1039]" pageId="125" pageNumber="122">
<collectingRegion box="[794,913,1018,1039]" country="Costa Rica" name="Puntarenas" pageId="125" pageNumber="122">Puntarenas</collectingRegion>
Prov.
</emphasis>
, Las Cruces,
<date box="[1101,1329,1018,1040]" pageId="125" pageNumber="122" value="1991-07-01" valueMax="1991-08-31" valueMin="1991-07-01">1 July - 31 August 1991</date>
(coll. J. Doubles) -
<specimenCount box="[926,996,1046,1068]" pageId="125" pageNumber="122" type="male">1 male</specimenCount>
,
<specimenCount box="[1004,1092,1046,1068]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[1103,1186,1046,1068]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="125" pageNumber="122">UMMZ</collectionCode>
);
<emphasis bold="true" box="[1202,1319,1046,1067]" pageId="125" pageNumber="122">
<collectingRegion box="[1202,1315,1046,1067]" country="Panama" name="Panama" pageId="125" pageNumber="122">
<collectingCountry box="[1202,1315,1046,1067]" name="Panama" pageId="125" pageNumber="122">PANAMA</collectingCountry>
</collectingRegion>
:
</emphasis>
Barro
<collectingRegion box="[794,891,1074,1096]" country="United States of America" name="Colorado" pageId="125" pageNumber="122">Colorado</collectingRegion>
Island,
<date box="[971,1097,1074,1096]" pageId="125" pageNumber="122" value="1933-06-28">28 June 1933</date>
(coll. H.H. Hood) -
<specimenCount box="[1296,1382,1074,1096]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[801,869,1102,1124]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="125" pageNumber="122">ANSP</collectionCode>
); Canal Zone, Barro
<collectingRegion box="[1087,1186,1102,1124]" country="United States of America" name="Colorado" pageId="125" pageNumber="122">Colorado</collectingRegion>
,
<date box="[1198,1324,1102,1124]" pageId="125" pageNumber="122" value="1924-07-18">18 July 1924</date>
(coll. N. Banks) -
<specimenCount box="[915,1003,1130,1152]" pageId="125" pageNumber="122" type="female">1 female</specimenCount>
(
<collectionCode box="[1015,1084,1130,1152]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="125" pageNumber="122">ANSP</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>