467 lines
75 KiB
XML
467 lines
75 KiB
XML
<document id="18D0AFA925AE8708CCD1A19ED51D81E9" ID-DOI="10.11646/zootaxa.3904.2.8" ID-GBIF-Dataset="594d467a-990e-4bf2-9bce-049b05e5748c" ID-ISSN="1175-5326" ID-Zenodo-Dep="244764" ID-ZooBank="FD269219-5572-44FB-8D42-6AE7A4CF3606" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460887222271" checkinUser="plazi" docAuthor="Malamel, Jobi J., Sankaran, Pradeep M., Joseph, Mathew M. & Sebastian, Pothalil A." docDate="2015" docId="03DF87CFB639FF8E8CBFFF258A7D76E7" docLanguage="en" docName="zt03904p297.pdf" docOrigin="Zootaxa 3904 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lysania prolixus Malamel, Sankaran, Joseph & Sebastian, 2015, sp. nov." docType="treatment" docVersion="10" lastPageNumber="296" masterDocId="FFE6FFB7B638FF8D8C28FFB28F5B7700" masterDocTitle="First record of the wolf spider genus Lysania Thorell, 1890 from India with the description of a new species (Araneae: Lycosidae: Zoicinae)" masterLastPageNumber="297" masterPageNumber="293" pageNumber="294" updateTime="1698359221022" updateUser="plazi">
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<mods:titleInfo id="B7ECDD99F18CC177060EE4314A4D9A44">
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<mods:title id="7F00B53E1C993E969B9D2E519E274E56">First record of the wolf spider genus Lysania Thorell, 1890 from India with the description of a new species (Araneae: Lycosidae: Zoicinae)</mods:title>
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<mods:name id="E550099707E3A3BBA199A4D8748CDE3A" type="personal">
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<mods:namePart id="DB43FFD3FF0DFFC9286B940093A6EFBD">Malamel, Jobi J.</mods:namePart>
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<mods:namePart id="44B65475DBF37A2C3AC7C6E18B140AB1">Sankaran, Pradeep M.</mods:namePart>
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</mods:name>
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<mods:name id="602941F2026D8841739EB1D3ABD9A900" type="personal">
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<mods:namePart id="B3DEB25C4A9A4B862C4611B9CFAB756B">Joseph, Mathew M.</mods:namePart>
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</mods:name>
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<mods:name id="E301E7B7441E865EB918D1B32777B613" type="personal">
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<mods:roleTerm id="619A07FA04024376F0282F9718C1C74A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9E1098D69D032D7371D00F56CA1ECE06">Sebastian, Pothalil A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0950053C2A60B018358C7BD043737FE6">text</mods:typeOfResource>
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<mods:title id="B4E72A422CEEBEAB98BFCAA5D44D55E1">Zootaxa</mods:title>
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<mods:part id="95351725D2E2D29A09E3CB0E4E5DE68C">
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<mods:date id="59AE2851DB63CBCF4AAE881210E4D458">2015</mods:date>
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<mods:detail id="F21C257AAF8CE59469E5AECB93793EE6" type="volume">
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<mods:number id="EAC3AFFEF5E3153ED8F94FC029EB25FC">3904</mods:number>
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<mods:detail id="A879003BAAA585659860986F644F5BBF" type="issue">
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<mods:number id="5AC0F5BAF630A5F815CE7758C97F2D73">2</mods:number>
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<mods:identifier id="FF61FE1A6D90F6C562AC73E7A82509BE" type="DOI">10.11646/zootaxa.3904.2.8</mods:identifier>
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<treatment id="03DF87CFB639FF8E8CBFFF258A7D76E7" ID-DOI="http://doi.org/10.5281/zenodo.6096892" ID-GBIF-Taxon="119586668" ID-Zenodo-Dep="6096892" LSID="urn:lsid:plazi:treatment:03DF87CFB639FF8E8CBFFF258A7D76E7" httpUri="http://treatment.plazi.org/id/03DF87CFB639FF8E8CBFFF258A7D76E7" lastPageId="3" lastPageNumber="296" pageId="1" pageNumber="294">
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<subSubSection id="C36C6552B639FF8C8CBFFF258E3277D1" pageId="1" pageNumber="294" type="nomenclature">
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<paragraph id="8BC936D9B639FF8C8CBFFF258E9E77B2" blockId="1.[151,453,151,209]" box="[151,453,151,178]" pageId="1" pageNumber="294">
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<heading id="D08181B5B639FF8C8CBFFF258E9E77B2" bold="true" box="[151,453,151,178]" fontSize="11" level="1" pageId="1" pageNumber="294" reason="1">
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<emphasis id="B902EACBB639FF8C8CBFFF258E9E77B2" bold="true" box="[151,453,151,178]" pageId="1" pageNumber="294">
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<taxonomicName id="4C764D5AB639FF8C8CBFFF258E3B77B1" box="[151,352,151,177]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="prolixus" status="sp. nov.">
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<emphasis id="B902EACBB639FF8C8CBFFF258E3B77B1" bold="true" box="[151,352,151,177]" italics="true" pageId="1" pageNumber="294">Lysania prolixus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23157B0B639FF8C8D40FF2A8E9E77B2" box="[360,453,152,178]" pageId="1" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC936D9B639FF8C8CBFFF088E3277D1" blockId="1.[151,453,151,209]" box="[151,361,186,209]" pageId="1" pageNumber="294">
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(
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<figureCitation id="134D2A5CB639FF8C8CB6FF088E4F77D1" box="[158,276,186,209]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1A–L</figureCitation>
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,
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<figureCitation id="134D2A5CB639FF8C8D08FF088E3877D1" box="[288,355,186,209]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2A–H</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C6552B639FF8C8CBFFEB08B467687" pageId="1" pageNumber="294" type="materials_examined">
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<paragraph id="8BC936D9B639FF8C8CBFFEB08B467687" blockId="1.[151,1436,256,2025]" pageId="1" pageNumber="294">
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<emphasis id="B902EACBB639FF8C8CBFFEB08E1A7619" bold="true" box="[151,321,258,281]" pageId="1" pageNumber="294">
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<typeStatus id="54CD887BB639FF8C8CBFFEB08F947619" box="[151,207,258,281]" pageId="1" pageNumber="294">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54CD887BB639FF8C8D64FEB08EE87619" box="[332,435,258,281]" pageId="1" pageNumber="294" type="holotype">Holotype</typeStatus>
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: Male (
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<collectionCode id="ED67AE1CB639FF8C8E23FEB08D0C7619" box="[523,599,258,281]" httpUri="http://grbio.org/cool/7csr-pk9c" name="Arachnology Division, Sacred Heart College" pageId="1" pageNumber="294">ADSH</collectionCode>
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510162):
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<collectingCountry id="F3617649B639FF8C8EE8FEB08DA27619" box="[704,761,258,281]" name="India" pageId="1" pageNumber="294">India</collectingCountry>
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, Kerala, Ernakulam, Cherukadu (
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<geoCoordinate id="EE42501EB639FF8C8859FEB08BB67619" box="[1137,1261,256,281]" direction="north" orientation="latitude" pageId="1" pageNumber="294" precision="15" value="10.139444">10o08'22''N</geoCoordinate>
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,
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<geoCoordinate id="EE42501EB639FF8C88D5FEB08A2F7619" box="[1277,1396,256,281]" direction="east" orientation="longitude" pageId="1" pageNumber="294" precision="15" value="76.66722">76o40'02''E</geoCoordinate>
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) in Bhoothathankettu Forest Reserve,
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<specimenCount id="9D70FD50B639FF8C8E3CFE978D0D763C" box="[532,598,293,316]" pageId="1" pageNumber="294" type="generic">
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<quantity id="4C8E9B3CB639FF8C8E3CFE978D09763C" box="[532,594,293,316]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="294" unit="m" value="37.0">37 m</quantity>
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.
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||
</specimenCount>
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alt.,
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<date id="FFC81019B639FF8C8EBBFE978C04763C" box="[659,863,293,316]" pageId="1" pageNumber="294" value="2013-11-06">6 November 2013</date>
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, Pradeep M. S. leg. by hand;
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||
<typeStatus id="54CD887BB639FF8C889EFE978A79763C" box="[1206,1314,293,316]" pageId="1" pageNumber="294" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="9D70FD50B639FF8C8918FE978AC7763C" box="[1328,1436,293,316]" pageId="1" pageNumber="294" type="female">2 females</specimenCount>
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||
(
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||
<collectionCode id="ED67AE1CB639FF8C8CB6FEF88FB27661" box="[158,233,330,353]" httpUri="http://grbio.org/cool/7csr-pk9c" name="Arachnology Division, Sacred Heart College" pageId="1" pageNumber="294">ADSH</collectionCode>
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510262), same data as
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<typeStatus id="54CD887BB639FF8C8DC5FEF88D157661" box="[493,590,330,353]" pageId="1" pageNumber="294" type="holotype">holotype</typeStatus>
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||
;
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||
<specimenCount id="9D70FD50B639FF8C8E75FEF88DE67661" box="[605,701,330,353]" pageId="1" pageNumber="294" type="female">1 female</specimenCount>
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||
(
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||
<collectionCode id="ED67AE1CB639FF8C8EF8FEF88C477661" box="[720,796,330,353]" httpUri="http://grbio.org/cool/7csr-pk9c" name="Arachnology Division, Sacred Heart College" pageId="1" pageNumber="294">ADSH</collectionCode>
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510362):
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<collectingCountry id="F3617649B639FF8C8FADFEF88CE57661" box="[901,958,330,353]" name="India" pageId="1" pageNumber="294">India</collectingCountry>
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, Kerala, Ernakulam, Illithodu (
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||
<geoCoordinate id="EE42501EB639FF8C8934FEF88ACC7661" box="[1308,1431,328,353]" direction="north" orientation="latitude" pageId="1" pageNumber="294" precision="15" value="10.198889">10o11'56''N</geoCoordinate>
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,
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<geoCoordinate id="EE42501EB639FF8C8CBFFEC28E567687" box="[151,269,366,391]" direction="east" orientation="longitude" pageId="1" pageNumber="294" precision="15" value="76.55028">76o33'01''E</geoCoordinate>
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||
) in Malayatoor,
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||
<quantity id="4C8E9B3CB639FF8C8D97FEC28EAE7687" box="[447,501,368,391]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="1" pageNumber="294" unit="m" value="25.0">25 m</quantity>
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alt.,
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<date id="FFC81019B639FF8C8E02FEC28DB17687" box="[554,746,368,391]" pageId="1" pageNumber="294" value="2014-02-25">25 February 2014</date>
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, Pradeep M. S. leg. by hand.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36C6552B639FF8C8CEFFE208C49759C" pageId="1" pageNumber="294" type="diagnosis">
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<paragraph id="8BC936D9B639FF8C8CEFFE208C49759C" blockId="1.[151,1436,256,2025]" pageId="1" pageNumber="294">
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<emphasis id="B902EACBB639FF8C8CEFFE208E6076A9" bold="true" box="[199,315,402,425]" pageId="1" pageNumber="294">Diagnosis.</emphasis>
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The male of
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<taxonomicName id="4C764D5AB639FF8C8DFEFE208D1476A9" box="[470,591,401,425]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="prolixus" status="sp. nov.">
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<emphasis id="B902EACBB639FF8C8DFEFE208D1476A9" box="[470,591,401,425]" italics="true" pageId="1" pageNumber="294">L. prolixus</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A23157B0B639FF8C8E75FE208DE276A9" box="[605,697,402,425]" pageId="1" pageNumber="294" rank="species">
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||
<emphasis id="B902EACBB639FF8C8E75FE208DE276A9" bold="true" box="[605,697,402,425]" pageId="1" pageNumber="294">sp. nov.</emphasis>
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</taxonomicNameLabel>
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can be separated from all other described congeners (except
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<taxonomicName id="4C764D5AB639FF8C89AEFE208E5776CC" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="sabahensis">
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<emphasis id="B902EACBB639FF8C89AEFE208E5776CC" italics="true" pageId="1" pageNumber="294">L. sabahensis</emphasis>
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||
</taxonomicName>
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, which is unknown) by the presence of an elongated embolus (
|
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<figureCitation id="134D2A5CB639FF8C8F8EFE078CAC76CC" box="[934,1015,437,460]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1F</figureCitation>
|
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,
|
||
<figureCitation id="134D2A5CB639FF8C882AFE078B7F76CC" box="[1026,1060,437,460]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2C</figureCitation>
|
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), (which is very short in
|
||
<taxonomicName id="4C764D5AB639FF8C8902FE078AC776CC" box="[1322,1436,436,460]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="deangia">
|
||
<emphasis id="B902EACBB639FF8C8902FE078AC776CC" box="[1322,1436,436,460]" italics="true" pageId="1" pageNumber="294">L. deangia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFE74B28B639FF8C8CB6FE6A8E6976EF" author="Li" box="[158,306,471,495]" pageId="1" pageNumber="294" refString="Li, Z., Wang, L. Y. & Zhang, Z. S. (2013) The first record of the wolf spider subfamily Zoicinae from China (Araneae: Lycosidae), with the description of two new species. Zootaxa, 3701 (1), 24 - 34. http: // dx. doi. org / 10.11646 / zootaxa. 3701.1.2" type="journal article" year="2013">
|
||
Li
|
||
<emphasis id="B902EACBB639FF8C8C96FE6A8FAF76EF" box="[190,244,471,495]" italics="true" pageId="1" pageNumber="294">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
, fig 1A) and in
|
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<taxonomicName id="4C764D5AB639FF8C8DF3FE6A8D0C76EF" box="[475,599,472,495]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="pygmaea">
|
||
<emphasis id="B902EACBB639FF8C8DF3FE6A8D0C76EF" box="[475,599,472,495]" italics="true" pageId="1" pageNumber="294">L. pygmaea</emphasis>
|
||
</taxonomicName>
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||
, it is long (
|
||
<bibRefCitation id="EFE74B28B639FF8C8EF8FE6A8C3F76EF" author="Li" box="[720,868,471,495]" pageId="1" pageNumber="294" refString="Li, Z., Wang, L. Y. & Zhang, Z. S. (2013) The first record of the wolf spider subfamily Zoicinae from China (Araneae: Lycosidae), with the description of two new species. Zootaxa, 3701 (1), 24 - 34. http: // dx. doi. org / 10.11646 / zootaxa. 3701.1.2" type="journal article" year="2013">
|
||
Li
|
||
<emphasis id="B902EACBB639FF8C8ED8FE6A8C7D76EF" box="[752,806,471,495]" italics="true" pageId="1" pageNumber="294">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
, fig 3A), but shorter than that of
|
||
<taxonomicName id="4C764D5AB639FF8C88ECFE6A8A6C76EF" box="[1220,1335,471,495]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="prolixus" status="sp. nov.">
|
||
<emphasis id="B902EACBB639FF8C88ECFE6A8A6C76EF" box="[1220,1335,471,495]" italics="true" pageId="1" pageNumber="294">L. prolixus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23157B0B639FF8C8916FE6A8AC876EF" box="[1342,1427,472,495]" pageId="1" pageNumber="294" rank="species">
|
||
<emphasis id="B902EACBB639FF8C8916FE6A8AC876EF" bold="true" box="[1342,1427,472,495]" pageId="1" pageNumber="294">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
) and a stout spine-like terminal apophysis situated below the distal half of the embolus (
|
||
<figureCitation id="134D2A5CB639FF8C8844FE488B9E7511" box="[1132,1221,506,529]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1G</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C88FDFE488BAD7511" box="[1237,1270,506,529]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2D</figureCitation>
|
||
) (which is not reported in both
|
||
<taxonomicName id="4C764D5AB639FF8C8D61FDAF8EE77534" box="[329,444,540,564]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="deangia">
|
||
<emphasis id="B902EACBB639FF8C8D61FDAF8EE77534" box="[329,444,540,564]" italics="true" pageId="1" pageNumber="294">L. deangia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C764D5AB639FF8C8DD9FDAF8D357534" box="[497,622,541,564]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="pygmaea">
|
||
<emphasis id="B902EACBB639FF8C8DD9FDAF8D357534" box="[497,622,541,564]" italics="true" pageId="1" pageNumber="294">L. pygmaea</emphasis>
|
||
</taxonomicName>
|
||
). Females can be recognised by very long, coiled copulatory ducts (
|
||
<figureCitation id="134D2A5CB639FF8C8962FDAF8ACF7534" box="[1354,1428,541,564]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">Fig 2F</figureCitation>
|
||
) (in
|
||
<taxonomicName id="4C764D5AB639FF8C8C94FDF28E757557" box="[188,302,575,599]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="deangia">
|
||
<emphasis id="B902EACBB639FF8C8C94FDF28E757557" box="[188,302,575,599]" italics="true" pageId="1" pageNumber="294">L. deangia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C764D5AB639FF8C8D4CFDF28EBB7557" box="[356,480,576,599]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="pygmaea">
|
||
<emphasis id="B902EACBB639FF8C8D4CFDF28EBB7557" box="[356,480,576,599]" italics="true" pageId="1" pageNumber="294">L. pygmaea</emphasis>
|
||
</taxonomicName>
|
||
, the copulatory ducts are straight (
|
||
<bibRefCitation id="EFE74B28B639FF8C8F7EFDF28CB77557" author="Li" box="[854,1004,575,599]" pageId="1" pageNumber="294" refString="Li, Z., Wang, L. Y. & Zhang, Z. S. (2013) The first record of the wolf spider subfamily Zoicinae from China (Araneae: Lycosidae), with the description of two new species. Zootaxa, 3701 (1), 24 - 34. http: // dx. doi. org / 10.11646 / zootaxa. 3701.1.2" type="journal article" year="2013">
|
||
Li
|
||
<emphasis id="B902EACBB639FF8C8F5EFDF28CF67557" box="[886,941,575,599]" italics="true" pageId="1" pageNumber="294">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
, figs 1D, 3D)) and the widely elongated median plate (
|
||
<figureCitation id="134D2A5CB639FF8C8D1EFDD08EDF7579" box="[310,388,610,633]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">Fig 2E</figureCitation>
|
||
) (in both
|
||
<taxonomicName id="4C764D5AB639FF8C8DD2FDD08D2B7579" box="[506,624,609,633]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="deangia">
|
||
<emphasis id="B902EACBB639FF8C8DD2FDD08D2B7579" box="[506,624,609,633]" italics="true" pageId="1" pageNumber="294">L. deangia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C764D5AB639FF8C8E84FDD08C777579" box="[684,812,610,633]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="pygmaea">
|
||
<emphasis id="B902EACBB639FF8C8E84FDD08C777579" box="[684,812,610,633]" italics="true" pageId="1" pageNumber="294">L. pygmaea</emphasis>
|
||
</taxonomicName>
|
||
, it is W-shaped (
|
||
<bibRefCitation id="EFE74B28B639FF8C8FD9FDD08BCA7579" author="Li" box="[1009,1169,609,633]" pageId="1" pageNumber="294" refString="Li, Z., Wang, L. Y. & Zhang, Z. S. (2013) The first record of the wolf spider subfamily Zoicinae from China (Araneae: Lycosidae), with the description of two new species. Zootaxa, 3701 (1), 24 - 34. http: // dx. doi. org / 10.11646 / zootaxa. 3701.1.2" type="journal article" year="2013">
|
||
Li
|
||
<emphasis id="B902EACBB639FF8C883CFDD08B147579" box="[1044,1103,609,633]" italics="true" pageId="1" pageNumber="294">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
, figs 1C, 3C) and in
|
||
<taxonomicName id="4C764D5AB639FF8C89AEFDD08E57759C" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="294" phylum="Arthropoda" rank="species" species="sabahensis">
|
||
<emphasis id="B902EACBB639FF8C89AEFDD08E57759C" italics="true" pageId="1" pageNumber="294">L. sabahensis</emphasis>
|
||
</taxonomicName>
|
||
, it is squarish
|
||
<bibRefCitation id="EFE74B28B639FF8C8D8EFD378C54759C" author="Lehtinen" box="[422,783,645,668]" pageId="1" pageNumber="294" refString="Lehtinen, P. T. & Hippa, H. (1979) Spiders of the Oriental-Australian region. I. Lycosidae: Venoniinae and Zoicinae. Annales Zoologici Fennici, 16, 1 - 22." type="journal article" year="1979">(Lehtinen & Hippa 1979, fig 55))</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C6552B639FF8F8CEFFD1A8DD27019" lastPageId="2" lastPageNumber="295" pageId="1" pageNumber="294" type="description">
|
||
<paragraph id="8BC936D9B639FF8C8CEFFD1A8E0E74B4" blockId="1.[151,1436,256,2025]" pageId="1" pageNumber="294">
|
||
<emphasis id="B902EACBB639FF8C8CEFFD1A8E0975BF" bold="true" box="[199,338,680,703]" pageId="1" pageNumber="294">Description.</emphasis>
|
||
Male (
|
||
<typeStatus id="54CD887BB639FF8C8D89FD1A8D5975BF" box="[417,514,680,703]" pageId="1" pageNumber="294" type="holotype">holotype</typeStatus>
|
||
, figs 1A–C): Prosoma uniformly yellowish-brown, except for a light posterior stripe behind fovea. Fovea longitudinal (0.23), reddish-brown. Eye field black; AER slightly recurved; PER strongly recurved (
|
||
<figureCitation id="134D2A5CB639FF8C8CB6FD5F8FB47404" box="[158,239,749,772]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">Fig 2H</figureCitation>
|
||
). Opisthosoma yellowish-green, hirsute; dorsally and laterally provided with numerous small chalk-white spots, mid-dorsally with 7 short straw-coloured transverse bands bordered on each side by longitudinal chalk-white bands and large spots; venter yellowish. Clypeus and chelicerae yellowish-brown. Cheliceral pro- and retromargins with three teeth each; 2nd promarginal tooth largest. Sternum, maxillae and labium yellowish; maxillae and labium without scopulae. Spinnerets yellowish. Legs yellowish with inconspicuous light annulations; all leg and pedipalp patellae disto-dorsally with a long spine.
|
||
</paragraph>
|
||
<paragraph id="8BC936D9B639FF8C8CEFFC728C0073C9" blockId="1.[151,1436,256,2025]" pageId="1" pageNumber="294">
|
||
Body length 3.52. Prosoma length 1.79, width 1.27, thickness 0.69. Opisthosoma length 1.73, width 0.99, thickness 0.92. Eyes diameter: AME 0.08. ALE 0.05. PME 0.13. PLE 0.10. Eye interdistance: AME–AME 0.02. PME–PME 0.06. AME–ALE 0.03. PME–PLE 0.06. AME–PME 0.05. PLE–PLE 0.21. Clypeus height at AME 0.08, at ALE 0.09. Chelicera length 0.75. Measurements of palp and legs: Palp 2.32 [0.77, 0.35, 0.45, 0.75], I 7.44 [1.97, 0.73, 1.83, 1.76, 1.15], II 6.70 [1.81, 0.67, 1.54, 1.63, 1.05], III 5.73 [1.66, 0.43, 1.14, 1.53, 0.97], IV 8.98 [2.29, 0.63, 2.19, 2.69, 1.18]. Leg formula: 4123. Spination. Palp: femur 1300, patella 0 200, tibia 0 120, tarsus 0002; legs: femur
|
||
<date id="FFC81019B639FF8C88EEFBDF8A577384" box="[1222,1292,1133,1156]" pageId="1" pageNumber="294" value="1310-01">I 1310</date>
|
||
(right 0310),
|
||
<date id="FFC81019B639FF8C8CBFFB228E4873A7" box="[151,275,1168,1191]" pageId="1" pageNumber="294" value="1310-02" valueMax="1310-04-31" valueMin="1310-02-00">II–IV 1310</date>
|
||
; patellae I–IV 0200; tibia I 0 204 (right 0104), II 0 212,
|
||
<date id="FFC81019B639FF8C8F4EFB228CE673A7" box="[870,957,1168,1191]" pageId="1" pageNumber="294" value="1313-03">III 1313</date>
|
||
(right 1213),
|
||
<date id="FFC81019B639FF8C887CFB228BEA73A7" box="[1108,1201,1168,1191]" pageId="1" pageNumber="294" value="2214-04">IV 2214</date>
|
||
; metatarsus I 0 0 0 6,
|
||
<date id="FFC81019B639FF8C89A1FB228F9473C9" pageId="1" pageNumber="294" value="1005-02">II 1005</date>
|
||
, III 4033 (right 4043), IV 5033 (right 5043); tarsi I–IV 0 0 0 0.
|
||
</paragraph>
|
||
<paragraph id="8BC936D9B639FF8C8CEFFB678D4A72BC" blockId="1.[151,1436,256,2025]" pageId="1" pageNumber="294">
|
||
Copulatory organ (
|
||
<figureCitation id="134D2A5CB639FF8C8DA7FB678D5D73EC" box="[399,518,1237,1260]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1D–G</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8E38FB678D0D73EC" box="[528,598,1237,1260]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2A–D</figureCitation>
|
||
): Palpal segments stout, yellowish-brown; cymbium disto-ventrally with a pair of spines (
|
||
<figureCitation id="134D2A5CB639FF8C8D2EFB4A8E03720F" box="[262,344,1272,1295]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1F</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8D4DFB4A8ED3720F" box="[357,392,1272,1295]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2C</figureCitation>
|
||
). Subtegulum small (
|
||
<figureCitation id="134D2A5CB639FF8C8E44FB4A8D86720F" box="[620,733,1272,1295]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1E–F</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8EC1FB4A8C70720F" box="[745,811,1272,1295]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2B–C</figureCitation>
|
||
). Embolus long (
|
||
<figureCitation id="134D2A5CB639FF8C8FCAFB4A8B70720F" box="[994,1067,1272,1295]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Fig 1F</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C881FFB4A8B01720F" box="[1079,1114,1272,1295]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2C</figureCitation>
|
||
), originating distally from the bulb (
|
||
<figureCitation id="134D2A5CB639FF8C8CFCFAA88E737231" box="[212,296,1306,1329]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1E</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8D1BFAA88E2D7231" box="[307,374,1306,1329]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2B–C</figureCitation>
|
||
); median part lies behind the anterior margin of tegulum (
|
||
<figureCitation id="134D2A5CB639FF8C8FFFFAA88B7E7231" box="[983,1061,1306,1329]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">Fig 2C</figureCitation>
|
||
); tip of embolus thin, directed at 11 o’clock position (
|
||
<figureCitation id="134D2A5CB639FF8C8D73FA8F8EE97254" box="[347,434,1341,1364]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1F</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8DEAFA8F8EBE7254" box="[450,485,1341,1364]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2C</figureCitation>
|
||
). Conductor wide and membranous (
|
||
<figureCitation id="134D2A5CB639FF8C8FAEFA8F8C827254" box="[902,985,1341,1364]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">Fig 2C</figureCitation>
|
||
). Lateral apophysis originates prolateromedially from the tegulum (
|
||
<figureCitation id="134D2A5CB639FF8C8DE6FAD28D787277" box="[462,547,1376,1399]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1F</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8E1AFAD28D0F7277" box="[562,596,1376,1399]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2C</figureCitation>
|
||
), with a broad base, while its tip narrow, hook-like (
|
||
<figureCitation id="134D2A5CB639FF8C88B7FAD28BAF7277" box="[1183,1268,1376,1399]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1F</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C892BFAD28A7D7277" box="[1283,1318,1376,1399]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2C</figureCitation>
|
||
). Terminal apophysis short, stout spine-like situated behind the distal half of the embolus (
|
||
<figureCitation id="134D2A5CB639FF8C8FC0FA308B657299" box="[1000,1086,1410,1433]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1G</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8862FA308B367299" box="[1098,1133,1410,1433]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2D</figureCitation>
|
||
), with a pointed tip directed at 1 o’clock position (
|
||
<figureCitation id="134D2A5CB639FF8C8D57FA178E8E72BC" box="[383,469,1445,1468]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1G</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8DC8FA178D5F72BC" box="[480,516,1445,1468]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC936D9B639FF8C8CEFFA7A8DA571D4" blockId="1.[151,1436,256,2025]" pageId="1" pageNumber="294">
|
||
Female (
|
||
<typeStatus id="54CD887BB639FF8C8D0CFA7A8EDD72DF" box="[292,390,1480,1503]" pageId="1" pageNumber="294" type="paratype">Paratype</typeStatus>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8DBAFA7A8D5872DF" box="[402,515,1480,1503]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="1" pageNumber="294">Figs 1H–J</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB639FF8C8E26FA7A8D6872DF" box="[526,563,1480,1503]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">2H</figureCitation>
|
||
): Prosoma uniformly yellowish-brown, except for three light stripes, two anterolaterally to and one behind the fovea. Fovea longitudinal (0.25), reddish-brown. Eye field black; AER slightly recurved; PER strongly recurved (
|
||
<figureCitation id="134D2A5CB639FF8C8DB4F9BF8EB67124" box="[412,493,1549,1572]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="1" pageNumber="294">Fig 2H</figureCitation>
|
||
). Opisthosoma yellowish-green, hirsute; dorsally and laterally provided with numerous small chalk-white spots, mid-dorsally with 7 short straw-coloured transverse bands bordered on each side by longitudinal chalk-white bands and large spots; venter yellowish. Clypeus, chelicerae, sternum, maxillae and labium yellowishbrown. Cheliceral pro- and retromargins with three teeth each; 2nd promarginal tooth largest. Maxillae and labium without scopulae. Spinnerets, palpal segments yellowish. Legs yellowish with conspicuous light greenish-black annulations; all leg and pedipalp patellae disto-dorsally with a long spine.
|
||
</paragraph>
|
||
<paragraph id="8BC936D9B639FF8C8CEFF9528D3D70E9" blockId="1.[151,1436,256,2025]" pageId="1" pageNumber="294">
|
||
Body length 3.58. Prosoma length 1.62, width 1.08, thickness 0.72. Opisthosoma length 1.96, width 1.32, thickness 1.37. Eyes diameter: AME 0.07. ALE 0.03. PME 0.14. PLE 0.11. Eye interdistance: AME–AME 0.01. PME–PME 0.08. AME–ALE 0.03. PME–PLE 0.07. AME–PME 0.05. PLE–PLE 0.29. Clypeus height at AME 0.07, at ALE 0.06. Chelicera length 0.64. Measurements of pedipalp and legs: Palp 1.86 [0.65, 0.29, 0.41, 0.51], I 5.50 [1.53, 0.57, 1.36, 1.22, 0.82], II 4.70 [1.33, 0.48, 1.10, 1.04, 0.75], III 4.49 [1.28, 0.43, 1.01, 1.12, 0.65], IV 6.89 [1.80, 0.50, 1.68, 1.93, 0.98]. Leg formula: 4123. Spination. Palp: femur 1300, patella 0 200, tibia 2100, tarsus 1012; legs: femora
|
||
<date id="FFC81019B639FF8C8927F83F8A2D70A4" box="[1295,1398,1933,1956]" pageId="1" pageNumber="294" value="1300-01" valueMax="1300-02-31" valueMin="1300-01-00">I–II 1300</date>
|
||
,
|
||
<date id="FFC81019B639FF8C89A9F83F8F9470C7" pageId="1" pageNumber="294" value="1310-03">III 1310</date>
|
||
,
|
||
<date id="FFC81019B639FF8C8CF2F8028E6970C7" box="[218,306,1968,1991]" pageId="1" pageNumber="294" value="1310-04">IV 1310</date>
|
||
(right 2310); patellae I–IV 0200; tibia I 0 204,
|
||
<date id="FFC81019B639FF8C8F00F8028C2270C7" box="[808,889,1968,1991]" pageId="1" pageNumber="294" value="2202-02">II 2202</date>
|
||
,
|
||
<date id="FFC81019B639FF8C8FACF8028C8170C7" box="[900,986,1968,1991]" pageId="1" pageNumber="294" value="2123-03">III 2123</date>
|
||
(right 2124),
|
||
<date id="FFC81019B639FF8C8847F8028B9770C7" box="[1135,1228,1968,1991]" pageId="1" pageNumber="294" value="2124-04">IV 2124</date>
|
||
; metatarsus I 0 0 0 6,
|
||
<date id="FFC81019B639FF8C8CBFF8608FBC70E9" box="[151,231,2002,2025]" pageId="1" pageNumber="294" value="2013-02">II 2013</date>
|
||
, III 4033, IV 4043; tarsi I–IV 0 0 0 0.
|
||
</paragraph>
|
||
<caption id="DF096651B63AFF8F8CBFF9F58CAB7197" httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="2" pageNumber="295" targetBox="[180,1436,197,1575]" targetPageId="2">
|
||
<paragraph id="8BC936D9B63AFF8F8CBFF9F58CAB7197" blockId="2.[151,1436,1607,1688]" pageId="2" pageNumber="295">
|
||
<emphasis id="B902EACBB63AFF8F8CBFF9F58E61715D" bold="true" box="[151,314,1607,1629]" pageId="2" pageNumber="295">FIGURE 1A–L.</emphasis>
|
||
<taxonomicName id="4C764D5AB63AFF8F8D17F9FA8E86715D" box="[319,477,1608,1629]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="prolixus" status="sp. nov.">
|
||
<emphasis id="B902EACBB63AFF8F8D17F9FA8E86715D" box="[319,477,1608,1629]" italics="true" pageId="2" pageNumber="295">Lysania prolixus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B902EACBB63AFF8F8DCCF9F58D15715D" bold="true" box="[484,590,1607,1629]" pageId="2" pageNumber="295">
|
||
<taxonomicNameLabel id="A23157B0B63AFF8F8DCCF9F58D6B715D" box="[484,560,1607,1629]" pageId="2" pageNumber="295" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A,
|
||
</emphasis>
|
||
male (holotype) habitus, dorsal view;
|
||
<emphasis id="B902EACBB63AFF8F8F97F9F58C8F715D" bold="true" box="[959,980,1607,1629]" pageId="2" pageNumber="295">B,</emphasis>
|
||
same, ventral view;
|
||
<emphasis id="B902EACBB63AFF8F88B4F9F58BE8715D" bold="true" box="[1180,1203,1607,1629]" pageId="2" pageNumber="295">C,</emphasis>
|
||
same, lateral view.
|
||
<emphasis id="B902EACBB63AFF8F8959F9F58FF6717A" bold="true" pageId="2" pageNumber="295">D & E,</emphasis>
|
||
male left pedipalp, prolateral view;
|
||
<emphasis id="B902EACBB63AFF8F8E27F9D68D44717A" bold="true" box="[527,543,1636,1658]" pageId="2" pageNumber="295">F,</emphasis>
|
||
same, ventral view;
|
||
<emphasis id="B902EACBB63AFF8F8EC4F9D68DA4717A" bold="true" box="[748,767,1636,1658]" pageId="2" pageNumber="295">G,</emphasis>
|
||
same retrolateral view.
|
||
<emphasis id="B902EACBB63AFF8F8FC3F9D68B5F717A" bold="true" box="[1003,1028,1636,1658]" pageId="2" pageNumber="295">H,</emphasis>
|
||
female (paratype) habitus, dorsal view;
|
||
<emphasis id="B902EACBB63AFF8F89A4F9D68AC7717A" bold="true" box="[1420,1436,1636,1658]" pageId="2" pageNumber="295">I,</emphasis>
|
||
same, ventral view;
|
||
<emphasis id="B902EACBB63AFF8F8D70F9308E317198" bold="true" box="[344,362,1666,1688]" pageId="2" pageNumber="295">J,</emphasis>
|
||
same, lateral view.
|
||
<emphasis id="B902EACBB63AFF8F8E00F9308D1A7198" bold="true" box="[552,577,1666,1688]" pageId="2" pageNumber="295">K,</emphasis>
|
||
epigyne, ventral view;
|
||
<emphasis id="B902EACBB63AFF8F8F0AF9308C637198" bold="true" box="[802,824,1666,1688]" pageId="2" pageNumber="295">L,</emphasis>
|
||
same, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC936D9B63AFF8F8CEFF90F8DD27019" blockId="2.[151,1436,1725,2026]" pageId="2" pageNumber="295">
|
||
Copulatory organ (
|
||
<figureCitation id="134D2A5CB63AFF8F8DBBF90F8D5071D4" box="[403,523,1725,1748]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="2" pageNumber="295">Figs 1K–L</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB63AFF8F8E30F90F8DD871D4" box="[536,643,1725,1748]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="2" pageNumber="295">2E–F, 2G</figureCitation>
|
||
): Median plate broad, hirsute with slightly convex posterior border (
|
||
<figureCitation id="134D2A5CB63AFF8F8944F90F8FE271F7" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="2" pageNumber="295">Figs 1K</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB63AFF8F8CEDF9528FBC71F7" box="[197,231,1760,1783]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="2" pageNumber="295">2E</figureCitation>
|
||
). Copulatory openings lie postero-laterally to the median plate (
|
||
<figureCitation id="134D2A5CB63AFF8F8FBDF9528CB671F7" box="[917,1005,1760,1783]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="2" pageNumber="295">Figs 1K</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB63AFF8F8FD2F9528B4371F7" box="[1018,1048,1760,1783]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="2" pageNumber="295">2E</figureCitation>
|
||
).Copulatory ducts long, coiled with small, ovate spermathecal heads (
|
||
<figureCitation id="134D2A5CB63AFF8F8DD4F8B08D0A7019" box="[508,593,1794,1817]" captionStart="FIGURE 1 A – L" captionStartId="2.[151,244,1607,1629]" captionTargetBox="[180,1436,197,1575]" captionTargetId="figure@2.[150,1436,191,1586]" captionTargetPageId="2" captionText="FIGURE 1 A – L. Lysania prolixus sp. nov. A, male (holotype) habitus, dorsal view; B, same, ventral view; C, same, lateral view. D & E, male left pedipalp, prolateral view; F, same, ventral view; G, same retrolateral view. H, female (paratype) habitus, dorsal view; I, same, ventral view; J, same, lateral view. K, epigyne, ventral view; L, same, dorsal view." httpUri="https://zenodo.org/record/244765/files/figure.png" pageId="2" pageNumber="295">Figs 1L</figureCitation>
|
||
,
|
||
<figureCitation id="134D2A5CB63AFF8F8E75F8B08D267019" box="[605,637,1794,1817]" captionStart="FIGURE 2 A – H" captionStartId="3.[151,244,1818,1840]" captionTargetBox="[163,1425,548,1784]" captionTargetId="figure@3.[150,1436,538,1797]" captionTargetPageId="3" captionText="FIGURE 2 A – H. Lysania prolixus sp. nov. A & B, male left pedipalp, prolateral view; C, same, ventral view; D, same, retrolateral view. E, epigyne, ventral view; F, same, dorsal view; G, schematic course of internal duct system, dorsal. H, female eye arrangement, frontal view. C—conductor, CO—copulatory opening, E—embolus, FD—fertilization duct, LA—lateral apophysis, MP—median plate of epigyne, SH—spermathecal head, SS—spermathecal stalk, ST—subtegulum, T—tegulum, TA—terminal apophysis. Scale bars: A, 0.67 mm; B – D, 0.24 mm; E – F, 0.03 mm; H, 0.15 mm." httpUri="https://zenodo.org/record/244766/files/figure.png" pageId="2" pageNumber="295">2F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C6552B63AFF8F8CEFF8978E09705F" pageId="2" pageNumber="295" type="etymology">
|
||
<paragraph id="8BC936D9B63AFF8F8CEFF8978E09705F" blockId="2.[151,1436,1725,2026]" pageId="2" pageNumber="295">
|
||
<emphasis id="B902EACBB63AFF8F8CEFF8978E1C703C" bold="true" box="[199,327,1829,1852]" pageId="2" pageNumber="295">Etymology.</emphasis>
|
||
The specific epithet is an adjective and is derived from the elongated embolus: Latin
|
||
<taxonomicName id="4C764D5AB63AFF8F88C7F8968A1E703C" box="[1263,1349,1828,1852]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="prolixus">
|
||
<emphasis id="B902EACBB63AFF8F88C7F8968A1E703C" box="[1263,1349,1828,1852]" italics="true" pageId="2" pageNumber="295">prolixus</emphasis>
|
||
</taxonomicName>
|
||
= long. Gender feminine.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36C6552B63AFF8E8CEFF8D88A7D76E7" lastPageId="3" lastPageNumber="296" pageId="2" pageNumber="295" type="distribution">
|
||
<paragraph id="8BC936D9B63AFF8F8CEFF8D88EC370A4" blockId="2.[151,1436,1725,2026]" pageId="2" pageNumber="295">
|
||
<emphasis id="B902EACBB63AFF8F8CEFF8D88E027081" bold="true" box="[199,345,1898,1921]" pageId="2" pageNumber="295">Distribution.</emphasis>
|
||
At present known only from Kerala,
|
||
<collectingCountry id="F3617649B63AFF8F8ECEF8D88C467081" box="[742,797,1898,1921]" name="India" pageId="2" pageNumber="295">India</collectingCountry>
|
||
and representing the first records of the genus
|
||
<taxonomicName id="4C764D5AB63AFF8F8926F8D88A397081" box="[1294,1378,1898,1921]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="295" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EACBB63AFF8F8926F8D88A397081" box="[1294,1378,1898,1921]" italics="true" pageId="2" pageNumber="295">Lysania</emphasis>
|
||
</taxonomicName>
|
||
from the Indian subcontinent.
|
||
</paragraph>
|
||
<paragraph id="8BC936D9B63AFF8F8CEFF8028E1D70EA" blockId="2.[151,1436,1725,2026]" pageId="2" pageNumber="295">
|
||
<emphasis id="B902EACBB63AFF8F8CEFF8028E2670C7" bold="true" box="[199,381,1968,1991]" pageId="2" pageNumber="295">Natural history.</emphasis>
|
||
Both male and females of
|
||
<taxonomicName id="4C764D5AB63AFF8F8E8DF8028C4270C7" box="[677,793,1967,1991]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="prolixus" status="sp. nov.">
|
||
<emphasis id="B902EACBB63AFF8F8E8DF8028C4270C7" box="[677,793,1967,1991]" italics="true" pageId="2" pageNumber="295">L. prolixus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23157B0B63AFF8F8F09F8028C2370C7" box="[801,888,1968,1991]" pageId="2" pageNumber="295" rank="species">
|
||
<emphasis id="B902EACBB63AFF8F8F09F8028C2370C7" bold="true" box="[801,888,1968,1991]" pageId="2" pageNumber="295">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
were collected from small sheet-webs constructed near the ground.
|
||
</paragraph>
|
||
<paragraph id="8BC936D9B63BFF8E8CEFFF2A8C5477F4" blockId="3.[151,1436,151,487]" pageId="3" pageNumber="296">
|
||
<emphasis id="B902EACBB63BFF8E8CEFFF2A8E5677AF" bold="true" box="[199,269,152,175]" pageId="3" pageNumber="296">Notes.</emphasis>
|
||
<bibRefCitation id="EFE74B28B63BFF8E8D3CFF2A8EE377AF" box="[276,440,151,175]" pageId="3" pageNumber="296" refString="Li, Z., Wang, L. Y. & Zhang, Z. S. (2013) The first record of the wolf spider subfamily Zoicinae from China (Araneae: Lycosidae), with the description of two new species. Zootaxa, 3701 (1), 24 - 34. http: // dx. doi. org / 10.11646 / zootaxa. 3701.1.2" type="journal article">
|
||
Li
|
||
<emphasis id="B902EACBB63BFF8E8D1CFF2A8E3177AF" box="[308,362,151,175]" italics="true" pageId="3" pageNumber="296">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
suggested the presence of metallic luster on the dorsal opisthosoma of male
|
||
<taxonomicName id="4C764D5AB63BFF8E88DCFF2A8A1377AF" box="[1268,1352,152,175]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="296" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EACBB63BFF8E88DCFF2A8A1377AF" box="[1268,1352,152,175]" italics="true" pageId="3" pageNumber="296">Lysania</emphasis>
|
||
</taxonomicName>
|
||
species as a diagnostic feature for the genus. However, the male of
|
||
<taxonomicName id="4C764D5AB63BFF8E8F6BFF088CE277D1" box="[835,953,185,209]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="296" phylum="Arthropoda" rank="species" species="prolixus" status="sp. nov.">
|
||
<emphasis id="B902EACBB63BFF8E8F6BFF088CE277D1" box="[835,953,185,209]" italics="true" pageId="3" pageNumber="296">L. prolixus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23157B0B63BFF8E8FEDFF088B4477D1" box="[965,1055,186,209]" pageId="3" pageNumber="296" rank="species">
|
||
<emphasis id="B902EACBB63BFF8E8FEDFF088B4477D1" bold="true" box="[965,1055,186,209]" pageId="3" pageNumber="296">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has no such feature on its dorsal opisthosoma. Instead, it is simply covered with black hairs.
|
||
</paragraph>
|
||
<paragraph id="8BC936D9B63BFF8E8CEFFEB28A7D76E7" blockId="3.[151,1436,151,487]" pageId="3" pageNumber="296">
|
||
We considered both
|
||
<taxonomicName id="4C764D5AB63BFF8E8D8AFEB28EAD7617" box="[418,502,256,279]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="296" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EACBB63BFF8E8D8AFEB28EAD7617" box="[418,502,256,279]" italics="true" pageId="3" pageNumber="296">Lysania</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C764D5AB63BFF8E8E02FEB28DAC7617" authority="Simon, 1898" authorityName="Simon" authorityYear="1898" box="[554,759,256,279]" class="Arachnida" family="Lycosidae" genus="Zoica" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="296" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EACBB63BFF8E8E02FEB28D3C7617" box="[554,615,256,279]" italics="true" pageId="3" pageNumber="296">Zoica</emphasis>
|
||
Simon, 1898
|
||
</taxonomicName>
|
||
as genera for this new species, however, it best fits the generic characters of the genus
|
||
<taxonomicName id="4C764D5AB63BFF8E8DB2FE908EB57639" box="[410,494,290,313]" class="Arachnida" family="Lycosidae" genus="Lysania" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="296" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EACBB63BFF8E8DB2FE908EB57639" box="[410,494,290,313]" italics="true" pageId="3" pageNumber="296">Lysania</emphasis>
|
||
</taxonomicName>
|
||
proposed by
|
||
<bibRefCitation id="EFE74B28B63BFF8E8EABFE908CC27639" author="Lehtinen" box="[643,921,290,313]" pageId="3" pageNumber="296" refString="Lehtinen, P. T. & Hippa, H. (1979) Spiders of the Oriental-Australian region. I. Lycosidae: Venoniinae and Zoicinae. Annales Zoologici Fennici, 16, 1 - 22." type="journal article" year="1979">Lehtinen & Hippa (1979)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFE74B28B63BFF8E8FF8FE908B277639" box="[976,1148,289,313]" pageId="3" pageNumber="296" refString="Li, Z., Wang, L. Y. & Zhang, Z. S. (2013) The first record of the wolf spider subfamily Zoicinae from China (Araneae: Lycosidae), with the description of two new species. Zootaxa, 3701 (1), 24 - 34. http: // dx. doi. org / 10.11646 / zootaxa. 3701.1.2" type="journal article">
|
||
Li
|
||
<emphasis id="B902EACBB63BFF8E8FD9FE908B737639" box="[1009,1064,289,313]" italics="true" pageId="3" pageNumber="296">et al.</emphasis>
|
||
(2013)
|
||
</bibRefCitation>
|
||
: opisthosoma dorsally and laterally with a pattern of yellowish-white spots and bands (see figs 1A, 1C, 1H, 1J); legs with annulations (see figs 1A, 1H); beak-like lateral apophysis (see figs 1F, 2C) (in
|
||
<taxonomicName id="4C764D5AB63BFF8E8EE4FEDA8C53767F" box="[716,776,360,383]" class="Arachnida" family="Lycosidae" genus="Zoica" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="296" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EACBB63BFF8E8EE4FEDA8C53767F" box="[716,776,360,383]" italics="true" pageId="3" pageNumber="296">Zoica</emphasis>
|
||
</taxonomicName>
|
||
, this kind of modification is absent); wider conductor (see fig 2C); spine-like embolus having distal origin (see figs 1F, 2C); long copulatory ducts (here it is coiled) (see fig 2F). In
|
||
<taxonomicName id="4C764D5AB63BFF8E8CBFFE1F8F8F76C4" box="[151,212,429,452]" class="Arachnida" family="Lycosidae" genus="Zoica" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="296" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EACBB63BFF8E8CBFFE1F8F8F76C4" box="[151,212,429,452]" italics="true" pageId="3" pageNumber="296">Zoica</emphasis>
|
||
</taxonomicName>
|
||
, the copulatory ducts are reduced in length (see
|
||
<bibRefCitation id="EFE74B28B63BFF8E8EF5FE1F8C2F76C4" author="Li" box="[733,884,428,452]" pageId="3" pageNumber="296" refString="Li, Z., Wang, L. Y. & Zhang, Z. S. (2013) The first record of the wolf spider subfamily Zoicinae from China (Araneae: Lycosidae), with the description of two new species. Zootaxa, 3701 (1), 24 - 34. http: // dx. doi. org / 10.11646 / zootaxa. 3701.1.2" type="journal article" year="2013">
|
||
Li
|
||
<emphasis id="B902EACBB63BFF8E8ED4FE1F8C6876C4" box="[764,819,428,452]" italics="true" pageId="3" pageNumber="296">et al.</emphasis>
|
||
2013
|
||
</bibRefCitation>
|
||
); moreover, the specimens (both male and females) were collected from small sheet webs, which are apparently absent in members of a vagrant genus like
|
||
<taxonomicName id="4C764D5AB63BFF8E88CBFE628A7B76E7" box="[1251,1312,464,487]" class="Arachnida" family="Lycosidae" genus="Zoica" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="296" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B902EACBB63BFF8E88CBFE628A7B76E7" box="[1251,1312,464,487]" italics="true" pageId="3" pageNumber="296">Zoica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |