187 lines
20 KiB
XML
187 lines
20 KiB
XML
<document id="882C0CAAFBDA81BE8A292B95C55E327B" ID-CLB-Dataset="4244" ID-DOI="10.11646/palaeoentomology.4.4.8" ID-GBIF-Dataset="cec45842-65e1-491b-b5db-054def0f46a8" ID-ISSN="2624-2834" ID-Zenodo-Dep="5507996" ID-ZooBank="E15981ED-8CDE-4815-9466-25AC86D0356F" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631646286413" checkinUser="carolina" docAuthor="WICHARD, WILFRIED, MÜLLER, PATRICK & XU, CHUN-PENG" docDate="2021" docId="03BE2125FFE3560FFF65C84BFD9EF93E" docLanguage="en" docName="palaeoentomology.4.4.8.pdf" docOrigin="Palaeoentomology 4 (4)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.4.8" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Palaeopsilotreta Wichard & Wang 2017" docType="treatment" docVersion="6" lastPageNumber="334" masterDocId="FF87595DFFE0560CFFEDC919FFFBFFFA" masterDocTitle="The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber" masterLastPageNumber="338" masterPageNumber="331" pageNumber="334" updateTime="1699448848240" updateUser="ExternalLinkService">
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<mods:title id="A712A40ADDD15A295A212F4C34821C20">The odontocerid genera Psilotreta and Palaeopsilotreta (Insecta, Trichoptera) embedded in mid-Cretaceous Burmese amber</mods:title>
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<mods:namePart id="74B08C985D3F499FDD2C86A946D21450">WICHARD, WILFRIED</mods:namePart>
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<mods:namePart id="23CB507D35562A8ECFA0619D6E96DDEA">MÜLLER, PATRICK</mods:namePart>
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<mods:roleTerm id="BA05A7A103BF1337BBE96DEE2A9A720B">Author</mods:roleTerm>
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<mods:namePart id="60814467CCB4CA17D46DB05CC5719A70">XU, CHUN-PENG</mods:namePart>
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<mods:typeOfResource id="659D425FCAFC5C4ED0FB4BA053DEBBC2">text</mods:typeOfResource>
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<mods:title id="6BF7D1DE88B86D8D203F3553FF6386A4">Palaeoentomology</mods:title>
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<mods:part id="98553A54D290EBD08E610DD30D547D82">
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<mods:date id="860B1984CC5CC58CFE9BF6F57CBA3D5E">2021</mods:date>
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<mods:number id="F0DEBE9A0AD49A63D41FA829EB3E1708">2021-08-31</mods:number>
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<mods:url id="1717883ED8E4C02E61E30BEC645F6F4A">http://dx.doi.org/10.11646/palaeoentomology.4.4.8</mods:url>
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<treatment id="03BE2125FFE3560FFF65C84BFD9EF93E" ID-DOI="http://doi.org/10.5281/zenodo.5508008" ID-GBIF-Taxon="188190116" ID-Zenodo-Dep="5508008" LSID="urn:lsid:plazi:treatment:03BE2125FFE3560FFF65C84BFD9EF93E" httpUri="http://treatment.plazi.org/id/03BE2125FFE3560FFF65C84BFD9EF93E" lastPageNumber="334" pageId="3" pageNumber="334">
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<subSubSection id="C30DC3B8FFE3560FFF65C84BFD4CFE96" box="[136,695,338,365]" pageId="3" pageNumber="334" type="nomenclature">
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<paragraph id="8BA89033FFE3560FFF65C84BFD4CFE96" blockId="3.[136,695,338,365]" box="[136,695,338,365]" pageId="3" pageNumber="334">
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<heading id="D0E0275FFFE3560FFF65C84BFD4CFE96" box="[136,695,338,365]" fontSize="10" level="2" pageId="3" pageNumber="334" reason="2">
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<emphasis id="B9634C21FFE3560FFF65C84BFD4CFE96" bold="true" box="[136,695,338,365]" pageId="3" pageNumber="334">
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Genus
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<taxonomicName id="4C17EBB0FFE3560FFF36C84BFD4CFE96" authority="Wichard & Wang, 2017" authorityName="Wichard & Wang" authorityYear="2017" box="[219,695,338,365]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
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<emphasis id="B9634C21FFE3560FFF36C84BFE6DFE97" bold="true" box="[219,406,338,365]" italics="true" pageId="3" pageNumber="334">Palaeopsilotreta</emphasis>
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<bibRefCitation id="EF86EDC2FFE3560FFE70C84BFD4CFE96" author="Wichard, W. & Wang, B." box="[413,695,338,364]" pageId="3" pageNumber="334" pagination="198 - 203" refId="ref4454" refString="Wichard, W. & Wang, B. (2017) A remarkable caddisfly with bipectinate antennae in Cretaceous Burmese amber (Insecta, Trichoptera). Cretaceous Research, 69, 198 - 203. https: // doi. org / 10.1016 / j. cretres. 2016.09.012" type="journal article" year="2017">Wichard & Wang, 2017</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30DC3B8FFE3560FFF65C883FF13FB53" pageId="3" pageNumber="334" type="type_taxon">
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<paragraph id="8BA89033FFE3560FFF65C883FF3BFE22" blockId="3.[136,778,410,1732]" pageId="3" pageNumber="334">
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<emphasis id="B9634C21FFE3560FFF65C883FED2FE4E" bold="true" box="[136,297,410,436]" pageId="3" pageNumber="334">
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<typeStatus id="54AC2E91FFE3560FFF65C883FF38FE4E" box="[136,195,410,436]" pageId="3" pageNumber="334">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C17EBB0FFE3560FFEDBC883FF3BFE22" authority="Wichard & Wang, 2017" authorityName="Wichard & Wang" authorityYear="2017" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="xiai">
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<emphasis id="B9634C21FFE3560FFEDBC883FDDCFE4E" box="[310,551,410,436]" italics="true" pageId="3" pageNumber="334">Palaeopsilotreta xiai</emphasis>
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<bibRefCitation id="EF86EDC2FFE3560FFDD9C883FF3BFE22" author="Wichard, W. & Wang, B." pageId="3" pageNumber="334" pagination="198 - 203" refId="ref4454" refString="Wichard, W. & Wang, B. (2017) A remarkable caddisfly with bipectinate antennae in Cretaceous Burmese amber (Insecta, Trichoptera). Cretaceous Research, 69, 198 - 203. https: // doi. org / 10.1016 / j. cretres. 2016.09.012" type="journal article" year="2017">Wichard & Wang, 2017</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="8BA89033FFE3560FFF50C8FBFF13FB53" blockId="3.[136,778,410,1732]" pageId="3" pageNumber="334">
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<emphasis id="B9634C21FFE3560FFF50C8FBFE62FE06" bold="true" box="[189,409,482,508]" pageId="3" pageNumber="334">Revised diagnosis.</emphasis>
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Head: Male antennae (partially) bipectinate,about as long as forewings.Rami of bipectinate flagellomeres slim and about as long as flagellomeres. Female antennae simple, not bipectinate. Maxillary palps five-segmented, terminal segment not flexible or annulate. Ocelli absent. Frontal, antennal and posterior setose warts present. Thorax: Pronotum with two pronotal setose warts, mesoscutum bearing pair of setose warts, mesoscutellum covered by a single wart, large, domelike and ovoid. Tibial spurs: 2/4/4. Wings: In fore- and hindwings forks I, II, (III) and V present; discoidal cells open or closed, medial cell absent. In forewings media M variable: Simple (M
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<subScript id="17939276FFE3560FFE82CABCFE56FC49" attach="left" box="[367,429,933,947]" fontSize="6" pageId="3" pageNumber="334">1, 2, 3, 4</subScript>
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) or two-branched in M
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<subScript id="17939276FFE3560FFD53CABCFD2CFC49" attach="left" box="[702,727,933,947]" fontSize="6" pageId="3" pageNumber="334">1+2</subScript>
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and M
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<subScript id="17939276FFE3560FFF4CCAD0FF40FC2D" attach="left" box="[161,187,969,983]" fontSize="6" pageId="3" pageNumber="334">3+4</subScript>
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or three-branched in M
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<subScript id="17939276FFE3560FFE2BCAD0FE35FC2D" attach="left" box="[454,462,969,983]" fontSize="6" pageId="3" pageNumber="334">1</subScript>
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, M
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<subScript id="17939276FFE3560FFE1BCAD0FE05FC2D" attach="left" box="[502,510,969,983]" fontSize="6" pageId="3" pageNumber="334">2</subScript>
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and M
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<subScript id="17939276FFE3560FFDBDCAD0FD92FC2D" attach="left" box="[592,617,969,983]" fontSize="6" pageId="3" pageNumber="334">3+4</subScript>
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. Cu
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<subScript id="17939276FFE3560FFD74CAD0FD5AFC2D" attach="left" box="[665,673,969,983]" fontSize="6" pageId="3" pageNumber="334">2</subScript>
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reaching wing margin and not being reduced or absent as in other odontocerid genera. Male genitalia: Inferior appendages two-segmented, harpagoes originating from apex of coxopodites (
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<taxonomicName id="4C17EBB0FFE3560FFECECD5EFE8BFB9A" authorityName="Wichard & Wang" authorityYear="2017" box="[291,368,1095,1121]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="xiai">
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<emphasis id="B9634C21FFE3560FFECECD5EFECFFB9A" box="[291,308,1095,1120]" italics="true" pageId="3" pageNumber="334">P</emphasis>
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.
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<emphasis id="B9634C21FFE3560FFEABCD5EFE8BFB9A" box="[326,368,1095,1120]" italics="true" pageId="3" pageNumber="334">xiai</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C17EBB0FFE3560FFE6FCD5EFDE5FB9A" box="[386,542,1094,1121]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="burmanica">
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<emphasis id="B9634C21FFE3560FFE6FCD5EFE68FB9A" box="[386,403,1095,1120]" italics="true" pageId="3" pageNumber="334">P</emphasis>
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.
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<emphasis id="B9634C21FFE3560FFE48CD5FFDE5FB9A" box="[421,542,1094,1120]" italics="true" pageId="3" pageNumber="334">burmanica</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C17EBB0FFE3560FFDB1CD5EFD25FB9A" box="[604,734,1095,1121]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="cretacea">
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<emphasis id="B9634C21FFE3560FFDB1CD5EFD96FB9A" box="[604,621,1095,1120]" italics="true" pageId="3" pageNumber="334">P</emphasis>
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.
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<emphasis id="B9634C21FFE3560FFD92CD5EFD25FB9A" box="[639,734,1095,1120]" italics="true" pageId="3" pageNumber="334">cretacea</emphasis>
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</taxonomicName>
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) or originating mesally on coxopodites (
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<taxonomicName id="4C17EBB0FFE3560FFDD4CD72FD4BFB7E" authorityName="WICHARD & MÜLLER & XU" authorityYear="2021" box="[569,688,1130,1157]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="kachini">
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<emphasis id="B9634C21FFE3560FFDD4CD72FDB1FB7E" box="[569,586,1131,1156]" italics="true" pageId="3" pageNumber="334">P</emphasis>
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.
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<emphasis id="B9634C21FFE3560FFDB3CD73FD4BFB7E" box="[606,688,1130,1156]" italics="true" pageId="3" pageNumber="334">kachini</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C17EBB0FFE3560FFD1FCD72FF2CFB52" authorityName="WICHARD & MÜLLER & XU" authorityYear="2021" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="species" species="succini">
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<emphasis id="B9634C21FFE3560FFD1FCD72FCF8FB7E" box="[754,771,1131,1156]" italics="true" pageId="3" pageNumber="334">P</emphasis>
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.
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<emphasis id="B9634C21FFE3560FFF65CD96FF2CFB52" box="[136,215,1167,1192]" italics="true" pageId="3" pageNumber="334">succini</emphasis>
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</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C30DC3B8FFE3560FFF50CDABFD9EF93E" pageId="3" pageNumber="334" type="discussion">
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<paragraph id="8BA89033FFE3560FFF50CDABFDB3FA7B" blockId="3.[136,778,410,1732]" pageId="3" pageNumber="334">
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<emphasis id="B9634C21FFE3560FFF50CDABFECAFB36" bold="true" box="[189,305,1202,1228]" pageId="3" pageNumber="334">Remarks.</emphasis>
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The affiliation of the genus
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<taxonomicName id="4C17EBB0FFE3560FFD96CDABFF47FB0A" authorityName="Wichard & Wang" authorityYear="2017" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
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<emphasis id="B9634C21FFE3560FFD96CDABFF47FB0A" italics="true" pageId="3" pageNumber="334">Palaeopsilotreta</emphasis>
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</taxonomicName>
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to the family
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<taxonomicName id="4C17EBB0FFE3560FFE94CDCFFDE5FB0A" authorityName="Wallengren" authorityYear="1891" box="[377,542,1238,1264]" class="Insecta" family="Odontoceridae" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="family">Odontoceridae</taxonomicName>
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was discussed and proposed in
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<bibRefCitation id="EF86EDC2FFE3560FFEF0CDE3FDDAFAEE" author="Wichard, W. & Espeland, M. & Muller, P. & Wang, B." box="[285,545,1274,1301]" pageId="3" pageNumber="334" pagination="1 - 17" refId="ref4505" refString="Wichard, W., Espeland, M., Muller, P. & Wang, B. (2020) New species of caddisflies with bipectinate antennae from Cretaceous Burmese amber (Insecta, Trichoptera: Odontoceratidae, Calamoceratidae). European Journal of Taxonomy, 653, 1 - 17. https: // doi. org / 10.5852 / ejt. 2020.653" type="journal article" year="2020">
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Wichard
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<emphasis id="B9634C21FFE3560FFE64CDE2FE44FAEE" box="[393,447,1274,1300]" italics="true" pageId="3" pageNumber="334">et al</emphasis>
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. (2020)
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</bibRefCitation>
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.
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<taxonomicName id="4C17EBB0FFE3560FFDDCCDE3FD10FAEE" authorityName="Wichard & Wang" authorityYear="2017" box="[561,747,1274,1300]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
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<emphasis id="B9634C21FFE3560FFDDCCDE3FD10FAEE" box="[561,747,1274,1300]" italics="true" pageId="3" pageNumber="334">Palaeopsilotreta</emphasis>
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</taxonomicName>
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is closely related to
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<taxonomicName id="4C17EBB0FFE3560FFEB4CC07FE3DFAC2" authorityName="Banks" authorityYear="1899" box="[345,454,1310,1336]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
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<emphasis id="B9634C21FFE3560FFEB4CC07FE3DFAC2" box="[345,454,1310,1336]" italics="true" pageId="3" pageNumber="334">Psilotreta</emphasis>
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</taxonomicName>
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and is distinguished by the apomorphic, bipectinated antennae in the male, which are not verified in any species of
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<taxonomicName id="4C17EBB0FFE3560FFE3ECC7FFDBBFA7A" authorityName="Banks" authorityYear="1899" box="[467,576,1382,1408]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
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<emphasis id="B9634C21FFE3560FFE3ECC7FFDBBFA7A" box="[467,576,1382,1408]" italics="true" pageId="3" pageNumber="334">Psilotreta</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BA89033FFE3560FFF50CC93FD9EF93E" blockId="3.[136,778,410,1732]" pageId="3" pageNumber="334">
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Based on the presence of five-segmented palpi, the spur formula being 2/4/4, the absence of the medial cell and the presence of wing forks I, II, (III), and V in both wings (as in
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<taxonomicName id="4C17EBB0FFE3560FFE8ECCEFFE2BF9EA" authorityName="Banks" authorityYear="1899" box="[355,464,1526,1552]" class="Insecta" family="Odontoceridae" genus="Psilotreta" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9634C21FFE3560FFE8ECCEFFE2BF9EA" box="[355,464,1526,1552]" italics="true" pageId="3" pageNumber="334">Psilotreta</emphasis>
|
|
</taxonomicName>
|
|
), we can assign the fossil genus
|
|
<taxonomicName id="4C17EBB0FFE3560FFF3ACF03FE6AF9CE" authorityName="Wichard & Wang" authorityYear="2017" box="[215,401,1562,1588]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9634C21FFE3560FFF3ACF03FE6AF9CE" box="[215,401,1562,1588]" italics="true" pageId="3" pageNumber="334">Palaeopsilotreta</emphasis>
|
|
</taxonomicName>
|
|
to
|
|
<taxonomicName id="4C17EBB0FFE3560FFE2CCF03FD91F9CE" authorityName="Wallengren" authorityYear="1891" box="[449,618,1562,1588]" class="Insecta" family="Odontoceridae" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="family">Odontoceridae</taxonomicName>
|
|
. Whether the genus
|
|
<taxonomicName id="4C17EBB0FFE3560FFF38CF27FE74F9A2" authorityName="Wichard & Wang" authorityYear="2017" box="[213,399,1598,1624]" class="Insecta" family="Odontoceridae" genus="Palaeopsilotreta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9634C21FFE3560FFF38CF27FE74F9A2" box="[213,399,1598,1624]" italics="true" pageId="3" pageNumber="334">Palaeopsilotreta</emphasis>
|
|
</taxonomicName>
|
|
is a monophyletic group of the family
|
|
<taxonomicName id="4C17EBB0FFE3560FFF34CF7BFE85F986" authorityName="Wallengren" authorityYear="1891" box="[217,382,1634,1660]" class="Insecta" family="Odontoceridae" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="family">Odontoceridae</taxonomicName>
|
|
or of a closely related family (
|
|
<emphasis id="B9634C21FFE3560FFD37CF7AFD1DF986" box="[730,742,1635,1660]" italics="true" pageId="3" pageNumber="334">e</emphasis>
|
|
.
|
|
<emphasis id="B9634C21FFE3560FFD03CF7AFD07F986" box="[750,764,1635,1660]" italics="true" pageId="3" pageNumber="334">g</emphasis>
|
|
.,
|
|
<taxonomicName id="4C17EBB0FFE3560FFF65CF9FFEC5F95B" authorityName="Mosely" authorityYear="1936" box="[136,318,1670,1697]" class="Insecta" family="Philorheithridae" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="334" phylum="Arthropoda" rank="family">Philorheithridae</taxonomicName>
|
|
) needs in time to be investigated with further fossil findings and taxonomic data.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
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</document> |