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<mods:title id="355D0397BA194493F7A06044D5383CFB">On some blattisociid mites (Acari: Mesostigmata: Lasioseius, Cheiroseius) from Slovakia, with notes on the genus Hyattella sensu Krantz, 1962</mods:title>
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<taxonomicName id="4C0EE0A3D669FFCCFF0CC633FDE7FD59" authority="Westerboer, 1963" authorityName="Westerboer" authorityYear="1963" box="[151,631,618,645]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="168" phylum="Arthropoda" rank="species" species="lacunosus">
<emphasis id="B97A4732D669FFCCFF0CC633FDE7FD59" bold="true" box="[151,631,618,645]" pageId="9" pageNumber="168">
<emphasis id="B97A4732D669FFCCFF0CC633FE08FD58" bold="true" box="[151,408,618,645]" italics="true" pageId="9" pageNumber="168">Lasioseius lacunosus</emphasis>
<bibRefCitation id="EF9FE6D1D669FFCCFE04C633FDE7FD59" author="Westerboer, I." box="[415,631,618,645]" pageId="9" pageNumber="168" pagination="179 - 450" refId="ref11510" refString="Westerboer, I. (1963) Die Familie Podocinidae Berlese 1916. Abschnitt IV. In: Stammer, H. J. (Ed.), Beitrage zur Systematik und Okologie mitteleuropaischer Acarina. Band II. Mesostigmata I. Akademische Verlagsgesellschaft Geest &amp; Portig K. - G., Leipzig, pp. 179 - 450." type="book chapter" year="1963">Westerboer, 1963</bibRefCitation>
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(
<figureCitation id="133587A5D669FFCCFF04C6D7FEDAFD74" box="[159,330,654,681]" captionStart-0="FIGURES 15–17" captionStart-1="FIGURE 18–22" captionStart-2="FIGURES 23–26" captionStartId-0="7.[151,264,1906,1930]" captionStartId-1="10.[151,250,1906,1930]" captionStartId-2="11.[151,264,1906,1930]" captionTargetBox-0="[151,1436,181,1882]" captionTargetBox-1="[151,1436,181,1881]" captionTargetBox-2="[151,1436,181,1881]" captionTargetId-0="figure-18@7.[151,1436,181,1882]" captionTargetId-1="figure-22@10.[151,1436,181,1882]" captionTargetId-2="figure-18@11.[151,1436,181,1881]" captionTargetPageId-0="7" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="FIGURES 15–17. Lasioseius spp., females. 15. L. cheiroseioides sp. nov., sperm-reception structures, lateral view; 16. L. lacunosus, dorsal idiosoma; 17. L. lacunosus, ventral gnathosoma. Scale bars: 25 µm (Fig. 17), 50 µm (Fig. 15), 100 µm (Fig. 16)." captionText-1="FIGURE 18–22. Lasioseius lacunosus. 18. Chelicera (male), lateral view; 19. Epistome (male); 20. Chelicera (female), lateral view; 21. Femur II (male), ventral view; 22. Tarsus II (male), ventral view. Scale bars: 25 µm (Figs 18‒20), 50 µm (Figs 21, 22)." captionText-2="FIGURES 23–26. Lasioseius spp., females. 23. L. lacunosus, ventral shields; 24. L. cheiroseioides sp. nov., dorsal shield sculpture of hexagonal area; 25. L. lacunosus, dorsal shield sculpture of hexagonal area; 26. L. epicrioides, dorsal shield sculpture of posteromedial area. Scale bars: 50 µm (Fig. 23), 25 µm (Figs 24‒26)." figureDoi-0="http://doi.org/10.5281/zenodo.10146695" figureDoi-1="http://doi.org/10.5281/zenodo.10146697" figureDoi-2="http://doi.org/10.5281/zenodo.10146699" httpUri-0="https://zenodo.org/record/10146695/files/figure.png" httpUri-1="https://zenodo.org/record/10146697/files/figure.png" httpUri-2="https://zenodo.org/record/10146699/files/figure.png" pageId="9" pageNumber="168">Figs 16‒23, 25</figureCitation>
,
<figureCitation id="133587A5D669FFCCFECDC6D7FE0DFD74" box="[342,413,655,681]" captionStart="FIGURES 27–31" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[151,1435,181,1881]" captionTargetId="figure-22@12.[151,1435,181,1881]" captionTargetPageId="12" captionText="FIGURES 27–31. Lasioseius spp., dorsoventral habitus. 27. L. cheiroseioides sp. nov. (female); 28. L. epicrioides (female); 29. L. diffindatus (female); 30. L. lacunosus (male); 31. L. lacunosus (female). Scale bar: 300 µm." figureDoi="http://doi.org/10.5281/zenodo.10146701" httpUri="https://zenodo.org/record/10146701/files/figure.png" pageId="9" pageNumber="168">30, 31</figureCitation>
)
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<taxonomicName id="4C0EE0A3D669FFCCFF0CC68CFD4DFD36" authority="Westerboer, 1963: 238" authorityName="Westerboer" authorityPageNumber="238" authorityYear="1963" box="[151,733,723,747]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="168" phylum="Arthropoda" rank="species" species="lacunosus" subGenus="Lasioseius">
<emphasis id="B97A4732D669FFCCFF0CC68CFE92FD36" box="[151,258,724,747]" italics="true" pageId="9" pageNumber="168">Lasioseius</emphasis>
(
<emphasis id="B97A4732D669FFCCFE8AC68CFEECFD36" box="[273,380,724,747]" italics="true" pageId="9" pageNumber="168">Lasioseius</emphasis>
)
<emphasis id="B97A4732D669FFCCFE10C68BFE60FD36" box="[395,496,723,747]" italics="true" pageId="9" pageNumber="168">lacunosus</emphasis>
<bibRefCitation id="EF9FE6D1D669FFCCFE6DC68BFD4DFD36" author="Westerboer, I." box="[502,733,723,747]" pageId="9" pageNumber="168" pagination="179 - 450" refId="ref11510" refString="Westerboer, I. (1963) Die Familie Podocinidae Berlese 1916. Abschnitt IV. In: Stammer, H. J. (Ed.), Beitrage zur Systematik und Okologie mitteleuropaischer Acarina. Band II. Mesostigmata I. Akademische Verlagsgesellschaft Geest &amp; Portig K. - G., Leipzig, pp. 179 - 450." type="book chapter" year="1963">Westerboer, 1963: 238</bibRefCitation>
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<taxonomicName id="4C0EE0A3D669FFCCFF0CC6AAFEE0FCD7" authorityName="Westerboer" authorityYear="1963" box="[151,368,754,778]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="168" phylum="Arthropoda" rank="species" species="lacunosus">
<emphasis id="B97A4732D669FFCCFF0CC6AAFEE0FCD7" box="[151,368,754,778]" italics="true" pageId="9" pageNumber="168">Lasioseius lacunosus</emphasis>
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.—
<treatmentCitation id="0AAFBD31D669FFCCFE14C6AAFDACFCD7" author="Karg, W." box="[399,572,754,778]" page="248" pageId="9" pageNumber="168" year="1971">
<bibRefCitation id="EF9FE6D1D669FFCCFE14C6AAFDACFCD7" author="Karg, W." box="[399,572,754,778]" pageId="9" pageNumber="168" pagination="1 - 475" refId="ref10128" refString="Karg, W. (1971) Acari (Acarina), Milben, Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands, 59, 1 - 475." type="journal article" year="1971">Karg, 1971: 248</bibRefCitation>
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;
<treatmentCitation id="0AAFBD31D669FFCCFDD2C6AAFD24FCD7" author="Karg, W." box="[585,692,754,778]" page="249" pageId="9" pageNumber="168" year="1993">
<bibRefCitation id="EF9FE6D1D669FFCCFDD2C6AAFD24FCD7" author="Karg, W." box="[585,692,754,778]" pageId="9" pageNumber="168" refId="ref10233" refString="Karg, W. (1993) Acari (Acarina), Milben Parasitiformes (Anactinochaeta) Cohors Gamasina Leach, Raubmilben. Die Tierwelt Deutschlands und der angrenzenden Meeresteile nach ihren Merkmalen und ihrer Lebensweise. 59. Teil. 2. Uberarbeitete Auflage, Gustav Fischer Verlag, Jena, Stuttgart and New York, 523 pp." type="book" year="1993">1993: 249</bibRefCitation>
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;
<treatmentCitation id="0AAFBD31D669FFCCFD5AC6AAFCE0FCD7" author="Treat, A. E." box="[705,880,754,778]" page="121" pageId="9" pageNumber="168" year="1975">
<bibRefCitation id="EF9FE6D1D669FFCCFD5AC6AAFCE0FCD7" author="Treat, A. E." box="[705,880,754,778]" pageId="9" pageNumber="168" refId="ref11424" refString="Treat, A. E. (1975) Mites of Moths and Butterflies. Comstock University Press, New York, New York, 362 pp." type="book" year="1975">Treat, 1975: 121</bibRefCitation>
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;
<treatmentCitation id="0AAFBD31D669FFCCFCE7C6AAFB50FCD7" author="Farrier, M. H. &amp; Hennessey, M. K." box="[892,1216,754,778]" page="36" pageId="9" pageNumber="168" year="1993">
<bibRefCitation id="EF9FE6D1D669FFCCFCE7C6AAFB50FCD7" author="Farrier, M. H. &amp; Hennessey, M. K." box="[892,1216,754,778]" pageId="9" pageNumber="168" pagination="1 - 408" refId="ref9830" refString="Farrier, M. H. &amp; Hennessey, M. K. (1993) Soil-inhabiting and free-living Mesostigmata (Acari-Parasitiformes) from North America. An annotated checklist with bibliography and index. North Carolina Agricultural Research Service, North Carolina State University, Raleigh, Technical Bulletin, 302, 1 - 408." type="journal article" year="1993">Farrier &amp; Hennessey, 1993: 36</bibRefCitation>
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<bibRefCitation id="EF9FE6D1D669FFCCFB56C6AAFEB3FCF5" author="Gwiazdowicz, D. J. &amp; Blaszak, C. &amp; Ehrnsberger, R." pageId="9" pageNumber="168" pagination="35 - 39" refId="ref9925" refString="Gwiazdowicz, D. J., Blaszak, C. &amp; Ehrnsberger, R. (2008) Die Milben in der Zoologischen Staatssammlung Munchen. Teil 12. Gattungen: Blattisocius Keegan, 1944, Hoploseius Berlese, 1914, Lasioseius Berlese, 1916, Platyseius Berlese, 1916. Spixiana, 31, 35 - 39." type="journal article" year="2008">
Gwiazdowicz
<emphasis id="B97A4732D669FFCCFAFAC6ABFA00FCD7" box="[1377,1424,754,778]" italics="true" pageId="9" pageNumber="168">et al</emphasis>
., 2008: 36
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<bibRefCitation id="EF9FE6D1D669FFCCFEB5C748FDA0FCF5" author="Cakmak, I. &amp; Faraji, F. &amp; Cobanoglu, S." box="[302,560,784,809]" pageId="9" pageNumber="168" pagination="575 - 586" refId="ref9304" refString="Cakmak, I., Faraji, F. &amp; Cobanoglu, S. (2011) A checklist and key to the Ascoidea and Phytoseioidea (except Phytoseiidae) species of Turkey with three new species records (Acari: Mesostigmata). Turkiye Entomoloji Dergisi, 35, 575 - 586." type="journal article" year="2011">
Çakmak
<emphasis id="B97A4732D669FFCCFE12C749FE25FCF5" box="[393,437,784,808]" italics="true" pageId="9" pageNumber="168">et al</emphasis>
., 2011: 583
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<bibRefCitation id="EF9FE6D1D669FFCCFDA1C749FCA5FCF4" author="Moraes, G. J. de &amp; Britto, E. P. J. &amp; Mineiro, J. L. de &amp; Halliday, B." box="[570,821,784,809]" pageId="9" pageNumber="168" pagination="1 - 299" refId="ref11200" refString="Moraes, G. J. de, Britto, E. P. J., Mineiro, J. L. de C. &amp; Halliday, B. (2016) Catalogue of the mite families Ascidae Voigts &amp; Oudemans, Blattisociidae Garman and Melicharidae Hirschmann (Acaria: Mesostigmata). Zootaxa, 4112 (1), 1 - 299. https: // doi. org / 10.11646 / zootaxa. 4112.1.1" type="journal article" year="2016">
Moraes
<emphasis id="B97A4732D669FFCCFD16C749FD29FCF5" box="[653,697,784,808]" italics="true" pageId="9" pageNumber="168">et al</emphasis>
., 2016: 173
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<taxonomicName id="4C0EE0A3D669FFCCFF0CC768FE79FC9A" authorityName="Westerboer" authorityYear="1963" box="[151,489,815,839]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="168" phylum="Arthropoda" rank="species" species="lacunosus" subGenus="Crinidens">
<emphasis id="B97A4732D669FFCCFF0CC768FE92FC9A" box="[151,258,816,839]" italics="true" pageId="9" pageNumber="168">Lasioseius</emphasis>
(
<emphasis id="B97A4732D669FFCCFE8AC777FEE5FC9A" box="[273,373,815,839]" italics="true" pageId="9" pageNumber="168">Crinidens</emphasis>
)
<emphasis id="B97A4732D669FFCCFE1FC777FE79FC9A" box="[388,489,815,839]" italics="true" pageId="9" pageNumber="168">lacunosus</emphasis>
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.—
<treatmentCitation id="0AAFBD31D669FFCCFD93C768FD21FC9A" author="Karg, W." box="[520,689,815,840]" page="349" pageId="9" pageNumber="168" year="1980">
<bibRefCitation id="EF9FE6D1D669FFCCFD93C768FD21FC9A" author="Karg, W." box="[520,689,815,840]" pageId="9" pageNumber="168" pagination="344 - 367" refId="ref10167" refString="Karg, W. (1980) Die Raubmilbengattung Lasioseius Berlese, 1916. Zoologische Jahrbucher. Abteilung fur Systematik, Okologie und Geographie der Tiere, 107, 344 - 367." type="journal article" year="1980">Karg, 1980: 349</bibRefCitation>
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<bibRefCitation id="EF9FE6D1D669FFCCFD20C777FC70FC9A" author="Christian, A. &amp; Karg, W." box="[699,992,815,840]" pageId="9" pageNumber="168" pagination="99 - 250" refId="ref9450" refString="Christian, A. &amp; Karg, W. (2006) The predatory mite genus Lasioseius Berlese, 1916 (Acari, Gamasina). Abhandlungen und Berichte des Naturkundemuseums Gorlitz, 77, 99 - 250." type="journal article" year="2006">Christian &amp; Karg, 2006: 156</bibRefCitation>
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</subSubSection>
<subSubSection id="C314C8ABD669FFCCFF0CC72DFD22F85D" pageId="9" pageNumber="168" type="materials_examined">
<paragraph id="8BB19B20D669FFCCFF0CC72DFEFBFC52" blockId="9.[151,1436,885,1055]" box="[151,363,885,911]" pageId="9" pageNumber="168">
<emphasis id="B97A4732D669FFCCFF0CC72DFEFBFC52" box="[151,363,885,911]" italics="true" pageId="9" pageNumber="168">Material examined</emphasis>
</paragraph>
<paragraph id="8BB19B20D669FFCCFF0CC7C1FE60FBC2" blockId="9.[151,1436,885,1055]" pageId="9" pageNumber="168">
<specimenCount id="9D0850A9D669FFCCFF0CC7C1FE8DFC6E" box="[151,285,921,947]" count="1" pageId="9" pageNumber="168" type="female">One female</specimenCount>
,
<specimenCount id="9D0850A9D669FFCCFEB2C7C1FE3FFC6E" box="[297,431,921,947]" count="3" pageId="9" pageNumber="168" type="male">three males</specimenCount>
: SW
<collectingCountry id="F319DBB0D669FFCCFE70C7C1FDC0FC6E" box="[491,592,921,947]" name="Slovakia" pageId="9" pageNumber="168">Slovakia</collectingCountry>
, Little Carpathians Mts.,
<taxonomicName id="4C0EE0A3D669FFCCFCF4C7C1FEBCFC05" authority="Capital, Zahorska Bystrica Settlement, Kacin Forest" authorityName="Capital, Zahorska Bystrica Settlement, Kacin Forest" class="Clitellata" family="Naididae" genus="Bratislava" kingdom="Animalia" order="Tubificida" pageId="9" pageNumber="168" phylum="Annelida" rank="genus">Bratislava Capital, Záhorská Bystrica Settlement, Kačín Forest</taxonomicName>
, elevation
<quantity id="4CF636C5D669FFCCFE33C7E5FE62FC05" box="[424,498,957,984]" metricMagnitude="2" metricUnit="m" metricValue="3.35" pageId="9" pageNumber="168" unit="m" value="335.0">335 m</quantity>
, decidouous forest, in bark crevices of a maple (
<taxonomicName id="4C0EE0A3D669FFCCFBBAC7E6FBEFFC0A" box="[1057,1151,957,984]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="168" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B97A4732D669FFCCFBBAC7E6FBC6FC05" box="[1057,1110,958,984]" italics="true" pageId="9" pageNumber="168">Acer</emphasis>
sp.
</taxonomicName>
) soaked with xylem sap,
<date id="FFB0BDE0D669FFCCFF0CC7B9FEA1FC26" box="[151,305,993,1020]" pageId="9" pageNumber="168" value="2019-06-27">27 June 2019</date>
(leg. author). All specimens were deposited in the Institute of Zoology of the Slovak Academy of Sciences,
<taxonomicName id="4C0EE0A3D669FFCCFE9CC05DFEEBFBC2" box="[263,379,1029,1055]" class="Clitellata" family="Naididae" genus="Bratislava" kingdom="Animalia" order="Tubificida" pageId="9" pageNumber="179" phylum="Annelida" rank="genus">Bratislava</taxonomicName>
,
<collectingCountry id="F319DBB0D669FFCCFE1CC05DFE7DFBC2" box="[391,493,1029,1055]" name="Slovakia" pageId="9" pageNumber="168">Slovakia</collectingCountry>
.
</paragraph>
<paragraph id="8BB19B20D669FFCCFF0CC016FEC3FBBA" blockId="9.[151,1437,1102,1488]" box="[151,339,1102,1127]" pageId="9" pageNumber="168">
<emphasis id="B97A4732D669FFCCFF0CC016FEC3FBBA" box="[151,339,1102,1127]" italics="true" pageId="9" pageNumber="168">Taxonomic notes</emphasis>
</paragraph>
<paragraph id="8BB19B20D669FFCCFF0CC029FB18FAE2" blockId="9.[151,1437,1102,1488]" pageId="9" pageNumber="168">
This is the first record of
<taxonomicName id="4C0EE0A3D669FFCCFE51C02AFDCBFB56" authorityName="Westerboer" authorityYear="1963" box="[458,603,1137,1164]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="168" phylum="Arthropoda" rank="species" species="lacunosus">
<emphasis id="B97A4732D669FFCCFE51C02AFE4AFB56" box="[458,474,1138,1163]" italics="true" pageId="9" pageNumber="168">L</emphasis>
.
<emphasis id="B97A4732D669FFCCFE70C029FDCBFB56" box="[491,603,1137,1163]" italics="true" pageId="9" pageNumber="168">lacunosus</emphasis>
</taxonomicName>
for
<collectingCountry id="F319DBB0D669FFCCFD0AC029FD67FB56" box="[657,759,1137,1163]" name="Slovakia" pageId="9" pageNumber="168">Slovakia</collectingCountry>
. Some specific morphological structures of a female from
<collectingCountry id="F319DBB0D669FFCCFF0CC0CDFF6CFB72" box="[151,252,1173,1199]" name="Slovakia" pageId="9" pageNumber="168">Slovakia</collectingCountry>
, which are important for verifying its identity, can be seen on the micrographs (
<figureCitation id="133587A5D669FFCCFB0BC0CDFA84FB72" box="[1168,1300,1173,1199]" captionStart="FIGURES 15–17" captionStartId="7.[151,264,1906,1930]" captionTargetBox="[151,1436,181,1882]" captionTargetId="figure-18@7.[151,1436,181,1882]" captionTargetPageId="7" captionText="FIGURES 15–17. Lasioseius spp., females. 15. L. cheiroseioides sp. nov., sperm-reception structures, lateral view; 16. L. lacunosus, dorsal idiosoma; 17. L. lacunosus, ventral gnathosoma. Scale bars: 25 µm (Fig. 17), 50 µm (Fig. 15), 100 µm (Fig. 16)." figureDoi="http://doi.org/10.5281/zenodo.10146695" httpUri="https://zenodo.org/record/10146695/files/figure.png" pageId="9" pageNumber="168">Figs 16, 17</figureCitation>
,
<figureCitation id="133587A5D669FFCCFAB9C0CDFAD1FB72" box="[1314,1345,1173,1199]" captionStart="FIGURE 18–22" captionStartId="10.[151,250,1906,1930]" captionTargetBox="[151,1436,181,1881]" captionTargetId="figure-22@10.[151,1436,181,1882]" captionTargetPageId="10" captionText="FIGURE 18–22. Lasioseius lacunosus. 18. Chelicera (male), lateral view; 19. Epistome (male); 20. Chelicera (female), lateral view; 21. Femur II (male), ventral view; 22. Tarsus II (male), ventral view. Scale bars: 25 µm (Figs 18‒20), 50 µm (Figs 21, 22)." figureDoi="http://doi.org/10.5281/zenodo.10146697" httpUri="https://zenodo.org/record/10146697/files/figure.png" pageId="9" pageNumber="168">20</figureCitation>
,
<figureCitation id="133587A5D669FFCCFAD6C0CDFA07FB6D" box="[1357,1431,1173,1200]" captionStart="FIGURES 23–26" captionStartId="11.[151,264,1906,1930]" captionTargetBox="[151,1436,181,1881]" captionTargetId="figure-18@11.[151,1436,181,1881]" captionTargetPageId="11" captionText="FIGURES 23–26. Lasioseius spp., females. 23. L. lacunosus, ventral shields; 24. L. cheiroseioides sp. nov., dorsal shield sculpture of hexagonal area; 25. L. lacunosus, dorsal shield sculpture of hexagonal area; 26. L. epicrioides, dorsal shield sculpture of posteromedial area. Scale bars: 50 µm (Fig. 23), 25 µm (Figs 24‒26)." figureDoi="http://doi.org/10.5281/zenodo.10146699" httpUri="https://zenodo.org/record/10146699/files/figure.png" pageId="9" pageNumber="168">23, 25</figureCitation>
,
<figureCitation id="133587A5D669FFCCFF0CC0E1FF26FB0E" box="[151,182,1209,1235]" captionStart="FIGURES 27–31" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[151,1435,181,1881]" captionTargetId="figure-22@12.[151,1435,181,1881]" captionTargetPageId="12" captionText="FIGURES 27–31. Lasioseius spp., dorsoventral habitus. 27. L. cheiroseioides sp. nov. (female); 28. L. epicrioides (female); 29. L. diffindatus (female); 30. L. lacunosus (male); 31. L. lacunosus (female). Scale bar: 300 µm." figureDoi="http://doi.org/10.5281/zenodo.10146701" httpUri="https://zenodo.org/record/10146701/files/figure.png" pageId="9" pageNumber="168">31</figureCitation>
). I also examined a fairly well preserved female, clearly conspecific with the female from
<collectingCountry id="F319DBB0D669FFCCFB26C0E1FAB2FB0E" box="[1213,1314,1209,1235]" name="Slovakia" pageId="9" pageNumber="168">Slovakia</collectingCountry>
, on a slide deposited in the Bavarian State Collection of Zoology in Munich and labeled ʻ
<taxonomicName id="4C0EE0A3D669FFCCFBBBC086FA80FB2A" authority="Mašán, 2023" authorityName="Mašán" authorityYear="2023" box="[1056,1296,1245,1271]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="168" phylum="Arthropoda" rank="species" species="lacunosus" status="sp. nov.">
<emphasis id="B97A4732D669FFCCFBBBC086FA80FB2A" box="[1056,1296,1245,1271]" italics="true" pageId="9" pageNumber="168">Lasioseius lacunosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A249FA49D669FFCCFA82C086FACBFB2A" box="[1305,1371,1245,1272]" pageId="9" pageNumber="168" rank="species">n. sp.</taxonomicNameLabel>
ʼ (No. W34/14). Unfortunately, this available specimen (slide) does not bear a
<typeStatus id="54B52582D669FFCCFC23C15AFC78FAC1" box="[952,1000,1282,1308]" pageId="9" pageNumber="168">type</typeStatus>
designation or specific collection data, but according to
<bibRefCitation id="EF9FE6D1D669FFCCFECFC17DFDEFFAE2" author="Gwiazdowicz, D. J. &amp; Blaszak, C. &amp; Ehrnsberger, R." box="[340,639,1317,1344]" pageId="9" pageNumber="168" pagination="35 - 39" refId="ref9925" refString="Gwiazdowicz, D. J., Blaszak, C. &amp; Ehrnsberger, R. (2008) Die Milben in der Zoologischen Staatssammlung Munchen. Teil 12. Gattungen: Blattisocius Keegan, 1944, Hoploseius Berlese, 1914, Lasioseius Berlese, 1916, Platyseius Berlese, 1916. Spixiana, 31, 35 - 39." type="journal article" year="2008">
Gwiazdowicz
<emphasis id="B97A4732D669FFCCFE6EC17EFDB6FAE2" box="[501,550,1317,1343]" italics="true" pageId="9" pageNumber="168">et al</emphasis>
. (2008)
</bibRefCitation>
it belongs to the original series of Westerboer.
</paragraph>
<paragraph id="8BB19B20D669FFCCFF5CC112FDF8FA0D" blockId="9.[151,1437,1102,1488]" pageId="9" pageNumber="168">
As also noted in the taxonomic notes on the new species above,
<taxonomicName id="4C0EE0A3D669FFCCFC10C112FB88FABE" authorityName="Mašán" authorityYear="2023" box="[907,1048,1353,1380]" class="Arachnida" family="Ascidae" genus="Lasioseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="168" phylum="Arthropoda" rank="species" species="lacunosus">
<emphasis id="B97A4732D669FFCCFC10C112FC0BFABE" box="[907,923,1354,1379]" italics="true" pageId="9" pageNumber="168">L</emphasis>
.
<emphasis id="B97A4732D669FFCCFC33C111FB88FABE" box="[936,1048,1353,1379]" italics="true" pageId="9" pageNumber="168">lacunosus</emphasis>
</taxonomicName>
has specially modified macrosetae on the tarsi
<collectionCode id="ED1F03E5D669FFCCFE84C136FEA1FA55" box="[287,305,1390,1416]" pageId="9" pageNumber="168">II</collectionCode>
‒
<collectionCode id="ED1F03E5D669FFCCFEA4C136FECCFA55" box="[319,348,1390,1416]" pageId="9" pageNumber="168">IV</collectionCode>
, and this fact was not illustrated or mentioned in the original description by
<bibRefCitation id="EF9FE6D1D669FFCCFB58C135FA09FA5A" author="Westerboer, I." box="[1219,1433,1389,1415]" pageId="9" pageNumber="168" pagination="179 - 450" refId="ref11510" refString="Westerboer, I. (1963) Die Familie Podocinidae Berlese 1916. Abschnitt IV. In: Stammer, H. J. (Ed.), Beitrage zur Systematik und Okologie mitteleuropaischer Acarina. Band II. Mesostigmata I. Akademische Verlagsgesellschaft Geest &amp; Portig K. - G., Leipzig, pp. 179 - 450." type="book chapter" year="1963">Westerboer (1963)</bibRefCitation>
. Similar to the legs, the males have not been described before, so I include the following description of some of the most important morphological structures:
</paragraph>
<paragraph id="8BB19B20D669FFCCFF0CC1A6FD08F9CA" blockId="9.[151,1437,1533,1920]" box="[151,664,1533,1560]" pageId="9" pageNumber="168">
<emphasis id="B97A4732D669FFCCFF0CC1A6FE8BF9CA" box="[151,283,1534,1559]" italics="true" pageId="9" pageNumber="168">Description</emphasis>
(
<emphasis id="B97A4732D669FFCCFEB7C1A5FEF6F9CA" box="[300,358,1533,1559]" italics="true" pageId="9" pageNumber="168">Male</emphasis>
) ‒ (
<figureCitation id="133587A5D669FFCCFE09C1A5FDF8F9CA" box="[402,616,1533,1559]" captionStart="FIGURE 18–22" captionStartId="10.[151,250,1906,1930]" captionTargetBox="[151,1436,181,1881]" captionTargetId="figure-22@10.[151,1436,181,1882]" captionTargetPageId="10" captionText="FIGURE 18–22. Lasioseius lacunosus. 18. Chelicera (male), lateral view; 19. Epistome (male); 20. Chelicera (female), lateral view; 21. Femur II (male), ventral view; 22. Tarsus II (male), ventral view. Scale bars: 25 µm (Figs 18‒20), 50 µm (Figs 21, 22)." figureDoi="http://doi.org/10.5281/zenodo.10146697" httpUri="https://zenodo.org/record/10146697/files/figure.png" pageId="9" pageNumber="168">Figs 18, 19, 21, 22</figureCitation>
,
<figureCitation id="133587A5D669FFCCFDE8C1A5FD00F9CA" box="[627,656,1533,1559]" captionStart="FIGURES 27–31" captionStartId="12.[151,264,1905,1929]" captionTargetBox="[151,1435,181,1881]" captionTargetId="figure-22@12.[151,1435,181,1881]" captionTargetPageId="12" captionText="FIGURES 27–31. Lasioseius spp., dorsoventral habitus. 27. L. cheiroseioides sp. nov. (female); 28. L. epicrioides (female); 29. L. diffindatus (female); 30. L. lacunosus (male); 31. L. lacunosus (female). Scale bar: 300 µm." figureDoi="http://doi.org/10.5281/zenodo.10146701" httpUri="https://zenodo.org/record/10146701/files/figure.png" pageId="9" pageNumber="168">30</figureCitation>
)
</paragraph>
<paragraph id="8BB19B20D669FFCCFF0CC279FD22F85D" blockId="9.[151,1437,1533,1920]" pageId="9" pageNumber="168">
Dorsal shield 500‒555 long, shorter than legs
<collectionCode id="ED1F03E5D669FFCCFD28C27AFD40F9E1" box="[691,720,1570,1596]" pageId="9" pageNumber="168">IV</collectionCode>
(525‒575), with coarse reticulate pattern and setae similar to those of female; all anteromarginal setae (
<emphasis id="B97A4732D669FFCCFDE0C21DFD2AF982" box="[635,698,1605,1632]" italics="true" pageId="9" pageNumber="168">r2‒r6</emphasis>
) on dorsal shield, but all posteromarginal setae (
<emphasis id="B97A4732D669FFCCFB75C21DFAABF982" box="[1262,1339,1605,1632]" italics="true" pageId="9" pageNumber="168">
<collectionCode id="ED1F03E5D669FFCCFB75C21DFA91F982" box="[1262,1281,1605,1631]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="168">R</collectionCode>
1‒
<collectionCode id="ED1F03E5D669FFCCFA80C21DFABCF982" box="[1307,1324,1605,1631]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="168">R</collectionCode>
6
</emphasis>
) on soft cuticle adjacent to shield margin;
<emphasis id="B97A4732D669FFCCFD97C232FDA1F95E" box="[524,561,1642,1667]" italics="true" pageId="9" pageNumber="168">
<collectionCode id="ED1F03E5D669FFCCFD97C232FDA1F95E" box="[524,561,1642,1667]" pageId="9" pageNumber="168">UR</collectionCode>
</emphasis>
setae of submarginal cuticle completely reduced, absent (four pairs in female); opisthogastric region normally with six pairs of pre-anal setae on ventrianal shield (
<emphasis id="B97A4732D669FFCCFBD0C2D5FB20F97A" box="[1099,1200,1677,1704]" italics="true" pageId="9" pageNumber="168">
<collectionCode id="ED1F03E5D669FFCCFBD0C2D5FBFCF97A" box="[1099,1132,1677,1703]" pageId="9" pageNumber="168">JV</collectionCode>
1‒
<collectionCode id="ED1F03E5D669FFCCFB1FC2D5FB33F97A" box="[1156,1187,1677,1703]" pageId="9" pageNumber="168">JV</collectionCode>
3
</emphasis>
,
<emphasis id="B97A4732D669FFCCFB25C2D6FB7AF975" box="[1214,1258,1678,1704]" italics="true" pageId="9" pageNumber="168">JV5</emphasis>
,
<emphasis id="B97A4732D669FFCCFB62C2D5FAB7F97A" box="[1273,1319,1677,1703]" italics="true" pageId="9" pageNumber="168">ZV1</emphasis>
,
<emphasis id="B97A4732D669FFCCFAADC2D5FAF5F97A" box="[1334,1381,1677,1703]" italics="true" pageId="9" pageNumber="168">ZV2</emphasis>
, and asymmetrically sometimes
<emphasis id="B97A4732D669FFCCFE53C2E9FE67F916" box="[456,503,1713,1739]" italics="true" pageId="9" pageNumber="168">
<collectionCode id="ED1F03E5D669FFCCFE53C2E9FE7AF916" box="[456,490,1713,1739]" country="Slovakia" lsid="urn:lsid:biocol.org:col:14185" name="Technical University" pageId="9" pageNumber="168" type="Herbarium">ZV</collectionCode>
4
</emphasis>
) and two pairs on soft cuticle (
<emphasis id="B97A4732D669FFCCFCD5C2E9FCEDF916" box="[846,893,1713,1739]" italics="true" pageId="9" pageNumber="168">
<collectionCode id="ED1F03E5D669FFCCFCD5C2E9FCE0F916" box="[846,880,1713,1739]" country="Slovakia" lsid="urn:lsid:biocol.org:col:14185" name="Technical University" pageId="9" pageNumber="168" type="Herbarium">ZV</collectionCode>
4
</emphasis>
and
<emphasis id="B97A4732D669FFCCFC29C2EAFC70F911" box="[946,992,1714,1740]" italics="true" pageId="9" pageNumber="168">
<collectionCode id="ED1F03E5D669FFCCFC29C2EAFC44F911" box="[946,980,1714,1740]" country="Slovakia" lsid="urn:lsid:biocol.org:col:14185" name="Technical University" pageId="9" pageNumber="168" type="Herbarium">ZV</collectionCode>
5
</emphasis>
), but setae
<emphasis id="B97A4732D669FFCCFBC4C2E9FB1BF916" box="[1119,1163,1713,1739]" italics="true" pageId="9" pageNumber="168">JV4</emphasis>
and
<emphasis id="B97A4732D669FFCCFB24C2E9FB7EF916" box="[1215,1262,1713,1739]" italics="true" pageId="9" pageNumber="168">ZV3</emphasis>
absent (present in female); chelicerae with elongate middle article, fixed digit with one small subapical tooth and a row of five teeth, movable digit with one distal tooth and an intricately formed spermatodactyl, with club-shaped proximal and medial parts and mushroom-shaped distal part (
<figureCitation id="133587A5D669FFCCFD35C345FC92F8EA" box="[686,770,1821,1847]" captionStart="FIGURE 18–22" captionStartId="10.[151,250,1906,1930]" captionTargetBox="[151,1436,181,1881]" captionTargetId="figure-22@10.[151,1436,181,1882]" captionTargetPageId="10" captionText="FIGURE 18–22. Lasioseius lacunosus. 18. Chelicera (male), lateral view; 19. Epistome (male); 20. Chelicera (female), lateral view; 21. Femur II (male), ventral view; 22. Tarsus II (male), ventral view. Scale bars: 25 µm (Figs 18‒20), 50 µm (Figs 21, 22)." figureDoi="http://doi.org/10.5281/zenodo.10146697" httpUri="https://zenodo.org/record/10146697/files/figure.png" pageId="9" pageNumber="168">Fig. 18</figureCitation>
); epistome as in female (
<figureCitation id="133587A5D669FFCCFB87C345FBE0F8EA" box="[1052,1136,1821,1847]" captionStart="FIGURE 18–22" captionStartId="10.[151,250,1906,1930]" captionTargetBox="[151,1436,181,1881]" captionTargetId="figure-22@10.[151,1436,181,1882]" captionTargetPageId="10" captionText="FIGURE 18–22. Lasioseius lacunosus. 18. Chelicera (male), lateral view; 19. Epistome (male); 20. Chelicera (female), lateral view; 21. Femur II (male), ventral view; 22. Tarsus II (male), ventral view. Scale bars: 25 µm (Figs 18‒20), 50 µm (Figs 21, 22)." figureDoi="http://doi.org/10.5281/zenodo.10146697" httpUri="https://zenodo.org/record/10146697/files/figure.png" pageId="9" pageNumber="168">Fig. 19</figureCitation>
); legs
<collectionCode id="ED1F03E5D669FFCCFB21C346FB5DF8E5" box="[1210,1229,1822,1848]" pageId="9" pageNumber="168">II</collectionCode>
slightly thickened and stouter than the others, and partially armed ‒ some anteroventral setae, namely
<emphasis id="B97A4732D669FFCCFBACC31AFBC1F886" box="[1079,1105,1858,1883]" italics="true" pageId="9" pageNumber="168">av</emphasis>
on femur
<collectionCode id="ED1F03E5D669FFCCFB5FC31AFB47F881" box="[1220,1239,1858,1884]" pageId="9" pageNumber="168">II</collectionCode>
(
<figureCitation id="133587A5D669FFCCFB7EC319FAA6F886" box="[1253,1334,1857,1883]" captionStart="FIGURE 18–22" captionStartId="10.[151,250,1906,1930]" captionTargetBox="[151,1436,181,1881]" captionTargetId="figure-22@10.[151,1436,181,1882]" captionTargetPageId="10" captionText="FIGURE 18–22. Lasioseius lacunosus. 18. Chelicera (male), lateral view; 19. Epistome (male); 20. Chelicera (female), lateral view; 21. Femur II (male), ventral view; 22. Tarsus II (male), ventral view. Scale bars: 25 µm (Figs 18‒20), 50 µm (Figs 21, 22)." figureDoi="http://doi.org/10.5281/zenodo.10146697" httpUri="https://zenodo.org/record/10146697/files/figure.png" pageId="9" pageNumber="168">Fig. 21</figureCitation>
) and
<emphasis id="B97A4732D669FFCCFAEFC319FA0CF886" box="[1396,1436,1857,1883]" italics="true" pageId="9" pageNumber="168">av2</emphasis>
on tarsus
<collectionCode id="ED1F03E5D669FFCCFE98C33EFE86F85D" box="[259,278,1894,1920]" pageId="9" pageNumber="168">II</collectionCode>
(
<figureCitation id="133587A5D669FFCCFEBFC33DFEE8F8A2" box="[292,376,1893,1919]" captionStart="FIGURE 18–22" captionStartId="10.[151,250,1906,1930]" captionTargetBox="[151,1436,181,1881]" captionTargetId="figure-22@10.[151,1436,181,1882]" captionTargetPageId="10" captionText="FIGURE 18–22. Lasioseius lacunosus. 18. Chelicera (male), lateral view; 19. Epistome (male); 20. Chelicera (female), lateral view; 21. Femur II (male), ventral view; 22. Tarsus II (male), ventral view. Scale bars: 25 µm (Figs 18‒20), 50 µm (Figs 21, 22)." figureDoi="http://doi.org/10.5281/zenodo.10146697" httpUri="https://zenodo.org/record/10146697/files/figure.png" pageId="9" pageNumber="168">Fig. 22</figureCitation>
), modified into a stout spur.
</paragraph>
</subSubSection>
</treatment>
</document>
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