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<document id="ADF120A9C45B8D80523542FF3C465D9F" ID-CLB-Dataset="20247" ID-DOI="10.1093/zoolinnean/zlab014" ID-GBIF-Dataset="5cbb504f-e207-44fa-ac62-807a6de2f2db" ID-ISSN="0024-4082" ID-Zenodo-Dep="6354284" ID-ZooBank="3E370CBF-7DD1-4E73-BC37-81BF12EDFEED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1647256456928" checkinUser="plazi" docAuthor="Klautau, Michelle, Lopes, Matheus Vieira, Tavares, Gabriela &amp; Pérez, Thierry" docDate="2022" docId="03E08916FFEB2D10FECCFD0BFA954446" docLanguage="en" docName="zlab014.pdf" docOrigin="Zoological Journal of the Linnean Society 194" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Leucandra ornata Klautau &amp; Lopes &amp; Tavares &amp; Pérez 2022, SP. NOV." docType="treatment" docUuid="2F2F2C38-A216-4B1F-8A04-7CAA93B8F0EC" docUuidSource="ZooBank" docVersion="7" lastPageNumber="710" masterDocId="FFD9F16EFFCC2D37FFEAFFE6FF9C4053" masterDocTitle="Integrative taxonomy of calcareous sponges (Porifera: Calcarea) from Réunion Island, Indian Ocean" masterLastPageNumber="725" masterPageNumber="671" pageNumber="710" updateTime="1699317944552" updateUser="ExternalLinkService">
<mods:mods id="983603217ED25FBAB6409B62DB0CDC73" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5FFF8B6B17243B23AE3D1B0830DB4F12">
<mods:title id="1E368948FCC7A82C4F669F404D5A2288">Integrative taxonomy of calcareous sponges (Porifera: Calcarea) from Réunion Island, Indian Ocean</mods:title>
</mods:titleInfo>
<mods:name id="92F7C899C52EEC5C9AEF92C41BFCE05F" type="personal">
<mods:role id="67176F543AFB8DF3DC8E0C1FC0953876">
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<mods:namePart id="520D19D29992F6C8952EC871DCAC4B6B">Klautau, Michelle</mods:namePart>
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<mods:name id="D7579BC1C88A29FC22A8911947B0DD77" type="personal">
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<mods:roleTerm id="63DF3BA9B2A3CB3309D7899D25E0EE15">Author</mods:roleTerm>
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<mods:namePart id="FE0857BD371509F584E5DB8BB90CAB33">Lopes, Matheus Vieira</mods:namePart>
</mods:name>
<mods:name id="915AD8D115E19CA0A4283256865C1880" type="personal">
<mods:role id="A9E8BD90A5CB82407B691547108EBAB9">
<mods:roleTerm id="E8C08C350F234DEC7F99AA32890308C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F85C64F3AD8B39A2551A386D6ADE9CF">Tavares, Gabriela</mods:namePart>
</mods:name>
<mods:name id="9F09551BB26A3296D7C5D179A67C73CA" type="personal">
<mods:role id="C26346E6EB079E8CA7D69A4D12AF8F69">
<mods:roleTerm id="AE5326ECAE6CBB49A51F62F5EAD08ED3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B2E712B7544FBAA26D241E836A40D2D1">Pérez, Thierry</mods:namePart>
</mods:name>
<mods:typeOfResource id="BBA670EE47742F241F02D01F377D51E3">text</mods:typeOfResource>
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<mods:title id="ABD6142DE3E5E9A83D6325856F2364BD">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="7E3640EE99B3A472A936BB83E4318423">
<mods:date id="0365300057AF65343B9023F53CE68E49">2022</mods:date>
<mods:detail id="AD0183B212E10B53121E5AF4E2710959" type="pubDate">
<mods:number id="3684E10A5318BAA234C579332E9A4D4B">2021-05-06</mods:number>
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<mods:detail id="C0FC6C6D11982A73912A7672EE404631" type="volume">
<mods:number id="A4E06CFD5255ABA3A9654C4C678D043F">194</mods:number>
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<mods:classification id="9B4D679B4007861CB30ED1705B64C89A">journal article</mods:classification>
<mods:identifier id="BB957EA3C7744EA7030ACCF94758CC96" type="CLB-Dataset">20247</mods:identifier>
<mods:identifier id="5323839F4AEBE7FBABA19E8DB9DF2A0F" type="DOI">10.1093/zoolinnean/zlab014</mods:identifier>
<mods:identifier id="CC9379BFEF9B59D2FF8AB0F0DEECBC42" type="GBIF-Dataset">5cbb504f-e207-44fa-ac62-807a6de2f2db</mods:identifier>
<mods:identifier id="5FC671275AC0E94C415FAC8FA56ED933" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="94C06F844230DAAB30E0257970CB8D93" type="Zenodo-Dep">6354284</mods:identifier>
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<treatment id="03E08916FFEB2D10FECCFD0BFA954446" ID-DOI="http://doi.org/10.5281/zenodo.6352004" ID-GBIF-Taxon="194018764" ID-Zenodo-Dep="6352004" LSID="urn:lsid:zoobank.org:act:2F2F2C38-A216-4B1F-8A04-7CAA93B8F0EC" httpUri="http://treatment.plazi.org/id/03E08916FFEB2D10FECCFD0BFA954446" lastPageNumber="710" pageId="39" pageNumber="710">
<subSubSection id="C3536B8BFFEB2D10FECCFD0BFD1B4356" box="[294,647,749,773]" pageId="39" pageNumber="710" type="nomenclature">
<paragraph id="8BF63800FFEB2D10FECCFD0BFD1B4356" blockId="39.[294,647,749,773]" box="[294,647,749,773]" pageId="39" pageNumber="710">
<heading id="D0BE8F6CFFEB2D10FECCFD0BFD1B4356" box="[294,647,749,773]" centered="true" fontSize="9" level="2" pageId="39" pageNumber="710" reason="2">
<taxonomicName id="4C494383FFEB2D10FECCFD0BFD814350" ID-CoL="8H7FR" authority="Klautau &amp; Lopes &amp; Tavares &amp; Pérez, 2022" authorityName="Klautau &amp; Lopes &amp; Tavares &amp; Pérez" authorityYear="2022" box="[294,541,749,772]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="39" pageNumber="714" phylum="Porifera" rank="species" species="ornata" status="sp. nov.">LEUCANDRA ORNATA</taxonomicName>
<emphasis id="B93DE412FFEB2D10FDCEFD16FD1B4356" bold="true" box="[548,647,749,773]" pageId="39" pageNumber="710">
<taxonomicNameLabel id="A20E5969FFEB2D10FDCEFD16FD1B4356" box="[548,647,749,773]" pageId="39" pageNumber="710" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3536B8BFFEB2D10FEBAFCF5FA954446" pageId="39" pageNumber="710" type="description">
<paragraph id="8BF63800FFEB2D10FEBAFCF5FDC1437F" blockId="39.[336,605,787,812]" box="[336,605,787,812]" pageId="39" pageNumber="710">
(
<figureCitation id="13722485FFEB2D10FEB2FCF5FE47437F" box="[344,475,787,812]" captionStart-0="Figure 22" captionStart-1="Figure 23" captionStart-2="Figure 24" captionStart-3="Figure 25" captionStartId-0="39.[163,241,1876,1898]" captionStartId-1="40.[145,224,1675,1697]" captionStartId-2="41.[163,242,1675,1697]" captionStartId-3="42.[145,227,1618,1640]" captionTargetBox-0="[243,1363,1376,1837]" captionTargetBox-1="[202,1368,195,1635]" captionTargetBox-2="[220,1386,195,1635]" captionTargetBox-3="[225,1345,195,1579]" captionTargetId-0="figure-531@39.[243,1363,1376,1837]" captionTargetId-1="figure-24@40.[202,1368,195,1635]" captionTargetId-2="figure-24@41.[220,1386,195,1635]" captionTargetId-3="figure-24@42.[225,1345,195,1579]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionTargetPageId-3="42" captionText-0="Figure 22. Leucandra ornata (holotype; UFRJPOR 8928). A, specimen in vivo. B, fixed specimen." captionText-1="Figure 23. Leucandra ornata (holotype; UFRJPOR 8928). A, oscular margin. B, cross-section. C, tangential section of the cortex. D, detail of the cortex showing diactines II organised in bouquet. E, detail of the choanosome showing a tetractine of the canals. F, cross-section of the atrium (microdiactines in the inset). cx = cortex, at = atrium." captionText-2="Figure 24. Leucandra ornata (holotype; UFRJPOR 8928). SEM: A, entire specimen. B, tangential section of the cortex. C, detail of the cortex showing a microdiactine (arrow) and a cortical triactine with elevated centre (asterisk). D, tangential section of the atrium. E, detail of an exhalant canal with microdiactines. F, detail of the microdiactines with spines. osc = osculum." captionText-3="Figure 25. Leucandra ornata (holotype; UFRJPOR 8928). A, diactines. B, microdiactines. C, cortical triactines. D, choanosomal large triactine. E, canal triactine. F, tetractine of the canals. G, atrial tetractines. H, tetractine of the osculum." figureDoi-0="http://doi.org/10.5281/zenodo.6354364" figureDoi-1="http://doi.org/10.5281/zenodo.6354366" figureDoi-2="http://doi.org/10.5281/zenodo.6354372" figureDoi-3="http://doi.org/10.5281/zenodo.6354374" httpUri-0="https://zenodo.org/record/6354364/files/figure.png" httpUri-1="https://zenodo.org/record/6354366/files/figure.png" httpUri-2="https://zenodo.org/record/6354372/files/figure.png" httpUri-3="https://zenodo.org/record/6354374/files/figure.png" pageId="39" pageNumber="710">FIGS 22–25</figureCitation>
;
<tableCitation id="C6CB0DBBFFEB2D10FE02FCF5FDC9437F" box="[488,597,787,812]" captionStart="Table 11" captionStartId="43.[143,207,665,687]" captionTargetPageId="43" captionText="Table 11. Spicule measurements of Leucandra ornata and the original measurements of L. riojai. H = holotype" pageId="39" pageNumber="710">TABLE 11</tableCitation>
)
</paragraph>
<paragraph id="8BF63800FFEB2D10FF49FCDBFC954323" blockId="39.[163,779,828,880]" pageId="39" pageNumber="710">
<emphasis id="B93DE412FFEB2D10FF49FCDBFE6C4301" box="[163,496,828,850]" italics="true" pageId="39" pageNumber="710">Z o o b a n k r e g i s t r a t i o n:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="FFEF02D5FFEB2D10FF18FCBCFC994323" box="[242,773,858,880]" pageId="39" pageNumber="710">2F2F2C38-A216-4B1F-8A04-7CAA93B8F0EC</uuid>
.
</paragraph>
<paragraph id="8BF63800FFEB2D10FF49FC72FE7C44D3" blockId="39.[163,778,916,1152]" pageId="39" pageNumber="710">
<emphasis id="B93DE412FFEB2D10FF49FC72FE3F43FA" box="[163,419,916,937]" italics="true" pageId="39" pageNumber="710">
Diagnosis:
<taxonomicName id="4C494383FFEB2D10FECFFC72FE3F43FA" authorityName="Haeckel" authorityYear="1872" box="[293,419,916,937]" class="Calcarea" family="Grantiidae" genus="Leucandra" kingdom="Animalia" order="Leucosolenida" pageId="39" pageNumber="710" phylum="Porifera" rank="genus">Leucandra</taxonomicName>
</emphasis>
with spined microdiactines in the cortical and in the atrial skeletons. The cortical microdiactines are frequently organized forming bouquets. Large cortical diactines are also present, along with cortical triactines. The choanosome contains regular triactines and sagittal triactines and tetractines. The atrial skeleton has microdiactines with spines and tetractines.
</paragraph>
<paragraph id="8BF63800FFEB2D10FF49FB47FE3E44A7" blockId="39.[163,778,1185,1268]" pageId="39" pageNumber="710">
<emphasis id="B93DE412FFEB2D10FF49FB47FEB344E5" box="[163,303,1185,1206]" italics="true" pageId="39" pageNumber="710">Etymology:</emphasis>
From the Latin
<emphasis id="B93DE412FFEB2D10FDF8FB44FDE244E4" box="[530,638,1186,1207]" italics="true" pageId="39" pageNumber="710">ornatum</emphasis>
, ornate or decorated, for the presence of many diactines on the surface of this species.
</paragraph>
<paragraph id="8BF63800FFEB2D10FCD1FF20FC4040AA" blockId="39.[827,1441,197,250]" pageId="39" pageNumber="710">
<materialsCitation id="3B21325DFFEB2D10FCD1FF20FC4440AA" ID-GBIF-Occurrence="3704837330" country="Réunion" location="Cap La Houssaye Reef" municipality="Indian Ocean" pageId="39" pageNumber="710" specimenCount="1" typeStatus="holotype">
<emphasis id="B93DE412FFEB2D10FCD1FF20FC414089" box="[827,989,197,219]" italics="true" pageId="39" pageNumber="710">
<typeStatus id="54F286A2FFEB2D10FCD1FF20FCEF4088" box="[827,883,198,219]" pageId="39" pageNumber="710">Type</typeStatus>
locality:
</emphasis>
<location id="8E966EDBFFEB2D10FC0DFF23FB634088" LSID="urn:lsid:plazi:treatment:03E08916FFEB2D10FECCFD0BFA954446:8E966EDBFFEB2D10FC0DFF23FB634088" box="[999,1279,197,219]" country="Réunion" municipality="Indian Ocean" name="Cap La Houssaye Reef" pageId="39" pageNumber="710">Cap La Houssaye Reef</location>
,
<collectingCountry id="F35E7890FFEB2D10FAE4FF20FA024088" box="[1294,1438,197,219]" name="Réunion" pageId="39" pageNumber="710">La Réunion</collectingCountry>
,
<collectingMunicipality id="6B92A27AFFEB2D10FCD1FF02FC4440AA" box="[827,984,228,250]" pageId="39" pageNumber="710">Indian Ocean</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF63800FFEB2D10FCD1FEC5FB6141C7" blockId="39.[827,1442,290,404]" pageId="39" pageNumber="710">
<emphasis id="B93DE412FFEB2D10FCD1FEC5FB95416B" box="[827,1033,290,312]" italics="true" pageId="39" pageNumber="710">Ty p e m a t e r i a l:</emphasis>
H o l o t y p e – U F R J P O
<materialsCitation id="3B21325DFFEB2D10FAAAFEC5FA3E4125" ID-GBIF-Occurrence="3704837326" collectionCode="R, CAH" collectorName="Indian Ocean" country="Réunion" location="Cap La Houssaye Reef" pageId="39" pageNumber="710" specimenCode="R 8, CAH2" specimenCount="1">
<specimenCode id="DBEF907BFFEB2D10FAAAFEC5FAED416B" box="[1344,1393,291,312]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="39" pageNumber="710">R 8</specimenCode>
9 2 8 (= 171108-
<specimenCode id="DBEF907BFFEB2D10FC57FEA7FB964105" box="[957,1034,321,342]" collectionCode="CAH" country="Zimbabwe" lsid="urn:lsid:biocol.org:col:14678" name="University of Zimbabwe" pageId="39" pageNumber="710" type="Herbarium">CAH2</specimenCode>
-TP2, Photos: TP5973–5975).
<location id="8E966EDBFFEB2D10FA99FEA7FB8A4125" LSID="urn:lsid:plazi:treatment:03E08916FFEB2D10FECCFD0BFA954446:8E966EDBFFEB2D10FA99FEA7FB8A4125" country="Réunion" name="Cap La Houssaye Reef" pageId="39" pageNumber="710">Cap La Houssaye Reef</location>
,
<collectingCountry id="F35E7890FFEB2D10FBC9FE86FB2E4125" box="[1059,1202,352,374]" name="Réunion" pageId="39" pageNumber="710">La Réunion</collectingCountry>
,
<collectorName id="26BC5DD6FFEB2D10FB2AFE86FAFA4126" box="[1216,1382,352,374]" pageId="39" pageNumber="710">Indian Ocean</collectorName>
, coll.
</materialsCitation>
<materialsCitation id="3B21325DFFEB2D10FCD1FE99FB6641C7" ID-GBIF-Occurrence="3704837331" box="[827,1274,382,404]" collectingDate="2017-11-08" collectionCode="T" country="Réunion" pageId="39" pageNumber="710" specimenCount="1">
<collectionCode id="ED58A0C5FFEB2D10FCD1FE99FCD041C7" box="[827,844,383,404]" name="Tavera, Department of Geology and Geophysics" pageId="39" pageNumber="710">T</collectionCode>
. Pérez,
<date id="FFF71EC0FFEB2D10FC48FE99FBF741C7" box="[930,1131,382,404]" pageId="39" pageNumber="710" value="2017-11-08">
<collectingDate id="EFB3E728FFEB2D10FC48FE99FBF741C7" box="[930,1131,382,404]" pageId="39" pageNumber="710" value="2017-11-08">8 November 2017</collectingDate>
</date>
, 10 m depth
</materialsCitation>
.
</paragraph>
<paragraph id="8BF63800FFEB2D10FCD1FE5EFC2D41BF" blockId="39.[827,1441,440,492]" pageId="39" pageNumber="710">
<emphasis id="B93DE412FFEB2D10FCD1FE5EFC08419E" box="[827,916,440,461]" italics="true" pageId="39" pageNumber="710">Colour:</emphasis>
White alive (
<figureCitation id="13722485FFEB2D10FBAFFE5EFB32419E" box="[1093,1198,440,462]" captionStart="Figure 22" captionStartId="39.[163,241,1876,1898]" captionTargetBox="[243,1363,1376,1837]" captionTargetId="figure-531@39.[243,1363,1376,1837]" captionTargetPageId="39" captionText="Figure 22. Leucandra ornata (holotype; UFRJPOR 8928). A, specimen in vivo. B, fixed specimen." figureDoi="http://doi.org/10.5281/zenodo.6354364" httpUri="https://zenodo.org/record/6354364/files/figure.png" pageId="39" pageNumber="710">Fig. 22A</figureCitation>
) and after fixation (
<figureCitation id="13722485FFEB2D10FCA9FE30FC3D41BF" box="[835,929,470,492]" captionStart="Figure 22" captionStartId="39.[163,241,1876,1898]" captionTargetBox="[243,1363,1376,1837]" captionTargetId="figure-531@39.[243,1363,1376,1837]" captionTargetPageId="39" captionText="Figure 22. Leucandra ornata (holotype; UFRJPOR 8928). A, specimen in vivo. B, fixed specimen." figureDoi="http://doi.org/10.5281/zenodo.6354364" httpUri="https://zenodo.org/record/6354364/files/figure.png" pageId="39" pageNumber="710">Fig. 22B</figureCitation>
).
</paragraph>
<paragraph id="8BF63800FFEB2D10FCD1FDF3FA954446" blockId="39.[827,1444,533,1046]" pageId="39" pageNumber="710">
<emphasis id="B93DE412FFEB2D10FCD1FDF3FBF34278" box="[827,1135,533,555]" italics="true" pageId="39" pageNumber="710">Morphology and anatomy:</emphasis>
Sponge vase-shaped with apical osculum surrounded by a crown of trichoxeas (
<figureCitation id="13722485FFEB2D10FCA9FDB4FC59423B" box="[835,965,594,616]" captionStart="Figure 22" captionStartId="39.[163,241,1876,1898]" captionTargetBox="[243,1363,1376,1837]" captionTargetId="figure-531@39.[243,1363,1376,1837]" captionTargetPageId="39" captionText="Figure 22. Leucandra ornata (holotype; UFRJPOR 8928). A, specimen in vivo. B, fixed specimen." figureDoi="http://doi.org/10.5281/zenodo.6354364" httpUri="https://zenodo.org/record/6354364/files/figure.png" pageId="39" pageNumber="710">Figs 22A, B</figureCitation>
,
<figureCitation id="13722485FFEB2D10FC3BFDB5FC62423B" box="[977,1022,595,616]" captionStart="Figure 24" captionStartId="41.[163,242,1675,1697]" captionTargetBox="[220,1386,195,1635]" captionTargetId="figure-24@41.[220,1386,195,1635]" captionTargetPageId="41" captionText="Figure 24. Leucandra ornata (holotype; UFRJPOR 8928). SEM: A, entire specimen. B, tangential section of the cortex. C, detail of the cortex showing a microdiactine (arrow) and a cortical triactine with elevated centre (asterisk). D, tangential section of the atrium. E, detail of an exhalant canal with microdiactines. F, detail of the microdiactines with spines. osc = osculum." figureDoi="http://doi.org/10.5281/zenodo.6354372" httpUri="https://zenodo.org/record/6354372/files/figure.png" pageId="39" pageNumber="710">24A</figureCitation>
) supported by sagittal triactines and tetractines (
<figureCitation id="13722485FFEB2D10FC29FD97FBBC42D5" box="[963,1056,625,647]" captionStart="Figure 23" captionStartId="40.[145,224,1675,1697]" captionTargetBox="[202,1368,195,1635]" captionTargetId="figure-24@40.[202,1368,195,1635]" captionTargetPageId="40" captionText="Figure 23. Leucandra ornata (holotype; UFRJPOR 8928). A, oscular margin. B, cross-section. C, tangential section of the cortex. D, detail of the cortex showing diactines II organised in bouquet. E, detail of the choanosome showing a tetractine of the canals. F, cross-section of the atrium (microdiactines in the inset). cx = cortex, at = atrium." figureDoi="http://doi.org/10.5281/zenodo.6354366" httpUri="https://zenodo.org/record/6354366/files/figure.png" pageId="39" pageNumber="710">Fig. 23A</figureCitation>
). Surface hispid because of several large diactines protruding through the surface (
<figureCitation id="13722485FFEB2D10FA9BFD76FC1A4297" captionStart="Figure 22" captionStartId="39.[163,241,1876,1898]" captionTargetBox="[243,1363,1376,1837]" captionTargetId="figure-531@39.[243,1363,1376,1837]" captionTargetPageId="39" captionText="Figure 22. Leucandra ornata (holotype; UFRJPOR 8928). A, specimen in vivo. B, fixed specimen." figureDoi="http://doi.org/10.5281/zenodo.6354364" httpUri="https://zenodo.org/record/6354364/files/figure.png" pageId="39" pageNumber="710">Figs 22A, B</figureCitation>
,
<figureCitation id="13722485FFEB2D10FC79FD49FC234297" box="[915,959,687,708]" captionStart="Figure 23" captionStartId="40.[145,224,1675,1697]" captionTargetBox="[202,1368,195,1635]" captionTargetId="figure-24@40.[202,1368,195,1635]" captionTargetPageId="40" captionText="Figure 23. Leucandra ornata (holotype; UFRJPOR 8928). A, oscular margin. B, cross-section. C, tangential section of the cortex. D, detail of the cortex showing diactines II organised in bouquet. E, detail of the choanosome showing a tetractine of the canals. F, cross-section of the atrium (microdiactines in the inset). cx = cortex, at = atrium." figureDoi="http://doi.org/10.5281/zenodo.6354366" httpUri="https://zenodo.org/record/6354366/files/figure.png" pageId="39" pageNumber="710">23C</figureCitation>
,
<figureCitation id="13722485FFEB2D10FC26FD49FC654297" box="[972,1017,687,708]" captionStart="Figure 24" captionStartId="41.[163,242,1675,1697]" captionTargetBox="[220,1386,195,1635]" captionTargetId="figure-24@41.[220,1386,195,1635]" captionTargetPageId="41" captionText="Figure 24. Leucandra ornata (holotype; UFRJPOR 8928). SEM: A, entire specimen. B, tangential section of the cortex. C, detail of the cortex showing a microdiactine (arrow) and a cortical triactine with elevated centre (asterisk). D, tangential section of the atrium. E, detail of an exhalant canal with microdiactines. F, detail of the microdiactines with spines. osc = osculum." figureDoi="http://doi.org/10.5281/zenodo.6354372" httpUri="https://zenodo.org/record/6354372/files/figure.png" pageId="39" pageNumber="710">24B</figureCitation>
). The aquiferous system is leuconoid. The cortical skeleton is composed of tangential triactines, large diactines and microdiactines protruding through the surface (
<figureCitation id="13722485FFEB2D10FB25FCECFAC24373" box="[1231,1374,778,800]" captionStart="Figure 22" captionStartId="39.[163,241,1876,1898]" captionTargetBox="[243,1363,1376,1837]" captionTargetId="figure-531@39.[243,1363,1376,1837]" captionTargetPageId="39" captionText="Figure 22. Leucandra ornata (holotype; UFRJPOR 8928). A, specimen in vivo. B, fixed specimen." figureDoi="http://doi.org/10.5281/zenodo.6354364" httpUri="https://zenodo.org/record/6354364/files/figure.png" pageId="39" pageNumber="710">Figs 22A, B</figureCitation>
,
<figureCitation id="13722485FFEB2D10FA84FCECFCD2436D" captionStart="Figure 23" captionStartId="40.[145,224,1675,1697]" captionTargetBox="[202,1368,195,1635]" captionTargetId="figure-24@40.[202,1368,195,1635]" captionTargetPageId="40" captionText="Figure 23. Leucandra ornata (holotype; UFRJPOR 8928). A, oscular margin. B, cross-section. C, tangential section of the cortex. D, detail of the cortex showing diactines II organised in bouquet. E, detail of the choanosome showing a tetractine of the canals. F, cross-section of the atrium (microdiactines in the inset). cx = cortex, at = atrium." figureDoi="http://doi.org/10.5281/zenodo.6354366" httpUri="https://zenodo.org/record/6354366/files/figure.png" pageId="39" pageNumber="710">23C, D</figureCitation>
,
<figureCitation id="13722485FFEB2D10FCB7FCCFFC17436D" box="[861,907,809,830]" captionStart="Figure 24" captionStartId="41.[163,242,1675,1697]" captionTargetBox="[220,1386,195,1635]" captionTargetId="figure-24@41.[220,1386,195,1635]" captionTargetPageId="41" captionText="Figure 24. Leucandra ornata (holotype; UFRJPOR 8928). SEM: A, entire specimen. B, tangential section of the cortex. C, detail of the cortex showing a microdiactine (arrow) and a cortical triactine with elevated centre (asterisk). D, tangential section of the atrium. E, detail of an exhalant canal with microdiactines. F, detail of the microdiactines with spines. osc = osculum." figureDoi="http://doi.org/10.5281/zenodo.6354372" httpUri="https://zenodo.org/record/6354372/files/figure.png" pageId="39" pageNumber="710">24C</figureCitation>
). The cortical microdiactines form bouquets or lie tangentially to the cortex (
<figureCitation id="13722485FFEB2D10FB51FCAEFAB8430E" box="[1211,1316,840,862]" captionStart="Figure 23" captionStartId="40.[145,224,1675,1697]" captionTargetBox="[202,1368,195,1635]" captionTargetId="figure-24@40.[202,1368,195,1635]" captionTargetPageId="40" captionText="Figure 23. Leucandra ornata (holotype; UFRJPOR 8928). A, oscular margin. B, cross-section. C, tangential section of the cortex. D, detail of the cortex showing diactines II organised in bouquet. E, detail of the choanosome showing a tetractine of the canals. F, cross-section of the atrium (microdiactines in the inset). cx = cortex, at = atrium." figureDoi="http://doi.org/10.5281/zenodo.6354366" httpUri="https://zenodo.org/record/6354366/files/figure.png" pageId="39" pageNumber="710">Figs 23D</figureCitation>
,
<figureCitation id="13722485FFEB2D10FAD8FCAEFAFC430E" box="[1330,1376,840,861]" captionStart="Figure 24" captionStartId="41.[163,242,1675,1697]" captionTargetBox="[220,1386,195,1635]" captionTargetId="figure-24@41.[220,1386,195,1635]" captionTargetPageId="41" captionText="Figure 24. Leucandra ornata (holotype; UFRJPOR 8928). SEM: A, entire specimen. B, tangential section of the cortex. C, detail of the cortex showing a microdiactine (arrow) and a cortical triactine with elevated centre (asterisk). D, tangential section of the atrium. E, detail of an exhalant canal with microdiactines. F, detail of the microdiactines with spines. osc = osculum." figureDoi="http://doi.org/10.5281/zenodo.6354372" httpUri="https://zenodo.org/record/6354372/files/figure.png" pageId="39" pageNumber="710">24C</figureCitation>
). The choanosome contains large and small triactines. These small triactines are found spread in the choanosome or surrounding the canals, although tetractines are the main spicules in the canals (
<figureCitation id="13722485FFEB2D10FB5DFC24FA844384" box="[1207,1304,962,984]" captionStart="Figure 23" captionStartId="40.[145,224,1675,1697]" captionTargetBox="[202,1368,195,1635]" captionTargetId="figure-24@40.[202,1368,195,1635]" captionTargetPageId="40" captionText="Figure 23. Leucandra ornata (holotype; UFRJPOR 8928). A, oscular margin. B, cross-section. C, tangential section of the cortex. D, detail of the cortex showing diactines II organised in bouquet. E, detail of the choanosome showing a tetractine of the canals. F, cross-section of the atrium (microdiactines in the inset). cx = cortex, at = atrium." figureDoi="http://doi.org/10.5281/zenodo.6354366" httpUri="https://zenodo.org/record/6354366/files/figure.png" pageId="39" pageNumber="710">Fig. 23E</figureCitation>
). The atrial skeleton is composed of tetractines and microdiactines tangential to the atrium (
<figureCitation id="13722485FFEB2D10FBB4FBE6FB224446" box="[1118,1214,1024,1046]" captionStart="Figure 23" captionStartId="40.[145,224,1675,1697]" captionTargetBox="[202,1368,195,1635]" captionTargetId="figure-24@40.[202,1368,195,1635]" captionTargetPageId="40" captionText="Figure 23. Leucandra ornata (holotype; UFRJPOR 8928). A, oscular margin. B, cross-section. C, tangential section of the cortex. D, detail of the cortex showing diactines II organised in bouquet. E, detail of the choanosome showing a tetractine of the canals. F, cross-section of the atrium (microdiactines in the inset). cx = cortex, at = atrium." figureDoi="http://doi.org/10.5281/zenodo.6354366" httpUri="https://zenodo.org/record/6354366/files/figure.png" pageId="39" pageNumber="710">Figs 23F</figureCitation>
,
<figureCitation id="13722485FFEB2D10FB21FBE6FB664446" box="[1227,1274,1024,1045]" captionStart="Figure 24" captionStartId="41.[163,242,1675,1697]" captionTargetBox="[220,1386,195,1635]" captionTargetId="figure-24@41.[220,1386,195,1635]" captionTargetPageId="41" captionText="Figure 24. Leucandra ornata (holotype; UFRJPOR 8928). SEM: A, entire specimen. B, tangential section of the cortex. C, detail of the cortex showing a microdiactine (arrow) and a cortical triactine with elevated centre (asterisk). D, tangential section of the atrium. E, detail of an exhalant canal with microdiactines. F, detail of the microdiactines with spines. osc = osculum." figureDoi="http://doi.org/10.5281/zenodo.6354372" httpUri="https://zenodo.org/record/6354372/files/figure.png" pageId="39" pageNumber="710">24D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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