1232 lines
150 KiB
XML
1232 lines
150 KiB
XML
<document id="F1EA7E3F8BB9E70CEF020CF4DD3811AA" ID-CLB-Dataset="32526" ID-DOI="10.11646/zootaxa.4303.2.2" ID-GBIF-Dataset="2e8b5af7-a7c0-4b2f-a02b-936e9a8584fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="840902" ID-ZooBank="6BB7825C-94F0-431E-A338-BFC9218076F4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502299197728" checkinUser="plazi" docAuthor="Grisales-Martínez, Freddy A., Velasco, Julián A., Bolívar, Wilmar, Williams, Ernest E. & Daza, Juan M." docDate="2017" docId="03F2E3179B578D3072CDFD30FCDFF8A1" docLanguage="en" docName="zootaxa.4303.2.2.pdf" docOrigin="Zootaxa 4303 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anolis urraoi Grisales-Martínez & Velasco & Bolívar & Williams & Daza 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="223" masterDocId="FFCB9B6F9B548D3A725AFF9EFF93FFE6" masterDocTitle="The taxonomic and phylogenetic status of some poorly known Anolis species from the Andes of Colombia with the description of a nomen nudum taxon" masterLastPageNumber="230" masterPageNumber="213" pageNumber="216" updateTime="1698466432041" updateUser="ExternalLinkService">
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<mods:title id="F6F19AA84A47C001ADBE2700DD0F8AB0">The taxonomic and phylogenetic status of some poorly known Anolis species from the Andes of Colombia with the description of a nomen nudum taxon</mods:title>
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<mods:namePart id="6E8D7961E296C3BDB97AC369BD11B499">Grisales-Martínez, Freddy A.</mods:namePart>
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<mods:namePart id="0C61A852C62122E6542D8FD378E84B6A">Velasco, Julián A.</mods:namePart>
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<mods:namePart id="2EA0999F0DAA66A5ABBF4C0F10B88447">Bolívar, Wilmar</mods:namePart>
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<mods:namePart id="1F6C682B6A895E01E3AEA6D6A5BFFF2A">Williams, Ernest E.</mods:namePart>
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<mods:namePart id="2785B966C5AF9DBD60DF322DC7752A5D">Daza, Juan M.</mods:namePart>
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<mods:title id="E2024C1587A7F52E6674D4D3F9ADEAF3">Zootaxa</mods:title>
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<treatment id="03F2E3179B578D3072CDFD30FCDFF8A1" ID-DOI="http://doi.org/10.5281/zenodo.6027712" ID-GBIF-Taxon="132648346" ID-Zenodo-Dep="6027712" LSID="urn:lsid:plazi:treatment:03F2E3179B578D3072CDFD30FCDFF8A1" httpUri="http://treatment.plazi.org/id/03F2E3179B578D3072CDFD30FCDFF8A1" lastPageId="10" lastPageNumber="223" pageId="3" pageNumber="216">
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<subSubSection id="C341018A9B578D3972CDFD30FEBEFD0C" pageId="3" pageNumber="216" type="nomenclature">
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<paragraph id="8BE452019B578D3972CDFD30FE0DFD2E" blockId="3.[151,414,686,746]" box="[151,414,686,712]" pageId="3" pageNumber="216">
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<heading id="D0ACE56D9B578D3972CDFD30FE0DFD2E" bold="true" box="[151,414,686,712]" fontSize="11" level="1" pageId="3" pageNumber="216" reason="1">
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<emphasis id="B92F8E139B578D3972CDFD30FE0DFD2E" bold="true" box="[151,414,686,712]" pageId="3" pageNumber="216">
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<taxonomicName id="4C5B29829B578D3972CDFD30FEAAFD2E" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[151,313,686,712]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="216" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
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<emphasis id="B92F8E139B578D3972CDFD30FEAAFD2E" bold="true" box="[151,313,686,712]" italics="true" pageId="3" pageNumber="216">Anolis urraoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21C33689B578D39731AFD30FE0DFD2E" box="[320,414,686,712]" pageId="3" pageNumber="216" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE452019B578D3972CDFD4FFEBEFD0C" blockId="3.[151,414,686,746]" box="[151,301,721,746]" pageId="3" pageNumber="216">
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(
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<figureCitation id="13604E849B578D3972C5FD4FFF7FFD0C" box="[159,236,721,746]" captionStart="FIGURE 1" captionStartId="3.[151,250,1778,1800]" captionTargetBox="[152,1435,1039,1753]" captionTargetId="figure@3.[151,1436,1038,1754]" captionTargetPageId="3" captionText="FIGURE 1. Anolis urraoi sp. nov. male holotype (MHUA-R 12733). Photograph by F. Grisales-Martínez." httpUri="https://zenodo.org/record/840904/files/figure.png" pageId="3" pageNumber="216">Figs. 1</figureCitation>
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,
|
||
<figureCitation id="13604E849B578D3972A0FD4CFE99FD0C" box="[250,266,722,746]" captionStart="FIGURE 3" captionStartId="5.[151,250,1947,1969]" captionTargetBox="[151,1436,733,1926]" captionTargetId="figure@5.[151,1436,733,1926]" captionTargetPageId="5" captionText="FIGURE 3. Holotype of Anolis urraoi sp. nov. (MHUA-R 12733, adult male). A, dorsal; B, lateral; C, ventral view of head; D, detail of dorsal scales at mid-dorsum of body; E, detail of flank scales at mid-dorsum of body; and F, detail of ventral scales at mid-dorsum of body. Scale bar = 5 mm." httpUri="https://zenodo.org/record/840908/files/figure.png" pageId="3" pageNumber="216">3</figureCitation>
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,
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||
<figureCitation id="13604E849B578D39734CFD4CFEB6FD0C" box="[278,293,722,746]" captionStart="FIGURE 4" captionStartId="7.[151,250,1604,1626]" captionTargetBox="[169,1454,411,1583]" captionTargetId="figure@7.[169,1454,411,1583]" captionTargetPageId="7" captionText="FIGURE 4. Dewlap coloration in Andean anoles. A, Anolis urraoi sp. nov., paratype, MHUA-R 12732, photo by F. A. Grisales-Martínez; B, A. urraoi sp. nov., holotype, MHUA-R 12733, photo by F. A. Grisales-Martínez; C, A. urraoi sp. nov., paratype, MHUA-R 12541, photo by F. A. Duarte; D, A. fuscoauratus, QCAZ 8326, Pastaza, Ecuador, photo by M. R. Bustamante, FaunaWebEcuador; E, A. antonii, El Dovio, Valle del Cauca, photo by D. Alzate-Estrada; F, A. antonii, Yotoco, Valle del Cauca, photo by W. Bolívar; G, A. tolimensis, MHUA-R 12770, photo by F. A. Grisales-Martínez; H, A. mariarum MHUA-R 13033, photo by F. A. Grisales-Martínez; and I, A. mariarum MHUA-R 12730, photo by F. A. Grisales-Martínez." httpUri="https://zenodo.org/record/840910/files/figure.png" pageId="3" pageNumber="216">4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C341018A9B578D3972CDFC8BFD01FCAC" pageId="3" pageNumber="216" type="description">
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<paragraph id="8BE452019B578D3972CDFC8BFDF4FCCD" blockId="3.[151,658,789,842]" box="[151,615,789,811]" pageId="3" pageNumber="216">Proposed standard English name: Urrao anole</paragraph>
|
||
<paragraph id="8BE452019B578D3972CDFCAAFD01FCAC" blockId="3.[151,658,789,842]" box="[151,658,820,842]" pageId="3" pageNumber="216">
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Proposed standard Spanish name:
|
||
<taxonomicName id="4C5B29829B578D3973A8FCAAFDA0FCAF" authorityName="Daudin" authorityYear="1802" box="[498,563,820,841]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="216" phylum="Chordata" rank="genus">
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||
<emphasis id="B92F8E139B578D3973A8FCAAFDA0FCAF" box="[498,563,820,841]" italics="true" pageId="3" pageNumber="216">Anolis</emphasis>
|
||
</taxonomicName>
|
||
de Urrao
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C341018A9B578D3F72CDFCE8FEBDFE4B" lastPageId="5" lastPageNumber="218" pageId="3" pageNumber="216" type="materials_examined">
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<paragraph id="8BE452019B578D3972CDFCE8FDA7FC31" blockId="3.[151,1437,886,983]" pageId="3" pageNumber="216">
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<materialsCitation id="3B33585C9B578D3972CDFCE8FDA7FC31" ID-GBIF-Occurrence="1585968388" collectingDate="1931-08" collectionCode="MHUA-R" collectorName="B. Rendon-Valencia & Grisales-Martinez & Figs & Table" elevation="1822" location="Rio Penderisco" municipality="Urrao" pageId="3" pageNumber="216" specimenCode="MHUA-R 12733" specimenCount="1" stateProvince="Antioquia" typeStatus="holotype">
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<emphasis id="B92F8E139B578D3972CDFCE8FE98FC69" bold="true" box="[151,267,886,911]" pageId="3" pageNumber="216">
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<typeStatus id="54E0ECA39B578D3972CDFCE8FE95FC69" box="[151,262,886,911]" pageId="3" pageNumber="216" type="holotype">Holotype</typeStatus>
|
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.
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</emphasis>
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<specimenCode id="DBFDFA7A9B578D39734BFCE9FE41FC69" box="[273,466,887,911]" collectionCode="MHUA-R" pageId="3" pageNumber="216">
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<collectionCode id="ED4ACAC49B578D39734BFCE9FE11FC69" box="[273,386,887,911]" pageId="3" pageNumber="216">MHUA-R</collectionCode>
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12733
|
||
</specimenCode>
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||
; adult male, from departament of
|
||
<collectingRegion id="499F9CE39B578D397111FCE8FC2DFC69" box="[843,958,886,911]" country="Colombia" name="Antioquia" pageId="3" pageNumber="216">Antioquia</collectingRegion>
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||
,
|
||
<collectingMunicipality id="6B80C87B9B578D397193FCE9FB99FC68" box="[969,1034,887,910]" pageId="3" pageNumber="216">Urrao</collectingMunicipality>
|
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municipality, paddock on the banks of
|
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<location id="8E8404DA9B578D3972EFFC05FEF0FC55" LSID="urn:lsid:plazi:treatment:03F2E3179B578D3072CDFD30FCDFF8A1:8E8404DA9B578D3972EFFC05FEF0FC55" box="[181,355,922,947]" municipality="Urrao" name="Rio Penderisco" pageId="3" pageNumber="216" stateProvince="Antioquia">Río Penderisco</location>
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: 6.314253, -76.138528,
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<elevation id="0076B5329B578D397028FC05FD5BFC54" box="[626,712,923,947]" metricMagnitude="3" metricUnit="m" metricValue="1.8219999999999998" pageId="3" pageNumber="216" unit="m" value="1822.0">
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<quantity id="4CA3FFE49B578D397028FC05FD5BFC54" box="[626,712,923,947]" metricMagnitude="3" metricUnit="m" metricValue="1.8219999999999998" pageId="3" pageNumber="216" unit="m" value="1822.0">1822 m</quantity>
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</elevation>
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, collected by
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<collectorName id="26AE37D79B578D397138FC05FBD7FC55" box="[866,1092,922,947]" pageId="3" pageNumber="216">B. Rendón-Valencia</collectorName>
|
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and F.A.
|
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<collectorName id="26AE37D79B578D3976F5FC04FAEAFC55" box="[1199,1401,922,947]" pageId="3" pageNumber="216">Grisales-Martínez</collectorName>
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on
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<collectingDate id="EFA18D299B578D3972CDFC21FE9DFC31" box="[151,270,959,983]" pageId="3" pageNumber="216" value="1931-08">
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<date id="FFE574C19B578D3972CDFC21FE9DFC31" box="[151,270,959,983]" pageId="3" pageNumber="216" value="1931-08">31 August</date>
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</collectingDate>
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, 2014 (
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<figureCitation id="13604E849B578D39733AFC20FE35FC31" box="[352,422,958,983]" captionStart="FIGURE 1" captionStartId="3.[151,250,1778,1800]" captionTargetBox="[152,1435,1039,1753]" captionTargetId="figure@3.[151,1436,1038,1754]" captionTargetPageId="3" captionText="FIGURE 1. Anolis urraoi sp. nov. male holotype (MHUA-R 12733). Photograph by F. Grisales-Martínez." httpUri="https://zenodo.org/record/840904/files/figure.png" pageId="3" pageNumber="216">
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<collectorName id="26AE37D79B578D39733AFC20FE01FC31" box="[352,402,958,983]" pageId="3" pageNumber="216">Figs</collectorName>
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1
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</figureCitation>
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,
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<figureCitation id="13604E849B578D3973EEFC21FE57FC31" box="[436,452,959,983]" captionStart="FIGURE 3" captionStartId="5.[151,250,1947,1969]" captionTargetBox="[151,1436,733,1926]" captionTargetId="figure@5.[151,1436,733,1926]" captionTargetPageId="5" captionText="FIGURE 3. Holotype of Anolis urraoi sp. nov. (MHUA-R 12733, adult male). A, dorsal; B, lateral; C, ventral view of head; D, detail of dorsal scales at mid-dorsum of body; E, detail of flank scales at mid-dorsum of body; and F, detail of ventral scales at mid-dorsum of body. Scale bar = 5 mm." httpUri="https://zenodo.org/record/840908/files/figure.png" pageId="3" pageNumber="216">3</figureCitation>
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;
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<tableCitation id="C6D967BA9B578D39738BFC20FDB0FC31" box="[465,547,958,983]" captionStart="TABLE 1" captionStartId="2.[151,239,580,603]" captionText="TABLE 1. Arithmetic mean and associated standard deviation from selected morphometric characters in four species of Andean anoles. See text for abbreviations of measurements." pageId="3" pageNumber="216">
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<collectorName id="26AE37D79B578D39738BFC20FD9CFC31" box="[465,527,958,983]" pageId="3" pageNumber="216">Table</collectorName>
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1
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</tableCitation>
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).
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</materialsCitation>
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</paragraph>
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<caption id="DF2402899B578D3972CDF96CFB43F8EE" box="[151,1232,1778,1800]" httpUri="https://zenodo.org/record/840904/files/figure.png" pageId="3" pageNumber="216" targetBox="[152,1435,1039,1753]" targetPageId="3">
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<paragraph id="8BE452019B578D3972CDF96CFB43F8EE" blockId="3.[151,1232,1778,1800]" box="[151,1232,1778,1800]" pageId="3" pageNumber="216">
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<emphasis id="B92F8E139B578D3972CDF96CFE80F8E1" bold="true" box="[151,275,1778,1800]" pageId="3" pageNumber="216">FIGURE 1.</emphasis>
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<taxonomicName id="4C5B29829B578D397343F96CFE32F8EE" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[281,417,1778,1800]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="216" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
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<emphasis id="B92F8E139B578D397343F96CFE32F8EE" box="[281,417,1778,1800]" italics="true" pageId="3" pageNumber="216">Anolis urraoi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21C33689B578D3973FDF96CFE64F8E1" box="[423,503,1778,1799]" pageId="3" pageNumber="216" rank="species">
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<emphasis id="B92F8E139B578D3973FDF96CFE64F8E1" bold="true" box="[423,503,1778,1799]" pageId="3" pageNumber="216">sp. nov.</emphasis>
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</taxonomicNameLabel>
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||
male holotype (MHUA-R 12733). Photograph by F. Grisales-Martínez.
|
||
</paragraph>
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</caption>
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<caption id="DF2402899B508D3E72CDF883FAD1F896" httpUri="https://zenodo.org/record/840906/files/figure.png" pageId="4" pageNumber="217" targetBox="[164,1423,206,1768]" targetPageId="4">
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<paragraph id="8BE452019B508D3E72CDF883FAD1F896" blockId="4.[151,1436,1821,1904]" pageId="4" pageNumber="217">
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<emphasis id="B92F8E139B508D3E72CDF883FE87F8D4" bold="true" box="[151,276,1821,1843]" pageId="4" pageNumber="217">FIGURE 2.</emphasis>
|
||
Adult males of the Andean species of the
|
||
<taxonomicName id="4C5B29829B508D3E7095F883FC08F8D4" authorityName="D'Orbigny" authorityYear="1837" box="[719,923,1821,1842]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="217" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B508D3E7095F883FC08F8D4" box="[719,923,1821,1842]" italics="true" pageId="4" pageNumber="217">Anolis fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
<emphasis id="B92F8E139B508D3E7666F883FBDDF8D4" bold="true" box="[1084,1102,1821,1842]" pageId="4" pageNumber="217">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B508D3E7607F883FAB4F8D4" authorityName="D'Orbigny" authorityYear="1837" box="[1117,1319,1821,1843]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="217" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B508D3E7607F883FBE0F8D4" box="[1117,1139,1821,1842]" italics="true" pageId="4" pageNumber="217">A.</emphasis>
|
||
aff.
|
||
<emphasis id="B92F8E139B508D3E76FEF883FAB4F8D4" box="[1188,1319,1821,1842]" italics="true" pageId="4" pageNumber="217">fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
, MHUA-R 12791, photo by Angelly Vasquez;
|
||
<emphasis id="B92F8E139B508D3E705AF8A2FD82F8B7" bold="true" box="[512,529,1852,1873]" pageId="4" pageNumber="217">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B508D3E7044F8A2FD31F8B7" authorityName="Werner" authorityYear="1916" box="[542,674,1852,1873]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="217" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="B92F8E139B508D3E7044F8A2FD31F8B7" box="[542,674,1852,1873]" italics="true" pageId="4" pageNumber="217">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
, MHUA-R 12770, photo by F.A. Grisales-Martínez;
|
||
<emphasis id="B92F8E139B508D3E7698F8A2FB47F8B4" bold="true" box="[1218,1236,1852,1874]" pageId="4" pageNumber="217">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B508D3E76BBF8A2FE9FF896" authority=", Bitaco, La Cumbre" authorityName="Bitaco, La Cumbre" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="217" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B508D3E76BBF8A2FAD5F8B7" box="[1249,1350,1852,1873]" italics="true" pageId="4" pageNumber="217">A. antonii</emphasis>
|
||
, Bitaco, La Cumbre
|
||
</taxonomicName>
|
||
, Valle del Cauca, photo by W. Bolívar; and
|
||
<emphasis id="B92F8E139B508D3E709DF8C4FD4AF889" bold="true" box="[711,729,1882,1903]" pageId="4" pageNumber="217">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B508D3E70BCF8C5FCFBF896" box="[742,872,1883,1904]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="217" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B508D3E70BCF8C5FCFBF896" box="[742,872,1883,1904]" italics="true" pageId="4" pageNumber="217">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
MHUA-R 12700, photo by M. Rivera-Correa.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE452019B508D3F729DF802FEBDFE4B" blockId="4.[151,1436,1948,2010]" lastBlockId="5.[151,1437,151,680]" lastPageId="5" lastPageNumber="218" pageId="4" pageNumber="217">
|
||
<materialsCitation id="3B33585C9B508D3F729DF802FDB6FF49" ID-GBIF-Occurrence="1585968389" collectionCode="MHUA-R" collectorName="El Maravillo & Daza" country="Colombia" county="Finca La Lucia" elevation="2120" lastPageId="5" lastPageNumber="218" location="Urrao" municipality="Corregimiento La Encarnacion" pageId="4" pageNumber="217" specimenCode="MHUA-R 12519-20-21" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<emphasis id="B92F8E139B508D3E729DF802FED5F853" bold="true" box="[199,326,1948,1973]" pageId="4" pageNumber="217">
|
||
<typeStatus id="54E0ECA39B508D3E729DF802FED2F853" box="[199,321,1948,1973]" pageId="4" pageNumber="217" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
n = 38: males = 21, females = 17 (all from
|
||
<collectingRegion id="499F9CE39B508D3E7174F802FC32F853" box="[814,929,1948,1973]" country="Colombia" name="Antioquia" pageId="4" pageNumber="217">Antioquia</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34C12919B508D3E71F6F802FB8DF853" box="[940,1054,1948,1973]" name="Colombia" pageId="4" pageNumber="217">Colombia</collectingCountry>
|
||
.
|
||
<tableCitation id="C6D967BA9B508D3E7673F802FBEAF853" box="[1065,1145,1948,1973]" captionStart="TABLE 1" captionStartId="2.[151,239,580,603]" captionText="TABLE 1. Arithmetic mean and associated standard deviation from selected morphometric characters in four species of Andean anoles. See text for abbreviations of measurements." pageId="4" pageNumber="217">Table1</tableCitation>
|
||
).
|
||
<specimenCode id="DBFDFA7A9B508D3E76D6F803FA04F853" box="[1164,1431,1948,1973]" pageId="4" pageNumber="217">
|
||
<collectionCode id="ED4ACAC49B508D3E76D6F803FB6EF853" box="[1164,1277,1949,1973]" pageId="4" pageNumber="217">MHUA-R</collectionCode>
|
||
12519-20-21
|
||
</specimenCode>
|
||
,
|
||
<location id="8E8404DA9B508D3E72CDF85CFF4BF83F" LSID="urn:lsid:plazi:treatment:03F2E3179B578D3072CDFD30FCDFF8A1:8E8404DA9B508D3E72CDF85CFF4BF83F" box="[151,216,1986,2009]" country="Colombia" county="Finca La Lucia" municipality="Corregimiento La Encarnacion" name="Urrao" pageId="4" pageNumber="217" stateProvince="Antioquia">Urrao</location>
|
||
municipality,
|
||
<collectingMunicipality id="6B80C87B9B508D3E7323F85FFD44F83C" box="[377,727,1985,2010]" pageId="4" pageNumber="217">Corregimiento La Encarnación</collectingMunicipality>
|
||
, vereda
|
||
<collectorName id="26AE37D79B508D3E716EF85FFC56F83C" box="[820,965,1985,2010]" pageId="4" pageNumber="217">El Maravillo</collectorName>
|
||
,
|
||
<collectingCounty id="62852A8D9B508D3E718BF85FFBEDF83F" box="[977,1150,1985,2010]" pageId="4" pageNumber="217">Finca La Lucía</collectingCounty>
|
||
, 6,51669, -76,1484,
|
||
<elevation id="0076B5329B508D3F773EF85FFF3CFF49" lastPageId="5" lastPageNumber="218" metricMagnitude="3" metricUnit="m" metricValue="2.12" pageId="4" pageNumber="217" unit="m" value="2120.0">
|
||
<quantity id="4CA3FFE49B508D3F773EF85FFF3CFF49" lastPageId="5" lastPageNumber="218" metricMagnitude="3" metricUnit="m" metricValue="2.12" pageId="4" pageNumber="217" unit="m" value="2120.0">2120 m</quantity>
|
||
</elevation>
|
||
, collected on 2012 by J.M.
|
||
<collectorName id="26AE37D79B518D3F73B2FF06FDB6FF49" box="[488,549,152,175]" pageId="5" pageNumber="218">Daza</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B33585C9B518D3F706BFF06FC56FF11" ID-GBIF-Occurrence="1585968404" collectionCode="MHUA-R" collectorName="El Maravillo & Daza" country="Colombia" elevation="2156" location="Urrao" municipality="Corregimiento La Encarnacion" pageId="5" pageNumber="218" specimenCode="MHUA-R 12521, 12523" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<specimenCode id="DBFDFA7A9B518D3F706BFF06FCD4FF56" box="[561,839,151,176]" pageId="5" pageNumber="218">
|
||
<collectionCode id="ED4ACAC49B518D3F706BFF06FD30FF56" box="[561,675,152,176]" pageId="5" pageNumber="218">MHUA-R</collectionCode>
|
||
12521, 12523
|
||
</specimenCode>
|
||
, 12525, 12526, 12528, 12529, 12530, 12532, 12535, 12536, 12539, 12540, 12541, 12544, 12545,
|
||
<location id="8E8404DA9B518D3F70E1FF23FD6FFF32" LSID="urn:lsid:plazi:treatment:03F2E3179B578D3072CDFD30FCDFF8A1:8E8404DA9B518D3F70E1FF23FD6FFF32" box="[699,764,189,212]" country="Colombia" municipality="Corregimiento La Encarnacion" name="Urrao" pageId="5" pageNumber="218" stateProvince="Antioquia">Urrao</location>
|
||
municipality,
|
||
<collectingMunicipality id="6B80C87B9B518D3F71F1FF22FA8BFF33" box="[939,1304,188,213]" pageId="5" pageNumber="218">Corregimiento La Encarnación</collectingMunicipality>
|
||
, vereda
|
||
<collectorName id="26AE37D79B518D3F77D9FF22FE9BFF1E" pageId="5" pageNumber="218">El Maravillo</collectorName>
|
||
, 6.52133, -76.14022,
|
||
<elevation id="0076B5329B518D3F73A6FF41FDC0FF11" box="[508,595,223,248]" metricMagnitude="3" metricUnit="m" metricValue="2.1559999999999997" pageId="5" pageNumber="218" unit="m" value="2156.0">
|
||
<quantity id="4CA3FFE49B518D3F73A6FF41FDC0FF11" box="[508,595,223,248]" metricMagnitude="3" metricUnit="m" metricValue="2.1559999999999997" pageId="5" pageNumber="218" unit="m" value="2156.0">2156 m</quantity>
|
||
</elevation>
|
||
, collected on 2012 by J.M.
|
||
<collectorName id="26AE37D79B518D3F71D2FF7EFC56FF11" box="[904,965,224,247]" pageId="5" pageNumber="218">Daza</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B33585C9B518D3F718AFF7EFAB6FE83" ID-GBIF-Occurrence="1585968385" collectingDate="1931-08" collectionCode="MHUA-R" collectorName="Urrao & Las Orquideas National Park & Calles & M. A. Serna & Echeverri" country="Colombia" elevation="1816" location="river Penderisco" pageId="5" pageNumber="218" specimenCount="1" stateProvince="Antioquia" typeStatus="paratype">
|
||
<collectionCode id="ED4ACAC49B518D3F718AFF7EFBD2FF1E" box="[976,1089,224,248]" pageId="5" pageNumber="218">MHUA-R</collectionCode>
|
||
12731-32,
|
||
<collectorName id="26AE37D79B518D3F769BFF7EFA91FF11" box="[1217,1282,224,247]" pageId="5" pageNumber="218">Urrao</collectorName>
|
||
municipality, in the urban area, paddock on the banks of
|
||
<location id="8E8404DA9B518D3F70FBFE9AFCCCFEFB" LSID="urn:lsid:plazi:treatment:03F2E3179B578D3072CDFD30FCDFF8A1:8E8404DA9B518D3F70FBFE9AFCCCFEFB" box="[673,863,260,285]" country="Colombia" name="river Penderisco" pageId="5" pageNumber="218" stateProvince="Antioquia">river Penderisco</location>
|
||
, 6.314434, -76.138444,
|
||
<elevation id="0076B5329B518D3F7627FE9AFB4BFEFA" box="[1149,1240,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.816" pageId="5" pageNumber="218" unit="m" value="1816.0">
|
||
<quantity id="4CA3FFE49B518D3F7627FE9AFB4BFEFA" box="[1149,1240,260,285]" metricMagnitude="3" metricUnit="m" metricValue="1.816" pageId="5" pageNumber="218" unit="m" value="1816.0">1816 m</quantity>
|
||
</elevation>
|
||
, collected on
|
||
<collectingDate id="EFA18D299B518D3F77DAFE9AFF78FEA6" pageId="5" pageNumber="218" value="1931-08">
|
||
<date id="FFE574C19B518D3F77DAFE9AFF78FEA6" pageId="5" pageNumber="218" value="1931-08">31 August</date>
|
||
</collectingDate>
|
||
, 2014 by B. Rendón-Valencia and F.A. Grisales-Martínez; CSJ 455, 467, 814, 819, 822-23, 837, 839,
|
||
<collectorName id="26AE37D79B518D3F7701FEB6FA0FFED9" box="[1371,1436,296,319]" pageId="5" pageNumber="218">Urrao</collectorName>
|
||
municipality,
|
||
<collectorName id="26AE37D79B518D3F7369FED3FD11FE83" box="[307,642,332,357]" pageId="5" pageNumber="218">Las Orquideas National Park</collectorName>
|
||
, vereda
|
||
<collectorName id="26AE37D79B518D3F70BFFED2FCBDFE83" box="[741,814,332,357]" pageId="5" pageNumber="218">Calles</collectorName>
|
||
, collected by
|
||
<collectorName id="26AE37D79B518D3F718AFED2FBC6FE82" box="[976,1109,332,356]" pageId="5" pageNumber="218">M.A. Serna</collectorName>
|
||
and H.
|
||
<collectorName id="26AE37D79B518D3F76EFFED2FAB6FE83" box="[1205,1317,332,357]" pageId="5" pageNumber="218">Echeverri</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B33585C9B518D3F7769FED2FEBDFE4B" ID-GBIF-Occurrence="1585968398" collectorName="Urrao & M. A. Serna" country="Colombia" location="Magdalena" pageId="5" pageNumber="218" specimenCount="1" stateProvince="Magdalena" typeStatus="paratype">
|
||
CSJ 635; 703,815, 843, 848, 856, 858-59, 1028,
|
||
<date id="FFE574C19B518D3F7009FEF1FD20FE6E" box="[595,691,367,392]" pageId="5" pageNumber="218" value="1074" valueMax="1075">1074-75</date>
|
||
,
|
||
<collectorName id="26AE37D79B518D3F709AFEEEFC92FE61" box="[704,769,368,391]" pageId="5" pageNumber="218">Urrao</collectorName>
|
||
municipality, La
|
||
<collectingRegion id="499F9CE39B518D3F7191FEF1FBD9FE6E" box="[971,1098,367,392]" country="Colombia" name="Magdalena" pageId="5" pageNumber="218">Magdalena</collectingRegion>
|
||
, collected by
|
||
<collectorName id="26AE37D79B518D3F76B2FEEEFAFFFE6E" box="[1256,1388,368,392]" pageId="5" pageNumber="218">M.A. Serna</collectorName>
|
||
and H. Echeverri.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C341018A9B518D3C729DFE29FF6CFD2B" lastPageId="6" lastPageNumber="219" pageId="5" pageNumber="218" type="diagnosis">
|
||
<paragraph id="8BE452019B518D3F729DFE29FE24FD4E" blockId="5.[151,1437,151,680]" pageId="5" pageNumber="218">
|
||
<emphasis id="B92F8E139B518D3F729DFE29FED2FE36" bold="true" box="[199,321,439,464]" pageId="5" pageNumber="218">Diagnosis.</emphasis>
|
||
An small brown-greenish
|
||
<taxonomicName id="4C5B29829B518D3F70DDFE26FD5DFE36" authorityName="Daudin" authorityYear="1802" box="[647,718,440,464]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="218" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92F8E139B518D3F70DDFE26FD5DFE36" box="[647,718,440,464]" italics="true" pageId="5" pageNumber="218">Anolis</emphasis>
|
||
</taxonomicName>
|
||
assigned to the
|
||
<taxonomicName id="4C5B29829B518D3F71C4FE26FB16FE36" authorityName="D'Orbigny" authorityYear="1837" box="[926,1157,440,464]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="218" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B518D3F71C4FE26FB16FE36" box="[926,1157,440,464]" italics="true" pageId="5" pageNumber="218">Anolis fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
group by phylogenetic evidence presented in this study (see phylogenetic results) and by having the following combination of morphological characters described by
|
||
<bibRefCitation id="EFCA2FF09B518D3F7009FE61FC98FDFE" author="Williams" box="[595,779,511,536]" pageId="5" pageNumber="218" refString="Williams, E. E. (1976) South American anoles; the species groups. Papeis Avulsos de Zoologia, 29, 259 - 268. https: // doi. org / 10.11606 / issn. 2176 - 7793. v 19 i 1 - 2 p 1 - 176" type="journal article" year="1976">Williams (1976)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFCA2FF09B518D3F7119FE61FBF5FDFE" author="Ayala" box="[835,1126,511,536]" pageId="5" pageNumber="218" refString="Ayala, S. C. & Williams, E. E. (1988) New or problematic Anolis from Colombia, VI. Two fuscoauratoid anoles from the Pacific lowlands, A. maculiventris Boulenger, 1898 and A. medemi, a new species from Gorgona Island. Breviora, 490, 1 - 16." type="journal article" year="1988">Ayala & Williams (1988)</bibRefCitation>
|
||
: 1) Head with small frontal depression and keeled scales not enlarged (similar in size to the dorsal scales); 2) small dorsal scales, uniform and usually keeled (smaller than ventral scales); 3) smooth ventral scales (sometimes with a low keeling); 4) broad and well-defined toes lamellae; 5) less than 20 lamellae on phalanges II and III of the fourth toe; and 6) round and slender tail without ridge.
|
||
</paragraph>
|
||
<caption id="DF2402899B518D3F72CDF805FDBFF809" httpUri="https://zenodo.org/record/840908/files/figure.png" pageId="5" pageNumber="218" targetBox="[151,1436,733,1926]" targetPageId="5">
|
||
<paragraph id="8BE452019B518D3F72CDF805FDBFF809" blockId="5.[151,1436,1947,2031]" pageId="5" pageNumber="218">
|
||
<emphasis id="B92F8E139B518D3F72CDF805FE81F856" bold="true" box="[151,274,1947,1969]" pageId="5" pageNumber="218">FIGURE 3.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C5B29829B518D3F73CDF805FD8DF857" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[407,542,1947,1969]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="218" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B518D3F73CDF805FD8DF857" box="[407,542,1947,1969]" italics="true" pageId="5" pageNumber="218">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B518D3F707EF802FDE0F857" box="[548,627,1948,1969]" pageId="5" pageNumber="218" rank="species">
|
||
<emphasis id="B92F8E139B518D3F707EF802FDE0F857" bold="true" box="[548,627,1948,1969]" pageId="5" pageNumber="218">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(MHUA-R 12733, adult male). A, dorsal; B, lateral; C, ventral view of head; D, detail of dorsal scales at mid-dorsum of body; E, detail of flank scales at mid-dorsum of body; and F, detail of ventral scales at mid-dorsum of body. Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE452019B528D3C729DFF06FF6CFD2B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="219">
|
||
Because the
|
||
<taxonomicName id="4C5B29829B528D3C7302FF06FDA9FF56" authorityName="D'Orbigny" authorityYear="1837" box="[344,570,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B528D3C7302FF06FDA9FF56" box="[344,570,152,176]" italics="true" pageId="6" pageNumber="219">Anolis fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
group is not recognized in the most comprehensive phylogeny to date (i.e.,
|
||
<bibRefCitation id="EFCA2FF09B528D3C72CDFF23FED7FF32" author="Poe" box="[151,324,188,212]" pageId="6" pageNumber="219" refString="Poe, S., Nieto-Montes de Oca, A., Torres-Carvajal, O., de Queiroz, K., Velasco, J. A., Truett, B., Gray, L. N., Ryan, M. J., Kohler, G., Ayala-Varela, F. & Latella, I. (2017) A Phylogenetic, Biogeographic, and Taxonomic study of all Extant Species of Anolis (Squamata; Iguanidae). Systematic Biology. [published online]" type="book" year="2017">
|
||
Poe
|
||
<emphasis id="B92F8E139B528D3C7292FF23FE92FF32" box="[200,257,188,212]" italics="true" pageId="6" pageNumber="219">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
), for our taxonomic account we compared the new taxon with species closely related or distributed in the same biogeographic regions (e.g., Central America, Pacific lowlands and the northern Andes).
|
||
<taxonomicName id="4C5B29829B528D3C775FFF7EFA0FFF11" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[1285,1436,224,248]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B528D3C775FFF7EFA0FFF11" box="[1285,1436,224,248]" italics="true" pageId="6" pageNumber="219">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B528D3C72CDFE9AFF61FEFB" box="[151,242,260,285]" pageId="6" pageNumber="219" rank="species">
|
||
<emphasis id="B92F8E139B528D3C72CDFE9AFF61FEFB" bold="true" box="[151,242,260,285]" pageId="6" pageNumber="219">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from other species in the
|
||
<taxonomicName id="4C5B29829B528D3C7029FE9AFCB6FEFA" authorityName="D'Orbigny" authorityYear="1837" box="[627,805,260,284]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B528D3C7029FE9AFCB6FEFA" box="[627,805,260,284]" italics="true" pageId="6" pageNumber="219">A. fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
group from Central America by having a male dewlap bright orange anteriorly and light rose color on the posterior third or half. In contrast, males of
|
||
<taxonomicName id="4C5B29829B528D3C7696FEB6FE6AFE83" authority="Poe, Scarpetta & Schaad 2015" authorityName="Poe, Scarpetta & Schaad" authorityYear="2015" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="elcopeensis">
|
||
<emphasis id="B92F8E139B528D3C7696FEB6FA0FFEA6" box="[1228,1436,296,320]" italics="true" pageId="6" pageNumber="219">Anolis elcopeensis</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C72CDFED3FE6AFE83" author="Poe" box="[151,505,332,357]" pageId="6" pageNumber="219" refString="Poe, S., Scarpetta, S. & Schaad, E. W. (2015) A new species of Anolis (Squamata: Iguanidae) from Panama. Amphibian & Reptile Conservation, 9 (1), 1 - 13." type="journal article" year="2015">Poe, Scarpetta & Schaad 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5B29829B528D3C705DFED2FBF1FE82" authority="Kohler, Ponce, Sunyer & Batista 2007" authorityName="Kohler, Ponce, Sunyer & Batista" authorityYear="2007" box="[519,1122,332,357]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="gruuo">
|
||
<emphasis id="B92F8E139B528D3C705DFED2FD08FE82" box="[519,667,332,356]" italics="true" pageId="6" pageNumber="219">Anolis gruuo</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C70FFFED2FBF1FE82" author="Kohler" box="[677,1122,332,357]" pageId="6" pageNumber="219" refString="Kohler, G., Ponce, M., Sunyer, J. & Batista, A. (2007) Four new species of anoles (Genus Anolis) from the Serrania de Tabasara, West- Central Panama (Squamata: Polychrotidae). Herpetologica, 63 (3), 375 - 391." type="journal article" year="2007">Köhler, Ponce, Sunyer & Batista 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5B29829B528D3C762BFED2FE1EFE6E" authority="Echelle, Echelle & Fitch 1971" authorityName="Echelle, Echelle & Fitch" authorityYear="1971" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="carpenteri">
|
||
<emphasis id="B92F8E139B528D3C762BFED2FAABFE82" box="[1137,1336,332,356]" italics="true" pageId="6" pageNumber="219">Anolis carpenteri</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C771BFED2FE1EFE6E" author="Echelle" pageId="6" pageNumber="219" refString="Echelle, A. A., Echelle, A. F. & Fitch, H. S. (1971) A new anole from Costa Rica. Herpetologica, 27 (3), 354 - 362." type="journal article" year="1971">Echelle, Echelle & Fitch 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B29829B528D3C739DFEEEFC99FE6E" authority="Peters 1874" authorityName="Peters" authorityYear="1874" box="[455,778,368,392]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="polylepis">
|
||
<emphasis id="B92F8E139B528D3C739DFEEEFDEFFE6E" box="[455,636,368,392]" italics="true" pageId="6" pageNumber="219">Anolis polylepis</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C70DFFEEEFC99FE6E" author="Peters" box="[645,778,368,392]" pageId="6" pageNumber="219" refString="Peters, W. C. H. (1874) Uber neue Saurier (Spaeriodactylus, Anolis, Phrynosoma, Tropidolepisma, Lygosoma, Ophioscincus) aus Centralamerica, Mexico und Australien. Monatsberichte der Koniglichen Preussische Akademie des Wissenschaften zu Berlin, 1874, 738 - 747." type="journal article" year="1874">Peters 1874</bibRefCitation>
|
||
</taxonomicName>
|
||
all have orange dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C7676FEEEFAF8FE6E" authority="Dunn 1940" authorityName="Dunn" authorityYear="1940" box="[1068,1387,368,392]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="kemptoni">
|
||
<emphasis id="B92F8E139B528D3C7676FEEEFB71FE6E" box="[1068,1250,368,392]" italics="true" pageId="6" pageNumber="219">Anolis kemptoni</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C76B6FEEEFAF8FE6E" author="Dunn" box="[1260,1387,368,392]" pageId="6" pageNumber="219" refString="Dunn, E. R. (1940) New and noteworthy herpetological material from Panama. Proceedings of the Academy of Natural Sciences of Philadelphia, 92, 105 - 122." type="journal article" year="1940">Dunn 1940</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B29829B528D3C72CDFE0AFCF3FE4A" authority="Kohler, Ponce, Sunyer & Batista 2007" authorityName="Kohler, Ponce, Sunyer & Batista" authorityYear="2007" box="[151,864,404,429]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="pseudokemptoni">
|
||
<emphasis id="B92F8E139B528D3C72CDFE0AFE0DFE4A" box="[151,414,404,428]" italics="true" pageId="6" pageNumber="219">Anolis pseudokemptoni</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C73FDFE0AFCF3FE4A" author="Kohler" box="[423,864,404,429]" pageId="6" pageNumber="219" refString="Kohler, G., Ponce, M., Sunyer, J. & Batista, A. (2007) Four new species of anoles (Genus Anolis) from the Serrania de Tabasara, West- Central Panama (Squamata: Polychrotidae). Herpetologica, 63 (3), 375 - 391." type="journal article" year="2007">Köhler, Ponce, Sunyer & Batista 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
have red-orange anterior and pink posterior male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C72A2FE26FD6CFE36" authority="Arosemena & Ibanez 1993" authorityName="Arosemena & Ibanez" authorityYear="1993" box="[248,767,439,464]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="fortunensis">
|
||
<emphasis id="B92F8E139B528D3C72A2FE26FE56FE36" box="[248,453,440,464]" italics="true" pageId="6" pageNumber="219">Anolis fortunensis</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C7397FE26FD6CFE36" author="Arosemena" box="[461,767,439,464]" pageId="6" pageNumber="219" refString="Arosemena, F. A. & Ibanez, R. (1993) Una especie nueva de Anolis (Squamata: Iguanidae) del grupo fuscoauratus de Fortuna, Panama. Revista de Biologia Tropical, 41 (2), 267 - 272." type="journal article" year="1993">Arosemena & Ibáñez 1993</bibRefCitation>
|
||
</taxonomicName>
|
||
has red anterior and orange posterior male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C770EFE26FE38FE12" authority="Kohler 2009" authorityName="Kohler" authorityYear="2009" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="monteverde">
|
||
<emphasis id="B92F8E139B528D3C770EFE26FE8AFE12" italics="true" pageId="6" pageNumber="219">Anolis monteverde</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C7345FE42FE38FE12" author="Kohler" box="[287,427,476,501]" pageId="6" pageNumber="219" refString="Kohler, G. (2009) New Species of Anolis Formerly Referred to as Anolis altae from Monteverde, Costa Rica (Squamata: Polychrotidae). Journal of Herpetology, 43 (1), 11 - 20." type="journal article" year="2009">Köhler 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
yellow ocher to orange yellow in the basal portion and reddish-orange in the distal portion of the male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C73DFFD9EFD04FDFE" authority="Dunn 1930" authorityName="Dunn" authorityYear="1930" box="[389,663,512,536]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="altae">
|
||
<emphasis id="B92F8E139B528D3C73DFFD9EFD9DFDFE" box="[389,526,512,536]" italics="true" pageId="6" pageNumber="219">Anolis altae</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C704DFD9EFD04FDFE" author="Dunn" box="[535,663,512,536]" pageId="6" pageNumber="219" refString="Dunn, E. R. (1930) Notes on Central American Anolis. Proceedings of the New England Zoological Club, 12, 15 - 24." type="journal article" year="1930">Dunn 1930</bibRefCitation>
|
||
</taxonomicName>
|
||
with a more or less uniformly reddish-orange male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C770EFD9EFEEEFDDB" authority="Cope 1862" authorityName="Cope" authorityYear="1862" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="limifrons">
|
||
<emphasis id="B92F8E139B528D3C770EFD9EFF68FDDA" italics="true" pageId="6" pageNumber="219">Anolis limifrons</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C7359FDBAFEEEFDDB" author="Cope" box="[259,381,548,573]" pageId="6" pageNumber="219" refString="Cope, E. D. (1862) Contributions to Neotropical saurology. Proceedings of the Academy of Natural Sciences of Philadelphia, 14, 176 - 188." type="journal article" year="1862">Cope 1862</bibRefCitation>
|
||
</taxonomicName>
|
||
with a dull white with a small basal orange blotch male dewlap,
|
||
<taxonomicName id="4C5B29829B528D3C7605FDBAFEDBFD86" authority="Kohler & Sunyer 2008" authorityName="Kohler & Sunyer" authorityYear="2008" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="apletophallus">
|
||
<emphasis id="B92F8E139B528D3C7605FDBAFAD5FDDA" box="[1119,1350,548,572]" italics="true" pageId="6" pageNumber="219">Anolis apletophallus</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C7714FDBAFEDBFD86" author="Kohler" pageId="6" pageNumber="219" refString="Kohler, G. & Sunyer, J (2008) Two new species of anoles formerly referred to as anolis limifrons (Squamata: Polychrotidae). Herpetologica, 64 (1), 92 - 108." type="journal article" year="2008">Köhler & Sunyer 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
with a uniformly dull white, almost uniformly orange male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C7630FDD6FF5DFD62" authority="Kohler 2011" authorityName="Kohler" authorityYear="2011" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="tenorioensis">
|
||
<emphasis id="B92F8E139B528D3C7630FDD6FAD7FDB9" box="[1130,1348,584,608]" italics="true" pageId="6" pageNumber="219">Anolis tenorioensis</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C7714FDD9FF5DFD62" author="Kohler" pageId="6" pageNumber="219" refString="Kohler, G. (2011) A new species of anole related to Anolis altae from Volcan Tenorio, Costa Rica (Reptilia, Squamata, Polychrotidae). Zootaxa, 3120, 29 - 42." type="journal article" year="2011">Köhler 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
with a dark red with brown blotches male dewlap. Furthermore, these species have a geographic distribution in the mid-highlands of lower Central America and
|
||
<taxonomicName id="4C5B29829B528D3C708DFD0EFCAEFD41" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[727,829,656,679]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B528D3C708DFD0EFCAEFD41" box="[727,829,656,679]" italics="true" pageId="6" pageNumber="219">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B528D3C711EFD11FC0EFD4E" box="[836,925,655,680]" pageId="6" pageNumber="219" rank="species">
|
||
<emphasis id="B92F8E139B528D3C711EFD11FC0EFD4E" bold="true" box="[836,925,655,680]" pageId="6" pageNumber="219">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distributed in northwestern Andes in South America.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C341018A9B528D3C729DFD46FEBCFA9E" pageId="6" pageNumber="219" type="reference_group">
|
||
<paragraph id="8BE452019B528D3C729DFD46FABEFBF6" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="219">
|
||
<taxonomicName id="4C5B29829B528D3C729DFD46FEF3FD09" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[199,352,728,752]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B528D3C729DFD46FEF3FD09" box="[199,352,728,752]" italics="true" pageId="6" pageNumber="219">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B528D3C7333FD49FE56FD16" box="[361,453,727,752]" pageId="6" pageNumber="219" rank="species">
|
||
<emphasis id="B92F8E139B528D3C7333FD49FE56FD16" bold="true" box="[361,453,727,752]" pageId="6" pageNumber="219">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distinguished from the South America members of
|
||
<taxonomicName id="4C5B29829B528D3C7660FD46FB7FFD16" authorityName="D'Orbigny" authorityYear="1837" box="[1082,1260,728,752]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B528D3C7660FD46FB7FFD16" box="[1082,1260,728,752]" italics="true" pageId="6" pageNumber="219">A. fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
group by male dewlap and meristic patterns.
|
||
<taxonomicName id="4C5B29829B528D3C73BEFD62FDD9FCF2" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[484,586,764,788]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B528D3C73BEFD62FDD9FCF2" box="[484,586,764,788]" italics="true" pageId="6" pageNumber="219">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B528D3C700BFD62FD39FCF3" box="[593,682,764,789]" pageId="6" pageNumber="219" rank="species">
|
||
<emphasis id="B92F8E139B528D3C700BFD62FD39FCF3" bold="true" box="[593,682,764,789]" pageId="6" pageNumber="219">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have a male dewlap bright orange anteriorly and light rose color on the posterior third or half (vs.
|
||
<taxonomicName id="4C5B29829B528D3C73A7FCBEFD01FCDE" authorityName="Werner" authorityYear="1916" box="[509,658,800,824]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="B92F8E139B528D3C73A7FCBEFD01FCDE" box="[509,658,800,824]" italics="true" pageId="6" pageNumber="219">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
with solid reddish-orange male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C762AFCBEFB71FCD1" box="[1136,1250,800,823]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B528D3C762AFCBEFB71FCD1" box="[1136,1250,800,823]" italics="true" pageId="6" pageNumber="219">A. antonii</emphasis>
|
||
</taxonomicName>
|
||
with a reddishorange anterior and pink posterior male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C708CFCDAFCF9FCBA" box="[726,874,836,860]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B528D3C708CFCDAFCF9FCBA" box="[726,874,836,860]" italics="true" pageId="6" pageNumber="219">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
with orange-red anterior and yellow posterior / orange-red anterior and pink posterior male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C70ACFCF6FC23FC66" authorityName="Boulenger" authorityYear="1898" box="[758,944,872,896]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="maculiventris">
|
||
<emphasis id="B92F8E139B528D3C70ACFCF6FC23FC66" box="[758,944,872,896]" italics="true" pageId="6" pageNumber="219">A. maculiventris</emphasis>
|
||
</taxonomicName>
|
||
with orange peripherally and red medially male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C736CFC12FE3DFC42" authorityName="Ayala & Williams" authorityYear="1988" box="[310,430,908,932]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="medemi">
|
||
<emphasis id="B92F8E139B528D3C736CFC12FE3DFC42" box="[310,430,908,932]" italics="true" pageId="6" pageNumber="219">A. medemi</emphasis>
|
||
</taxonomicName>
|
||
with pink anterior and orange posterior male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C7640FC12FADEFC43" authority="Cope 1875" authorityName="Cope" authorityYear="1875" box="[1050,1357,908,933]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="trachyderma">
|
||
<emphasis id="B92F8E139B528D3C7640FC12FB59FC42" box="[1050,1226,908,932]" italics="true" pageId="6" pageNumber="219">A. trachyderma</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C7688FC12FADEFC43" author="Cope" box="[1234,1357,908,933]" pageId="6" pageNumber="219" refString="Cope, E. D. (1875) Report on the Reptiles brought by Professor James Orton from the middle and upper Amazon and western Peru. Journal of the Academy of Natural Sciences of Philadelphia, 2 (8), 159 - 183" type="journal article" year="1875">Cope 1875</bibRefCitation>
|
||
</taxonomicName>
|
||
with a reddish-orange with black scales male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C70E6FC2EFCFCFC2E" authorityName="D'Orbigny" authorityYear="1837" box="[700,879,944,968]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B528D3C70E6FC2EFCFCFC2E" box="[700,879,944,968]" italics="true" pageId="6" pageNumber="219">A. fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
with solid pink male dewlap;
|
||
<taxonomicName id="4C5B29829B528D3C768FFC2EFF5CFC0B" authority="Cope 1875" authorityName="Cope" authorityYear="1875" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="bocourti">
|
||
<emphasis id="B92F8E139B528D3C768FFC2EFAC4FC2E" box="[1237,1367,944,968]" italics="true" pageId="6" pageNumber="219">A. bocourti</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C773BFC2EFF5CFC0B" author="Cope" pageId="6" pageNumber="219" refString="Cope, E. D. (1875) Report on the Reptiles brought by Professor James Orton from the middle and upper Amazon and western Peru. Journal of the Academy of Natural Sciences of Philadelphia, 2 (8), 159 - 183" type="journal article" year="1875">Cope 1875</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B29829B528D3C735FFC4AFDC7FC0A" authority="Boulenger 1908" authorityName="Boulenger" authorityYear="1908" box="[261,596,980,1005]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="scapularis">
|
||
<emphasis id="B92F8E139B528D3C735FFC4AFE0AFC0A" box="[261,409,980,1004]" italics="true" pageId="6" pageNumber="219">A. scapularis</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B528D3C73FAFC4AFDC7FC0A" author="Boulenger" box="[416,596,980,1005]" pageId="6" pageNumber="219" refString="Boulenger, G. A. (1908) Descriptions of new batrachians and reptiles discoverd by Mr. M. G. Palmer in south-western Colombia. Annals and Magazine of Natural History, 8 (2), 515 - 522." type="journal article" year="1908">Boulenger 1908</bibRefCitation>
|
||
</taxonomicName>
|
||
have a white male dewlap (the validity of this species has been questioned and it has been considered as a synonym of
|
||
<taxonomicName id="4C5B29829B528D3C70D9FC66FCA0FBF6" authorityName="D'Orbigny" authorityYear="1837" box="[643,819,1016,1040]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B528D3C70D9FC66FCA0FBF6" box="[643,819,1016,1040]" italics="true" pageId="6" pageNumber="219">A. fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<bibRefCitation id="EFCA2FF09B528D3C7136FC66FB34FBF6" author="Poe" box="[876,1191,1015,1040]" pageId="6" pageNumber="219" refString="Poe, S. & Yanez-Miranda, C. (2008) Another New Species of Green Anolis (Squamata: Iguania) from the Eastern Andes of Peru. Journal of Herpetology, 42 (3), 564 - 571. https: // doi. org / 10.1670 / 07 - 264.1" type="journal article" year="2008">Poe & Yañez-Miranda 2008</bibRefCitation>
|
||
) (
|
||
<figureCitation id="13604E849B528D3C76E7FC69FAB3FBF6" box="[1213,1312,1015,1040]" captionStart="FIGURE 4" captionStartId="7.[151,250,1604,1626]" captionTargetBox="[169,1454,411,1583]" captionTargetId="figure@7.[169,1454,411,1583]" captionTargetPageId="7" captionText="FIGURE 4. Dewlap coloration in Andean anoles. A, Anolis urraoi sp. nov., paratype, MHUA-R 12732, photo by F. A. Grisales-Martínez; B, A. urraoi sp. nov., holotype, MHUA-R 12733, photo by F. A. Grisales-Martínez; C, A. urraoi sp. nov., paratype, MHUA-R 12541, photo by F. A. Duarte; D, A. fuscoauratus, QCAZ 8326, Pastaza, Ecuador, photo by M. R. Bustamante, FaunaWebEcuador; E, A. antonii, El Dovio, Valle del Cauca, photo by D. Alzate-Estrada; F, A. antonii, Yotoco, Valle del Cauca, photo by W. Bolívar; G, A. tolimensis, MHUA-R 12770, photo by F. A. Grisales-Martínez; H, A. mariarum MHUA-R 13033, photo by F. A. Grisales-Martínez; and I, A. mariarum MHUA-R 12730, photo by F. A. Grisales-Martínez." httpUri="https://zenodo.org/record/840910/files/figure.png" pageId="6" pageNumber="219">Figure 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE452019B528D3C729DFB82FEBCFA9E" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="219">
|
||
<taxonomicName id="4C5B29829B528D3C729DFB82FECEFBD2" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[199,349,1052,1076]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B528D3C729DFB82FECEFBD2" box="[199,349,1052,1076]" italics="true" pageId="6" pageNumber="219">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B528D3C733FFB82FE53FBD3" box="[357,448,1052,1077]" pageId="6" pageNumber="219" rank="species">
|
||
<emphasis id="B92F8E139B528D3C733FFB82FE53FBD3" bold="true" box="[357,448,1052,1077]" pageId="6" pageNumber="219">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have three to four scales between supraocular semicircles (vs.
|
||
<taxonomicName id="4C5B29829B528D3C76D3FB82FB6BFBD2" box="[1161,1272,1052,1076]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B528D3C76D3FB82FB6BFBD2" box="[1161,1272,1052,1076]" italics="true" pageId="6" pageNumber="219">A. antonii</emphasis>
|
||
</taxonomicName>
|
||
one scale), 10 to 18 scales across the snout in the second canthal (vs.
|
||
<taxonomicName id="4C5B29829B528D3C714CFBDEFC34FBB1" box="[790,935,1088,1111]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B528D3C714CFBDEFC34FBB1" box="[790,935,1088,1111]" italics="true" pageId="6" pageNumber="219">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
eight to 10 scales;
|
||
<taxonomicName id="4C5B29829B528D3C76D1FBDEFA8DFBBE" authorityName="Werner" authorityYear="1916" box="[1163,1310,1088,1112]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="B92F8E139B528D3C76D1FBDEFA8DFBBE" box="[1163,1310,1088,1112]" italics="true" pageId="6" pageNumber="219">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
nine to 10 scales;
|
||
<taxonomicName id="4C5B29829B528D3C72B1FBFAFE35FB9A" authorityName="Boulenger" authorityYear="1898" box="[235,422,1124,1148]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="maculiventris">
|
||
<emphasis id="B92F8E139B528D3C72B1FBFAFE35FB9A" box="[235,422,1124,1148]" italics="true" pageId="6" pageNumber="219">A. maculiventris</emphasis>
|
||
</taxonomicName>
|
||
10 to 14 scales), six or eight loreal rows (vs.
|
||
<taxonomicName id="4C5B29829B528D3C7198FBFAFBA0FB9A" box="[962,1075,1124,1148]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B528D3C7198FBFAFBA0FB9A" box="[962,1075,1124,1148]" italics="true" pageId="6" pageNumber="219">A. antonii</emphasis>
|
||
</taxonomicName>
|
||
five); ventral scales slightly to strongly keeled (vs. ventral scales smooth to faintly keeled in
|
||
<emphasis id="B92F8E139B528D3C7100FB16FBF8FB79" box="[858,1131,1160,1184]" italics="true" pageId="6" pageNumber="219">
|
||
<taxonomicName id="4C5B29829B528D3C7100FB16FC7DFB46" authorityName="Werner" authorityYear="1916" box="[858,1006,1160,1184]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="tolimensis">A. tolimensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5B29829B528D3C71A1FB16FBF8FB79" box="[1019,1131,1160,1183]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="antonii">A. antonii</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C5B29829B528D3C76F1FB16FAAFFB79" box="[1195,1340,1160,1183]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B528D3C76F1FB16FAAFFB79" box="[1195,1340,1160,1183]" italics="true" pageId="6" pageNumber="219">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C5B29829B528D3C770EFB16FF4CFB22" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B528D3C770EFB16FF4CFB22" italics="true" pageId="6" pageNumber="219">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B528D3C72BCFB32FED3FB23" box="[230,320,1196,1221]" pageId="6" pageNumber="219" rank="species">
|
||
<emphasis id="B92F8E139B528D3C72BCFB32FED3FB23" bold="true" box="[230,320,1196,1221]" pageId="6" pageNumber="219">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distributed in the northwestern Andes on the western slope of the Cordillera Occidental between
|
||
<date id="FFE574C19B528D3C72CDFB4EFEADFB0E" box="[151,318,1231,1256]" pageId="6" pageNumber="219" value="1700" valueMax="2220">
|
||
1700 and
|
||
<quantity id="4CA3FFE49B528D3C735CFB4EFEADFB0E" box="[262,318,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="6" pageNumber="219" unit="m" value="2220.0">2220</quantity>
|
||
</date>
|
||
m meters above sea level. In contrast
|
||
<taxonomicName id="4C5B29829B528D3C70B4FB4EFCF7FB0E" authorityName="Ayala & Williams" authorityYear="1988" box="[750,868,1232,1256]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="medemi">
|
||
<emphasis id="B92F8E139B528D3C70B4FB4EFCF7FB0E" box="[750,868,1232,1256]" italics="true" pageId="6" pageNumber="219">A. medemi</emphasis>
|
||
</taxonomicName>
|
||
is distributed in the Gorgona Island on the Pacific coast of Colombia;
|
||
<taxonomicName id="4C5B29829B528D3C73D3FB6AFDDBFAEA" authorityName="Boulenger" authorityYear="1898" box="[393,584,1268,1292]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="maculiventris">
|
||
<emphasis id="B92F8E139B528D3C73D3FB6AFDDBFAEA" box="[393,584,1268,1292]" italics="true" pageId="6" pageNumber="219">A. maculiventris</emphasis>
|
||
</taxonomicName>
|
||
in the Pacific lowlands and foothills of the Cordillera Occidental;
|
||
<taxonomicName id="4C5B29829B528D3C77DEFB6AFEBBFAD6" authorityName="D'Orbigny" authorityYear="1837" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B528D3C77DEFB6AFEBBFAD6" italics="true" pageId="6" pageNumber="219">A. fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
in the Magdalena river valley and the Amazonian region;
|
||
<taxonomicName id="4C5B29829B528D3C71B5FA86FB30FAD6" authorityName="Cope" authorityYear="1875" box="[1007,1187,1304,1328]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="trachyderma">
|
||
<emphasis id="B92F8E139B528D3C71B5FA86FB30FAD6" box="[1007,1187,1304,1328]" italics="true" pageId="6" pageNumber="219">A. trachyderma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5B29829B528D3C76EDFA86FAA8FAD6" authorityName="Cope" authorityYear="1875" box="[1207,1339,1304,1328]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="bocourti">
|
||
<emphasis id="B92F8E139B528D3C76EDFA86FAA8FAD6" box="[1207,1339,1304,1328]" italics="true" pageId="6" pageNumber="219">A. bocourti</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5B29829B528D3C77DEFA86FE9FFAB2" authorityName="Boulenger" authorityYear="1908" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="219" phylum="Chordata" rank="species" species="scapularis">
|
||
<emphasis id="B92F8E139B528D3C77DEFA86FE9FFAB2" italics="true" pageId="6" pageNumber="219">A. scapularis</emphasis>
|
||
</taxonomicName>
|
||
in the Amazonian region. Additional characteristics from the Andean fuscoauratoid group are described in the table 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C341018A9B528D30729DFA1AFC95F919" lastPageId="10" lastPageNumber="223" pageId="6" pageNumber="219" type="description">
|
||
<paragraph id="8BE452019B528D3C729DFA1AFF3FFA26" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="219">
|
||
<emphasis id="B92F8E139B528D3C729DFA1AFDFDFA7B" bold="true" box="[199,622,1412,1437]" pageId="6" pageNumber="219">Description of external morphology</emphasis>
|
||
(
|
||
<typeStatus id="54E0ECA39B528D3C7026FA1AFD73FA7B" box="[636,736,1412,1437]" pageId="6" pageNumber="219" type="holotype">holotype</typeStatus>
|
||
in parenthesis): Morphological dimensions are shown in
|
||
<tableCitation id="C6D967BA9B528D3C7704FA1AFF35FA26" captionStart="TABLE 1" captionStartId="2.[151,239,580,603]" captionText="TABLE 1. Arithmetic mean and associated standard deviation from selected morphometric characters in four species of Andean anoles. See text for abbreviations of measurements." pageId="6" pageNumber="219">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE452019B528D3C729DFA52FE7CF8CE" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="219">
|
||
<emphasis id="B92F8E139B528D3C729DFA52FE9DFA03" bold="true" box="[199,270,1484,1509]" pageId="6" pageNumber="219">Head.</emphasis>
|
||
Snout scales strongly keeled; keeling can be uni- or multicarinate; moderate prefrontal depression; shallow parietal depression. Circumnasal separated from the rostral scale by upper prenasal, lower prenasal; 6-7 (7) internasal scales; 6-8 (7) postrostral scales; distinct canthal ridge; usually 10-17 (11) scales between second canthals. Supraocular scales weakly keeled; one complete row of circumorbital scales to two rows of circumorbital scales with one incomplete; 3-4 (3) scales between supraorbital semicircles; 4-5 (4) scales between supraorbital semicircles and interparietal plate. Scales anterior to ear opening slightly larger than posterior; 6-8 (6) loreal scale rows; 40-65 (42) loreal scales; three subocular scales in narrow contact with three supralabial scales. Supralabials scales elongated with 6-7 (6) supralabial scales to level below center of eye; six (6) infralabial scales to level below center of eye; the first pair of sublabials is identical with the outer pair of postmentals; slightly enlarged outer postmental scales with 6-8 (6).
|
||
</paragraph>
|
||
<paragraph id="8BE452019B528D3C729DF8AAFE9AF83B" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="219">
|
||
<emphasis id="B92F8E139B528D3C729DF8AAFE89F8AB" bold="true" box="[199,282,1844,1869]" pageId="6" pageNumber="219">Trunk.</emphasis>
|
||
Dorsal scales keeled-conical; subimbricate scales with rounded posterior margins the 7-10 (9) median rows moderately enlarged; 80-106 (88) dorsal scales between levels of axilla and groin; lateral body scales homogeneous conical without axillary depression; ventral scales slightly to strongly keeled, subimbricate scales with mostly rounded posterior margins, some slightly mucronate; 58-71 (60) ventral scales between levels of axilla and groin.
|
||
</paragraph>
|
||
<paragraph id="8BE452019B538D3D729DFF09FD46FF1E" blockId="7.[151,1436,151,357]" pageId="7" pageNumber="220">
|
||
<emphasis id="B92F8E139B538D3D729DFF09FEBBFF56" bold="true" box="[199,296,151,176]" pageId="7" pageNumber="220">Dewlap.</emphasis>
|
||
Dewlap large-sized, the anterior insertion is below level of eye, the posterior insertion onto chest; lateral scales subequal to ventrals in 8-10 (8) rows descending posteriorly, each a single scale wide; edge scales small, imbricate, keeled, closely peaked (
|
||
<figureCitation id="13604E849B538D3D703FFF41FD5BFF1E" box="[613,712,223,248]" captionStart="FIGURE 4" captionStartId="7.[151,250,1604,1626]" captionTargetBox="[169,1454,411,1583]" captionTargetId="figure@7.[169,1454,411,1583]" captionTargetPageId="7" captionText="FIGURE 4. Dewlap coloration in Andean anoles. A, Anolis urraoi sp. nov., paratype, MHUA-R 12732, photo by F. A. Grisales-Martínez; B, A. urraoi sp. nov., holotype, MHUA-R 12733, photo by F. A. Grisales-Martínez; C, A. urraoi sp. nov., paratype, MHUA-R 12541, photo by F. A. Duarte; D, A. fuscoauratus, QCAZ 8326, Pastaza, Ecuador, photo by M. R. Bustamante, FaunaWebEcuador; E, A. antonii, El Dovio, Valle del Cauca, photo by D. Alzate-Estrada; F, A. antonii, Yotoco, Valle del Cauca, photo by W. Bolívar; G, A. tolimensis, MHUA-R 12770, photo by F. A. Grisales-Martínez; H, A. mariarum MHUA-R 13033, photo by F. A. Grisales-Martínez; and I, A. mariarum MHUA-R 12730, photo by F. A. Grisales-Martínez." httpUri="https://zenodo.org/record/840910/files/figure.png" pageId="7" pageNumber="220">Figure 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE452019B538D3D729DFE9AFC7BFE83" blockId="7.[151,1436,151,357]" pageId="7" pageNumber="220">
|
||
<emphasis id="B92F8E139B538D3D729DFE9AFE89FEFB" bold="true" box="[199,282,260,285]" pageId="7" pageNumber="220">Limbs.</emphasis>
|
||
Limb scales unicarinate dorsally, granular posteriorly and ventrally; supradigital scales strongly keeled; 13-16 (14) lamellae under phalanges II and III of fourth toe; subdigital pad is clearly dilated, while the width of the subdigital scales of the distal phalanx does not show dilatation.
|
||
</paragraph>
|
||
<caption id="DF2402899B538D3D72CDF9DAFAE5F915" httpUri="https://zenodo.org/record/840910/files/figure.png" pageId="7" pageNumber="220" targetBox="[169,1454,411,1583]" targetPageId="7">
|
||
<paragraph id="8BE452019B538D3D72CDF9DAFAE5F915" blockId="7.[151,1436,1604,1779]" pageId="7" pageNumber="220">
|
||
<emphasis id="B92F8E139B538D3D72CDF9DAFE84F9BF" bold="true" box="[151,279,1604,1626]" pageId="7" pageNumber="220">FIGURE 4.</emphasis>
|
||
Dewlap coloration in Andean anoles.
|
||
<emphasis id="B92F8E139B538D3D70E3F9DAFD58F9BF" bold="true" box="[697,715,1604,1625]" pageId="7" pageNumber="220">A</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B538D3D7087F9DAFCF9F9BF" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[733,874,1604,1625]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B538D3D7087F9DAFCF9F9BF" box="[733,874,1604,1625]" italics="true" pageId="7" pageNumber="220">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B538D3D712FF9DAFC59F9BF" box="[885,970,1604,1625]" pageId="7" pageNumber="220" rank="species">
|
||
<emphasis id="B92F8E139B538D3D712FF9DAFC59F9BF" bold="true" box="[885,970,1604,1625]" pageId="7" pageNumber="220">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, MHUA-R 12732, photo by F.A. Grisales-Martínez;
|
||
<emphasis id="B92F8E139B538D3D7306F9FDFEFEF99E" bold="true" box="[348,365,1635,1656]" pageId="7" pageNumber="220">B</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B538D3D7321F9FDFE4AF99E" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[379,473,1635,1656]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B538D3D7321F9FDFE4AF99E" box="[379,473,1635,1656]" italics="true" pageId="7" pageNumber="220">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B538D3D73BBF9FDFDA1F99E" box="[481,562,1635,1656]" pageId="7" pageNumber="220" rank="species">
|
||
<emphasis id="B92F8E139B538D3D73BBF9FDFDA1F99E" bold="true" box="[481,562,1635,1656]" pageId="7" pageNumber="220">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype, MHUA-R 12733, photo by F.A. Grisales-Martínez;
|
||
<emphasis id="B92F8E139B538D3D76E7F9FCFB5CF99E" bold="true" box="[1213,1231,1634,1656]" pageId="7" pageNumber="220">C</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B538D3D7687F9FDFAAFF99E" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[1245,1340,1635,1656]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B538D3D7687F9FDFAAFF99E" box="[1245,1340,1635,1656]" italics="true" pageId="7" pageNumber="220">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B538D3D7719F9FDFA06F99E" box="[1347,1429,1635,1656]" pageId="7" pageNumber="220" rank="species">
|
||
<emphasis id="B92F8E139B538D3D7719F9FDFA06F99E" bold="true" box="[1347,1429,1635,1656]" pageId="7" pageNumber="220">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, MHUA-R 12541, photo by F.A. Duarte;
|
||
<emphasis id="B92F8E139B538D3D7098F91FFD47F970" bold="true" box="[706,724,1665,1686]" pageId="7" pageNumber="220">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B538D3D70B2F91CFC1EF970" authorityName="D'Orbigny" authorityYear="1837" box="[744,909,1665,1687]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B538D3D70B2F91CFC1EF970" box="[744,909,1665,1687]" italics="true" pageId="7" pageNumber="220">A. fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
, QCAZ 8326, Pastaza, Ecuador, photo by M.R. Bustamante, FaunaWebEcuador;
|
||
<emphasis id="B92F8E139B538D3D73B1F93EFE6FF953" bold="true" box="[491,508,1696,1717]" pageId="7" pageNumber="220">E</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B538D3D7050F93EFD4EF953" authority=", El Dovio" authorityName="El Dovio" box="[522,733,1695,1717]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B538D3D7050F93EFDFCF953" box="[522,623,1696,1717]" italics="true" pageId="7" pageNumber="220">A. antonii</emphasis>
|
||
, El Dovio
|
||
</taxonomicName>
|
||
, Valle del Cauca, photo by D. Alzate-Estrada;
|
||
<emphasis id="B92F8E139B538D3D76E5F93EFB5FF953" bold="true" box="[1215,1228,1696,1717]" pageId="7" pageNumber="220">F</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B538D3D7681F93EFA0BF953" authority=", Yotoco" authorityName="Yotoco" box="[1243,1432,1696,1717]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B538D3D7681F93EFAD3F953" box="[1243,1344,1696,1717]" italics="true" pageId="7" pageNumber="220">A. antonii</emphasis>
|
||
, Yotoco
|
||
</taxonomicName>
|
||
, Valle del Cauca, photo by W. Bolívar;
|
||
<emphasis id="B92F8E139B538D3D7071F920FDAFF932" bold="true" box="[555,572,1726,1748]" pageId="7" pageNumber="220">G</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B538D3D7010F921FD5DF935" authorityName="Werner" authorityYear="1916" box="[586,718,1726,1748]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="B92F8E139B538D3D7010F921FD5DF935" box="[586,718,1726,1748]" italics="true" pageId="7" pageNumber="220">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
, MHUA-R 12770, photo by F.A. Grisales-Martínez;
|
||
<emphasis id="B92F8E139B538D3D76ACF921FA99F932" bold="true" box="[1270,1290,1727,1748]" pageId="7" pageNumber="220">H</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B538D3D7743F921FA0FF932" box="[1305,1436,1727,1748]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B538D3D7743F921FA0FF932" box="[1305,1436,1727,1748]" italics="true" pageId="7" pageNumber="220">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
MHUA-R 13033, photo by F.A. Grisales-Martínez; and
|
||
<emphasis id="B92F8E139B538D3D708AF943FD49F914" bold="true" box="[720,730,1757,1778]" pageId="7" pageNumber="220">I</emphasis>
|
||
,
|
||
<taxonomicName id="4C5B29829B538D3D70BCF943FCF4F915" box="[742,871,1757,1779]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B538D3D70BCF943FCF4F915" box="[742,871,1757,1779]" italics="true" pageId="7" pageNumber="220">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
MHUA-R 12730, photo by F.A. Grisales-Martínez.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE452019B538D3D729DF881FBADF8BA" blockId="7.[151,1437,1823,2029]" pageId="7" pageNumber="220">
|
||
<emphasis id="B92F8E139B538D3D729DF881FF69F8DE" bold="true" box="[199,250,1823,1848]" pageId="7" pageNumber="220">Tail.</emphasis>
|
||
Tail slender, cylindrical or very slightly compressed, somewhat swollen at the base in males, all scales keeled, median dorsal scales slightly enlarged; postcloacal scales not differentiated.
|
||
</paragraph>
|
||
<paragraph id="8BE452019B538D3D729DF8F9FC8DF80B" blockId="7.[151,1437,1823,2029]" pageId="7" pageNumber="220">
|
||
<emphasis id="B92F8E139B538D3D729DF8F9FED3F866" bold="true" box="[199,320,1895,1920]" pageId="7" pageNumber="220">Hemipenis</emphasis>
|
||
. Preserved males did not have well everted hemipenis and detailed description of relevant characters could not be recovered. In general, the hemipenis in
|
||
<taxonomicName id="4C5B29829B538D3D70E4F812FCD8F845" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[702,843,1932,1956]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="220" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B538D3D70E4F812FCD8F845" box="[702,843,1932,1956]" italics="true" pageId="7" pageNumber="220">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B538D3D710BF815FC36F842" box="[849,933,1931,1956]" pageId="7" pageNumber="220" rank="species">
|
||
<emphasis id="B92F8E139B538D3D710BF815FC36F842" bold="true" box="[849,933,1931,1956]" pageId="7" pageNumber="220">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is small, tubular and slightly bifurcated distally.
|
||
<emphasis id="B92F8E139B538D3D729DF831FECCF82E" bold="true" box="[199,351,1967,1992]" pageId="7" pageNumber="220">Color in life.</emphasis>
|
||
Some specimens are brown with a black stripe on the back, while others have a greenish color. The colors are somewhat sexually dimorphic (see below).
|
||
</paragraph>
|
||
<paragraph id="8BE452019B5C8D32729DFF06FC29FE6E" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="221">
|
||
<emphasis id="B92F8E139B5C8D32729DFF06FEB5FF56" box="[199,294,152,176]" italics="true" pageId="8" pageNumber="221">Females</emphasis>
|
||
: Head dark greenish brown to black; a brown or yellow crossbar between eyes sometimes present. Body brown or yellow-brown to yellowish or olive green, usually black or very dark brown middorsally, occasionally interrupted with brown spots; sides brown or yellow-brown, occasionally with darker vertical shadows or small rufous spots. In some females a slender light brown vertebral stripe is bordered laterally by thin dark brown lines extending from head to tail. Gular and belly regions white, usually maculate, with dark brown spots or striations. Legs brown, sometimes with narrow pale crossbands. Tail brown, usually with broad alternating dark brown and yellow brown bands, ventrally white on proximal third.
|
||
</paragraph>
|
||
<paragraph id="8BE452019B5C8D32729DFE0AFF72FDDB" blockId="8.[151,1437,151,573]" pageId="8" pageNumber="221">
|
||
<emphasis id="B92F8E139B5C8D32729DFE0AFE98FE4A" box="[199,267,404,428]" italics="true" pageId="8" pageNumber="221">Males</emphasis>
|
||
: Head and body similar to females, more often greenish brown, black or dark brown middorsal region usually broader than in females, sometimes crossed with this greenish bands. Sides brown, tan or yellowish green, sometimes with small white spots. Gular and belly areas cream, infrequently spotted or striated with dark brown. Dewlap is bright orange anteriorly and light rose color on the posterior third or half, longitudinal rows of white scales.
|
||
</paragraph>
|
||
<caption id="DF2402899B5C8D3272CDFDF2FB67FD65" box="[151,1268,619,643]" pageId="8" pageNumber="221">
|
||
<paragraph id="8BE452019B5C8D3272CDFDF2FB67FD65" blockId="8.[151,1268,619,643]" box="[151,1268,619,643]" pageId="8" pageNumber="221">
|
||
<emphasis id="B92F8E139B5C8D3272CDFDF2FE9AFD65" bold="true" box="[151,265,620,643]" pageId="8" pageNumber="221">TABLE 2.</emphasis>
|
||
Comparison of morphological traits among species in the Andean
|
||
<taxonomicName id="4C5B29829B5C8D32718EFDF5FB34FD65" authorityName="D'Orbigny" authorityYear="1837" box="[980,1191,619,643]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="221" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B5C8D32718EFDF5FB34FD65" box="[980,1191,619,643]" italics="true" pageId="8" pageNumber="221">Anolis fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE452019B5C8D3272C5FD02FABEF8E0" pageId="8" pageNumber="221">
|
||
<table id="F95BA0A19B5C72C572CDFD05FA01F8C2" box="[151,1426,667,1828]" gridcols="5" gridrows="10" pageId="8" pageNumber="221">
|
||
<tr id="356B50439B5C72C572CDFD05FA01FD52" box="[151,1426,667,692]" gridrow="0" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDFD05FE36FD52" box="[151,421,667,692]" gridcol="0" gridrow="0" pageId="8" pageNumber="221">Character</th>
|
||
<th id="76BA393F9B5C72C573E0FD05FD08FD52" box="[442,667,667,692]" gridcol="1" gridrow="0" pageId="8" pageNumber="221">
|
||
<taxonomicName id="4C5B29829B5C8D3273E0FD00FDB1FD55" box="[442,546,670,691]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="221" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B5C8D3273E0FD00FDB1FD55" bold="true" box="[442,546,670,691]" italics="true" pageId="8" pageNumber="221">A. antonii</emphasis>
|
||
</taxonomicName>
|
||
1
|
||
</th>
|
||
<th id="76BA393F9B5C72C570EDFD05FC25FD52" box="[695,950,667,692]" gridcol="2" gridrow="0" pageId="8" pageNumber="221">
|
||
<taxonomicName id="4C5B29829B5C8D3270EDFD00FCACFD55" box="[695,831,670,691]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="221" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B5C8D3270EDFD00FCACFD55" bold="true" box="[695,831,670,691]" italics="true" pageId="8" pageNumber="221">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
2
|
||
</th>
|
||
<th id="76BA393F9B5C72C57190FD05FB55FD52" box="[970,1222,667,692]" gridcol="3" gridrow="0" pageId="8" pageNumber="221">
|
||
<taxonomicName id="4C5B29829B5C8D327191FD00FBC1FD52" authorityName="Werner" authorityYear="1916" box="[971,1106,669,692]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="221" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="B92F8E139B5C8D327191FD00FBC1FD52" bold="true" box="[971,1106,669,692]" italics="true" pageId="8" pageNumber="221">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
3
|
||
</th>
|
||
<th id="76BA393F9B5C72C57680FD05FA01FD52" box="[1242,1426,667,692]" gridcol="4" gridrow="0" pageId="8" pageNumber="221">
|
||
<taxonomicName id="4C5B29829B5C8D327681FD01FAABFD52" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[1243,1336,671,692]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="221" phylum="Chordata" rank="species" species="urraoi">
|
||
<emphasis id="B92F8E139B5C8D327681FD01FAABFD52" box="[1243,1336,671,692]" italics="true" pageId="8" pageNumber="221">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
4
|
||
</th>
|
||
</tr>
|
||
<tr id="356B50439B5C72C572CDFD5CFA01FD11" box="[151,1426,706,759]" gridrow="1" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDFD5CFE36FD11" box="[151,421,706,759]" gridcol="0" gridrow="1" pageId="8" pageNumber="221">Scales between semicircles</th>
|
||
<td id="76BA393F9B5C72C573E0FD5CFD08FD11" box="[442,667,706,759]" gridcol="1" gridrow="1" pageId="8" pageNumber="221">1</td>
|
||
<td id="76BA393F9B5C72C570EDFD5CFC25FD11" box="[695,950,706,759]" gridcol="2" gridrow="1" pageId="8" pageNumber="221">1–2</td>
|
||
<td id="76BA393F9B5C72C57190FD5CFB55FD11" box="[970,1222,706,759]" gridcol="3" gridrow="1" pageId="8" pageNumber="221">2–3</td>
|
||
<td id="76BA393F9B5C72C57680FD5CFA01FD11" box="[1242,1426,706,759]" gridcol="4" gridrow="1" pageId="8" pageNumber="221">3–4</td>
|
||
</tr>
|
||
<tr id="356B50439B5C72C572CDFC9AFA01FCDC" box="[151,1426,772,826]" gridrow="2" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDFC9AFE36FCDC" box="[151,421,772,826]" gridcol="0" gridrow="2" pageId="8" pageNumber="221">Scales between second canthals</th>
|
||
<td id="76BA393F9B5C72C573E0FC9AFD08FCDC" box="[442,667,772,826]" gridcol="1" gridrow="2" pageId="8" pageNumber="221">Undescribed</td>
|
||
<td id="76BA393F9B5C72C570EDFC9AFC25FCDC" box="[695,950,772,826]" gridcol="2" gridrow="2" pageId="8" pageNumber="221">8–10</td>
|
||
<td id="76BA393F9B5C72C57190FC9AFB55FCDC" box="[970,1222,772,826]" gridcol="3" gridrow="2" pageId="8" pageNumber="221">9–10</td>
|
||
<td id="76BA393F9B5C72C57680FC9AFA01FCDC" box="[1242,1426,772,826]" gridcol="4" gridrow="2" pageId="8" pageNumber="221">10–18</td>
|
||
</tr>
|
||
<tr id="356B50439B5C72C572CDFCD9FA01FCB8" box="[151,1426,839,862]" gridrow="3" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDFCD9FE36FCB8" box="[151,421,839,862]" gridcol="0" gridrow="3" pageId="8" pageNumber="221">Loreal rows</th>
|
||
<td id="76BA393F9B5C72C573E0FCD9FD08FCB8" box="[442,667,839,862]" gridcol="1" gridrow="3" pageId="8" pageNumber="221">5</td>
|
||
<td id="76BA393F9B5C72C570EDFCD9FC25FCB8" box="[695,950,839,862]" gridcol="2" gridrow="3" pageId="8" pageNumber="221">5–7</td>
|
||
<td id="76BA393F9B5C72C57190FCD9FB55FCB8" box="[970,1222,839,862]" gridcol="3" gridrow="3" pageId="8" pageNumber="221">6–7</td>
|
||
<td id="76BA393F9B5C72C57680FCD9FA01FCB8" box="[1242,1426,839,862]" gridcol="4" gridrow="3" pageId="8" pageNumber="221">6–8</td>
|
||
</tr>
|
||
<tr id="356B50439B5C72C572CDFCF5FA01FC46" box="[151,1426,875,928]" gridrow="4" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDFCF5FE36FC46" box="[151,421,875,928]" gridcol="0" gridrow="4" pageId="8" pageNumber="221">Supralabials to below the centre of the eye</th>
|
||
<td id="76BA393F9B5C72C573E0FCF5FD08FC46" box="[442,667,875,928]" gridcol="1" gridrow="4" pageId="8" pageNumber="221">6–7</td>
|
||
<td id="76BA393F9B5C72C570EDFCF5FC25FC46" box="[695,950,875,928]" gridcol="2" gridrow="4" pageId="8" pageNumber="221">6–7</td>
|
||
<td id="76BA393F9B5C72C57190FCF5FB55FC46" box="[970,1222,875,928]" gridcol="3" gridrow="4" pageId="8" pageNumber="221">6–7</td>
|
||
<td id="76BA393F9B5C72C57680FCF5FA01FC46" box="[1242,1426,875,928]" gridcol="4" gridrow="4" pageId="8" pageNumber="221">6–7</td>
|
||
</tr>
|
||
<tr id="356B50439B5C72C572CDFC30FA01FBE4" box="[151,1426,942,1026]" gridrow="5" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDFC30FE36FBE4" box="[151,421,942,1026]" gridcol="0" gridrow="5" pageId="8" pageNumber="221">Gular scales</th>
|
||
<td id="76BA393F9B5C72C573E0FC30FD08FBE4" box="[442,667,942,1026]" gridcol="1" gridrow="5" pageId="8" pageNumber="221">Feebly keeled</td>
|
||
<td id="76BA393F9B5C72C570EDFC30FC25FBE4" box="[695,950,942,1026]" gridcol="2" gridrow="5" pageId="8" pageNumber="221">Feebly keeled</td>
|
||
<td id="76BA393F9B5C72C57190FC30FB55FBE4" box="[970,1222,942,1026]" gridcol="3" gridrow="5" pageId="8" pageNumber="221">Feebly keeled</td>
|
||
<td id="76BA393F9B5C72C57680FC30FA01FBE4" box="[1242,1426,942,1026]" gridcol="4" gridrow="5" pageId="8" pageNumber="221">Edge scales small, imbricate, keeled, closely peaked</td>
|
||
</tr>
|
||
<tr id="356B50439B5C72C572CDFB91FA01FAFD" box="[151,1426,1039,1307]" gridrow="6" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDFB91FE36FAFD" box="[151,421,1039,1307]" gridcol="0" gridrow="6" pageId="8" pageNumber="221">Dorsal scales</th>
|
||
<td id="76BA393F9B5C72C573E0FB91FD08FAFD" box="[442,667,1039,1307]" gridcol="1" gridrow="6" pageId="8" pageNumber="221">Dorsal scales small, flat, keeled, becoming gradually smaller towards the sides, where they are minute and granular</td>
|
||
<td id="76BA393F9B5C72C570EDFB91FC25FAFD" box="[695,950,1039,1307]" gridcol="2" gridrow="6" pageId="8" pageNumber="221">Dorsal scales small, conical granules, the median rows being slightly enlarged and keeled, laterals becoming very small.</td>
|
||
<td id="76BA393F9B5C72C57190FB91FB55FAFD" box="[970,1222,1039,1307]" gridcol="3" gridrow="6" pageId="8" pageNumber="221">Dorsal scales small, conical granules, the median rows being slightly enlarged and keeled, laterals becoming very small.</td>
|
||
<td id="76BA393F9B5C72C57680FB91FA01FAFD" box="[1242,1426,1039,1307]" gridcol="4" gridrow="6" pageId="8" pageNumber="221">Dorsal scales keeled-conical; subimbricate scales with rounded posterior margins the 7-10 median rows moderately enlarged.</td>
|
||
</tr>
|
||
<tr id="356B50439B5C72C572CDFAA1FA01F9CA" box="[151,1426,1343,1580]" gridrow="7" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDFAA1FE36F9CA" box="[151,421,1343,1580]" gridcol="0" gridrow="7" pageId="8" pageNumber="221">Belly and chest scales</th>
|
||
<td id="76BA393F9B5C72C573E0FAA1FD08F9CA" box="[442,667,1343,1580]" gridcol="1" gridrow="7" pageId="8" pageNumber="221">Ventral scales larger than dorsals, flat, juxtaposed, smooth to faintly keeled</td>
|
||
<td id="76BA393F9B5C72C570EDFAA1FC25F9CA" box="[695,950,1343,1580]" gridcol="2" gridrow="7" pageId="8" pageNumber="221">Ventral scales much larger than dorsals, flat, subimbricate, cycloid, smooth to faintly keeled.</td>
|
||
<td id="76BA393F9B5C72C57190FAA1FB55F9CA" box="[970,1222,1343,1580]" gridcol="3" gridrow="7" pageId="8" pageNumber="221">Ventral scales larger than dorsals, flat, subimbricate, cycloid, smooth.</td>
|
||
<td id="76BA393F9B5C72C57680FAA1FA01F9CA" box="[1242,1426,1343,1580]" gridcol="4" gridrow="7" pageId="8" pageNumber="221">Ventral scales larger than dorsals, some slightly mucronate, subimbricate, slightly to strongly keeled.</td>
|
||
</tr>
|
||
<tr id="356B50439B5C72C572CDF9CFFA01F904" box="[151,1426,1617,1762]" gridrow="8" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDF9CFFE36F904" box="[151,421,1617,1762]" gridcol="0" gridrow="8" pageId="8" pageNumber="221">Dewlap male</th>
|
||
<td id="76BA393F9B5C72C573E0F9CFFD08F904" box="[442,667,1617,1762]" gridcol="1" gridrow="8" pageId="8" pageNumber="221">Reddish-orange anterior and pink posterior.</td>
|
||
<td id="76BA393F9B5C72C570EDF9CFFC25F904" box="[695,950,1617,1762]" gridcol="2" gridrow="8" pageId="8" pageNumber="221">Orange-red anterior and yellow posterior/ Orange-red anterior and pink posterior.</td>
|
||
<td id="76BA393F9B5C72C57190F9CFFB55F904" box="[970,1222,1617,1762]" gridcol="3" gridrow="8" pageId="8" pageNumber="221">Solid reddish-orange</td>
|
||
<td id="76BA393F9B5C72C57680F9CFFA01F904" box="[1242,1426,1617,1762]" gridcol="4" gridrow="8" pageId="8" pageNumber="221">Yellowish-orange anteriorly and opaque pink on the posterior third or half.</td>
|
||
</tr>
|
||
<tr id="356B50439B5C72C572CDF96EFA01F8C2" box="[151,1426,1776,1828]" gridrow="9" pageId="8" pageNumber="221">
|
||
<th id="76BA393F9B5C72C572CDF96EFE36F8C2" box="[151,421,1776,1828]" gridcol="0" gridrow="9" pageId="8" pageNumber="221">Lamellae under phalanges II and III of fourth toe</th>
|
||
<td id="76BA393F9B5C72C573E0F96EFD08F8C2" box="[442,667,1776,1828]" gridcol="1" gridrow="9" pageId="8" pageNumber="221">16</td>
|
||
<td id="76BA393F9B5C72C570EDF96EFC25F8C2" box="[695,950,1776,1828]" gridcol="2" gridrow="9" pageId="8" pageNumber="221">15 to 16</td>
|
||
<td id="76BA393F9B5C72C57190F96EFB55F8C2" box="[970,1222,1776,1828]" gridcol="3" gridrow="9" pageId="8" pageNumber="221">13 to 16</td>
|
||
<td id="76BA393F9B5C72C57680F96EFA01F8C2" box="[1242,1426,1776,1828]" gridcol="4" gridrow="9" pageId="8" pageNumber="221">13 to 16</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BE452019B5C8D3272CDF8DDFCF9F847" blockId="8.[151,1410,1859,1988]" pageId="8" pageNumber="221">
|
||
1 Based on the description of
|
||
<taxonomicName id="4C5B29829B5C8D327395F8DBFD7AF8BA" authority="Boulenger 1908" authorityName="Boulenger" authorityYear="1908" box="[463,745,1861,1884]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="221" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B5C8D327395F8DBFDABF8BA" box="[463,568,1861,1884]" italics="true" pageId="8" pageNumber="221">A. antonii</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B5C8D327065F8DBFD7AF8BA" author="Boulenger" box="[575,745,1861,1884]" pageId="8" pageNumber="221" refString="Boulenger, G. A. (1908) Descriptions of new batrachians and reptiles discoverd by Mr. M. G. Palmer in south-western Colombia. Annals and Magazine of Natural History, 8 (2), 515 - 522." type="journal article" year="1908">Boulenger 1908</bibRefCitation>
|
||
</taxonomicName>
|
||
and specimen photographs. 2 Based on the description of
|
||
<taxonomicName id="4C5B29829B5C8D327395F8F9FD62F898" authority="Barbour 1932" authorityName="Barbour" authorityYear="1932" box="[463,753,1895,1919]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="221" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B5C8D327395F8F9FDCBF899" box="[463,600,1895,1919]" italics="true" pageId="8" pageNumber="221">A. mariarum</emphasis>
|
||
<bibRefCitation id="EFCA2FF09B5C8D327004F8F9FD62F898" author="Barbour" box="[606,753,1895,1918]" pageId="8" pageNumber="221" refString="Barbour, T. (1932) New Anoles. Proceedings of the New England Zoological Club, 12, 97 - 102" type="journal article" year="1932">Barbour 1932</bibRefCitation>
|
||
</taxonomicName>
|
||
and fresh specimens from the type locality (see Appendix 1). 3 Based on fresh specimens from the type locality (see Appendix 1).
|
||
</paragraph>
|
||
<paragraph id="8BE452019B5C8D3272CDF835FE15F822" blockId="8.[151,1410,1859,1988]" box="[151,390,1963,1988]" pageId="8" pageNumber="221">4 Based on series type.</paragraph>
|
||
<caption id="DF2402899B5D8D3372CDFF06FD21FF09" pageId="9" pageNumber="222">
|
||
<paragraph id="8BE452019B5D8D3372CDFF06FD21FF09" blockId="9.[151,1436,152,239]" pageId="9" pageNumber="222">
|
||
<emphasis id="B92F8E139B5D8D3372CDFF06FE9CFF49" bold="true" box="[151,271,152,175]" pageId="9" pageNumber="222">TABLE 3.</emphasis>
|
||
Loadings of the SVL and 22 traits corrected for body size (see main text for details) in the principal component analysis (PCA) for adult specimens of the four Andean species of the
|
||
<taxonomicName id="4C5B29829B5D8D33765DFF29FB4FFF29" authorityName="D'Orbigny" authorityYear="1837" box="[1031,1244,183,207]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B5D8D33765DFF29FB4FFF29" box="[1031,1244,183,207]" italics="true" pageId="9" pageNumber="222">Anolis fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
species group:
|
||
<taxonomicName id="4C5B29829B5D8D3377DFFF29FF71FF09" authorityName="Boulenger" authorityYear="1908" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B5D8D3377DFFF29FF71FF09" italics="true" pageId="9" pageNumber="222">A. antonii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5B29829B5D8D3372AAFF49FEE4FF09" authorityName="Barbour" authorityYear="1932" box="[240,375,215,239]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B5D8D3372AAFF49FEE4FF09" box="[240,375,215,239]" italics="true" pageId="9" pageNumber="222">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5B29829B5D8D3373DFFF49FD9CFF09" authorityName="Werner" authorityYear="1916" box="[389,527,215,239]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="B92F8E139B5D8D3373DFFF49FD9CFF09" box="[389,527,215,239]" italics="true" pageId="9" pageNumber="222">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5B29829B5D8D337013FF49FD38FF09" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[585,683,215,239]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="urraoi">
|
||
<emphasis id="B92F8E139B5D8D337013FF49FD38FF09" box="[585,683,215,239]" italics="true" pageId="9" pageNumber="222">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE452019B5D8D3372C5FE94FA95FAE6" pageId="9" pageNumber="222">
|
||
<table id="F95BA0A19B5D72C572C5FE94FA9CFAE6" box="[159,1295,266,1280]" gridcols="5" gridrows="25" pageId="9" pageNumber="222">
|
||
<tr id="356B50439B5D72C572C5FE94FA9CFEC6" box="[159,1295,266,288]" gridrow="0" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FE94FEE7FEC6" box="[159,372,266,288]" gridcol="0" gridrow="0" pageId="9" pageNumber="222">Variable</th>
|
||
<th id="76BA393F9B5D72C5700AFE94FD02FEC6" box="[592,657,266,288]" gridcol="1" gridrow="0" pageId="9" pageNumber="222">PC1</th>
|
||
<th id="76BA393F9B5D72C5717EFE94FCF5FEC6" box="[804,870,266,288]" gridcol="2" gridrow="0" pageId="9" pageNumber="222">PC2</th>
|
||
<th id="76BA393F9B5D72C571A3FE94FBA8FEC6" box="[1017,1083,266,288]" gridcol="3" gridrow="0" pageId="9" pageNumber="222">PC3</th>
|
||
<th id="76BA393F9B5D72C57697FE94FA9CFEC6" box="[1229,1295,266,288]" gridcol="4" gridrow="0" pageId="9" pageNumber="222">PC4</th>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FEAAFA9CFEAC" box="[159,1295,308,330]" gridrow="1" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FEAAFEE7FEAC" box="[159,372,308,330]" gridcol="0" gridrow="1" pageId="9" pageNumber="222">SVL</th>
|
||
<td id="76BA393F9B5D72C5700AFEAAFD02FEAC" box="[592,657,308,330]" gridcol="1" gridrow="1" pageId="9" pageNumber="222">-0.240</td>
|
||
<td id="76BA393F9B5D72C5717EFEAAFCF5FEAC" box="[804,870,308,330]" gridcol="2" gridrow="1" pageId="9" pageNumber="222">0.024</td>
|
||
<td id="76BA393F9B5D72C571A3FEAAFBA8FEAC" box="[1017,1083,308,330]" gridcol="3" gridrow="1" pageId="9" pageNumber="222">-0.037</td>
|
||
<td id="76BA393F9B5D72C57697FEAAFA9CFEAC" box="[1229,1295,308,330]" gridcol="4" gridrow="1" pageId="9" pageNumber="222">0.009</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FEC3FA9CFE95" box="[159,1295,349,371]" gridrow="2" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FEC3FEE7FE95" box="[159,372,349,371]" gridcol="0" gridrow="2" pageId="9" pageNumber="222">TL</th>
|
||
<td id="76BA393F9B5D72C5700AFEC3FD02FE95" box="[592,657,349,371]" gridcol="1" gridrow="2" pageId="9" pageNumber="222">-0.165</td>
|
||
<td id="76BA393F9B5D72C5717EFEC3FCF5FE95" box="[804,870,349,371]" gridcol="2" gridrow="2" pageId="9" pageNumber="222">0.250</td>
|
||
<td id="76BA393F9B5D72C571A3FEC3FBA8FE95" box="[1017,1083,349,371]" gridcol="3" gridrow="2" pageId="9" pageNumber="222">0.334</td>
|
||
<td id="76BA393F9B5D72C57697FEC3FA9CFE95" box="[1229,1295,349,371]" gridcol="4" gridrow="2" pageId="9" pageNumber="222">-0.534</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FE18FA9CFE7A" box="[159,1295,390,412]" gridrow="3" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FE18FEE7FE7A" box="[159,372,390,412]" gridcol="0" gridrow="3" pageId="9" pageNumber="222">HL</th>
|
||
<td id="76BA393F9B5D72C5700AFE18FD02FE7A" box="[592,657,390,412]" gridcol="1" gridrow="3" pageId="9" pageNumber="222">-0.237</td>
|
||
<td id="76BA393F9B5D72C5717EFE18FCF5FE7A" box="[804,870,390,412]" gridcol="2" gridrow="3" pageId="9" pageNumber="222">0.045</td>
|
||
<td id="76BA393F9B5D72C571A3FE18FBA8FE7A" box="[1017,1083,390,412]" gridcol="3" gridrow="3" pageId="9" pageNumber="222">-0.009</td>
|
||
<td id="76BA393F9B5D72C57697FE18FA9CFE7A" box="[1229,1295,390,412]" gridcol="4" gridrow="3" pageId="9" pageNumber="222">0.055</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FE2EFA9CFE20" box="[159,1295,432,454]" gridrow="4" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FE2EFEE7FE20" box="[159,372,432,454]" gridcol="0" gridrow="4" pageId="9" pageNumber="222">HW</th>
|
||
<td id="76BA393F9B5D72C5700AFE2EFD02FE20" box="[592,657,432,454]" gridcol="1" gridrow="4" pageId="9" pageNumber="222">-0.210</td>
|
||
<td id="76BA393F9B5D72C5717EFE2EFCF5FE20" box="[804,870,432,454]" gridcol="2" gridrow="4" pageId="9" pageNumber="222">0.117</td>
|
||
<td id="76BA393F9B5D72C571A3FE2EFBA8FE20" box="[1017,1083,432,454]" gridcol="3" gridrow="4" pageId="9" pageNumber="222">-0.057</td>
|
||
<td id="76BA393F9B5D72C57697FE2EFA9CFE20" box="[1229,1295,432,454]" gridcol="4" gridrow="4" pageId="9" pageNumber="222">0.157</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FE47FA9CFE09" box="[159,1295,473,495]" gridrow="5" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FE47FEE7FE09" box="[159,372,473,495]" gridcol="0" gridrow="5" pageId="9" pageNumber="222">JL</th>
|
||
<td id="76BA393F9B5D72C5700AFE47FD02FE09" box="[592,657,473,495]" gridcol="1" gridrow="5" pageId="9" pageNumber="222">-0.210</td>
|
||
<td id="76BA393F9B5D72C5717EFE47FCF5FE09" box="[804,870,473,495]" gridcol="2" gridrow="5" pageId="9" pageNumber="222">0.016</td>
|
||
<td id="76BA393F9B5D72C571A3FE47FBA8FE09" box="[1017,1083,473,495]" gridcol="3" gridrow="5" pageId="9" pageNumber="222">-0.121</td>
|
||
<td id="76BA393F9B5D72C57697FE47FA9CFE09" box="[1229,1295,473,495]" gridcol="4" gridrow="5" pageId="9" pageNumber="222">-0.062</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FD9CFA9CFDFE" box="[159,1295,514,536]" gridrow="6" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FD9CFEE7FDFE" box="[159,372,514,536]" gridcol="0" gridrow="6" pageId="9" pageNumber="222">IoD</th>
|
||
<td id="76BA393F9B5D72C5700AFD9CFD02FDFE" box="[592,657,514,536]" gridcol="1" gridrow="6" pageId="9" pageNumber="222">-0.194</td>
|
||
<td id="76BA393F9B5D72C5717EFD9CFCF5FDFE" box="[804,870,514,536]" gridcol="2" gridrow="6" pageId="9" pageNumber="222">0.207</td>
|
||
<td id="76BA393F9B5D72C571A3FD9CFBA8FDFE" box="[1017,1083,514,536]" gridcol="3" gridrow="6" pageId="9" pageNumber="222">-0.248</td>
|
||
<td id="76BA393F9B5D72C57697FD9CFA9CFDFE" box="[1229,1295,514,536]" gridcol="4" gridrow="6" pageId="9" pageNumber="222">0.219</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FDB2FA9CFDA4" box="[159,1295,556,578]" gridrow="7" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FDB2FEE7FDA4" box="[159,372,556,578]" gridcol="0" gridrow="7" pageId="9" pageNumber="222">ED</th>
|
||
<td id="76BA393F9B5D72C5700AFDB2FD02FDA4" box="[592,657,556,578]" gridcol="1" gridrow="7" pageId="9" pageNumber="222">-0.240</td>
|
||
<td id="76BA393F9B5D72C5717EFDB2FCF5FDA4" box="[804,870,556,578]" gridcol="2" gridrow="7" pageId="9" pageNumber="222">0.035</td>
|
||
<td id="76BA393F9B5D72C571A3FDB2FBA8FDA4" box="[1017,1083,556,578]" gridcol="3" gridrow="7" pageId="9" pageNumber="222">-0.024</td>
|
||
<td id="76BA393F9B5D72C57697FDB2FA9CFDA4" box="[1229,1295,556,578]" gridcol="4" gridrow="7" pageId="9" pageNumber="222">0.032</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FDCBFA9CFD8D" box="[159,1295,597,619]" gridrow="8" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FDCBFEE7FD8D" box="[159,372,597,619]" gridcol="0" gridrow="8" pageId="9" pageNumber="222">EL</th>
|
||
<td id="76BA393F9B5D72C5700AFDCBFD02FD8D" box="[592,657,597,619]" gridcol="1" gridrow="8" pageId="9" pageNumber="222">-0.241</td>
|
||
<td id="76BA393F9B5D72C5717EFDCBFCF5FD8D" box="[804,870,597,619]" gridcol="2" gridrow="8" pageId="9" pageNumber="222">0.023</td>
|
||
<td id="76BA393F9B5D72C571A3FDCBFBA8FD8D" box="[1017,1083,597,619]" gridcol="3" gridrow="8" pageId="9" pageNumber="222">-0.035</td>
|
||
<td id="76BA393F9B5D72C57697FDCBFA9CFD8D" box="[1229,1295,597,619]" gridcol="4" gridrow="8" pageId="9" pageNumber="222">0.021</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FDE0FA9CFD72" box="[159,1295,638,660]" gridrow="9" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FDE0FEE7FD72" box="[159,372,638,660]" gridcol="0" gridrow="9" pageId="9" pageNumber="222">AGL</th>
|
||
<td id="76BA393F9B5D72C5700AFDE0FD02FD72" box="[592,657,638,660]" gridcol="1" gridrow="9" pageId="9" pageNumber="222">-0.166</td>
|
||
<td id="76BA393F9B5D72C5717EFDE0FCF5FD72" box="[804,870,638,660]" gridcol="2" gridrow="9" pageId="9" pageNumber="222">0.163</td>
|
||
<td id="76BA393F9B5D72C571A3FDE0FBA8FD72" box="[1017,1083,638,660]" gridcol="3" gridrow="9" pageId="9" pageNumber="222">0.525</td>
|
||
<td id="76BA393F9B5D72C57697FDE0FA9CFD72" box="[1229,1295,638,660]" gridcol="4" gridrow="9" pageId="9" pageNumber="222">-0.236</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FD36FA9CFD58" box="[159,1295,680,702]" gridrow="10" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FD36FEE7FD58" box="[159,372,680,702]" gridcol="0" gridrow="10" pageId="9" pageNumber="222">InD</th>
|
||
<td id="76BA393F9B5D72C5700AFD36FD02FD58" box="[592,657,680,702]" gridcol="1" gridrow="10" pageId="9" pageNumber="222">-0.208</td>
|
||
<td id="76BA393F9B5D72C5717EFD36FCF5FD58" box="[804,870,680,702]" gridcol="2" gridrow="10" pageId="9" pageNumber="222">0.244</td>
|
||
<td id="76BA393F9B5D72C571A3FD36FBA8FD58" box="[1017,1083,680,702]" gridcol="3" gridrow="10" pageId="9" pageNumber="222">-0.128</td>
|
||
<td id="76BA393F9B5D72C57697FD36FA9CFD58" box="[1229,1295,680,702]" gridcol="4" gridrow="10" pageId="9" pageNumber="222">0.226</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FD4FFA9CFD01" box="[159,1295,721,743]" gridrow="11" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FD4FFEE7FD01" box="[159,372,721,743]" gridcol="0" gridrow="11" pageId="9" pageNumber="222">NRD</th>
|
||
<td id="76BA393F9B5D72C5700AFD4FFD02FD01" box="[592,657,721,743]" gridcol="1" gridrow="11" pageId="9" pageNumber="222">-0.223</td>
|
||
<td id="76BA393F9B5D72C5717EFD4FFCF5FD01" box="[804,870,721,743]" gridcol="2" gridrow="11" pageId="9" pageNumber="222">0.200</td>
|
||
<td id="76BA393F9B5D72C571A3FD4FFBA8FD01" box="[1017,1083,721,743]" gridcol="3" gridrow="11" pageId="9" pageNumber="222">0.029</td>
|
||
<td id="76BA393F9B5D72C57697FD4FFA9CFD01" box="[1229,1295,721,743]" gridcol="4" gridrow="11" pageId="9" pageNumber="222">0.143</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FD64FA9CFCF6" box="[159,1295,762,784]" gridrow="12" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FD64FEE7FCF6" box="[159,372,762,784]" gridcol="0" gridrow="12" pageId="9" pageNumber="222">EJL</th>
|
||
<td id="76BA393F9B5D72C5700AFD64FD02FCF6" box="[592,657,762,784]" gridcol="1" gridrow="12" pageId="9" pageNumber="222">-0.209</td>
|
||
<td id="76BA393F9B5D72C5717EFD64FCF5FCF6" box="[804,870,762,784]" gridcol="2" gridrow="12" pageId="9" pageNumber="222">0.017</td>
|
||
<td id="76BA393F9B5D72C571A3FD64FBA8FCF6" box="[1017,1083,762,784]" gridcol="3" gridrow="12" pageId="9" pageNumber="222">0.089</td>
|
||
<td id="76BA393F9B5D72C57697FD64FA9CFCF6" box="[1229,1295,762,784]" gridcol="4" gridrow="12" pageId="9" pageNumber="222">0.168</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FCBAFA9CFCDC" box="[159,1295,804,826]" gridrow="13" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FCBAFEE7FCDC" box="[159,372,804,826]" gridcol="0" gridrow="13" pageId="9" pageNumber="222">ND</th>
|
||
<td id="76BA393F9B5D72C5700AFCBAFD02FCDC" box="[592,657,804,826]" gridcol="1" gridrow="13" pageId="9" pageNumber="222">-0.226</td>
|
||
<td id="76BA393F9B5D72C5717EFCBAFCF5FCDC" box="[804,870,804,826]" gridcol="2" gridrow="13" pageId="9" pageNumber="222">0.100</td>
|
||
<td id="76BA393F9B5D72C571A3FCBAFBA8FCDC" box="[1017,1083,804,826]" gridcol="3" gridrow="13" pageId="9" pageNumber="222">0.023</td>
|
||
<td id="76BA393F9B5D72C57697FCBAFA9CFCDC" box="[1229,1295,804,826]" gridcol="4" gridrow="13" pageId="9" pageNumber="222">0.259</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FCD3FA9CFC85" box="[159,1295,845,867]" gridrow="14" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FCD3FEE7FC85" box="[159,372,845,867]" gridcol="0" gridrow="14" pageId="9" pageNumber="222">SL</th>
|
||
<td id="76BA393F9B5D72C5700AFCD3FD02FC85" box="[592,657,845,867]" gridcol="1" gridrow="14" pageId="9" pageNumber="222">-0.184</td>
|
||
<td id="76BA393F9B5D72C5717EFCD3FCF5FC85" box="[804,870,845,867]" gridcol="2" gridrow="14" pageId="9" pageNumber="222">0.187</td>
|
||
<td id="76BA393F9B5D72C571A3FCD3FBA8FC85" box="[1017,1083,845,867]" gridcol="3" gridrow="14" pageId="9" pageNumber="222">-0.143</td>
|
||
<td id="76BA393F9B5D72C57697FCD3FA9CFC85" box="[1229,1295,845,867]" gridcol="4" gridrow="14" pageId="9" pageNumber="222">-0.241</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FCE8FA9CFC6A" box="[159,1295,886,908]" gridrow="15" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FCE8FEE7FC6A" box="[159,372,886,908]" gridcol="0" gridrow="15" pageId="9" pageNumber="222">IpL</th>
|
||
<td id="76BA393F9B5D72C5700AFCE8FD02FC6A" box="[592,657,886,908]" gridcol="1" gridrow="15" pageId="9" pageNumber="222">-0.239</td>
|
||
<td id="76BA393F9B5D72C5717EFCE8FCF5FC6A" box="[804,870,886,908]" gridcol="2" gridrow="15" pageId="9" pageNumber="222">0.019</td>
|
||
<td id="76BA393F9B5D72C571A3FCE8FBA8FC6A" box="[1017,1083,886,908]" gridcol="3" gridrow="15" pageId="9" pageNumber="222">-0.004</td>
|
||
<td id="76BA393F9B5D72C57697FCE8FA9CFC6A" box="[1229,1295,886,908]" gridcol="4" gridrow="15" pageId="9" pageNumber="222">0.005</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FC3EFA9CFC50" box="[159,1295,928,950]" gridrow="16" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FC3EFEE7FC50" box="[159,372,928,950]" gridcol="0" gridrow="16" pageId="9" pageNumber="222">IpW</th>
|
||
<td id="76BA393F9B5D72C5700AFC3EFD02FC50" box="[592,657,928,950]" gridcol="1" gridrow="16" pageId="9" pageNumber="222">-0.239</td>
|
||
<td id="76BA393F9B5D72C5717EFC3EFCF5FC50" box="[804,870,928,950]" gridcol="2" gridrow="16" pageId="9" pageNumber="222">0.012</td>
|
||
<td id="76BA393F9B5D72C571A3FC3EFBA8FC50" box="[1017,1083,928,950]" gridcol="3" gridrow="16" pageId="9" pageNumber="222">-0.031</td>
|
||
<td id="76BA393F9B5D72C57697FC3EFA9CFC50" box="[1229,1295,928,950]" gridcol="4" gridrow="16" pageId="9" pageNumber="222">-0.008</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FC57FA9CFC39" box="[159,1295,969,991]" gridrow="17" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FC57FEE7FC39" box="[159,372,969,991]" gridcol="0" gridrow="17" pageId="9" pageNumber="222">MW</th>
|
||
<td id="76BA393F9B5D72C5700AFC57FD02FC39" box="[592,657,969,991]" gridcol="1" gridrow="17" pageId="9" pageNumber="222">-0.217</td>
|
||
<td id="76BA393F9B5D72C5717EFC57FCF5FC39" box="[804,870,969,991]" gridcol="2" gridrow="17" pageId="9" pageNumber="222">-0.058</td>
|
||
<td id="76BA393F9B5D72C571A3FC57FBA8FC39" box="[1017,1083,969,991]" gridcol="3" gridrow="17" pageId="9" pageNumber="222">-0.263</td>
|
||
<td id="76BA393F9B5D72C57697FC57FA9CFC39" box="[1229,1295,969,991]" gridcol="4" gridrow="17" pageId="9" pageNumber="222">-0.140</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FC6CFA9CFBEE" box="[159,1295,1010,1032]" gridrow="18" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FC6CFEE7FBEE" box="[159,372,1010,1032]" gridcol="0" gridrow="18" pageId="9" pageNumber="222">HuL</th>
|
||
<td id="76BA393F9B5D72C5700AFC6CFD02FBEE" box="[592,657,1010,1032]" gridcol="1" gridrow="18" pageId="9" pageNumber="222">-0.175</td>
|
||
<td id="76BA393F9B5D72C5717EFC6CFCF5FBEE" box="[804,870,1010,1032]" gridcol="2" gridrow="18" pageId="9" pageNumber="222">-0.082</td>
|
||
<td id="76BA393F9B5D72C571A3FC6CFBA8FBEE" box="[1017,1083,1010,1032]" gridcol="3" gridrow="18" pageId="9" pageNumber="222">-0.417</td>
|
||
<td id="76BA393F9B5D72C57697FC6CFA9CFBEE" box="[1229,1295,1010,1032]" gridcol="4" gridrow="18" pageId="9" pageNumber="222">-0.424</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FB82FA9CFBD4" box="[159,1295,1052,1074]" gridrow="19" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FB82FEE7FBD4" box="[159,372,1052,1074]" gridcol="0" gridrow="19" pageId="9" pageNumber="222">UIL</th>
|
||
<td id="76BA393F9B5D72C5700AFB82FD02FBD4" box="[592,657,1052,1074]" gridcol="1" gridrow="19" pageId="9" pageNumber="222">-0.186</td>
|
||
<td id="76BA393F9B5D72C5717EFB82FCF5FBD4" box="[804,870,1052,1074]" gridcol="2" gridrow="19" pageId="9" pageNumber="222">0.028</td>
|
||
<td id="76BA393F9B5D72C571A3FB82FBA8FBD4" box="[1017,1083,1052,1074]" gridcol="3" gridrow="19" pageId="9" pageNumber="222">0.379</td>
|
||
<td id="76BA393F9B5D72C57697FB82FA9CFBD4" box="[1229,1295,1052,1074]" gridcol="4" gridrow="19" pageId="9" pageNumber="222">0.017</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FBDBFA9CFBBD" box="[159,1295,1093,1115]" gridrow="20" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FBDBFEE7FBBD" box="[159,372,1093,1115]" gridcol="0" gridrow="20" pageId="9" pageNumber="222">FL</th>
|
||
<td id="76BA393F9B5D72C5700AFBDBFD02FBBD" box="[592,657,1093,1115]" gridcol="1" gridrow="20" pageId="9" pageNumber="222">-0.182</td>
|
||
<td id="76BA393F9B5D72C5717EFBDBFCF5FBBD" box="[804,870,1093,1115]" gridcol="2" gridrow="20" pageId="9" pageNumber="222">-0.390</td>
|
||
<td id="76BA393F9B5D72C571A3FBDBFBA8FBBD" box="[1017,1083,1093,1115]" gridcol="3" gridrow="20" pageId="9" pageNumber="222">-0.134</td>
|
||
<td id="76BA393F9B5D72C57697FBDBFA9CFBBD" box="[1229,1295,1093,1115]" gridcol="4" gridrow="20" pageId="9" pageNumber="222">-0.277</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FBF0FA9CFB62" box="[159,1295,1134,1156]" gridrow="21" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FBF0FEE7FB62" box="[159,372,1134,1156]" gridcol="0" gridrow="21" pageId="9" pageNumber="222">TiL</th>
|
||
<td id="76BA393F9B5D72C5700AFBF0FD02FB62" box="[592,657,1134,1156]" gridcol="1" gridrow="21" pageId="9" pageNumber="222">-0.197</td>
|
||
<td id="76BA393F9B5D72C5717EFBF0FCF5FB62" box="[804,870,1134,1156]" gridcol="2" gridrow="21" pageId="9" pageNumber="222">-0.328</td>
|
||
<td id="76BA393F9B5D72C571A3FBF0FBA8FB62" box="[1017,1083,1134,1156]" gridcol="3" gridrow="21" pageId="9" pageNumber="222">0.244</td>
|
||
<td id="76BA393F9B5D72C57697FBF0FA9CFB62" box="[1229,1295,1134,1156]" gridcol="4" gridrow="21" pageId="9" pageNumber="222">0.265</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FB06FA9CFB48" box="[159,1295,1176,1198]" gridrow="22" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FB06FEE7FB48" box="[159,372,1176,1198]" gridcol="0" gridrow="22" pageId="9" pageNumber="222">FoL</th>
|
||
<td id="76BA393F9B5D72C5700AFB06FD02FB48" box="[592,657,1176,1198]" gridcol="1" gridrow="22" pageId="9" pageNumber="222">-0.180</td>
|
||
<td id="76BA393F9B5D72C5717EFB06FCF5FB48" box="[804,870,1176,1198]" gridcol="2" gridrow="22" pageId="9" pageNumber="222">-0.498</td>
|
||
<td id="76BA393F9B5D72C571A3FB06FBA8FB48" box="[1017,1083,1176,1198]" gridcol="3" gridrow="22" pageId="9" pageNumber="222">0.121</td>
|
||
<td id="76BA393F9B5D72C57697FB06FA9CFB48" box="[1229,1295,1176,1198]" gridcol="4" gridrow="22" pageId="9" pageNumber="222">0.075</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FB5FFA9CFB31" box="[159,1295,1217,1239]" gridrow="23" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FB5FFEE7FB31" box="[159,372,1217,1239]" gridcol="0" gridrow="23" pageId="9" pageNumber="222">4tL</th>
|
||
<td id="76BA393F9B5D72C5700AFB5FFD02FB31" box="[592,657,1217,1239]" gridcol="1" gridrow="23" pageId="9" pageNumber="222">-0.193</td>
|
||
<td id="76BA393F9B5D72C5717EFB5FFCF5FB31" box="[804,870,1217,1239]" gridcol="2" gridrow="23" pageId="9" pageNumber="222">-0.432</td>
|
||
<td id="76BA393F9B5D72C571A3FB5FFBA8FB31" box="[1017,1083,1217,1239]" gridcol="3" gridrow="23" pageId="9" pageNumber="222">0.070</td>
|
||
<td id="76BA393F9B5D72C57697FB5FFA9CFB31" box="[1229,1295,1217,1239]" gridcol="4" gridrow="23" pageId="9" pageNumber="222">-0.023</td>
|
||
</tr>
|
||
<tr id="356B50439B5D72C572C5FB74FA9CFAE6" box="[159,1295,1258,1280]" gridrow="24" pageId="9" pageNumber="222">
|
||
<th id="76BA393F9B5D72C572C5FB74FEE7FAE6" box="[159,372,1258,1280]" gridcol="0" gridrow="24" pageId="9" pageNumber="222">Cumulative Variance</th>
|
||
<td id="76BA393F9B5D72C5700AFB74FD02FAE6" box="[592,657,1258,1280]" gridcol="1" gridrow="24" pageId="9" pageNumber="222">0.732</td>
|
||
<td id="76BA393F9B5D72C5717EFB74FCF5FAE6" box="[804,870,1258,1280]" gridcol="2" gridrow="24" pageId="9" pageNumber="222">0.786</td>
|
||
<td id="76BA393F9B5D72C571A3FB74FBA8FAE6" box="[1017,1083,1258,1280]" gridcol="3" gridrow="24" pageId="9" pageNumber="222">0.822</td>
|
||
<td id="76BA393F9B5D72C57697FB74FA9CFAE6" box="[1229,1295,1258,1280]" gridcol="4" gridrow="24" pageId="9" pageNumber="222">0.850</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BE452019B5D8D33729DFADFFEFEF9D4" blockId="9.[151,1436,1345,2018]" pageId="9" pageNumber="222">
|
||
<emphasis id="B92F8E139B5D8D33729DFADFFE43FABC" bold="true" box="[199,464,1345,1370]" pageId="9" pageNumber="222">Color in preservative.</emphasis>
|
||
Preserved specimens dark brown or black middorsally (dark area broader in males), flanks with light brown, but in some few specimens, flanks are covered with a light orange or green (although this last condition is very rare). However, this green coloration in life is lost in preservative. Fore and hind legs may have light crossbars. No dark shoulder or occipital spot. Belly white or brownish white, darker toward sides, speckled (especially in females) with brown spots. Gular region in females white or white speckled with brown spots or vermiculations. Dewlap pale, often yellowish in anterior half, sometimes pale pink posteriorly, scale rows usually pigmented.
|
||
</paragraph>
|
||
<paragraph id="8BE452019B5D8D33729DF9A3FBFDF99C" blockId="9.[151,1436,1345,2018]" pageId="9" pageNumber="222">
|
||
<materialsCitation id="3B33585C9B5D8D33729DF9A3FBFDF99C" ID-GBIF-Occurrence="1585968392" country="Colombia" county="northwestern Andes" elevation="2220" location="Geographic" municipality="Urrao" pageId="9" pageNumber="222" specimenCount="1" stateProvince="Antioquia department">
|
||
<emphasis id="B92F8E139B5D8D33729DF9A3FE7CF9B0" bold="true" box="[199,495,1597,1622]" pageId="9" pageNumber="222">
|
||
<location id="8E8404DA9B5D8D33729DF9A3FEC0F9B0" LSID="urn:lsid:plazi:treatment:03F2E3179B578D3072CDFD30FCDFF8A1:8E8404DA9B5D8D33729DF9A3FEC0F9B0" box="[199,339,1597,1622]" country="Colombia" county="northwestern Andes" municipality="Urrao" name="Geographic" pageId="9" pageNumber="222" stateProvince="Antioquia department">Geographic</location>
|
||
distribution.
|
||
</emphasis>
|
||
<taxonomicName id="4C5B29829B5D8D3373ACF9A3FD1FF9B3" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[502,652,1597,1621]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B5D8D3373ACF9A3FD1FF9B3" box="[502,652,1597,1621]" italics="true" pageId="9" pageNumber="222">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B5D8D3370CEF9A3FD7DF9B0" box="[660,750,1597,1622]" pageId="9" pageNumber="222" rank="species">
|
||
<emphasis id="B92F8E139B5D8D3370CEF9A3FD7DF9B0" bold="true" box="[660,750,1597,1622]" pageId="9" pageNumber="222">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is known from
|
||
<collectingMunicipality id="6B80C87B9B5D8D3371FAF9A0FC72F9B3" box="[928,993,1598,1621]" pageId="9" pageNumber="222">Urrao</collectingMunicipality>
|
||
municipality,
|
||
<collectingRegion id="499F9CE39B5D8D3376D9F9A3FAE8F9B0" box="[1155,1403,1597,1622]" country="Colombia" name="Antioquia" pageId="9" pageNumber="222">Antioquia department</collectingRegion>
|
||
, in the
|
||
<collectingCounty id="62852A8D9B5D8D33729AF9FFFE33F99C" box="[192,416,1633,1658]" pageId="9" pageNumber="222">northwestern Andes</collectingCounty>
|
||
of
|
||
<collectingCountry id="F34C12919B5D8D33739CF9FFFDA7F99C" box="[454,564,1633,1658]" name="Colombia" pageId="9" pageNumber="222">Colombia</collectingCountry>
|
||
between
|
||
<date id="FFE574C19B5D8D3370FAF9FCFCD5F99C" box="[672,838,1633,1658]" pageId="9" pageNumber="222" value="1700" valueMax="2220">
|
||
1700 and
|
||
<elevation id="0076B5329B5D8D337154F9FCFCD5F99C" box="[782,838,1634,1658]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="9" pageNumber="222" unit="m" value="2220.0">
|
||
<quantity id="4CA3FFE49B5D8D337154F9FCFCD5F99C" box="[782,838,1634,1658]" metricMagnitude="3" metricUnit="m" metricValue="2.2199999999999998" pageId="9" pageNumber="222" unit="m" value="2220.0">2220</quantity>
|
||
</elevation>
|
||
</date>
|
||
m of elevation (
|
||
<figureCitation id="13604E849B5D8D3371A4F9FFFBF3F99C" box="[1022,1120,1633,1658]" captionStart="FIGURE 5" captionStartId="10.[151,250,1386,1408]" captionTargetBox="[161,1429,351,1355]" captionTargetId="figure@10.[151,1436,337,1365]" captionTargetPageId="10" captionText="FIGURE 5. Distribution map of the species in the Anolis fuscoauratus group included in this study. S and M indicate sequenced specimens and measured specimens respectively." httpUri="https://zenodo.org/record/840912/files/figure.png" pageId="9" pageNumber="222">Figure 5</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE452019B5D8D33729DF91BFBA8F8B4" blockId="9.[151,1436,1345,2018]" pageId="9" pageNumber="222">
|
||
<materialsCitation id="3B33585C9B5D8D33729DF91BFBA8F8B4" ID-GBIF-Occurrence="1585968416" collectingDate="2014-10-31" collectionCode="MHUA-R" county="Individuals" location="Specimens" municipality="Males" pageId="9" pageNumber="222" specimenCode="MHUA-R 12731-32-33" specimenCount="2" typeStatus="holotype">
|
||
<emphasis id="B92F8E139B5D8D33729DF91BFE04F978" bold="true" box="[199,407,1669,1694]" pageId="9" pageNumber="222">Ecological Notes.</emphasis>
|
||
<taxonomicName id="4C5B29829B5D8D3373F9F91BFDACF97B" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[419,575,1669,1693]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B5D8D3373F9F91BFDACF97B" box="[419,575,1669,1693]" italics="true" pageId="9" pageNumber="222">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B5D8D337010F91BFD3AF978" box="[586,681,1669,1694]" pageId="9" pageNumber="222" rank="species">
|
||
<emphasis id="B92F8E139B5D8D337010F91BFD3AF978" bold="true" box="[586,681,1669,1694]" pageId="9" pageNumber="222">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is very common in the
|
||
<typeStatus id="54E0ECA39B5D8D337194F918FC6DF978" box="[974,1022,1670,1694]" pageId="9" pageNumber="222">type</typeStatus>
|
||
locality.
|
||
<location id="8E8404DA9B5D8D337635F91BFB7BF978" LSID="urn:lsid:plazi:treatment:03F2E3179B578D3072CDFD30FCDFF8A1:8E8404DA9B5D8D337635F91BFB7BF978" box="[1135,1256,1669,1694]" county="Individuals" municipality="Males" name="Specimens" pageId="9" pageNumber="222">Specimens</location>
|
||
were collected mainly in open areas with shrubs and fences near human buildings, along sidewalks and roadways, and along the banks of the
|
||
<location id="8E8404DA9B5D8D33737CF953FE4DF900" LSID="urn:lsid:plazi:treatment:03F2E3179B578D3072CDFD30FCDFF8A1:8E8404DA9B5D8D33737CF953FE4DF900" box="[294,478,1741,1766]" county="Individuals" municipality="Males" name="Penderisco river" pageId="9" pageNumber="222">Penderisco river</location>
|
||
.
|
||
<location id="8E8404DA9B5D8D3373B2F953FD87F900" LSID="urn:lsid:plazi:treatment:03F2E3179B578D3072CDFD30FCDFF8A1:8E8404DA9B5D8D3373B2F953FD87F900" box="[488,532,1741,1766]" county="Individuals" municipality="Males" name="The" pageId="9" pageNumber="222">The</location>
|
||
habitat is similar where other congeners are commonly found (e.g.,
|
||
<taxonomicName id="4C5B29829B5D8D337757F950FA0FF903" authorityName="Barbour" authorityYear="1932" box="[1293,1436,1742,1765]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B5D8D337757F950FA0FF903" box="[1293,1436,1742,1765]" italics="true" pageId="9" pageNumber="222">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B29829B5D8D33729CF96CFEC4F8EF" authorityName="Werner" authorityYear="1916" box="[198,343,1777,1801]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="B92F8E139B5D8D33729CF96CFEC4F8EF" box="[198,343,1777,1801]" italics="true" pageId="9" pageNumber="222">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<collectingMunicipality id="6B80C87B9B5D8D337334F96FFE21F8EC" box="[366,434,1777,1802]" pageId="9" pageNumber="222">Males</collectingMunicipality>
|
||
are commonly seen on bushes and stakes (up to 3 meters high) performing display, while females are usually elusive and occupying low perches.
|
||
<collectingCounty id="62852A8D9B5D8D33717EF88BFC31F8C8" box="[804,930,1813,1838]" pageId="9" pageNumber="222">Individuals</collectingCounty>
|
||
of the
|
||
<typeStatus id="54E0ECA39B5D8D3371A3F888FBBAF8C8" box="[1017,1065,1814,1838]" pageId="9" pageNumber="222">type</typeStatus>
|
||
series (
|
||
<specimenCode id="DBFDFA7A9B5D8D3376DFF888FA07F8C8" box="[1157,1428,1814,1838]" pageId="9" pageNumber="222">
|
||
<collectionCode id="ED4ACAC49B5D8D3376DFF888FB6AF8C8" box="[1157,1273,1814,1838]" pageId="9" pageNumber="222">MHUA-R</collectionCode>
|
||
12731-32-33
|
||
</specimenCode>
|
||
) were found between 10:00 and 11:00 in a sunny day (
|
||
<collectingDate id="EFA18D299B5D8D3370B4F8A7FC26F8B4" box="[750,949,1849,1874]" pageId="9" pageNumber="222" value="2014-10-31">
|
||
<date id="FFE574C19B5D8D3370B4F8A7FC26F8B4" box="[750,949,1849,1874]" pageId="9" pageNumber="222" value="2014-10-31">October 31, 2014</date>
|
||
</collectingDate>
|
||
) (
|
||
<figureCitation id="13604E849B5D8D337191F8A7FBBEF8B4" box="[971,1069,1849,1874]" captionStart="FIGURE 6" captionStartId="11.[151,250,1315,1337]" captionTargetBox="[204,1382,409,1293]" captionTargetId="figure@11.[204,1382,409,1293]" captionTargetPageId="11" captionText="FIGURE 6. Habitat of Anolis urraoi sp. nov. on the banks of the Penderisco river, Urrao, Antioquia, Colombia." httpUri="https://zenodo.org/record/840914/files/figure.png" pageId="9" pageNumber="222">Figure 6</figureCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE452019B5D8D30729DF8C3FB9FFEFB" blockId="9.[151,1436,1345,2018]" lastBlockId="10.[151,1437,151,285]" lastPageId="10" lastPageNumber="223" pageId="9" pageNumber="222">
|
||
<emphasis id="B92F8E139B5D8D33729DF8C3FE2FF890" bold="true" box="[199,444,1885,1910]" pageId="9" pageNumber="222">Phylogenetic results.</emphasis>
|
||
<taxonomicName id="4C5B29829B5D8D337399F8C3FD11F893" authorityName="Barbour" authorityYear="1932" box="[451,642,1885,1909]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B5D8D337399F8C3FD11F893" box="[451,642,1885,1909]" italics="true" pageId="9" pageNumber="222">Anolis mariarum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5B29829B5D8D3370CAF8C0FD6CF893" authorityName="Boulenger" authorityYear="1908" box="[656,767,1886,1909]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B5D8D3370CAF8C0FD6CF893" box="[656,767,1886,1909]" italics="true" pageId="9" pageNumber="222">A. antonii</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5B29829B5D8D337166F8C0FC5DF893" authorityName="Werner" authorityYear="1916" box="[828,974,1885,1909]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="B92F8E139B5D8D337166F8C0FC5DF893" box="[828,974,1885,1909]" italics="true" pageId="9" pageNumber="222">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
form a monophyletic clade with a strong support (99%;
|
||
<figureCitation id="13604E849B5D8D337366F81FFE09F87C" box="[316,410,1921,1946]" captionStart="FIGURE 7" captionStartId="12.[151,250,1859,1881]" captionTargetBox="[246,1339,207,1830]" captionTargetId="figure@12.[246,1340,203,1838]" captionTargetPageId="12" captionText="FIGURE 7. Maximum likelihood tree (- lnL = - 13314.977) obtained with IQ-TREE depicting the phylogenetic relationships in the Andean species of the Anolis fuscoauratus species group. Inference based on 1465 characters of the ND 2 gene and intervening regions. Nodal support was estimated using the ultrafast bootstrap method using 10000 pseudoreplicates (Minh et al., 2013)." httpUri="https://zenodo.org/record/840916/files/figure.png" pageId="9" pageNumber="222">Figure 7</figureCitation>
|
||
) related to
|
||
<taxonomicName id="4C5B29829B5D8D337042F81CFDFCF87F" authorityName="Dunn" authorityYear="1930" box="[536,623,1921,1945]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="altae">
|
||
<emphasis id="B92F8E139B5D8D337042F81CFDFCF87F" box="[536,623,1921,1945]" italics="true" pageId="9" pageNumber="222">A. altae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F34C12919B5D8D3370E8F81CFCBCF87C" box="[690,815,1921,1946]" name="Costa Rica" pageId="9" pageNumber="222">Costa Rica</collectingCountry>
|
||
. By contrast,
|
||
<taxonomicName id="4C5B29829B5D8D33719CF81CFBBFF87F" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[966,1068,1922,1945]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B5D8D33719CF81CFBBFF87F" box="[966,1068,1922,1945]" italics="true" pageId="9" pageNumber="222">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B5D8D337669F81FFB1FF87C" box="[1075,1164,1921,1946]" pageId="9" pageNumber="222" rank="species">
|
||
<emphasis id="B92F8E139B5D8D337669F81FFB1FF87C" bold="true" box="[1075,1164,1921,1946]" pageId="9" pageNumber="222">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is more related to forms from the Pacific lowlands as
|
||
<taxonomicName id="4C5B29829B5D8D3373BCF838FDCDF85B" authorityName="Ayala & Williams" authorityYear="1988" box="[486,606,1957,1981]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="medemi">
|
||
<emphasis id="B92F8E139B5D8D3373BCF838FDCDF85B" box="[486,606,1957,1981]" italics="true" pageId="9" pageNumber="222">A. medemi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13604E849B5D8D337034F83BFD42F858" box="[622,721,1957,1982]" captionStart="FIGURE 7" captionStartId="12.[151,250,1859,1881]" captionTargetBox="[246,1339,207,1830]" captionTargetId="figure@12.[246,1340,203,1838]" captionTargetPageId="12" captionText="FIGURE 7. Maximum likelihood tree (- lnL = - 13314.977) obtained with IQ-TREE depicting the phylogenetic relationships in the Andean species of the Anolis fuscoauratus species group. Inference based on 1465 characters of the ND 2 gene and intervening regions. Nodal support was estimated using the ultrafast bootstrap method using 10000 pseudoreplicates (Minh et al., 2013)." httpUri="https://zenodo.org/record/840916/files/figure.png" pageId="9" pageNumber="222">Figure 7</figureCitation>
|
||
) and probably
|
||
<taxonomicName id="4C5B29829B5D8D3371DAF838FBAAF85B" authorityName="Boulenger" authorityYear="1898" box="[896,1081,1957,1981]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="maculiventris">
|
||
<emphasis id="B92F8E139B5D8D3371DAF838FBAAF85B" box="[896,1081,1957,1981]" italics="true" pageId="9" pageNumber="222">A. maculiventris</emphasis>
|
||
</taxonomicName>
|
||
(not included in this analysis). Also, this other clade is related with species from Central America (i.e.,
|
||
<taxonomicName id="4C5B29829B5D8D337190F857FBECF807" authorityName="Dunn" authorityYear="1940" box="[970,1151,1993,2017]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="species" species="kemptoni">
|
||
<emphasis id="B92F8E139B5D8D337190F857FBECF807" box="[970,1151,1993,2017]" italics="true" pageId="9" pageNumber="222">Anolis kemptoni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C5B29829B5D8D3376EDF857FB6CF807" authorityName="Daudin" authorityYear="1802" box="[1207,1279,1993,2017]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="222" phylum="Chordata" rank="genus">
|
||
<emphasis id="B92F8E139B5D8D3376EDF857FB6CF807" box="[1207,1279,1993,2017]" italics="true" pageId="9" pageNumber="222">Anolis</emphasis>
|
||
</taxonomicName>
|
||
sp. JGP2014, see
|
||
<bibRefCitation id="EFCA2FF09B5E8D307299FF09FE0CFF56" author="Phillips" box="[195,415,151,176]" pageId="10" pageNumber="223" refString="Phillips, J. G., Deitloff, J., Guyer, C., Huetteman, S. & Nicholson, K. E. (2015). Biogeography and evolution of a widespread Central American lizard species complex: Norops humilis, (Squamata: Dactyloidae). BMC Evolutionary Biology, 15 (1), 143. https: // doi. org / 10.1186 / s 12862 - 015 - 0391 - 4" type="journal article" year="2015">
|
||
Phillips
|
||
<emphasis id="B92F8E139B5E8D30737BFF07FEC9FF56" box="[289,346,152,176]" italics="true" pageId="10" pageNumber="223">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
). Although we did not have an extensive sampling within the large
|
||
<taxonomicName id="4C5B29829B5E8D3076FBFF06FAC0FF56" authorityName="D'Orbigny" authorityYear="1837" box="[1185,1363,152,176]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B5E8D3076FBFF06FAC0FF56" box="[1185,1363,152,176]" italics="true" pageId="10" pageNumber="223">A. fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
group (see
|
||
<bibRefCitation id="EFCA2FF09B5E8D307297FF23FEEEFF32" author="Poe" box="[205,381,188,212]" pageId="10" pageNumber="223" refString="Poe, S., Nieto-Montes de Oca, A., Torres-Carvajal, O., de Queiroz, K., Velasco, J. A., Truett, B., Gray, L. N., Ryan, M. J., Kohler, G., Ayala-Varela, F. & Latella, I. (2017) A Phylogenetic, Biogeographic, and Taxonomic study of all Extant Species of Anolis (Squamata; Iguanidae). Systematic Biology. [published online]" type="book" year="2017">
|
||
Poe
|
||
<emphasis id="B92F8E139B5E8D30735BFF23FEAFFF32" box="[257,316,188,212]" italics="true" pageId="10" pageNumber="223">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
for a complete phylogeny of all known anole species), the apparent geographic disjunction between the sister species
|
||
<taxonomicName id="4C5B29829B5E8D307398FF7EFDBAFF11" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[450,553,224,247]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B5E8D307398FF7EFDBAFF11" box="[450,553,224,247]" italics="true" pageId="10" pageNumber="223">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B5E8D307068FF41FD1FFF1E" box="[562,652,223,248]" pageId="10" pageNumber="223" rank="species">
|
||
<emphasis id="B92F8E139B5E8D307068FF41FD1FFF1E" bold="true" box="[562,652,223,248]" pageId="10" pageNumber="223">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C5B29829B5E8D30709EFF7EFCA8FF1E" authorityName="Ayala & Williams" authorityYear="1988" box="[708,827,224,248]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="medemi">
|
||
<emphasis id="B92F8E139B5E8D30709EFF7EFCA8FF1E" box="[708,827,224,248]" italics="true" pageId="10" pageNumber="223">A. medemi</emphasis>
|
||
</taxonomicName>
|
||
is most likely the result of lack of samples of species distributed in the Pacific lowlands of
|
||
<collectingCountry id="F34C12919B5E8D307061FE9AFD3FFEFB" box="[571,684,260,285]" name="Colombia" pageId="10" pageNumber="223">Colombia</collectingCountry>
|
||
,
|
||
<collectingCountry id="F34C12919B5E8D3070EDFE9AFC87FEFB" box="[695,788,260,285]" name="Ecuador" pageId="10" pageNumber="223">Ecuador</collectingCountry>
|
||
and Central America.
|
||
</paragraph>
|
||
<caption id="DF2402899B5E8D3072CDFAF4FD6BFA79" httpUri="https://zenodo.org/record/840912/files/figure.png" pageId="10" pageNumber="223" targetBox="[161,1429,351,1355]" targetPageId="10">
|
||
<paragraph id="8BE452019B5E8D3072CDFAF4FD6BFA79" blockId="10.[151,1436,1386,1439]" pageId="10" pageNumber="223">
|
||
<emphasis id="B92F8E139B5E8D3072CDFAF4FE84FA66" bold="true" box="[151,279,1386,1408]" pageId="10" pageNumber="223">FIGURE 5.</emphasis>
|
||
Distribution map of the species in the
|
||
<taxonomicName id="4C5B29829B5E8D307092FAF5FC04FA66" authorityName="D'Orbigny" authorityYear="1837" box="[712,919,1387,1408]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="fuscoauratus">
|
||
<emphasis id="B92F8E139B5E8D307092FAF5FC04FA66" box="[712,919,1387,1408]" italics="true" pageId="10" pageNumber="223">Anolis fuscoauratus</emphasis>
|
||
</taxonomicName>
|
||
group included in this study. S and M indicate sequenced specimens and measured specimens respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE452019B5E8D30729DFA58FC95F919" blockId="10.[151,1436,1478,1863]" pageId="10" pageNumber="223">
|
||
<emphasis id="B92F8E139B5E8D30729DFA58FE70FA39" bold="true" box="[199,483,1478,1503]" pageId="10" pageNumber="223">Morphometric analysis.</emphasis>
|
||
The PCA based on SVL and residual traits shown that Andean fuscoauratoid species tend to be grouped by body size (SVL) and shape traits (
|
||
<figureCitation id="13604E849B5E8D307153FA74FCF9F9E5" box="[777,874,1514,1539]" captionStart="FIGURE 8" captionStartId="13.[151,250,1921,1943]" captionTargetBox="[166,1426,769,1897]" captionTargetId="figure@13.[162,1426,769,1901]" captionTargetPageId="13" captionText="FIGURE 8. PCA plot of morphometric measurements from specimens of Anolis urraoi sp. nov. (red squares), A. mariarum (magenta circles), A. antonii (blue triangles), A. tolimensis (green diamonds) and some individuals tentatively assigned to A. mariarum (from Western Cordillera at Frontino National Park; black circles). PCA was performed with SVL (snout-vent length) and 22 morphological traits corrected for size (main text for details)." httpUri="https://zenodo.org/record/840918/files/figure.png" pageId="10" pageNumber="223">Figure 8</figureCitation>
|
||
). The loadings of each trait are shown in the
|
||
<tableCitation id="C6D967BA9B5E8D307704FA74FF34F9C1" captionStart="TABLE 3" captionStartId="9.[151,239,152,175]" captionText="TABLE 3. Loadings of the SVL and 22 traits corrected for body size (see main text for details) in the principal component analysis (PCA) for adult specimens of the four Andean species of the Anolis fuscoauratus species group: A. antonii, A. mariarum, A. tolimensis, and A. urraoi." pageId="10" pageNumber="223">Table 3</tableCitation>
|
||
. In the first PC axis, the SVL and ED exhibited the highest loads. In the second PC axis, the TL exhibited the highest load (
|
||
<tableCitation id="C6D967BA9B5E8D307377F9ACFE10F9AD" box="[301,387,1586,1611]" captionStart="TABLE 3" captionStartId="9.[151,239,152,175]" captionText="TABLE 3. Loadings of the SVL and 22 traits corrected for body size (see main text for details) in the principal component analysis (PCA) for adult specimens of the four Andean species of the Anolis fuscoauratus species group: A. antonii, A. mariarum, A. tolimensis, and A. urraoi." pageId="10" pageNumber="223">Table 3</tableCitation>
|
||
). Although the overlap between all forms is very high,
|
||
<taxonomicName id="4C5B29829B5E8D3071B5F9ACFB16F9AC" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[1007,1157,1586,1610]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B5E8D3071B5F9ACFB16F9AC" box="[1007,1157,1586,1610]" italics="true" pageId="10" pageNumber="223">Anolis urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B5E8D3076D6F9ACFB76F9AD" box="[1164,1253,1586,1611]" pageId="10" pageNumber="223" rank="species">
|
||
<emphasis id="B92F8E139B5E8D3076D6F9ACFB76F9AD" bold="true" box="[1164,1253,1586,1611]" pageId="10" pageNumber="223">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
tend to separate in the morphospace from the rest of species in the second PC axis (hindlimb dimensions). The discriminant analysis revealed that specimens from
|
||
<taxonomicName id="4C5B29829B5E8D307012F9E5FD3CF974" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[584,687,1659,1682]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B5E8D307012F9E5FD3CF974" box="[584,687,1659,1682]" italics="true" pageId="10" pageNumber="223">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B5E8D3070ECF9E4FC9CF975" box="[694,783,1658,1683]" pageId="10" pageNumber="223" rank="species">
|
||
<emphasis id="B92F8E139B5E8D3070ECF9E4FC9CF975" bold="true" box="[694,783,1658,1683]" pageId="10" pageNumber="223">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C5B29829B5E8D307147F9E5FC3FF974" authorityName="Barbour" authorityYear="1932" box="[797,940,1659,1682]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B5E8D307147F9E5FC3FF974" box="[797,940,1659,1682]" italics="true" pageId="10" pageNumber="223">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C5B29829B5E8D3071E0F9E5FBBAF974" authorityName="Boulenger" authorityYear="1908" box="[954,1065,1659,1682]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B5E8D3071E0F9E5FBBAF974" box="[954,1065,1659,1682]" italics="true" pageId="10" pageNumber="223">A. antonii</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5B29829B5E8D30763CF9E5FB64F974" authorityName="Werner" authorityYear="1916" box="[1126,1271,1658,1682]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="tolimensis">
|
||
<emphasis id="B92F8E139B5E8D30763CF9E5FB64F974" box="[1126,1271,1658,1682]" italics="true" pageId="10" pageNumber="223">A. tolimensis</emphasis>
|
||
</taxonomicName>
|
||
were assigned correctly to each species with a high probability (p-value> 0.5). Some specimens assigned tentatively to
|
||
<taxonomicName id="4C5B29829B5E8D3077DEF901FE94F93C" authorityName="Barbour" authorityYear="1932" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B5E8D3077DEF901FE94F93C" italics="true" pageId="10" pageNumber="223">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
from Frontino National Park (at Western Cordillera from the Andes) seems to overlap strongly with
|
||
<taxonomicName id="4C5B29829B5E8D3077DEF95DFF4CF918" authority="Grisales-Martínez & Velasco & Bolívar & Williams & Daza, 2017" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="urraoi" status="sp. nov.">
|
||
<emphasis id="B92F8E139B5E8D3077DEF95DFF4CF918" italics="true" pageId="10" pageNumber="223">A. urraoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21C33689B5E8D3072BFF978FEACF919" box="[229,319,1766,1791]" pageId="10" pageNumber="223" rank="species">
|
||
<emphasis id="B92F8E139B5E8D3072BFF978FEACF919" bold="true" box="[229,319,1766,1791]" pageId="10" pageNumber="223">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C5B29829B5E8D307317F979FE28F918" authorityName="Boulenger" authorityYear="1908" box="[333,443,1767,1790]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="antonii">
|
||
<emphasis id="B92F8E139B5E8D307317F979FE28F918" box="[333,443,1767,1790]" italics="true" pageId="10" pageNumber="223">A. antonii</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C5B29829B5E8D3073A3F979FD1BF918" authorityName="Barbour" authorityYear="1932" box="[505,648,1767,1790]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="mariarum">
|
||
<emphasis id="B92F8E139B5E8D3073A3F979FD1BF918" box="[505,648,1767,1790]" italics="true" pageId="10" pageNumber="223">A. mariarum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13604E849B5E8D3070CCF978FD64F919" box="[662,759,1766,1791]" captionStart="FIGURE 8" captionStartId="13.[151,250,1921,1943]" captionTargetBox="[166,1426,769,1897]" captionTargetId="figure@13.[162,1426,769,1901]" captionTargetPageId="13" captionText="FIGURE 8. PCA plot of morphometric measurements from specimens of Anolis urraoi sp. nov. (red squares), A. mariarum (magenta circles), A. antonii (blue triangles), A. tolimensis (green diamonds) and some individuals tentatively assigned to A. mariarum (from Western Cordillera at Frontino National Park; black circles). PCA was performed with SVL (snout-vent length) and 22 morphological traits corrected for size (main text for details)." httpUri="https://zenodo.org/record/840918/files/figure.png" pageId="10" pageNumber="223">Figure 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C341018A9B5E8D30729DF894FCDFF8A1" pageId="10" pageNumber="223" type="etymology">
|
||
<paragraph id="8BE452019B5E8D30729DF894FCDFF8A1" blockId="10.[151,1436,1478,1863]" pageId="10" pageNumber="223">
|
||
<emphasis id="B92F8E139B5E8D30729DF894FEDDF8C5" bold="true" box="[199,334,1802,1827]" pageId="10" pageNumber="223">Etymology.</emphasis>
|
||
The specific name
|
||
<taxonomicName id="4C5B29829B5E8D307066F895FD17F8C4" authorityName="Grisales-Martínez & Velasco & Bolívar & Williams & Daza" authorityYear="2017" box="[572,644,1803,1826]" class="Reptilia" family="Dactyloidae" genus="Anolis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="223" phylum="Chordata" rank="species" species="urraoi">
|
||
<emphasis id="B92F8E139B5E8D307066F895FD17F8C4" box="[572,644,1803,1826]" italics="true" pageId="10" pageNumber="223">urraoi</emphasis>
|
||
</taxonomicName>
|
||
refers to the locality where the new species was found, the Urrao municipality, a town on the northwestern Andes in
|
||
<collectingRegion id="499F9CE39B5E8D30708EF8B0FCDBF8A1" box="[724,840,1838,1863]" country="Colombia" name="Antioquia" pageId="10" pageNumber="223">Antioquia</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |