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<mods:title id="618E73C74B1255F7BFA8CB15E4FF9050">Cladistic analysis and taxonomic revision of the genus Karos Goodnight &amp; Goodnight, 1944 (Opiliones, Laniatores, Stygnopsidae)</mods:title>
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<mods:namePart id="7F88F87C3EE0150ED457F582E77443A2">Cruz-López, Jesús A.</mods:namePart>
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<mods:date id="F58EB4B4554094E8D06A87E70965BDC9">2015</mods:date>
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<paragraph id="8BDC36EFA24CFFB5FC65FECBC680FE8A" blockId="25.[945,1323,305,359]" box="[991,1276,305,327]" pageId="25" pageNumber="852">
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<taxonomicName id="4C634D6CA24CFFB5FC65FECBC6EAFE8A" authority="Cruz-López &amp; Francke, 2015" authorityName="Cruz-López &amp; Francke" authorityYear="2015" box="[991,1174,306,327]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="25" pageNumber="887" phylum="Arthropoda" rank="species" species="tersum" status="sp. nov.">KAROS TERSUM</taxonomicName>
<emphasis id="B917EAFDA24CFFB5FB25FECCC680FE8A" bold="true" box="[1183,1276,305,326]" pageId="25" pageNumber="852">
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<subSubSection id="C3796564A24CFFB5FC0BFEA8C757FEAB" box="[945,1323,337,359]" pageId="25" pageNumber="852" type="description">
<paragraph id="8BDC36EFA24CFFB5FC0BFEA8C757FEAB" blockId="25.[945,1323,305,359]" box="[945,1323,337,359]" pageId="25" pageNumber="852">
(
<figureCitation id="13582A6AA24CFFB5FC03FEA8C642FEAA" box="[953,1086,337,358]" captionStart-0="Figure 21" captionStart-1="Figure 22" captionStart-2="Figure 23" captionStartId-0="25.[164,243,1853,1872]" captionStartId-1="26.[144,223,624,643]" captionStartId-2="26.[144,223,1365,1384]" captionTargetBox-0="[166,1441,1226,1819]" captionTargetBox-1="[305,1261,200,591]" captionTargetBox-2="[144,1425,720,1336]" captionTargetId-0="figure-432@25.[164,1444,1223,1823]" captionTargetId-1="figure-222@26.[303,1264,197,594]" captionTargetId-2="figure-265@26.[144,1425,720,1336]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="26" captionText-0="Figure 21. Karos tersum sp. nov. A, habitus, dorsal view. B, scutum, dorsal view. C, habitus, lateral view. Scale bars: A = 3.8 mm, B = 1.5 mm, C = 1 mm." captionText-1="Figure 22. Karos tersum sp. nov. A, legs IV, mesal view. B, habitus, ventral view. C, ocularium, frontal view. Scale bars: A = 3.4 mm, B = 2.0 mm, C = 0.5 mm. The dark lines on (B) indicate the stigmatic area." captionText-2="Figure 23. Karos tersum sp. nov. male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C)." figureDoi-0="http://doi.org/10.5281/zenodo.5340017" figureDoi-1="http://doi.org/10.5281/zenodo.5340019" figureDoi-2="http://doi.org/10.5281/zenodo.5340021" httpUri-0="https://zenodo.org/record/5340017/files/figure.png" httpUri-1="https://zenodo.org/record/5340019/files/figure.png" httpUri-2="https://zenodo.org/record/5340021/files/figure.png" pageId="25" pageNumber="852">FIGS 2123</figureCitation>
,
<figureCitation id="13582A6AA24CFFB5FBF1FEA8C616FEAB" box="[1099,1130,337,359]" captionStart="Figure 59" captionStartId="56.[144,223,1207,1226]" captionTargetBox="[144,1425,196,1177]" captionTargetId="figure-239@56.[144,1425,196,1177]" captionTargetPageId="56" captionText="Figure 59. Distribution map of the species of the Karos genus-group, continued. White circles with black outline, Karos tersum sp. nov.; white circles with blue outline, Karos projectus; white circles with red outline, Karos singularis sp. nov.; black squares, Huasteca gratiosa comb. nov.; white square with black outline, Huasteca silhavyi sp. nov.; white square with red outline, Huasteca rugosa comb. nov.; black circles, Karos barbarikos." figureDoi="http://doi.org/10.5281/zenodo.5340095" httpUri="https://zenodo.org/record/5340095/files/figure.png" pageId="25" pageNumber="852">59</figureCitation>
,
<figureCitation id="13582A6AA24CFFB5FBCDFEA8C6DBFEAB" box="[1143,1191,337,359]" captionStart="Figure 61" captionStartId="58.[144,223,1345,1364]" captionTargetBox="[144,1425,194,1315]" captionTargetId="figure-0@58.[144,1425,194,1315]" captionTargetPageId="58" captionText="Figure 61. A, detail of one of the lateral clear areas forming tubercles on scutum of Crettaros santibanezi sp. nov. B, detail of one of the lateral clear areas projected in tubercles on scutum of Karos singularis sp. nov. C, detail of one of the lateral clear areas projected in tubercles on scutum of Karos tersum sp. nov. D, detail of one of the lateral clear areas projected in tubercles on scutum of Huasteca silhavyi sp. nov. E, detail of one of the lateral clear areas projected in tubercles on scutum of Montabunus foliorum. F, detail of one of the lateral clear areas projected in tubercles on scutum of Monterella tuberculata. G, extra row of pegs in Chapulobunus poblano sp. nov., indicated by arrows. H, dorsal apophyses on trochanter II in male of Karos singularis sp. nov. I, detail of mesotergal areas III and IV in Montabunus foliorum; dotted lines indicate the sulcus between these areas. J, detail of mesotergal areas III and IV on Chapulobunus unispinosus; dotted lines indicate the sulcus between these areas. These images are not at the same scale." figureDoi="http://doi.org/10.5281/zenodo.5340099" httpUri="https://zenodo.org/record/5340099/files/figure.png" pageId="25" pageNumber="852">61C</figureCitation>
,
<figureCitation id="13582A6AA24CFFB5FB0EFEA8C69AFEAB" box="[1204,1254,337,359]" captionStart="Figure 62" captionStartId="59.[164,243,1363,1382]" captionTargetBox="[244,1365,194,1334]" captionTargetId="figure-0@59.[244,1365,194,1334]" captionTargetPageId="59" captionText="Figure 62. A, dorsal view of lateral clear areas forming tubercles of Karos tersum sp. nov.; arrows indicate the clear areas. B, dorsal view of lateral clear areas projected in tubercles of Huasteca silhavyi sp. nov.; arrows indicate the clear areas. C, lateral view of posterior scutum of Karos singularis sp. nov., showing the spiniform tubercles. D, lateral view of posterior scutum of Montabunus foliorum, showing the rounded tubercles. E, dorsal view of row of pegs in Karos barbarikos; some pegs are indicated by black dots. F, dorsal view of row of pegs in Hoplobunus boneti; some pegs are indicated by black dots. These images are not at the same scale." figureDoi="http://doi.org/10.5281/zenodo.5340101" httpUri="https://zenodo.org/record/5340101/files/figure.png" pageId="25" pageNumber="852">62A</figureCitation>
,
<figureCitation id="13582A6AA24CFFB5FB48FEA8C75FFEAB" box="[1266,1315,337,359]" captionStart="Figure 63" captionStartId="60.[144,223,1276,1295]" captionTargetBox="[144,1425,196,1246]" captionTargetId="figure-0@60.[144,1425,196,1246]" captionTargetPageId="60" captionText="Figure 63. A, dorsal view of the anal bulge on Chapulobunus poblano sp. nov.; arrow indicates the bulge. B, lateral view of Huasteca silhavyi sp. nov., line indicates the very high lateral channel area. C, laterodorsal view of scutum of Crettaros valdezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. D, posterodorsal view of scutum of Crettaros santibanezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. E, dorsal view of trochanter IV of Karos barbarikos; arrow indicates the tridentate apophysis. F, anterodorsal view of scutum of Karos barbarikos, the black points mark the tubercles forming a V. G, dorsal ornamentation of Karos tersum sp. nov. showing the transversal row of tubercles on mesotergal areas; transversal row II indicated by black dots. H, detail of the ventral ornamentation on femur IV of Karos barbarikos. I, detail of the ectal ornamentation on tibia IV of Karos barbarikos; arrows indicate the spiniform tubercles. J, detail of the ventral ornamentation on leg IV in Chapulobunus poblano sp. nov. K, detail of the two apical spines on femur IV in Chapulobunus poblano sp. nov. These images are not at the same scale." figureDoi="http://doi.org/10.5281/zenodo.5340103" httpUri="https://zenodo.org/record/5340103/files/figure.png" pageId="25" pageNumber="852">63G</figureCitation>
)
</paragraph>
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<subSubSection id="C3796564A24CFFB5FC80FE80C106FE60" pageId="25" pageNumber="852" type="reference_group">
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<treatmentCitationGroup id="AB7311C1A24CFFB5FC80FE80C106FE60" pageId="25" pageNumber="852">
<treatmentCitation id="0AC210FEA24CFFB5FC80FE80C70AFE42" author="Goodnight CJ &amp; Goodnight ML" box="[826,1398,376,398]" page="35" pageId="25" pageNumber="852" year="1971">
<taxonomicName id="4C634D6CA24CFFB5FC80FE80C70AFE42" ID-CoL="7WT78" authority=": Goodnight &amp; Goodnight, 1971: 35" authorityName=": Goodnight &amp; Goodnight" authorityPageNumber="35" authorityYear="1971" box="[826,1398,376,398]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="25" pageNumber="852" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B917EAFDA24CFFB5FC80FE80C1A1FE42" box="[826,989,377,398]" italics="true" pageId="25" pageNumber="852">Karos parvus:</emphasis>
<bibRefCitation id="EFF24B1EA24CFFB5FC52FE81C70AFE42" author="Goodnight CJ &amp; Goodnight ML" box="[1000,1398,376,398]" pageId="25" pageNumber="852" pagination="33 - 45" refId="ref38192" refString="Goodnight CJ, Goodnight ML. 1971. Opilionids (Phalangida) of the family Phalangodidae from Mexican caves. Association for Mexican Cave Studies, Bulletin 4: 33 - 45." type="journal article" year="1971">Goodnight &amp; Goodnight, 1971: 35</bibRefCitation>
</taxonomicName>
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(in part).
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</paragraph>
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<subSubSection id="C3796564A24CFFB7FC80FE2EC061FBA0" lastPageId="27" lastPageNumber="854" pageId="25" pageNumber="852" type="materials_examined">
<paragraph id="8BDC36EFA24CFFB5FC80FE2EC1FCFD68" blockId="25.[826,1441,470,676]" pageId="25" pageNumber="852">
<emphasis id="B917EAFDA24CFFB5FC80FE2EC1A1FE27" box="[826,989,470,492]" italics="true" pageId="25" pageNumber="852">Type material:</emphasis>
<materialsCitation id="3B0B3CB2A24CFFB5FC5FFE2FC65AFDC7" country="Mexico" location="San Luis Potosi" pageId="25" pageNumber="852" specimenCode="CNAN-T0725" specimenCount="1" specimenCount-male="1" stateProvince="San Luis Potosi" typeStatus="holotype">
<collectingCountry id="F374767FA24CFFB5FC5FFE2FC632FE27" box="[997,1102,470,491]" name="Mexico" pageId="25" pageNumber="852">MEXICO</collectingCountry>
:
<emphasis id="B917EAFDA24CFFB5FBE7FE2FC764FE20" box="[1117,1304,470,492]" italics="true" pageId="25" pageNumber="852">
<collectingRegion id="49A7F80DA24CFFB5FBE7FE2FC76EFE20" box="[1117,1298,470,492]" country="Mexico" name="San Luis Potosi" pageId="25" pageNumber="852">San Luís Potosí</collectingRegion>
:
</emphasis>
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(
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) and
</materialsCitation>
<materialsCitation id="3B0B3CB2A24CFFB5FB89FE0CC100FD68" collectingDate="1970-07" collectionCode="TMM" collectorName="D. McKenzie &amp; Municipio Ciudad Valles &amp; La Cueva de la Llanura" country="Mexico" location="Micos" pageId="25" pageNumber="852" specimenCode="TMM 37.129" specimenCount="3" specimenCount-female="2" stateProvince="San Luis Potosi" typeStatus="paratype">
<specimenCount id="9D65FD66A24CFFB5FB89FE0CC61FFDC6" box="[1075,1123,500,522]" pageId="25" pageNumber="852" type="female">2 ♀</specimenCount>
<typeStatus id="54D8884DA24CFFB5FBD5FE0CC69AFDC6" box="[1135,1254,501,522]" pageId="25" pageNumber="852" type="paratype">paratypes</typeStatus>
(
<specimenCode id="DBC59E94A24CFFB5FB40FE0CC7E5FDC7" box="[1274,1433,501,523]" collectionCode="TMM" country="0" httpUri="http://grbio.org/cool/22jz-cx2d" name="Texas Memorial Museum" pageId="25" pageNumber="852">TMM 37.129</specimenCode>
) (39.
<date id="FFDD102FA24CFFB5FCD2FDEAC1ADFDE5" box="[872,977,531,553]" pageId="25" pageNumber="852" value="1970-07">
<collectingDate id="EF99E9C7A24CFFB5FCD2FDEAC1ADFDE5" box="[872,977,531,553]" pageId="25" pageNumber="852" value="1970-07">vii.1970</collectingDate>
</date>
;
<collectorName id="26965339A24CFFB5FC5CFDEDC6EDFDE5" box="[998,1169,531,553]" pageId="25" pageNumber="852">D. McKenzie</collectorName>
), (lat 22.11639°, lon 99.16917°).
<collectorName id="26965339A24CFFB5FC75FDCBC694FD84" box="[975,1256,562,584]" pageId="25" pageNumber="852">Municipio Ciudad Valles</collectorName>
,
<collectorName id="26965339A24CFFB5FB4EFDCBC1E5FDAA" pageId="25" pageNumber="852">La Cueva de la Llanura</collectorName>
,
<quantity id="4C9B9B0AA24CFFB5FC13FDA8C187FDAA" box="[937,1019,592,614]" metricMagnitude="3" metricUnit="m" metricValue="4.8" pageId="25" pageNumber="852" unit="km" value="4.8">4.8 km</quantity>
W
<location id="8EBC6034A24CFFB5FB90FDA9C610FDAA" LSID="urn:lsid:plazi:treatment:03CA87F9A24CFFB7FC65FECBC061FBA0:8EBC6034A24CFFB5FB90FDA9C610FDAA" box="[1066,1132,592,614]" country="Mexico" name="Micos" pageId="25" pageNumber="852" stateProvince="San Luis Potosi">Micos</location>
(previously designated as
<typeStatus id="54D8884DA24CFFB5FC80FD89C1D5FD49" box="[826,937,624,645]" pageId="25" pageNumber="852" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C634D6CA24CFFB5FC77FD89C618FD49" authorityName=": Goodnight &amp; Goodnight" authorityYear="1971" box="[973,1124,624,645]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="25" pageNumber="852" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B917EAFDA24CFFB5FC77FD89C618FD49" box="[973,1124,624,645]" italics="true" pageId="25" pageNumber="852">Karos parvus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF24B1EA24CFFB5FB34FD96C108FD68" author="Goodnight CJ &amp; Goodnight ML" pageId="25" pageNumber="852" pagination="33 - 45" refId="ref38192" refString="Goodnight CJ, Goodnight ML. 1971. Opilionids (Phalangida) of the family Phalangodidae from Mexican caves. Association for Mexican Cave Studies, Bulletin 4: 33 - 45." type="journal article" year="1971">Goodnight &amp; Goodnight, 1971</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36EFA24CFFB5FC80FD34C1B0FC92" blockId="25.[826,1441,717,862]" pageId="25" pageNumber="852">
<emphasis id="B917EAFDA24CFFB5FC80FD34C660FD2E" box="[826,1052,717,738]" italics="true" pageId="25" pageNumber="852">Material examined:</emphasis>
<materialsCitation id="3B0B3CB2A24CFFB5FB92FD34C1B4FC92" collectingDate="2011-02-13" collectionCode="CNAN" collectorName="O. Francke &amp; A. Valdez &amp; C. Santibanez &amp; J. Cruz &amp; G. Contreras &amp; Municipio Aquismon &amp; Mantetzulel" country="Mexico" latitude="21.630001" location="Cueva de El Sol" longLatPrecision="15" longitude="-99.06076" pageId="25" pageNumber="852" specimenCount="1" specimenCount-female="1" stateProvince="San Luis Potosi">
<collectingCountry id="F374767FA24CFFB5FB92FD34C6EEFD2E" box="[1064,1170,717,738]" name="Mexico" pageId="25" pageNumber="852">MEXICO</collectingCountry>
:
<emphasis id="B917EAFDA24CFFB5FB1EFD37C71BFD2F" box="[1188,1383,718,739]" italics="true" pageId="25" pageNumber="852">
<collectingRegion id="49A7F80DA24CFFB5FB1EFD37C71DFD2F" box="[1188,1377,718,739]" country="Mexico" name="San Luis Potosi" pageId="25" pageNumber="852">San Luís Potosí</collectingRegion>
:
</emphasis>
<specimenCount id="9D65FD66A24CFFB5FAC9FD34C7DDFD2F" box="[1395,1441,717,739]" pageId="25" pageNumber="852" type="female">1 ♀</specimenCount>
[
<collectionCode id="ED72AE2AA24CFFB5FCFAFD15C1F7FCCD" box="[832,907,748,769]" pageId="25" pageNumber="852">CNAN</collectionCode>
(
<date id="FFDD102FA24CFFB5FC23FD15C67EFCCE" box="[921,1026,748,770]" pageId="25" pageNumber="852" value="2011-02-13">
<collectingDate id="EF99E9C7A24CFFB5FC23FD15C67EFCCE" box="[921,1026,748,770]" pageId="25" pageNumber="852" value="2011-02-13">13.ii.2011</collectingDate>
</date>
;
<collectorName id="26965339A24CFFB5FBB7FD15C6FBFCCE" box="[1037,1159,748,770]" pageId="25" pageNumber="852">O. Francke</collectorName>
,
<collectorName id="26965339A24CFFB5FB2BFD15C685FCCE" box="[1169,1273,748,770]" pageId="25" pageNumber="852">A. Valdez</collectorName>
,
<collectorName id="26965339A24CFFB5FAB9FD15C7E1FCCE" box="[1283,1437,748,770]" pageId="25" pageNumber="852">C. Santibáñez</collectorName>
,
<collectorName id="26965339A24CFFB5FC80FCF2C1E4FCEC" box="[826,920,779,800]" pageId="25" pageNumber="852">J. Cruz</collectorName>
and
<collectorName id="26965339A24CFFB5FC5AFCF2C6F6FCED" box="[992,1162,778,801]" pageId="25" pageNumber="852">G. Contreras</collectorName>
)], (
<geoCoordinate id="EE575028A24CFFB5FB0EFCF2C73EFCEC" box="[1204,1346,778,800]" degrees="21" direction="north" minutes="37" orientation="latitude" pageId="25" pageNumber="852" precision="15" seconds="48" value="21.630001">21°3748″N</geoCoordinate>
,
<geoCoordinate id="EE575028A24CFFB5FAEFFCF2C1DFFCF3" degrees="99" direction="west" minutes="03" orientation="longitude" pageId="25" pageNumber="852" precision="1" seconds="38.73" value="-99.06076">99°03 38.73″W</geoCoordinate>
).
<collectorName id="26965339A24CFFB5FC06FCD0C6C0FCF3" box="[956,1212,809,831]" pageId="25" pageNumber="852">Municipio Aquismón</collectorName>
,
<location id="8EBC6034A24CFFB5FB77FCD0C7E0FCF3" LSID="urn:lsid:plazi:treatment:03CA87F9A24CFFB7FC65FECBC061FBA0:8EBC6034A24CFFB5FB77FCD0C7E0FCF3" box="[1229,1436,809,831]" country="Mexico" latitude="21.630001" longLatPrecision="15" longitude="-99.06076" name="Cueva de El Sol" pageId="25" pageNumber="852" stateProvince="San Luis Potosi">Cueva de El Sol</location>
,
<collectorName id="26965339A24CFFB5FC80FCB1C1B4FC92" box="[826,968,840,862]" pageId="25" pageNumber="852">Mantetzulel</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDC36EFA24CFFB5FC80FC7EC1E9FC16" blockId="25.[826,1441,903,986]" pageId="25" pageNumber="852">
<emphasis id="B917EAFDA24CFFB5FC80FC7EC1C6FC50" box="[826,954,903,924]" italics="true" pageId="25" pageNumber="852">Etymology:</emphasis>
From the Latin
<taxonomicName id="4C634D6CA24CFFB5FB34FC71C6A2FC51" authorityName="Cruz-López &amp; Francke" authorityYear="2015" box="[1166,1246,904,925]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="25" pageNumber="852" phylum="Arthropoda" rank="species" species="tersum">
<emphasis id="B917EAFDA24CFFB5FB34FC71C6A2FC51" box="[1166,1246,904,925]" italics="true" pageId="25" pageNumber="852">tersum</emphasis>
</taxonomicName>
(smooth), referring to the reduction in dorsal ornamentation of the scutum.
</paragraph>
<paragraph id="8BDC36EFA24CFFB6FC80FBFCC0E0F91A" blockId="25.[826,1441,1028,1173]" lastBlockId="26.[144,759,1483,1750]" lastPageId="26" lastPageNumber="853" pageId="25" pageNumber="852">
<emphasis id="B917EAFDA24CFFB5FC80FBFCC1C7FBD6" box="[826,955,1029,1050]" italics="true" pageId="25" pageNumber="852">Diagnosis:</emphasis>
Troglomorphic species. Sexual dimorphism of scutum unremarkable (
<figureCitation id="13582A6AA24CFFB5FB01FBDAC762FBF5" box="[1211,1310,1059,1081]" captionStart="Figure 21" captionStartId="25.[164,243,1853,1872]" captionTargetBox="[166,1441,1226,1819]" captionTargetId="figure-432@25.[164,1444,1223,1823]" captionTargetPageId="25" captionText="Figure 21. Karos tersum sp. nov. A, habitus, dorsal view. B, scutum, dorsal view. C, habitus, lateral view. Scale bars: A = 3.8 mm, B = 1.5 mm, C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340017" httpUri="https://zenodo.org/record/5340017/files/figure.png" pageId="25" pageNumber="852">Fig. 21B</figureCitation>
). Tubercles of transversal rows
<collectionCode id="ED72AE2AA24CFFB5FBA5FBBBC641FB9B" box="[1055,1085,1090,1111]" pageId="25" pageNumber="852">III</collectionCode>
and
<collectionCode id="ED72AE2AA24CFFB5FBC2FBBBC6E8FB9B" box="[1144,1172,1090,1111]" pageId="25" pageNumber="852">IV</collectionCode>
very small, mesotergal area
<collectionCode id="ED72AE2AA24CFFB5FCC2FB99C1F0FBB9" box="[888,908,1120,1141]" pageId="25" pageNumber="852">II</collectionCode>
with only a few central tubercles (
<figureCitation id="13582A6AA24CFFB5FA8FFB99C7E1FBB9" box="[1333,1437,1120,1142]" captionStart="Figure 21" captionStartId="25.[164,243,1853,1872]" captionTargetBox="[166,1441,1226,1819]" captionTargetId="figure-432@25.[164,1444,1223,1823]" captionTargetPageId="25" captionText="Figure 21. Karos tersum sp. nov. A, habitus, dorsal view. B, scutum, dorsal view. C, habitus, lateral view. Scale bars: A = 3.8 mm, B = 1.5 mm, C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340017" httpUri="https://zenodo.org/record/5340017/files/figure.png" pageId="25" pageNumber="852">Figs 21C</figureCitation>
,
<figureCitation id="13582A6AA24CFFB5FC80FB86C117FB59" box="[826,875,1151,1173]" captionStart="Figure 63" captionStartId="60.[144,223,1276,1295]" captionTargetBox="[144,1425,196,1246]" captionTargetId="figure-0@60.[144,1425,196,1246]" captionTargetPageId="60" captionText="Figure 63. A, dorsal view of the anal bulge on Chapulobunus poblano sp. nov.; arrow indicates the bulge. B, lateral view of Huasteca silhavyi sp. nov., line indicates the very high lateral channel area. C, laterodorsal view of scutum of Crettaros valdezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. D, posterodorsal view of scutum of Crettaros santibanezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. E, dorsal view of trochanter IV of Karos barbarikos; arrow indicates the tridentate apophysis. F, anterodorsal view of scutum of Karos barbarikos, the black points mark the tubercles forming a V. G, dorsal ornamentation of Karos tersum sp. nov. showing the transversal row of tubercles on mesotergal areas; transversal row II indicated by black dots. H, detail of the ventral ornamentation on femur IV of Karos barbarikos. I, detail of the ectal ornamentation on tibia IV of Karos barbarikos; arrows indicate the spiniform tubercles. J, detail of the ventral ornamentation on leg IV in Chapulobunus poblano sp. nov. K, detail of the two apical spines on femur IV in Chapulobunus poblano sp. nov. These images are not at the same scale." figureDoi="http://doi.org/10.5281/zenodo.5340103" httpUri="https://zenodo.org/record/5340103/files/figure.png" pageId="25" pageNumber="852">63G</figureCitation>
). Spines of the ocularium small, close to each other but not fused at their bases (
<figureCitation id="13582A6AA24FFFB6FE5AFA32C038FA2D" box="[480,580,1483,1505]" captionStart="Figure 22" captionStartId="26.[144,223,624,643]" captionTargetBox="[305,1261,200,591]" captionTargetId="figure-222@26.[303,1264,197,594]" captionTargetPageId="26" captionText="Figure 22. Karos tersum sp. nov. A, legs IV, mesal view. B, habitus, ventral view. C, ocularium, frontal view. Scale bars: A = 3.4 mm, B = 2.0 mm, C = 0.5 mm. The dark lines on (B) indicate the stigmatic area." figureDoi="http://doi.org/10.5281/zenodo.5340019" httpUri="https://zenodo.org/record/5340019/files/figure.png" pageId="26" pageNumber="853">Fig. 22C</figureCitation>
). Sexual length of femur ratio
<collectionCode id="ED72AE2AA24FFFB6FE89FA13C335FA33" box="[307,329,1514,1535]" pageId="26" pageNumber="853">II</collectionCode>
: 1.42 and
<collectionCode id="ED72AE2AA24FFFB6FE07FA13C3A6FA33" box="[445,474,1514,1535]" pageId="26" pageNumber="853">IV</collectionCode>
: 1.47. Metatarsus
<collectionCode id="ED72AE2AA24FFFB6FD13FA13C0C1FA33" box="[681,701,1514,1535]" pageId="26" pageNumber="853">II</collectionCode>
with three clear annuli in males, two in females. Dorsoectal apophyses of coxa
<collectionCode id="ED72AE2AA24FFFB6FED0F9DEC3FAF9F0" box="[362,390,1575,1596]" pageId="26" pageNumber="853">IV</collectionCode>
present, increasing in size distally, similar in both sexes. Male genitalia: apical margin convex. Lateral setae with cylindrical bases, flattened distally. Two pairs of parastylar setae, lateral to follis, basal pair slightly separated from follis base. Spiniform projections barely visible (
<figureCitation id="13582A6AA24FFFB6FD84F939C0F3F91A" box="[574,655,1728,1750]" captionStart="Figure 23" captionStartId="26.[144,223,1365,1384]" captionTargetBox="[144,1425,720,1336]" captionTargetId="figure-265@26.[144,1425,720,1336]" captionTargetPageId="26" captionText="Figure 23. Karos tersum sp. nov. male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C)." figureDoi="http://doi.org/10.5281/zenodo.5340021" httpUri="https://zenodo.org/record/5340021/files/figure.png" pageId="26" pageNumber="853">Fig. 23</figureCitation>
).
</paragraph>
<caption id="DF1C6667A24CFFB5FF1EF8C4C05CF8A2" ID-DOI="http://doi.org/10.5281/zenodo.5340017" ID-Zenodo-Dep="5340017" httpUri="https://zenodo.org/record/5340017/files/figure.png" pageId="25" pageNumber="852" startId="25.[164,243,1853,1872]" targetBox="[166,1441,1226,1819]" targetPageId="25">
<paragraph id="8BDC36EFA24CFFB5FF1EF8C4C05CF8A2" blockId="25.[164,1442,1853,1902]" pageId="25" pageNumber="852">
<emphasis id="B917EAFDA24CFFB5FF1EF8C4C05DF89C" bold="true" box="[164,545,1853,1873]" pageId="25" pageNumber="852">
Figure 21.
<taxonomicName id="4C634D6CA24CFFB5FE9EF8C7C3BDF89D" authority="Cruz-López &amp; Francke, 2015" authorityName="Cruz-López &amp; Francke" authorityYear="2015" box="[292,449,1854,1873]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="25" pageNumber="852" phylum="Arthropoda" rank="species" species="tersum" status="sp. nov.">
<emphasis id="B917EAFDA24CFFB5FE9EF8C7C3BDF89D" bold="true" box="[292,449,1854,1873]" italics="true" pageId="25" pageNumber="852">Karos tersum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2245786A24CFFB5FE73F8C4C05DF89C" box="[457,545,1853,1872]" pageId="25" pageNumber="852" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, habitus, dorsal view. B, scutum, dorsal view. C, habitus, lateral view. Scale bars: A = 3.8 mm, B = 1.5 mm, C = 1 mm.
</paragraph>
</caption>
<caption id="DF1C6667A24FFFB6FF2AFD89C607FD6D" ID-DOI="http://doi.org/10.5281/zenodo.5340019" ID-Zenodo-Dep="5340019" httpUri="https://zenodo.org/record/5340019/files/figure.png" pageId="26" pageNumber="853" startId="26.[144,223,624,643]" targetBox="[305,1261,200,591]" targetPageId="26">
<paragraph id="8BDC36EFA24FFFB6FF2AFD89C607FD6D" blockId="26.[144,1421,624,673]" pageId="26" pageNumber="853">
<emphasis id="B917EAFDA24FFFB6FF2AFD89C06DFD4F" bold="true" box="[144,529,624,643]" pageId="26" pageNumber="853">
Figure 22.
<taxonomicName id="4C634D6CA24FFFB6FEAAFD89C3D2FD4F" authority="Cruz-López &amp; Francke, 2015" authorityName="Cruz-López &amp; Francke" authorityYear="2015" box="[272,430,624,643]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="26" pageNumber="853" phylum="Arthropoda" rank="species" species="tersum" status="sp. nov.">
<emphasis id="B917EAFDA24FFFB6FEAAFD89C3D2FD4F" bold="true" box="[272,430,624,643]" italics="true" pageId="26" pageNumber="853">Karos tersum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2245786A24FFFB6FE02FD89C06DFD4F" box="[440,529,624,643]" pageId="26" pageNumber="853" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, legs IV, mesal view. B, habitus, ventral view. C, ocularium, frontal view. Scale bars: A = 3.4 mm, B = 2.0 mm, C = 0.5 mm. The dark lines on (B) indicate the stigmatic area.
</paragraph>
</caption>
<caption id="DF1C6667A24FFFB6FF2AFAACC17FFA4A" ID-DOI="http://doi.org/10.5281/zenodo.5340021" ID-Zenodo-Dep="5340021" httpUri="https://zenodo.org/record/5340021/files/figure.png" pageId="26" pageNumber="853" startId="26.[144,223,1365,1384]" targetBox="[144,1425,720,1336]" targetPageId="26">
<paragraph id="8BDC36EFA24FFFB6FF2AFAACC17FFA4A" blockId="26.[144,1421,1365,1414]" pageId="26" pageNumber="853">
<emphasis id="B917EAFDA24FFFB6FF2AFAACC06CFAA5" bold="true" box="[144,528,1365,1385]" pageId="26" pageNumber="853">
Figure 23.
<taxonomicName id="4C634D6CA24FFFB6FEAAFAAFC3D1FAA5" authority="Cruz-López &amp; Francke, 2015" authorityName="Cruz-López &amp; Francke" authorityYear="2015" box="[272,429,1366,1385]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="26" pageNumber="853" phylum="Arthropoda" rank="species" species="tersum" status="sp. nov.">
<emphasis id="B917EAFDA24FFFB6FEAAFAAFC3D1FAA5" bold="true" box="[272,429,1366,1385]" italics="true" pageId="26" pageNumber="853">Karos tersum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2245786A24FFFB6FE0DFAAFC06CFAA5" box="[439,528,1366,1385]" pageId="26" pageNumber="853" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C).
</paragraph>
</caption>
<paragraph id="8BDC36EFA24FFFB7FF2AF907C2B1FD44" blockId="26.[144,759,1789,1903]" lastBlockId="27.[164,779,198,648]" lastPageId="27" lastPageNumber="854" pageId="26" pageNumber="853">
<emphasis id="B917EAFDA24FFFB6FF2AF907C3A8F8DE" box="[144,468,1789,1811]" italics="true" pageId="26" pageNumber="853">
Description: Male
<typeStatus id="54D8884DA24FFFB6FED7F904C3B2F8DE" box="[365,462,1789,1810]" pageId="26" pageNumber="853" type="holotype">holotype</typeStatus>
:
</emphasis>
Measurements: scutum length 3.3, maximum scutum width 2.7. Legs. I 2.00/ 0.75/1.55/2.10/,
<collectionCode id="ED72AE2AA24FFFB6FEF6F8C2C31CF89C" box="[332,352,1851,1872]" pageId="26" pageNumber="853">II</collectionCode>
3.55/1.35/3.20/3.50,
<collectionCode id="ED72AE2AA24FFFB6FDE1F8C2C006F89C" box="[603,634,1851,1872]" pageId="26" pageNumber="853">III</collectionCode>
2.45/0.80/ 2.00/2.55,
<collectionCode id="ED72AE2AA24FFFB6FEBBF8A0C360F8A2" box="[257,284,1881,1902]" pageId="26" pageNumber="853">IV</collectionCode>
3.90/1.25/3.45/3.80.
<emphasis id="B917EAFDA24FFFB6FDBAF8A0C024F8A2" box="[512,600,1881,1902]" italics="true" pageId="26" pageNumber="853">Dorsum</emphasis>
. Eyes slightly pigmented (
<figureCitation id="13582A6AA24FFFB6FC15FA32C66DFA2D" box="[943,1041,1483,1505]" captionStart="Figure 21" captionStartId="25.[164,243,1853,1872]" captionTargetBox="[166,1441,1226,1819]" captionTargetId="figure-432@25.[164,1444,1223,1823]" captionTargetPageId="25" captionText="Figure 21. Karos tersum sp. nov. A, habitus, dorsal view. B, scutum, dorsal view. C, habitus, lateral view. Scale bars: A = 3.8 mm, B = 1.5 mm, C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340017" httpUri="https://zenodo.org/record/5340017/files/figure.png" pageId="26" pageNumber="853">Fig. 21B</figureCitation>
). Dorsum almost smooth, area
<collectionCode id="ED72AE2AA24FFFB6FAC3FA32C7F1FA2C" box="[1401,1421,1483,1504]" pageId="26" pageNumber="853">II</collectionCode>
with only a few, small central tubercles, complete rows only on areas
<collectionCode id="ED72AE2AA24FFFB6FC74F9F0C190F9D2" box="[974,1004,1545,1566]" pageId="26" pageNumber="853">III</collectionCode>
and
<collectionCode id="ED72AE2AA24FFFB6FB91F9F0C63BF9D2" box="[1067,1095,1545,1566]" pageId="26" pageNumber="853">IV</collectionCode>
, tubercles small, except the central, which is spiniform (
<figureCitation id="13582A6AA24FFFB6FB29F9DEC77CF9F1" box="[1171,1280,1575,1597]" captionStart="Figure 63" captionStartId="60.[144,223,1276,1295]" captionTargetBox="[144,1425,196,1246]" captionTargetId="figure-0@60.[144,1425,196,1246]" captionTargetPageId="60" captionText="Figure 63. A, dorsal view of the anal bulge on Chapulobunus poblano sp. nov.; arrow indicates the bulge. B, lateral view of Huasteca silhavyi sp. nov., line indicates the very high lateral channel area. C, laterodorsal view of scutum of Crettaros valdezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. D, posterodorsal view of scutum of Crettaros santibanezi sp. nov., showing the median humps on areas II and III indicated by dotted circles. E, dorsal view of trochanter IV of Karos barbarikos; arrow indicates the tridentate apophysis. F, anterodorsal view of scutum of Karos barbarikos, the black points mark the tubercles forming a V. G, dorsal ornamentation of Karos tersum sp. nov. showing the transversal row of tubercles on mesotergal areas; transversal row II indicated by black dots. H, detail of the ventral ornamentation on femur IV of Karos barbarikos. I, detail of the ectal ornamentation on tibia IV of Karos barbarikos; arrows indicate the spiniform tubercles. J, detail of the ventral ornamentation on leg IV in Chapulobunus poblano sp. nov. K, detail of the two apical spines on femur IV in Chapulobunus poblano sp. nov. These images are not at the same scale." figureDoi="http://doi.org/10.5281/zenodo.5340103" httpUri="https://zenodo.org/record/5340103/files/figure.png" pageId="26" pageNumber="853">Fig. 63G</figureCitation>
). Spines of ocularium short, close to each other (
<figureCitation id="13582A6AA24FFFB6FB7CF9BCC754F997" box="[1222,1320,1605,1627]" captionStart="Figure 22" captionStartId="26.[144,223,624,643]" captionTargetBox="[305,1261,200,591]" captionTargetId="figure-222@26.[303,1264,197,594]" captionTargetPageId="26" captionText="Figure 22. Karos tersum sp. nov. A, legs IV, mesal view. B, habitus, ventral view. C, ocularium, frontal view. Scale bars: A = 3.4 mm, B = 2.0 mm, C = 0.5 mm. The dark lines on (B) indicate the stigmatic area." figureDoi="http://doi.org/10.5281/zenodo.5340019" httpUri="https://zenodo.org/record/5340019/files/figure.png" pageId="26" pageNumber="853">Fig. 22C</figureCitation>
). Lateral clear areas on scutum triangular-shaped, clear areas on mesotergal area
<collectionCode id="ED72AE2AA24FFFB6FBA8F97AC658F954" box="[1042,1060,1667,1688]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="26" pageNumber="853" type="Museum">V</collectionCode>
apices and free tergites I and
<collectionCode id="ED72AE2AA24FFFB6FC9CF95BC141F97B" box="[806,829,1698,1719]" pageId="26" pageNumber="853">II</collectionCode>
, small, spiniform.
<emphasis id="B917EAFDA24FFFB6FB91F95BC605F97B" box="[1067,1145,1698,1719]" italics="true" pageId="26" pageNumber="853">Venter.</emphasis>
Densely covered with spiniform tubercles. Row of tubercles on coxa I longer than those of the other coxae. Coxa
<collectionCode id="ED72AE2AA24FFFB6FB6AF926C690F938" box="[1232,1260,1759,1780]" pageId="26" pageNumber="853">IV</collectionCode>
most densely covered by tubercles. Dorsoectal apophyses of coxa increasing in size distally.
<emphasis id="B917EAFDA24FFFB6FBF0F8E2C6BDF8FC" box="[1098,1217,1819,1840]" italics="true" pageId="26" pageNumber="853">Pedipalps.</emphasis>
Patella with two mesodistal tubercles.
<emphasis id="B917EAFDA24FFFB6FBA5F8C2C626F89C" box="[1055,1114,1851,1872]" italics="true" pageId="26" pageNumber="853">Legs.</emphasis>
Ornamentation similar on all legs, covered by small spiniform tubercles (
<figureCitation id="13582A6AA24FFFB6FA99F8A0C7FFF8A3" box="[1315,1411,1881,1903]" captionStart="Figure 22" captionStartId="26.[144,223,624,643]" captionTargetBox="[305,1261,200,591]" captionTargetId="figure-222@26.[303,1264,197,594]" captionTargetPageId="26" captionText="Figure 22. Karos tersum sp. nov. A, legs IV, mesal view. B, habitus, ventral view. C, ocularium, frontal view. Scale bars: A = 3.4 mm, B = 2.0 mm, C = 0.5 mm. The dark lines on (B) indicate the stigmatic area." figureDoi="http://doi.org/10.5281/zenodo.5340019" httpUri="https://zenodo.org/record/5340019/files/figure.png" pageId="26" pageNumber="853">Fig. 22A</figureCitation>
). Posterior legs slightly thicker than anterior ones. Metatarsus
<collectionCode id="ED72AE2AA24EFFB7FF48FF1CC37AFF36" box="[242,262,229,250]" pageId="27" pageNumber="854">II</collectionCode>
with three inconspicuous clear annuli. Femur
<collectionCode id="ED72AE2AA24EFFB7FF1EFEFAC2BEFED4" box="[164,194,259,280]" pageId="27" pageNumber="854">III</collectionCode>
curved. Tarsal count 4(2):8(3):6:6.
<emphasis id="B917EAFDA24EFFB7FDE7FEFAC177FED4" box="[605,779,259,280]" italics="true" pageId="27" pageNumber="854">Male genitalia.</emphasis>
Apical margin convex. Lateral setae basally cylindrical, distally flattened, basal pair small and ventrally displaced. Parastylar setae lateral to follis. Spiniform projections not exposed, but visible (
<figureCitation id="13582A6AA24EFFB7FDEAFE84C0DEFE58" box="[592,674,381,404]" captionStart="Figure 23" captionStartId="26.[144,223,1365,1384]" captionTargetBox="[144,1425,720,1336]" captionTargetId="figure-265@26.[144,1425,720,1336]" captionTargetPageId="26" captionText="Figure 23. Karos tersum sp. nov. male genitalia. A, dorsal view. B, ventral view. C, lateral view. White arrows indicate ventral microsetae in (B) and parastylar setae in (C)." figureDoi="http://doi.org/10.5281/zenodo.5340021" httpUri="https://zenodo.org/record/5340021/files/figure.png" pageId="27" pageNumber="854">Fig. 23</figureCitation>
).
<emphasis id="B917EAFDA24EFFB7FD02FE84C377FE7E" italics="true" pageId="27" pageNumber="854">
Female
<typeStatus id="54D8884DA24EFFB7FF1EFE64C379FE7E" box="[164,261,413,434]" pageId="27" pageNumber="854" type="paratype">paratype</typeStatus>
:
</emphasis>
Measurements: scutum length 3.2, maximum scutum width 2.5. Similar to male, with the following differences: mid-bulge shorter, giving scutum appearance of being slightly rectangular, sexual proportion of femora to metatarsi:
<collectionCode id="ED72AE2AA24EFFB7FE07FDEEC3ADFDE0" box="[445,465,535,556]" pageId="27" pageNumber="854">II</collectionCode>
1.42/1.42/1.56/1.52 and
<collectionCode id="ED72AE2AA24EFFB7FD55FDEEC177FDE0" box="[751,779,535,556]" pageId="27" pageNumber="854">IV</collectionCode>
1.47/1.78/1.56/1.40 (
<figureCitation id="13582A6AA24EFFB7FE38FDCCC39BFD86" box="[386,487,565,587]" captionStart="Figure 21" captionStartId="25.[164,243,1853,1872]" captionTargetBox="[166,1441,1226,1819]" captionTargetId="figure-432@25.[164,1444,1223,1823]" captionTargetPageId="25" captionText="Figure 21. Karos tersum sp. nov. A, habitus, dorsal view. B, scutum, dorsal view. C, habitus, lateral view. Scale bars: A = 3.8 mm, B = 1.5 mm, C = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5340017" httpUri="https://zenodo.org/record/5340017/files/figure.png" pageId="27" pageNumber="854">Figs 21A</figureCitation>
,
<figureCitation id="13582A6AA24EFFB7FE49FDCCC05FFD87" box="[499,547,565,587]" captionStart="Figure 22" captionStartId="26.[144,223,624,643]" captionTargetBox="[305,1261,200,591]" captionTargetId="figure-222@26.[303,1264,197,594]" captionTargetPageId="26" captionText="Figure 22. Karos tersum sp. nov. A, legs IV, mesal view. B, habitus, ventral view. C, ocularium, frontal view. Scale bars: A = 3.4 mm, B = 2.0 mm, C = 0.5 mm. The dark lines on (B) indicate the stigmatic area." figureDoi="http://doi.org/10.5281/zenodo.5340019" httpUri="https://zenodo.org/record/5340019/files/figure.png" pageId="27" pageNumber="854">22A</figureCitation>
). Metatarsus
<collectionCode id="ED72AE2AA24EFFB7FD07FDCCC0ADFD86" box="[701,721,565,586]" pageId="27" pageNumber="854">II</collectionCode>
with two inconspicuous clear annuli. Tarsal count 4(2):7(3):5 6:6.
</paragraph>
<paragraph id="8BDC36EFA24EFFB7FF1EFD45C36AFCE2" blockId="27.[164,779,700,814]" pageId="27" pageNumber="854">
<emphasis id="B917EAFDA24EFFB7FF1EFD45C321FD1D" box="[164,349,700,721]" italics="true" pageId="27" pageNumber="854">Natural history:</emphasis>
The female collected in Cueva de El Sol, Mantetzulel did not exhibit thanatosis behaviour, unlike other stygnopsids (
<bibRefCitation id="EFF24B1EA24EFFB7FDB8FD00C297FCE1" author="Cruz-Lopez JA &amp; Francke OF" pageId="27" pageNumber="854" pagination="481 - 490" refId="ref37717" refString="Cruz-Lopez JA, Francke OF. 2013 a. Two new species of the genus Paramitraceras Pickard-Cambridge, 1905 (Opiliones: Laniatores: Stygnopsidae) from Chiapas, Mexico. Zootaxa 3641: 481 - 490." type="journal article" year="2013">Cruz-López &amp; Francke, 2013a</bibRefCitation>
, b).
</paragraph>
<paragraph id="8BDC36EFA24EFFB7FF1EFC9BC061FBA0" blockId="27.[164,779,865,1132]" pageId="27" pageNumber="854">
<emphasis id="B917EAFDA24EFFB7FF1EFC9BC3FFFCBB" box="[164,387,866,887]" italics="true" pageId="27" pageNumber="854">Taxonomic accounts:</emphasis>
<bibRefCitation id="EFF24B1EA24EFFB7FE30FC98C0A4FCBB" author="Goodnight CJ &amp; Goodnight ML" box="[394,728,865,887]" pageId="27" pageNumber="854" pagination="33 - 45" refId="ref38192" refString="Goodnight CJ, Goodnight ML. 1971. Opilionids (Phalangida) of the family Phalangodidae from Mexican caves. Association for Mexican Cave Studies, Bulletin 4: 33 - 45." type="journal article" year="1971">Goodnight &amp; Goodnight (1971)</bibRefCitation>
designated some specimens as
<typeStatus id="54D8884DA24EFFB7FE65FC79C033FC59" box="[479,591,896,917]" pageId="27" pageNumber="854" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C634D6CA24EFFB7FDCEFC79C177FC59" authorityName=": Goodnight &amp; Goodnight" authorityYear="1971" box="[628,779,896,917]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="854" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B917EAFDA24EFFB7FDCEFC79C177FC59" box="[628,779,896,917]" italics="true" pageId="27" pageNumber="854">Karos parvus</emphasis>
</taxonomicName>
from localities other than the type locality, which proved to be incorrect.
<taxonomicName id="4C634D6CA24EFFB7FEE6FC47C38BFC1F" authority="Cruz-López &amp; Francke, 2015" authorityName="Cruz-López &amp; Francke" authorityYear="2015" box="[348,503,958,979]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="854" phylum="Arthropoda" rank="species" species="tersum" status="sp. nov.">
<emphasis id="B917EAFDA24EFFB7FEE6FC47C38BFC1F" box="[348,503,958,979]" italics="true" pageId="27" pageNumber="854">Karos tersum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2245786A24EFFB7FDBBFC44C024FC1E" box="[513,600,957,978]" pageId="27" pageNumber="854" rank="species">sp. nov.</taxonomicNameLabel>
is one of these incorrectly paratyped samples, and it is clearly different from
<taxonomicName id="4C634D6CA24EFFB7FEF3FC02C389FBDC" authorityName=": Goodnight &amp; Goodnight" authorityYear="1971" box="[329,501,1019,1040]" class="Arachnida" family="Stygnopsidae" genus="Karos" kingdom="Animalia" order="Opiliones" pageId="27" pageNumber="854" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B917EAFDA24EFFB7FEF3FC02C389FBDC" box="[329,501,1019,1040]" italics="true" pageId="27" pageNumber="854">Karos parvus</emphasis>
</taxonomicName>
. The new species is troglomorphic and quite pale in colour, has an almost smooth dorsum, and has the spines of the ocularium close to each other at the bases.
</paragraph>
</subSubSection>
</treatment>
</document>