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<document id="9EE4C88F6DEC5F88EDAA6FBF84617888" ID-DOI="10.5281/zenodo.196812" ID-GBIF-Dataset="7e7c2466-8c95-4e7c-90fb-b7ba92ccd8fe" ID-ISSN="1175-5326" ID-Zenodo-Dep="196812" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460253291746" checkinUser="plazi" docAuthor="Anstis, Marion, Tyler, Michael J., Roberts, Dale, Price, Luke C. &amp; Doughty, Paul" docDate="2010" docId="03E32F48FF86FFEEFF736597FD91FEBF" docLanguage="en" docName="zt02550p057.pdf" docOrigin="Zootaxa 2550" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Litoria aurifera Anstis, Tyler, Roberts, Price &amp; Doughty, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="FFDA5730FF84FFE8FFE46735FF88FFF8" masterDocTitle="A new species of Litoria (Anura: Hylidae) with a highly distinctive tadpole from the north-western Kimberley region of Western Australia" masterLastPageNumber="57" masterPageNumber="39" pageNumber="41" updateTime="1698243060239" updateUser="plazi">
<mods:mods id="18A79FA7DC3E3DB2AEC3D3585D4706C3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="984F765CB256C0A1DE94EEE5EDA72603">
<mods:title id="2778B7629EF1928DF528ED221A75F6B6">A new species of Litoria (Anura: Hylidae) with a highly distinctive tadpole from the north-western Kimberley region of Western Australia</mods:title>
</mods:titleInfo>
<mods:name id="69AAFB0AFFAB058EB822167E1235560B" type="personal">
<mods:role id="6346B9D38343D1558EEFFF0124443D9B">
<mods:roleTerm id="BB39173C208EFAEB47903CE7B9BD7BF7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BC698CD9BDC5A9D788B29F2C9C441DB0">Anstis, Marion</mods:namePart>
</mods:name>
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<mods:namePart id="D70B79C52D0719F405B92DBC059166E1">Tyler, Michael J.</mods:namePart>
</mods:name>
<mods:name id="66E3B42E040FB42D640B11C6C246CF55" type="personal">
<mods:role id="D53197B75EADB45ADE0B709310EBB9DF">
<mods:roleTerm id="91EB6A2E86012086AB201948D570F786">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="364395BC611DE9CCA7766F3479A9C30D">Roberts, Dale</mods:namePart>
</mods:name>
<mods:name id="30294B0D3C789DB074EED0B24E505F21" type="personal">
<mods:role id="A55E1A320917C277F96B07A09393E7C4">
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</mods:role>
<mods:namePart id="DF31D6AE70A22DB8DF00992D7D59A53D">Price, Luke C.</mods:namePart>
</mods:name>
<mods:name id="F5B159D71C0FDFC62957511E2138A467" type="personal">
<mods:role id="A581E41E324B11178836F82590527A2F">
<mods:roleTerm id="7FEBE7947A015CB309D1F85600596167">Author</mods:roleTerm>
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<mods:namePart id="E68BA82F2E554C24184DBAE08575AF85">Doughty, Paul</mods:namePart>
</mods:name>
<mods:typeOfResource id="793320DF59BDE27510BBC05E4691F238">text</mods:typeOfResource>
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<mods:title id="EEF793D17FFF3B2D8425589D9268E8C0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F9349A2E69B8DAC7EDAF0BE1516BF247">
<mods:date id="ACF44CAF10CBEBC882FBB9E013C1C85F">2010</mods:date>
<mods:detail id="D9C2CBBCCA53B74C050FA3C3D7712E22" type="volume">
<mods:number id="DC657A6E726037486720A83E58B5FB56">2550</mods:number>
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<mods:start id="9C398D9FE9D1DAA9F324FC2566E0A8E4">39</mods:start>
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<mods:classification id="3C67DE818A1772E06D9DEE96768AFCFB">journal article</mods:classification>
<mods:identifier id="C96E46C1CBFA8FE0F077831655178820" type="DOI">10.5281/zenodo.196812</mods:identifier>
<mods:identifier id="FC130F3ECAE84EF2270754E2D804E6F5" type="GBIF-Dataset">7e7c2466-8c95-4e7c-90fb-b7ba92ccd8fe</mods:identifier>
<mods:identifier id="63AD20E800F3657C6FC9B27B7BD75A76" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E38DBF74D18D3EF984C07DE436433F77" type="Zenodo-Dep">196812</mods:identifier>
</mods:mods>
<treatment id="03E32F48FF86FFEEFF736597FD91FEBF" ID-DOI="http://doi.org/10.5281/zenodo.5615769" ID-GBIF-Taxon="119392092" ID-Zenodo-Dep="5615769" LSID="urn:lsid:plazi:treatment:03E32F48FF86FFEEFF736597FD91FEBF" httpUri="http://treatment.plazi.org/id/03E32F48FF86FFEEFF736597FD91FEBF" lastPageId="6" lastPageNumber="43" pageId="2" pageNumber="41">
<subSubSection id="C350CDD5FF86FFEAFF736597FE42FD27" pageId="2" pageNumber="41" type="nomenclature">
<paragraph id="8BF59E5EFF86FFEAFF736597FE31FD44" blockId="2.[151,458,674,735]" box="[151,441,674,700]" pageId="2" pageNumber="41">
<heading id="D0BD2932FF86FFEAFF736597FE31FD44" bold="true" box="[151,441,674,700]" fontSize="11" level="1" pageId="2" pageNumber="41" reason="1">
<emphasis id="B93E424CFF86FFEAFF736597FE31FD44" bold="true" box="[151,441,674,700]" pageId="2" pageNumber="41">
<taxonomicName id="4C4AE5DDFF86FFEAFF736597FEDCFD44" box="[151,340,674,700]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="aurifera" status="sp. nov.">
<emphasis id="B93E424CFF86FFEAFF736597FEDCFD44" bold="true" box="[151,340,674,700]" italics="true" pageId="2" pageNumber="41">Litoria aurifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DFF37FF86FFEAFEB86597FE31FD44" box="[348,441,674,700]" pageId="2" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF59E5EFF86FFEAFF7365F0FE42FD27" blockId="2.[151,458,674,735]" box="[151,458,709,735]" pageId="2" pageNumber="41">Kimberley Rockhole Frog</paragraph>
</subSubSection>
<subSubSection id="C350CDD5FF86FFEAFF73643AFEBFFAF7" pageId="2" pageNumber="41" type="materials_examined">
<paragraph id="8BF59E5EFF86FFEAFF73643AFBDEFC8C" blockId="2.[151,1437,783,1967]" pageId="2" pageNumber="41">
<emphasis id="B93E424CFF86FFEAFF73643AFE9BFCD1" bold="true" box="[151,275,783,809]" pageId="2" pageNumber="41">
<typeStatus id="54F120FCFF86FFEAFF73643AFE86FCD1" box="[151,270,783,809]" pageId="2" pageNumber="41" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED5B069BFF86FFEAFEFF6425FEEBFCD2" LSID="urn:lsid:biocol.org:col:34981" box="[283,355,784,810]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="41">WAM</collectionCode>
R169913, an adult female collected as a tadpole from a creek near Bachsten Creek, Prince Regent River Nature Reserve, Western
<collectingCountry id="F35DDECEFF86FFEAFD646400FD79FCB7" box="[640,753,821,847]" name="Australia" pageId="2" pageNumber="41">Australia</collectingCountry>
(elevation
<quantity id="4CB233BBFF86FFEAFC9B6400FC40FCB7" box="[895,968,821,847]" metricMagnitude="2" metricUnit="m" metricValue="3.4299999999999997" pageId="2" pageNumber="41" unit="m" value="343.0">343m</quantity>
;
<geoCoordinate id="EE7EF899FF86FFEAFC326400FBF7FCB7" box="[982,1151,821,847]" direction="south" orientation="latitude" pageId="2" pageNumber="41" precision="1" value="-15.988389">15º59’18.2”S</geoCoordinate>
,
<geoCoordinate id="EE7EF899FF86FFEAFB686400FACDFCB7" box="[1164,1349,821,847]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="1" value="125.32725">125º19’38.1”E</geoCoordinate>
) on
<date id="FFF4B89EFF86FFEAFA9A6400FEBCFC8C" pageId="2" pageNumber="41" value="2007-01-23">23 January 2007</date>
by J.D. Roberts and P. Doughty and raised to an adult by M. Anstis.
</paragraph>
<paragraph id="8BF59E5EFF86FFEAFF22644AFC98FC1C" blockId="2.[151,1437,783,1967]" pageId="2" pageNumber="41">
<typeStatus id="54F120FCFF86FFEAFF22644AFECBFC61" box="[198,323,895,921]" pageId="2" pageNumber="41" type="paratype">
<emphasis id="B93E424CFF86FFEAFF22644AFECBFC61" bold="true" box="[198,323,895,921]" pageId="2" pageNumber="41">Paratypes</emphasis>
</typeStatus>
(prefixes excluded).
<typeStatus id="54F120FCFF86FFEAFDA364B5FD31FC62" box="[583,697,896,922]" pageId="2" pageNumber="41" type="paratype">Paratypes</typeStatus>
include a mix of field-collected adults, males with recorded calls, and two adults reared from tadpoles.
<collectionCode id="ED5B069BFF86FFEAFD7A6490FD6EFC47" LSID="urn:lsid:biocol.org:col:34981" box="[670,742,933,959]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="2" pageNumber="41">WAM</collectionCode>
specimens are all taken from smaller unnamed creeks off Bachsten Creek, Prince Regent River Nature Reserve.
</paragraph>
<paragraph id="8BF59E5EFF86FFEAFF2164C5FCE7FB67" blockId="2.[151,1437,783,1967]" pageId="2" pageNumber="41">
R168004 (female), 168006 (male), 168008 (female), 168009 (male), 168010 (female), 168013–4 (males), 168016–8 (males): elevation
<quantity id="4CB233BBFF86FFEAFDE56320FDC1FBD7" box="[513,585,1045,1071]" metricMagnitude="2" metricUnit="m" metricValue="3.4899999999999998" pageId="2" pageNumber="41" unit="m" value="349.0">349m</quantity>
,
<geoCoordinate id="EE7EF899FF86FFEAFDB36320FD6DFBD7" box="[599,741,1045,1071]" direction="south" orientation="latitude" pageId="2" pageNumber="41" precision="15" value="-15.989445">15°59'22&quot;S</geoCoordinate>
,
<geoCoordinate id="EE7EF899FF86FFEAFD176320FC1BFBD7" box="[755,915,1045,1071]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="15" value="125.32667">125°19'36&quot;E</geoCoordinate>
; 168059 (male), 168062 (male): elevation
<quantity id="4CB233BBFF86FFEAFF73630FFF55FBAC" box="[151,221,1082,1108]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="41" unit="m" value="260.0">260m</quantity>
,
<geoCoordinate id="EE7EF899FF86FFEAFF0E630FFEFAFBAC" box="[234,370,1082,1108]" direction="south" orientation="latitude" pageId="2" pageNumber="41" precision="15" value="-15.748334">15°44'54&quot;S</geoCoordinate>
,
<geoCoordinate id="EE7EF899FF86FFEAFE64630FFD91FBAC" box="[384,537,1082,1108]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="15" value="125.37361">125°22'25&quot;E</geoCoordinate>
; 168095 (male): elevation
<quantity id="4CB233BBFF86FFEAFCBD630FFC17FBAC" box="[857,927,1082,1108]" metricMagnitude="2" metricUnit="m" metricValue="2.32" pageId="2" pageNumber="41" unit="m" value="232.0">232m</quantity>
,
<geoCoordinate id="EE7EF899FF86FFEAFC4F630FFBBCFBAC" box="[939,1076,1082,1108]" direction="south" orientation="latitude" pageId="2" pageNumber="41" precision="15" value="-15.428333">15°25'42&quot;S</geoCoordinate>
,
<geoCoordinate id="EE7EF899FF86FFEAFBA5630FFB53FBAC" box="[1089,1243,1082,1108]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="15" value="125.31167">125°18'42&quot;E</geoCoordinate>
; 168114 (male), 168116 (male), 168118 (female), 168129 (male), 168134 (female): elevation
<quantity id="4CB233BBFF86FFEAFBC36355FBD6FB82" box="[1063,1118,1120,1146]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="41" unit="m" value="46.0">46m</quantity>
,
<geoCoordinate id="EE7EF899FF86FFEAFB8F6355FB7AFB82" box="[1131,1266,1120,1146]" direction="south" orientation="latitude" pageId="2" pageNumber="41" precision="15" value="-15.594444">15°35'40&quot;S</geoCoordinate>
,
<geoCoordinate id="EE7EF899FF86FFEAFB1B6355FA1FFB82" box="[1279,1431,1120,1146]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="15" value="125.187225">125°11'14&quot;E</geoCoordinate>
; 168182–5 (males): elevation
<quantity id="4CB233BBFF86FFEAFE0963B0FDBBFB67" box="[493,563,1157,1183]" metricMagnitude="2" metricUnit="m" metricValue="4.08" pageId="2" pageNumber="41" unit="m" value="408.0">408m</quantity>
,
<geoCoordinate id="EE7EF899FF86FFEAFDDB63B0FD4EFB67" box="[575,710,1157,1183]" direction="south" orientation="latitude" pageId="2" pageNumber="41" precision="15" value="-15.859445">15°51'34&quot;S</geoCoordinate>
,
<geoCoordinate id="EE7EF899FF86FFEAFD3763B0FCE3FB67" box="[723,875,1157,1183]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="15" value="125.69639">125°41'47&quot;E</geoCoordinate>
.
</paragraph>
<paragraph id="8BF59E5EFF86FFEAFF21639FFEBFFAF7" blockId="2.[151,1437,783,1967]" pageId="2" pageNumber="41">
<collectionCode id="ED5B069BFF86FFEAFF21639FFE94FB3C" LSID="urn:lsid:biocol.org:col:34982" box="[197,284,1194,1220]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="2" pageNumber="41">SAMA</collectionCode>
specimens were reared from tadpoles collected at
<quantity id="4CB233BBFF86FFEAFC6B639FFC6DFB3C" box="[911,997,1194,1220]" metricMagnitude="3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="41" unit="km" value="5.1">5.1 km</quantity>
ESE of junction of Pitta Creek and Prince Regent River, Western
<collectingCountry id="F35DDECEFF86FFEAFE1F63E5FDE3FB12" box="[507,619,1232,1258]" name="Australia" pageId="2" pageNumber="41">Australia</collectingCountry>
: 63001 and 63002 (male and female): elevation
<quantity id="4CB233BBFF86FFEAFB4063E5FB62FB12" box="[1188,1258,1232,1258]" metricMagnitude="2" metricUnit="m" metricValue="4.08" pageId="2" pageNumber="41" unit="m" value="408.0">408m</quantity>
,
<geoCoordinate id="EE7EF899FF86FFEAFB1263E5FA1FFB12" box="[1270,1431,1232,1258]" direction="south" orientation="latitude" pageId="2" pageNumber="41" precision="1" value="-15.8596115">15º51’34.6”S</geoCoordinate>
,
<geoCoordinate id="EE7EF899FF86FFEAFF7363C0FEBAFAF7" box="[151,306,1269,1295]" direction="east" orientation="longitude" pageId="2" pageNumber="41" precision="15" value="125.69639">125º41’47”E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C350CDD5FF86FFEAFF22622FFBFFF937" pageId="2" pageNumber="41" type="diagnosis">
<paragraph id="8BF59E5EFF86FFEAFF22622FFDCBFA32" blockId="2.[151,1437,783,1967]" pageId="2" pageNumber="41">
<emphasis id="B93E424CFF86FFEAFF22622FFECEFACC" bold="true" box="[198,326,1306,1332]" pageId="2" pageNumber="41">Diagnosis.</emphasis>
<collectingCountry id="F35DDECEFF86FFEAFEB4622FFE34FACC" box="[336,444,1306,1332]" name="Australia" pageId="2" pageNumber="41">Australia</collectingCountry>
has three hylid genera of which
<taxonomicName id="4C4AE5DDFF86FFEAFCA9622EFC41FACC" box="[845,969,1307,1332]" class="Amphibia" family="Pelodryadidae" genus="Cyclorana" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="B93E424CFF86FFEAFCA9622EFC41FACC" box="[845,969,1307,1332]" italics="true" pageId="2" pageNumber="41">Cyclorana</emphasis>
</taxonomicName>
is fossorial,
<taxonomicName id="4C4AE5DDFF86FFEAFB82622EFB7AFACC" box="[1126,1266,1307,1332]" class="Amphibia" family="Pelodryadidae" genus="Nyctimystes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="B93E424CFF86FFEAFB82622EFB7AFACC" box="[1126,1266,1307,1332]" italics="true" pageId="2" pageNumber="41">Nyctimystes</emphasis>
</taxonomicName>
has a vertical pupil and palpebral venation, and
<emphasis id="B93E424CFF86FFEAFDCB6275FD02FAA1" box="[559,650,1344,1369]" italics="true" pageId="2" pageNumber="41">
<taxonomicName id="4C4AE5DDFF86FFEAFDCB6275FD0DFAA1" box="[559,645,1344,1369]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">Litoria</taxonomicName>
,
</emphasis>
which has a horizontal pupil and no palpebral venation. The new taxon is assigned to
<taxonomicName id="4C4AE5DDFF86FFEAFE6F6250FE56FA86" box="[395,478,1381,1406]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="B93E424CFF86FFEAFE6F6250FE56FA86" box="[395,478,1381,1406]" italics="true" pageId="2" pageNumber="41">Litoria</emphasis>
</taxonomicName>
because of its obvious morphological affinity to
<taxonomicName id="4C4AE5DDFF86FFEAFBCE6250FB78FA86" box="[1066,1264,1381,1406]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="meiriana">
<emphasis id="B93E424CFF86FFEAFBCE6250FB78FA86" box="[1066,1264,1381,1406]" italics="true" pageId="2" pageNumber="41">Litoria meiriana</emphasis>
</taxonomicName>
in body form, size, skin colour and texture and its ability to bounce lightly across the water surface without immersion, an attribute shared only by
<taxonomicName id="4C4AE5DDFF86FFEAFE506285FDB5FA31" box="[436,573,1456,1481]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="meiriana">
<emphasis id="B93E424CFF86FFEAFE506285FDB5FA31" box="[436,573,1456,1481]" italics="true" pageId="2" pageNumber="41">L. meiriana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF59E5EFF86FFEAFF2262E0FEF0F97C" blockId="2.[151,1437,783,1967]" pageId="2" pageNumber="41">
<taxonomicName id="4C4AE5DDFF86FFEAFF2262E0FE0AFA16" box="[198,386,1493,1518]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="aurifera">
<emphasis id="B93E424CFF86FFEAFF2262E0FE0AFA16" box="[198,386,1493,1518]" italics="true" pageId="2" pageNumber="41">Litoria aurifera</emphasis>
</taxonomicName>
is a small (to
<quantity id="4CB233BBFF86FFEAFDD162E0FD06FA17" box="[565,654,1493,1519]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="41" unit="mm" value="22.0">22 mm</quantity>
), saxicoline species, characterized by a somewhat dorsoventrally compressed, slight body and numerous scattered tubercles over the dorsal surface. The snout is moderately pointed in dorsal view, canthus rostralis slightly curved, tympanum distinct and toes almost fully webbed. It can be distinguished from
<taxonomicName id="4C4AE5DDFF86FFEAFE2A6170FDD0F9A6" box="[462,600,1605,1630]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="meiriana">
<emphasis id="B93E424CFF86FFEAFE2A6170FDD0F9A6" box="[462,600,1605,1630]" italics="true" pageId="2" pageNumber="41">L. meiriana</emphasis>
</taxonomicName>
primarily by its vividly pigmented gold and black tadpole and by adult call characteristics.
</paragraph>
<paragraph id="8BF59E5EFF86FFEAFF2261A5FBFFF937" blockId="2.[151,1437,783,1967]" pageId="2" pageNumber="41">
While very similar to
<taxonomicName id="4C4AE5DDFF86FFEAFE2161A5FDC6F951" box="[453,590,1680,1705]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="meiriana">
<emphasis id="B93E424CFF86FFEAFE2161A5FDC6F951" box="[453,590,1680,1705]" italics="true" pageId="2" pageNumber="41">L. meiriana</emphasis>
</taxonomicName>
, the snout of adult
<taxonomicName id="4C4AE5DDFF86FFEAFCC861A5FC23F951" box="[812,939,1680,1705]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="2" pageNumber="41" phylum="Chordata" rank="species" species="aurifera">
<emphasis id="B93E424CFF86FFEAFCC861A5FC23F951" box="[812,939,1680,1705]" italics="true" pageId="2" pageNumber="41">L. aurifera</emphasis>
</taxonomicName>
tends to be slightly more pointed in dorsal view (
<figureCitation id="137182DBFF86FFEAFF3B6180FEC8F937" box="[223,320,1717,1743]" captionStart="FIGURE 1" captionStartId="3.[151,255,1696,1720]" captionTargetBox="[224,1365,188,1680]" captionTargetId="figure@3.[217,1369,172,1705]" captionTargetPageId="3" captionText="FIGURE 1. Head of holotype of Litoria meiriana (SAMA R 9082, female) compared with head of holotype of Litoria aurifera sp. nov. (WAM R 169913, female) and hand and foot of holotype of Litoria aurifera (WAM R 169913, female). A, B = holotype of L. meiriana dorsal and lateral views; C, D = holotype of L. aurifera dorsal and lateral views; E, F = hand and foot of L. aurifera. Bar represents 5 mm." httpUri="https://zenodo.org/record/196813/files/figure.png" pageId="2" pageNumber="41">Figure 1</figureCitation>
C), and SVL, TL and HW are slightly greater than the former species.
</paragraph>
</subSubSection>
<subSubSection id="C350CDD5FF86FFECFF2261EFFDA8F8F2" lastPageId="4" lastPageNumber="43" pageId="2" pageNumber="41" type="description">
<paragraph id="8BF59E5EFF86FFEAFF2261EFFD19F857" blockId="2.[151,1437,783,1967]" pageId="2" pageNumber="41">
<emphasis id="B93E424CFF86FFEAFF2261EFFE7AF90C" bold="true" box="[198,498,1754,1780]" pageId="2" pageNumber="41">
Description of
<typeStatus id="54F120FCFF86FFEAFE9B61EFFE65F90C" box="[383,493,1754,1780]" pageId="2" pageNumber="41" type="holotype">holotype</typeStatus>
.
</emphasis>
Size small (SVL
<quantity id="4CB233BBFF86FFEAFD2161EFFCBBF90C" box="[709,819,1754,1780]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="2" pageNumber="41" unit="mm" value="21.8">21.8 mm</quantity>
). Head slightly longer (
<quantity id="4CB233BBFF86FFEAFBAC61EFFB13F90C" box="[1096,1179,1754,1780]" metricMagnitude="-2" metricUnit="m" metricValue="7.7" pageId="2" pageNumber="41" unit="cm" value="7.7">7.7 cm</quantity>
) than broad (
<quantity id="4CB233BBFF86FFEAFADD61EFFA07F90C" box="[1337,1423,1754,1780]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="41" unit="cm" value="6.4">6.4 cm</quantity>
), HL/HW 1.203. Canthus rostralis very slightly curved. Snout moderately pointed when viewed from above; obtusely rounded in profile. Eye prominent, horizontal diameter
<quantity id="4CB233BBFF86FFEAFC446010FC77F8C7" box="[928,1023,1829,1855]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="41" unit="mm" value="2.7">2.7 mm</quantity>
. Eye to naris distance greater than internarial span (EN/IN 1.7). Tympanum distinct with sharply defined annulus; horizontal diameter 2.0 mm. Small circular gland at angle of jaws and poorly developed supratympanic fold. Vomerine teeth oblique, slope slightly downwards towards medial corner.
</paragraph>
<caption id="DF35CED6FF87FFEBFF736195FD3BF8EF" httpUri="https://zenodo.org/record/196813/files/figure.png" pageId="3" pageNumber="42" targetBox="[224,1365,188,1680]" targetPageId="3">
<paragraph id="8BF59E5EFF87FFEBFF736195FD3BF8EF" blockId="3.[151,1437,1696,1816]" pageId="3" pageNumber="42">
<emphasis id="B93E424CFF87FFEBFF736195FE93F940" bold="true" box="[151,283,1696,1720]" pageId="3" pageNumber="42">FIGURE 1.</emphasis>
Head of holotype of
<taxonomicName id="4C4AE5DDFF87FFEBFDE06194FD3FF940" box="[516,695,1697,1720]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="meiriana">
<emphasis id="B93E424CFF87FFEBFDE06194FD3FF940" box="[516,695,1697,1720]" italics="true" pageId="3" pageNumber="42">Litoria meiriana</emphasis>
</taxonomicName>
(SAMA R9082, female) compared with head of holotype of
<taxonomicName id="4C4AE5DDFF87FFEBFAB56194FF66F920" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="aurifera" status="sp. nov.">
<emphasis id="B93E424CFF87FFEBFAB56194FF66F920" italics="true" pageId="3" pageNumber="42">Litoria aurifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DFF37FF87FFEBFF1161F5FEC3F920" box="[245,331,1728,1752]" pageId="3" pageNumber="42" rank="species">
<emphasis id="B93E424CFF87FFEBFF1161F5FEC3F920" bold="true" box="[245,331,1728,1752]" pageId="3" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
(WAM R169913, female) and hand and foot of holotype of
<taxonomicName id="4C4AE5DDFF87FFEBFC3761F4FBF3F920" box="[979,1147,1729,1752]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="aurifera">
<emphasis id="B93E424CFF87FFEBFC3761F4FBF3F920" box="[979,1147,1729,1752]" italics="true" pageId="3" pageNumber="42">Litoria aurifera</emphasis>
</taxonomicName>
(WAM R169913, female). A, B = holotype of
<taxonomicName id="4C4AE5DDFF87FFEBFE8E61D4FE6FF900" box="[362,487,1761,1784]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="meiriana">
<emphasis id="B93E424CFF87FFEBFE8E61D4FE6FF900" box="[362,487,1761,1784]" italics="true" pageId="3" pageNumber="42">L. meiriana</emphasis>
</taxonomicName>
dorsal and lateral views; C, D = holotype of
<taxonomicName id="4C4AE5DDFF87FFEBFC2B61D4FBCCF900" box="[975,1092,1761,1784]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="aurifera">
<emphasis id="B93E424CFF87FFEBFC2B61D4FBCCF900" box="[975,1092,1761,1784]" italics="true" pageId="3" pageNumber="42">L. aurifera</emphasis>
</taxonomicName>
dorsal and lateral views; E, F = hand and foot of
<taxonomicName id="4C4AE5DDFF87FFEBFEA96034FE48F8E0" box="[333,448,1793,1816]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="3" pageNumber="42" phylum="Chordata" rank="species" species="aurifera">
<emphasis id="B93E424CFF87FFEBFEA96034FE48F8E0" box="[333,448,1793,1816]" italics="true" pageId="3" pageNumber="42">L. aurifera</emphasis>
</taxonomicName>
. Bar represents 5 mm.
</paragraph>
</caption>
<paragraph id="8BF59E5EFF87FFECFF226073FCFDFEDA" blockId="3.[151,1437,1862,2037]" lastBlockId="4.[151,1437,152,700]" lastPageId="4" lastPageNumber="43" pageId="3" pageNumber="42">
Fingers long and slender, in decreasing order of length 3&gt;4&gt;2&gt;1, with small, transversely oval terminal discs that are distinctly wider than digits; a trace of webbing between third and fourth fingers. Subarticular tubercle at base of terminal phalanx elongate and oval. Palmar tubercle broad and flattened (
<figureCitation id="137182DBFF87FFEBFB3460A4FAB9F853" box="[1232,1329,1937,1963]" captionStart="FIGURE 1" captionStartId="3.[151,255,1696,1720]" captionTargetBox="[224,1365,188,1680]" captionTargetId="figure@3.[217,1369,172,1705]" captionTargetPageId="3" captionText="FIGURE 1. Head of holotype of Litoria meiriana (SAMA R 9082, female) compared with head of holotype of Litoria aurifera sp. nov. (WAM R 169913, female) and hand and foot of holotype of Litoria aurifera (WAM R 169913, female). A, B = holotype of L. meiriana dorsal and lateral views; C, D = holotype of L. aurifera dorsal and lateral views; E, F = hand and foot of L. aurifera. Bar represents 5 mm." httpUri="https://zenodo.org/record/196813/files/figure.png" pageId="3" pageNumber="42">Figure 1</figureCitation>
E). Tibia length slightly more than one-half SVL (TL/SVL 0.55). Toes long and slender, in decreasing order of length 4&gt;5&gt;3&gt;2&gt;1, discs slightly more rounded than oval and wider than digits. Interdigital webbing extensive, reaching the base of the discs of all digits except the third and fourth (
<figureCitation id="137182DBFF80FFECFC0667ADFBCEFF4A" box="[994,1094,152,178]" captionStart="FIGURE 1" captionStartId="3.[151,255,1696,1720]" captionTargetBox="[224,1365,188,1680]" captionTargetId="figure@3.[217,1369,172,1705]" captionTargetPageId="3" captionText="FIGURE 1. Head of holotype of Litoria meiriana (SAMA R 9082, female) compared with head of holotype of Litoria aurifera sp. nov. (WAM R 169913, female) and hand and foot of holotype of Litoria aurifera (WAM R 169913, female). A, B = holotype of L. meiriana dorsal and lateral views; C, D = holotype of L. aurifera dorsal and lateral views; E, F = hand and foot of L. aurifera. Bar represents 5 mm." httpUri="https://zenodo.org/record/196813/files/figure.png" pageId="4" pageNumber="43">Figure 1</figureCitation>
F). Inner metatarsal tubercle small and elongate; outer metatarsal tubercle extremely small and circular. The dorsum bears numerous scattered slightly raised tubercles, middorsal tubercles more flattened. The throat is smooth; abdomen is weakly granular with granulation larger towards sides of body.
</paragraph>
<paragraph id="8BF59E5EFF80FFECFF226618FB26FDDF" blockId="4.[151,1437,152,700]" pageId="4" pageNumber="43">
The colour in life of the dorsal surface and limbs is mottled with shades of brown and reddish brown (Pantone 464M to 465M). Dorsal tubercles are surrounded by darker brown (462M), which is also in patches upon the hind limbs. The upper lip is often edged with dark brown and silver-white bars, and the sides of the body have scattered whitish tubercles on a dark brown background. The iris is reddish above and grey-gold below (
<figureCitation id="137182DBFF80FFECFF0A66F7FEDCFE24" box="[238,340,450,476]" captionStart="FIGURE 2" captionStartId="5.[151,255,1848,1872]" captionTargetBox="[240,1370,191,1833]" captionTargetId="figure@5.[212,1396,168,1857]" captionTargetPageId="5" captionText="FIGURE 2. Comparative adult frogs in life: Litoria meiriana and Litoria aurifera sp. nov. A = Litoria meiriana, active by day, Nawurlandja Rock, Kakadu National Park, Northern Territory. B = Litoria aurifera, at night in habitat, Prince Regent Nature Reserve, Western Australia. C = Litoria aurifera, holotype (WAM R 169913, female) dorsolateral view, during daytime, raised from tadpole in captivity. D = Litoria aurifera, holotype (WAM R 169913, female) dorsal view. E = Litoria aurifera, holotype (WAM R 169913, female) ventral view. F = Litoria aurifera, anterior view, at night in habitat, Prince Regent Nature Reserve, Western Australia. Bar represents 5 mm." httpUri="https://zenodo.org/record/196814/files/figure.png" pageId="4" pageNumber="43">Figure 2</figureCitation>
C). In ventral view, the submandibular area and throat are warm grey (1M), and the abdomen is white tinged with pale copper-pink (
<figureCitation id="137182DBFF80FFECFDBF66DDFD49FDFA" box="[603,705,488,514]" captionStart="FIGURE 2" captionStartId="5.[151,255,1848,1872]" captionTargetBox="[240,1370,191,1833]" captionTargetId="figure@5.[212,1396,168,1857]" captionTargetPageId="5" captionText="FIGURE 2. Comparative adult frogs in life: Litoria meiriana and Litoria aurifera sp. nov. A = Litoria meiriana, active by day, Nawurlandja Rock, Kakadu National Park, Northern Territory. B = Litoria aurifera, at night in habitat, Prince Regent Nature Reserve, Western Australia. C = Litoria aurifera, holotype (WAM R 169913, female) dorsolateral view, during daytime, raised from tadpole in captivity. D = Litoria aurifera, holotype (WAM R 169913, female) dorsal view. E = Litoria aurifera, holotype (WAM R 169913, female) ventral view. F = Litoria aurifera, anterior view, at night in habitat, Prince Regent Nature Reserve, Western Australia. Bar represents 5 mm." httpUri="https://zenodo.org/record/196814/files/figure.png" pageId="4" pageNumber="43">Figure 2</figureCitation>
E). The undersurface of the limbs is dark flesh colour (436M). In preservative, each of the pigments is paler and all red, gold and copper pigment is lost.
</paragraph>
<paragraph id="8BF59E5EFF80FFECFF226507FADAFD44" blockId="4.[151,1437,152,700]" pageId="4" pageNumber="43">
<emphasis id="B93E424CFF80FFECFF226507FEC9FDB4" bold="true" box="[198,321,562,588]" pageId="4" pageNumber="43">Variation.</emphasis>
Males (N = 20) range in SVL from
<quantity id="4CB233BBFF80FFECFD1D6507FC2DFDB4" box="[761,933,562,588]" metricMagnitude="-2" metricUnit="m" metricValue="1.92" metricValueMax="2.02" metricValueMin="1.8199999999999998" pageId="4" pageNumber="43" unit="mm" value="19.2" valueMax="20.2" valueMin="18.2">18.2–20.2 mm</quantity>
and females (N = 8) from
<quantity id="4CB233BBFF80FFECFB0D6507FA10FDB4" box="[1257,1432,562,588]" metricMagnitude="-2" metricUnit="m" metricValue="2.095" metricValueMax="2.16" metricValueMin="2.03" pageId="4" pageNumber="43" unit="mm" value="20.95" valueMax="21.6" valueMin="20.3">20.3–21.6 mm</quantity>
. The TL/SVL range is 0.479 to 0.594 (
<tableCitation id="C6C8ABE5FF80FFECFD83656DFD4AFD8A" box="[615,706,600,626]" captionStart="TABLE 1" captionStartId="4.[151,239,743,767]" captionTargetBox="[151,1436,890,1452]" captionTargetPageId="4" captionText="TABLE 1. Morphological measurements of Litoria aurifera sp. nov. and Litoria meiriana. Mean ± S. D. (range). SVL =" httpUri="http://table.plazi.org/id/DF35CED6FF80FFECFF7365D2FA14FD07" pageId="4" pageNumber="43" tableUuid="DF35CED6FF80FFECFF7365D2FA14FD07">Table 1</tableCitation>
). The nasal and maxillary bones are extremely thin and the anterior portion of the skull is therefore subject to distortion by compression.
<collectingCountry id="F35DDECEFF80FFECFBA26548FBEFFD6F" box="[1094,1127,637,663]" name="American Samoa" pageId="4" pageNumber="43">As</collectingCountry>
a result, some specimens were excluded from the analysis because they are slightly distorted rendering measurements unreliable.
</paragraph>
<caption id="DF35CED6FF80FFECFF7365D2FA14FD07" ID-Table-UUID="DF35CED6FF80FFECFF7365D2FA14FD07" box="[151,1436,743,767]" httpUri="http://table.plazi.org/id/DF35CED6FF80FFECFF7365D2FA14FD07" pageId="4" pageNumber="43" targetBox="[151,1436,890,1452]" targetIsTable="true" targetPageId="4">
<paragraph id="8BF59E5EFF80FFECFF7365D2FA14FD07" blockId="4.[151,1436,743,863]" box="[151,1436,743,767]" pageId="4" pageNumber="43">
<emphasis id="B93E424CFF80FFECFF7365D2FE81FD07" bold="true" box="[151,265,743,767]" pageId="4" pageNumber="43">TABLE 1.</emphasis>
Morphological measurements of
<taxonomicName id="4C4AE5DDFF80FFECFD9765DDFC93FD07" box="[627,795,744,767]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="aurifera" status="sp. nov.">
<emphasis id="B93E424CFF80FFECFD9765DDFC93FD07" box="[627,795,744,767]" italics="true" pageId="4" pageNumber="43">Litoria aurifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DFF37FF80FFECFCC665D2FCFFFD07" box="[802,887,743,767]" pageId="4" pageNumber="43" rank="species">
<emphasis id="B93E424CFF80FFECFCC665D2FCFFFD07" bold="true" box="[802,887,743,767]" pageId="4" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C4AE5DDFF80FFECFC4F65DDFBD4FD07" box="[939,1116,744,767]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="meiriana">
<emphasis id="B93E424CFF80FFECFC4F65DDFBD4FD07" box="[939,1116,744,767]" italics="true" pageId="4" pageNumber="43">Litoria meiriana</emphasis>
</taxonomicName>
. Mean ± S.D. (range). SVL =
</paragraph>
</caption>
<paragraph id="8BF59E5EFF80FFECFF73643DFA14FCE7" blockId="4.[151,1436,743,863]" box="[151,1436,776,799]" pageId="4" pageNumber="43">snout-vent length; TL = tibia length; HW = head width; HL = head length; IO = interorbital span; IN = internarial span;</paragraph>
<paragraph id="8BF59E5EFF80FFECFF73641DFA14FCC7" blockId="4.[151,1436,743,863]" box="[151,1436,808,831]" pageId="4" pageNumber="43">ED = horizontal eye diameter; TD = horizontal tympanum diameter; ET = eye to tympanum; EN = distance between eye</paragraph>
<paragraph id="8BF59E5EFF80FFECFF73647DFF77FCA7" blockId="4.[151,1436,743,863]" box="[151,255,840,863]" pageId="4" pageNumber="43">and naris.</paragraph>
<paragraph id="8BF59E5EFF80FFECFF7B644FFCD7FA54" pageId="4" pageNumber="43">
<table id="F94A6CFEFF800017FF73644FFA14FA54" box="[151,1436,890,1452]" gridcols="4" gridrows="11" pageId="4" pageNumber="43">
<tr id="357A9C1CFF800017FF73644FFA14FC49" box="[151,1436,890,945]" gridrow="0" pageId="4" pageNumber="43">
<th id="76ABF560FF800017FF73644FFE8FFC49" box="[151,263,890,945]" gridcol="0" gridrow="0" pageId="4" pageNumber="43">Character</th>
<th id="76ABF560FF800017FE89644FFDC7FC49" box="[365,591,890,945]" gridcol="1" gridrow="0" pageId="4" pageNumber="43">
<taxonomicName id="4C4AE5DDFF80FFECFE89644FFD9DFC69" box="[365,533,890,913]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="aurifera">
<emphasis id="B93E424CFF80FFECFE89644FFD9DFC69" box="[365,533,890,913]" italics="true" pageId="4" pageNumber="43">Litoria aurifera</emphasis>
</taxonomicName>
N = 27
</th>
<th id="76ABF560FF800017FD41644FFC00FC49" box="[677,904,890,945]" gridcol="2" gridrow="0" pageId="4" pageNumber="43">
<taxonomicName id="4C4AE5DDFF80FFECFD41644FFCDEFC69" box="[677,854,890,913]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="meiriana">
<emphasis id="B93E424CFF80FFECFD41644FFCDEFC69" box="[677,854,890,913]" italics="true" pageId="4" pageNumber="43">Litoria meiriana</emphasis>
</taxonomicName>
N = 27
</th>
<th id="76ABF560FF800017FC04644FFA14FC49" box="[992,1436,890,945]" gridcol="3" gridrow="0" pageId="4" pageNumber="43">Statistical Results</th>
</tr>
<tr id="357A9C1CFF800017FF7364F0FA14FC19" box="[151,1436,965,993]" gridrow="1" pageId="4" pageNumber="43">
<th id="76ABF560FF800017FF7364F0FE8FFC19" box="[151,263,965,993]" gridcol="0" gridrow="1" pageId="4" pageNumber="43">SVL</th>
<td id="76ABF560FF800017FE8964F0FDC7FC19" box="[365,591,965,993]" gridcol="1" gridrow="1" pageId="4" pageNumber="43">19.9±1.0 (18.2–22.3)</td>
<td id="76ABF560FF800017FD4164F0FC00FC19" box="[677,904,965,993]" gridcol="2" gridrow="1" pageId="4" pageNumber="43">18.8±1.3 (17.1–22.4)</td>
<td id="76ABF560FF800017FC0464F0FA14FC19" box="[992,1436,965,993]" gridcol="3" gridrow="1" pageId="4" pageNumber="43">
F1,52 = 14.63,
<emphasis id="B93E424CFF80FFECFB8A64F0FBF6FC24" box="[1134,1150,965,988]" italics="true" pageId="4" pageNumber="43">P</emphasis>
&lt;0.001
</td>
</tr>
<tr id="357A9C1CFF800017FF7364C7FA14FBF6" box="[151,1436,1010,1038]" gridrow="2" pageId="4" pageNumber="43">
<th id="76ABF560FF800017FF7364C7FE8FFBF6" box="[151,263,1010,1038]" gridcol="0" gridrow="2" pageId="4" pageNumber="43">TL</th>
<td id="76ABF560FF800017FE8964C7FDC7FBF6" box="[365,591,1010,1038]" gridcol="1" gridrow="2" pageId="4" pageNumber="43">11.0±0.7 (10.0–12.7)</td>
<td id="76ABF560FF800017FD4164C7FC00FBF6" box="[677,904,1010,1038]" gridcol="2" gridrow="2" pageId="4" pageNumber="43">10.3±0.5 (9.3–11.8)</td>
<td id="76ABF560FF800017FC0464C7FA14FBF6" box="[992,1436,1010,1038]" gridcol="3" gridrow="2" pageId="4" pageNumber="43">
F1,52 = 16.00,
<emphasis id="B93E424CFF80FFECFB8A64C7FBF6FBF1" box="[1134,1150,1010,1033]" italics="true" pageId="4" pageNumber="43">P</emphasis>
&lt;0.001
</td>
</tr>
<tr id="357A9C1CFF800017FF736315FA14FBA6" box="[151,1436,1056,1118]" gridrow="3" pageId="4" pageNumber="43">
<th id="76ABF560FF800017FF736315FE8FFBA6" box="[151,263,1056,1118]" gridcol="0" gridrow="3" pageId="4" pageNumber="43">HW</th>
<td id="76ABF560FF800017FE896315FDC7FBA6" box="[365,591,1056,1118]" gridcol="1" gridrow="3" pageId="4" pageNumber="43">7.4±0.3 (6.9–7.9)</td>
<td id="76ABF560FF800017FD416315FC00FBA6" box="[677,904,1056,1118]" gridcol="2" gridrow="3" pageId="4" pageNumber="43">6.9±0.6 (5.5–8.1)</td>
<td id="76ABF560FF800017FC046315FA14FBA6" box="[992,1436,1056,1118]" gridcol="3" gridrow="3" pageId="4" pageNumber="43">
F1,51 = 7.27,
<emphasis id="B93E424CFF80FFECFBBA6315FBE6FBCF" box="[1118,1134,1056,1079]" italics="true" pageId="4" pageNumber="43">P</emphasis>
&lt;0.01 SVL: F1,51 = 4.53,
<emphasis id="B93E424CFF80FFECFB7A6377FB26FBA1" box="[1182,1198,1090,1113]" italics="true" pageId="4" pageNumber="43">P</emphasis>
= 0.038
</td>
</tr>
<tr id="357A9C1CFF800017FF736345FA14FB56" box="[151,1436,1136,1198]" gridrow="4" pageId="4" pageNumber="43">
<th id="76ABF560FF800017FF736345FE8FFB56" box="[151,263,1136,1198]" gridcol="0" gridrow="4" pageId="4" pageNumber="43">HL</th>
<td id="76ABF560FF800017FE896345FDC7FB56" box="[365,591,1136,1198]" gridcol="1" gridrow="4" pageId="4" pageNumber="43">6.9±0.5 (6.3–8.1)</td>
<td id="76ABF560FF800017FD416345FC00FB56" box="[677,904,1136,1198]" gridcol="2" gridrow="4" pageId="4" pageNumber="43">6.8±0.4 (5.6–7.7)</td>
<td id="76ABF560FF800017FC046345FA14FB56" box="[992,1436,1136,1198]" gridcol="3" gridrow="4" pageId="4" pageNumber="43">
F1,51 = 0.28,
<emphasis id="B93E424CFF80FFECFBBA6345FBE6FB7F" box="[1118,1134,1136,1159]" italics="true" pageId="4" pageNumber="43">P</emphasis>
&lt;0.598 SVL: F1,51 = 5.04,
<emphasis id="B93E424CFF80FFECFB7A63A7FB26FB51" box="[1182,1198,1170,1193]" italics="true" pageId="4" pageNumber="43">P</emphasis>
= 0.029
</td>
</tr>
<tr id="357A9C1CFF800017FF7363F5FA14FB2F" box="[151,1436,1216,1239]" gridrow="5" pageId="4" pageNumber="43" rowspan-3="1">
<th id="76ABF560FF800017FF7363F5FE8FFB2F" box="[151,263,1216,1239]" gridcol="0" gridrow="5" pageId="4" pageNumber="43">IO</th>
<td id="76ABF560FF800017FE8963F5FDC7FB2F" box="[365,591,1216,1239]" gridcol="1" gridrow="5" pageId="4" pageNumber="43">4.0±0.5 (2.1–4.7)</td>
<td id="76ABF560FF800017FD4163F5FC00FB2F" box="[677,904,1216,1239]" gridcol="2" gridrow="5" pageId="4" pageNumber="43">3.9±0.5 (2.0–4.3)</td>
</tr>
<tr id="357A9C1CFF800017FF7363DFFA14FAF9" box="[151,1436,1258,1281]" gridrow="6" pageId="4" pageNumber="43" rowspan-3="1">
<th id="76ABF560FF800017FF7363DFFE8FFAF9" box="[151,263,1258,1281]" gridcol="0" gridrow="6" pageId="4" pageNumber="43">IN</th>
<td id="76ABF560FF800017FE8963DFFDC7FAF9" box="[365,591,1258,1281]" gridcol="1" gridrow="6" pageId="4" pageNumber="43">1.5±0.2 (1.0–1.8)</td>
<td id="76ABF560FF800017FD4163DFFC00FAF9" box="[677,904,1258,1281]" gridcol="2" gridrow="6" pageId="4" pageNumber="43">1.4±0.1 (1.0–1.7)</td>
</tr>
<tr id="357A9C1CFF800017FF736220FA14FAD4" box="[151,1436,1301,1324]" gridrow="7" pageId="4" pageNumber="43" rowspan-3="1">
<th id="76ABF560FF800017FF736220FE8FFAD4" box="[151,263,1301,1324]" gridcol="0" gridrow="7" pageId="4" pageNumber="43">ED</th>
<td id="76ABF560FF800017FE896220FDC7FAD4" box="[365,591,1301,1324]" gridcol="1" gridrow="7" pageId="4" pageNumber="43">2.2±0.2 (2.0–2.6)</td>
<td id="76ABF560FF800017FD416220FC00FAD4" box="[677,904,1301,1324]" gridcol="2" gridrow="7" pageId="4" pageNumber="43">2.3±0.2 (1.9–2.7)</td>
</tr>
<tr id="357A9C1CFF800017FF736275FA14FAAF" box="[151,1436,1344,1367]" gridrow="8" pageId="4" pageNumber="43" rowspan-3="1">
<th id="76ABF560FF800017FF736275FE8FFAAF" box="[151,263,1344,1367]" gridcol="0" gridrow="8" pageId="4" pageNumber="43">TD</th>
<td id="76ABF560FF800017FE896275FDC7FAAF" box="[365,591,1344,1367]" gridcol="1" gridrow="8" pageId="4" pageNumber="43">1.7±0.1 (1.4–2.1)</td>
<td id="76ABF560FF800017FD416275FC00FAAF" box="[677,904,1344,1367]" gridcol="2" gridrow="8" pageId="4" pageNumber="43">1.6±0.1 (1.3–1.8)</td>
</tr>
<tr id="357A9C1CFF800017FF73625FFA14FA79" box="[151,1436,1386,1409]" gridrow="9" pageId="4" pageNumber="43" rowspan-3="1">
<th id="76ABF560FF800017FF73625FFE8FFA79" box="[151,263,1386,1409]" gridcol="0" gridrow="9" pageId="4" pageNumber="43">ET</th>
<td id="76ABF560FF800017FE89625FFDC7FA79" box="[365,591,1386,1409]" gridcol="1" gridrow="9" pageId="4" pageNumber="43">0.6±0.1 (0.5–0.7)</td>
<td id="76ABF560FF800017FD41625FFC00FA79" box="[677,904,1386,1409]" gridcol="2" gridrow="9" pageId="4" pageNumber="43">0.6±0.1 (0.3–0.9)</td>
</tr>
<tr id="357A9C1CFF800017FF7362A0FA14FA54" box="[151,1436,1429,1452]" gridrow="10" pageId="4" pageNumber="43" rowspan-3="1">
<th id="76ABF560FF800017FF7362A0FE8FFA54" box="[151,263,1429,1452]" gridcol="0" gridrow="10" pageId="4" pageNumber="43">EN</th>
<td id="76ABF560FF800017FE8962A0FDC7FA54" box="[365,591,1429,1452]" gridcol="1" gridrow="10" pageId="4" pageNumber="43">2.0±0.1 (1.6–2.5)</td>
<td id="76ABF560FF800017FD4162A0FC00FA54" box="[677,904,1429,1452]" gridcol="2" gridrow="10" pageId="4" pageNumber="43">2.0±0.2 (1.6–2.4)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BF59E5EFF80FFECFF2262DFFDCCF947" blockId="4.[151,1437,1514,1988]" pageId="4" pageNumber="43">
<emphasis id="B93E424CFF80FFECFF2262DFFEF3F9FC" bold="true" box="[198,379,1514,1540]" pageId="4" pageNumber="43">Colour in life:</emphasis>
Two specimens reared in captivity had an iridescent reddish tinge over mottling on the dorsum. Specimens photographed before collection at night in the field were reddish brown to almost translucent beige over the limbs, where faint darker patches were visible beneath (
<figureCitation id="137182DBFF80FFECFB976100FB54F9B7" box="[1139,1244,1589,1615]" captionStart="FIGURE 2" captionStartId="5.[151,255,1848,1872]" captionTargetBox="[240,1370,191,1833]" captionTargetId="figure@5.[212,1396,168,1857]" captionTargetPageId="5" captionText="FIGURE 2. Comparative adult frogs in life: Litoria meiriana and Litoria aurifera sp. nov. A = Litoria meiriana, active by day, Nawurlandja Rock, Kakadu National Park, Northern Territory. B = Litoria aurifera, at night in habitat, Prince Regent Nature Reserve, Western Australia. C = Litoria aurifera, holotype (WAM R 169913, female) dorsolateral view, during daytime, raised from tadpole in captivity. D = Litoria aurifera, holotype (WAM R 169913, female) dorsal view. E = Litoria aurifera, holotype (WAM R 169913, female) ventral view. F = Litoria aurifera, anterior view, at night in habitat, Prince Regent Nature Reserve, Western Australia. Bar represents 5 mm." httpUri="https://zenodo.org/record/196814/files/figure.png" pageId="4" pageNumber="43">Figure 2</figureCitation>
B). Some others were darker reddish-brown with black mottling over dorsum and limbs (M. Barrett pers. comm.). The throat of calling males was dusky grey. Ventrally, the subgular, pectoral and abdominal regions were white and the limbs faint purple (Pantone 5155M).
</paragraph>
<paragraph id="8BF59E5EFF80FFECFF2261FFFDA8F8F2" blockId="4.[151,1437,1514,1988]" pageId="4" pageNumber="43">
<emphasis id="B93E424CFF80FFECFF2261FFFE63F91C" bold="true" box="[198,491,1738,1764]" pageId="4" pageNumber="43">Colour in preservative:</emphasis>
All iridophore pigment is lost. Ventrally, the limbs fade to the same tone as the remainder of the ventral surfaces.
</paragraph>
</subSubSection>
<subSubSection id="C350CDD5FF80FFECFF226020FC8BF8AC" pageId="4" pageNumber="43" type="etymology">
<paragraph id="8BF59E5EFF80FFECFF226020FC8BF8AC" blockId="4.[151,1437,1514,1988]" pageId="4" pageNumber="43">
<emphasis id="B93E424CFF80FFECFF226020FEDBF8D7" bold="true" box="[198,339,1813,1839]" pageId="4" pageNumber="43">Etymology.</emphasis>
Derived from the latin
<emphasis id="B93E424CFF80FFECFD926020FD4DF8D6" box="[630,709,1813,1838]" italics="true" pageId="4" pageNumber="43">aureus</emphasis>
(gold) and
<emphasis id="B93E424CFF80FFECFCB16020FC0BF8D6" box="[853,899,1813,1838]" italics="true" pageId="4" pageNumber="43">fero</emphasis>
(to carry or bear) alluding to the distinctive gold patches characterising the colour of the tadpole.
</paragraph>
</subSubSection>
<subSubSection id="C350CDD5FF80FFEEFF22606AFD91FEBF" lastPageId="6" lastPageNumber="45" pageId="4" pageNumber="43" type="distribution">
<paragraph id="8BF59E5EFF80FFEEFF22606AFD91FEBF" blockId="4.[151,1437,1514,1988]" lastBlockId="6.[151,1436,152,327]" lastPageId="6" lastPageNumber="45" pageId="4" pageNumber="43">
<emphasis id="B93E424CFF80FFECFF22606AFD8BF881" bold="true" box="[198,515,1887,1913]" pageId="4" pageNumber="43">Distribution and habitat.</emphasis>
<figureCitation id="137182DBFF80FFECFDE96055FDFBF882" box="[525,627,1888,1914]" captionStart="FIGURE 3" captionStartId="6.[151,255,1073,1097]" captionTargetBox="[396,1176,373,1056]" captionTargetId="figure@6.[396,1191,367,1067]" captionTargetPageId="6" captionText="FIGURE 3. Distributions of Litoria aurifera sp. nov. and Litoria meiriana in north-western Australia." httpUri="https://zenodo.org/record/196815/files/figure.png" pageId="4" pageNumber="43">Figure 3</figureCitation>
shows the distributions of
<taxonomicName id="4C4AE5DDFF80FFECFC586055FBB7F881" box="[956,1087,1888,1913]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="aurifera">
<emphasis id="B93E424CFF80FFECFC586055FBB7F881" box="[956,1087,1888,1913]" italics="true" pageId="4" pageNumber="43">L. aurifera</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4AE5DDFF80FFECFB986055FA80F881" box="[1148,1288,1888,1913]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="meiriana">
<emphasis id="B93E424CFF80FFECFB986055FA80F881" box="[1148,1288,1888,1913]" italics="true" pageId="4" pageNumber="43">L. meiriana</emphasis>
</taxonomicName>
. Adults and tadpoles of
<taxonomicName id="4C4AE5DDFF80FFECFEC760B0FE2CF866" box="[291,420,1925,1950]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="4" pageNumber="43" phylum="Chordata" rank="species" species="aurifera">
<emphasis id="B93E424CFF80FFECFEC760B0FE2CF866" box="[291,420,1925,1950]" italics="true" pageId="4" pageNumber="43">L. aurifera</emphasis>
</taxonomicName>
were collected from creeks where tadpoles of the new species were found, across an altitudinal range of about
<quantity id="4CB233BBFF80FFECFE58609FFDA2F83C" box="[444,554,1962,1988]" metricMagnitude="2" metricUnit="m" metricValue="2.27" metricValueMax="4.08" metricValueMin="0.45999999999999996" pageId="4" pageNumber="43" unit="m" value="227.0" valueMax="408.0" valueMin="46.0">46–408m</quantity>
within about
<quantity id="4CB233BBFF80FFECFD22609FFC95F83C" box="[710,797,1962,1988]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="43" unit="km" value="100.0">100 km</quantity>
S. of the Prince Regent River. All sites were first order, fast flowing creeks which collect water from rain on sandstone rock escarpments. Tadpoles were collected from pools within the creeks, but were never found in larger creeks or water that carries silt from flooding (M. Barrett, pers. comm.). Water depth in the pools ranged from a few cm to
<quantity id="4CB233BBFF82FFEEFBFE67D7FBC1FF04" box="[1050,1097,226,252]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="45" unit="m" value="1.0">1 m</quantity>
and substrate was irregular sandstone rock. In some sites the yellow sandstone rock was covered with patches of black algae, on which tadpoles were well camouflaged.
</paragraph>
</subSubSection>
</treatment>
</document>
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