treatments-xml/data/03/F2/DE/03F2DE0CA936C02CFF5B3785FE0AE3AB.xml
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<document ID-DOI="10.15560/16.4.821" ID-ISSN="1809-127X" approvalRequired="71" approvalRequired_for_taxonomicNames="9" approvalRequired_for_textStreams="60" approvalRequired_for_treatments="2" checkinTime="1630327227302" checkinUser="marcus" docAuthor="Cárdenas-Bautista, Johann Stephens, Parada-Alfonso, Jenny Andrea &amp; CarvajalCogollo, Juan E." docDate="2020" docId="03F2DE0CA936C02CFF5B3785FE0AE3AB" docLanguage="en" docName="CheckList.2020.16.4.821-839.pdf" docOrigin="Check List 16 (4)" docSource="http://dx.doi.org/10.15560/16.4.821" docStyle="DocumentStyle:BB30DA0EB5D11E1D6FE264CD32192DE2.1:CheckList.2017-.journal_article.annotatedlist" docStyleId="BB30DA0EB5D11E1D6FE264CD32192DE2" docStyleName="CheckList.2017-.journal_article.annotatedlist" docStyleVersion="1" docTitle="Anisocanthon villosus" docType="treatment" docVersion="1" lastPageNumber="824" masterDocId="FFCBA674A935C02FFFF33644FFE2E676" masterDocTitle="Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia" masterLastPageNumber="839" masterPageNumber="821" pageId="3" pageNumber="824" updateTime="1630327251151" updateUser="marcus" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Dung beetles (Scarabaeidae, Scarabaeinae) of the Foothills-Andean Forest strip of Villavicencio, Colombia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cárdenas-Bautista, Johann Stephens</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Parada-Alfonso, Jenny Andrea</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CarvajalCogollo, Juan E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Check List</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-07-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>16</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>821</mods:start>
<mods:end>839</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.15560/16.4.821</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.15560/16.4.821</mods:identifier>
<mods:identifier type="ISSN">1809-127X</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03F2DE0CA936C02CFF5B3785FE0AE3AB" httpUri="http://treatment.plazi.org/id/03F2DE0CA936C02CFF5B3785FE0AE3AB" lastPageNumber="824" pageId="3" pageNumber="824">
<subSubSection box="[168,589,449,473]" pageId="3" pageNumber="824" type="nomenclature">
<paragraph blockId="3.[168,589,449,473]" box="[168,589,449,473]" pageId="3" pageNumber="824">
<heading box="[168,589,449,473]" fontSize="10" level="2" pageId="3" pageNumber="824" reason="2">
<taxonomicName authority="(Harold, 1868)" baseAuthorityName="Harold" baseAuthorityYear="1868" box="[168,589,449,473]" class="Insecta" family="Scarabaeidae" genus="Anisocanthon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="824" phylum="Arthropoda" rank="species" species="villosus">
<emphasis bold="true" box="[168,589,449,473]" inLineHeading="true" pageId="3" pageNumber="824" reason="1">
<emphasis bold="true" box="[168,416,449,473]" italics="true" pageId="3" pageNumber="824">Anisocanthon villosus</emphasis>
(Harold, 1868)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="824" type="materials_examined">
<paragraph blockId="3.[165,776,491,785]" pageId="3" pageNumber="824">
<emphasis bold="true" box="[168,403,491,515]" pageId="3" pageNumber="824">Materials examined.</emphasis>
<materialsCitation collectingDate="2018-10-18" collectorName="Vereda El Carmen &amp; Parada Jenny" country="Colombia" county="Villavicencio" elevation="648" latitude="4.1401668" location="Farm" longLatPrecision="1" longitude="-73.66136" municipality="Vereda El Carmen" pageId="3" pageNumber="824" specimenCount="1" stateProvince="Meta">
<collectingCountry box="[408,551,491,515]" name="Colombia" pageId="3" pageNumber="824">COLOMBIA</collectingCountry>
• 4,
<quantity box="[596,688,491,515]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="824" unit="mm" value="5.6">5.60 mm</quantity>
(mean);
<collectingRegion box="[168,226,525,548]" country="Colombia" name="Meta" pageId="3" pageNumber="824">Meta</collectingRegion>
,
<collectingCounty box="[234,381,525,549]" pageId="3" pageNumber="824">Villavicencio</collectingCounty>
,
<collectorName box="[389,586,525,549]" pageId="3" pageNumber="824">
<collectingMunicipality box="[389,586,525,549]" pageId="3" pageNumber="824">Vereda El Carmen</collectingMunicipality>
</collectorName>
, “Las Pesebreras”
<location LSID="urn:lsid:plazi:treatment:03F2DE0CA936C02CFF5B3785FE0AE3AB:8E8439C1A936C02CFF56346BFF03E430" box="[165,225,559,582]" country="Colombia" county="Villavicencio" latitude="4.1401668" longLatPrecision="1" longitude="-73.66136" municipality="Vereda El Carmen" name="Farm" pageId="3" pageNumber="824" stateProvince="Meta">Farm</location>
;
<geoCoordinate box="[237,400,557,582]" degrees="04" direction="north" minutes="08" orientation="latitude" pageId="3" pageNumber="824" precision="1" seconds="24.60" value="4.1401668">04°0824.60″N</geoCoordinate>
,
<geoCoordinate box="[411,596,557,582]" degrees="073" direction="west" minutes="39" orientation="longitude" pageId="3" pageNumber="824" precision="1" seconds="40.89" value="-73.66136">073°3940.89″ W</geoCoordinate>
;
<elevation box="[607,733,558,583]" metricMagnitude="2" metricUnit="m" metricValue="6.4799999999999995" pageId="3" pageNumber="824" unit="m" value="648.0">
<quantity box="[607,676,558,582]" metricMagnitude="2" metricUnit="m" metricValue="6.4799999999999995" pageId="3" pageNumber="824" unit="m" value="648.0">648 m</quantity>
a.s.l.
</elevation>
;
<date pageId="3" pageNumber="824" value="2018-10-18">
<collectingDate pageId="3" pageNumber="824" value="2018-10-18">18 Oct. 2018</collectingDate>
</date>
; Cárdenas Johann and
<collectorName box="[520,664,592,617]" pageId="3" pageNumber="824">Parada Jenny</collectorName>
leg.; open pasture; Crplt4; UPTC-In-00018
</materialsCitation>
. •
<materialsCitation collectingDate="2019-04-29" collectorName="Vereda El Carmen &amp; Cardenas Johann &amp; Parada Jenny &amp; Crpat" country="Colombia" county="Villavicencio" elevation="672" latitude="4.139436" location="Farm" longLatPrecision="1" longitude="-73.6635" municipality="Vereda El Carmen" pageId="3" pageNumber="824" specimenCount="1" specimenCount-female="1" stateProvince="Meta">
<specimenCount box="[549,579,626,650]" pageId="3" pageNumber="824" type="female">1♀</specimenCount>
,
<quantity box="[592,687,626,650]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="824" unit="mm" value="5.6">5.60 mm</quantity>
(mean);
<collectingRegion box="[168,226,660,683]" country="Colombia" name="Meta" pageId="3" pageNumber="824">Meta</collectingRegion>
,
<collectingCounty box="[240,387,660,684]" pageId="3" pageNumber="824">Villavicencio</collectingCounty>
,
<collectorName box="[400,609,660,684]" pageId="3" pageNumber="824">
<collectingMunicipality box="[400,609,660,684]" pageId="3" pageNumber="824">Vereda El Carmen</collectingMunicipality>
</collectorName>
, “Las Pesebreras”
<location LSID="urn:lsid:plazi:treatment:03F2DE0CA936C02CFF5B3785FE0AE3AB:8E8439C1A936C02CFF2934F2FEF5E4BB" box="[218,279,694,717]" country="Colombia" county="Villavicencio" latitude="4.139436" longLatPrecision="1" longitude="-73.6635" municipality="Vereda El Carmen" name="Farm" pageId="3" pageNumber="824" stateProvince="Meta">Farm</location>
;
<geoCoordinate box="[288,449,692,717]" degrees="04" direction="north" minutes="08" orientation="latitude" pageId="3" pageNumber="824" precision="1" seconds="21.97" value="4.139436">04°0821.97″N</geoCoordinate>
,
<geoCoordinate box="[458,637,692,717]" degrees="073" direction="west" minutes="39" orientation="longitude" pageId="3" pageNumber="824" precision="1" seconds="48.60" value="-73.6635">073°3948.60″W</geoCoordinate>
;
<elevation box="[647,769,693,717]" metricMagnitude="2" metricUnit="m" metricValue="6.72" pageId="3" pageNumber="824" unit="m" value="672.0">
<quantity box="[647,713,693,717]" metricMagnitude="2" metricUnit="m" metricValue="6.72" pageId="3" pageNumber="824" unit="m" value="672.0">672 m</quantity>
a.s.l.
</elevation>
;
<date box="[167,314,727,751]" pageId="3" pageNumber="824" value="2019-04-29">
<collectingDate box="[167,314,727,751]" pageId="3" pageNumber="824" value="2019-04-29">29 Apr. 2019</collectingDate>
</date>
;
<collectorName box="[327,515,727,751]" pageId="3" pageNumber="824">Cárdenas Johann</collectorName>
and
<collectorName box="[573,719,727,752]" pageId="3" pageNumber="824">Parada Jenny</collectorName>
leg.; wooded pasture;
<collectorName box="[352,417,761,785]" pageId="3" pageNumber="824">Crpat</collectorName>
8; UPTC-In-00019
</materialsCitation>
.
</paragraph>
<paragraph blockId="3.[168,775,802,1028]" pageId="3" pageNumber="824">
<emphasis bold="true" box="[168,330,802,826]" pageId="3" pageNumber="824">Identification.</emphasis>
This species can be distinguished from other
<taxonomicName box="[238,478,836,860]" class="Insecta" family="Scarabaeidae" genus="Anisocanthon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="824" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[238,386,836,860]" italics="true" pageId="3" pageNumber="824">Anisocanthon</emphasis>
species
</taxonomicName>
by the bidentate clypeus with a pair of protuberances between the eyes; lateral edge of the pronotum angled towards its anterior medial parts; reddish to copper body with golden and strong setae from head to pygidium in the dorsal view (
<bibRefCitation author="Harold EV" pageId="3" pageNumber="824" pagination="1 - 144" refId="ref17077" refString="Harold EV (1868 a) Monographie der gattung Canthon. Berliner Entomologische Zeitschrift 12: 1 - 144." type="journal article" year="1868">Harold 1868a</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="824" type="distribution">
<paragraph blockId="3.[167,776,1046,1441]" pageId="3" pageNumber="824">
<emphasis bold="true" box="[168,317,1046,1070]" pageId="3" pageNumber="824">Distribution.</emphasis>
This species is widely distributed. It has been found in
<collectingCountry box="[323,438,1080,1104]" name="Venezuela" pageId="3" pageNumber="824">Venezuela</collectingCountry>
,
<collectingCountry box="[449,557,1080,1104]" name="Colombia" pageId="3" pageNumber="824">Colombia</collectingCountry>
,
<collectingCountry box="[567,659,1080,1104]" name="Ecuador" pageId="3" pageNumber="824">Ecuador</collectingCountry>
,
<collectingCountry box="[670,723,1080,1103]" name="Peru" pageId="3" pageNumber="824">Peru</collectingCountry>
,
<collectingCountry name="Bolivia" pageId="3" pageNumber="824">Bolivia</collectingCountry>
,
<collectingCountry box="[235,347,1114,1138]" name="Argentina" pageId="3" pageNumber="824">Argentina</collectingCountry>
, and
<collectingCountry box="[414,515,1114,1138]" name="Paraguay" pageId="3" pageNumber="824">Paraguay</collectingCountry>
(
<bibRefCitation author="Pereira FS &amp; Martinez A" pageId="3" pageNumber="824" pagination="91 - 192" refId="ref18349" refString="Pereira FS, Martinez A (1956) Os generos de Canthonini americanos (Col. Scarabaeidae). Revista Brasiliense Entomologia 6: 91 - 192." type="journal article" year="1956">Pereira and Martínez 1956</bibRefCitation>
). In
<collectingCountry box="[274,378,1147,1171]" name="Colombia" pageId="3" pageNumber="824">Colombia</collectingCountry>
it is known from the department of
<collectingRegion box="[170,225,1181,1204]" country="Colombia" name="Meta" pageId="3" pageNumber="824">Meta</collectingRegion>
(
<bibRefCitation author="Medina CA &amp; Lopera-Toro A &amp; Vitolo A &amp; Gill B" box="[244,464,1181,1205]" pageId="3" pageNumber="824" pagination="131 - 144" refId="ref17513" refString="Medina CA, Lopera-Toro A, Vitolo A, Gill B (2001) Escarabajos Coprofagos (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. Biota Colombiana 2 (2): 131 - 144." type="journal article" year="2001">Medina et al. 2001</bibRefCitation>
). This species is associated with disturbed habitats, found in forest remnants, gallery forests, forest edges and cattle pastures with grasses of variable height. The beetles were previously recorded in areas of up to
<quantity box="[348,414,1316,1340]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="824" unit="m" value="650.0">650 m</quantity>
a.s.l., and for this study area they were found at heights above
<quantity box="[469,535,1350,1374]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="824" unit="m" value="700.0">700 m</quantity>
a.s.l. This species was rare in the area (55 individuals), and it was mostly found in grasslands that were used for cattle grazing.
</paragraph>
<paragraph blockId="3.[168,488,1477,1501]" box="[168,488,1477,1501]" pageId="3" pageNumber="824">
<taxonomicName authority="Hoffmannsegg, 1817" authorityName="Hoffmannsegg" authorityYear="1817" box="[168,488,1477,1501]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="824" phylum="Arthropoda" rank="genus">
<emphasis box="[168,261,1477,1501]" italics="true" pageId="3" pageNumber="824">Canthon</emphasis>
Hoffmannsegg, 1817
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>