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<document id="EB1B63709D38162BC6B29B453C59FAA0" ID-CLB-Dataset="56137" ID-DOI="10.37828/em.2022.59.4" ID-GBIF-Dataset="41cd47cf-b847-49c3-92ec-bf06c3317d35" ID-ISSN="2336-9744" ID-Zenodo-Dep="8044286" ID-ZooBank="urn:lsid:zoobank.org:pub:943334E8-20EA-4399-831F-9D3564952364" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1686850161303" checkinUser="felipe" docAuthor="Ustjuzhanin, Petr Ya. &amp; Kovtunovich, Vasily N." docDate="2022" docId="038DF00DFFE6856EFF212F1C2E5A19C1" docLanguage="en" docName="EcolMontenegrina.59.38-45.pdf" docOrigin="Ecologica Montenegrina 59" docSource="http://dx.doi.org/10.37828/em.2022.59.4" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Emmelina vera Ustjuzhanin &amp; Kovtunovich 2022, sp. n." docType="treatment" docUuid="E742FEC4-E5BB-49F9-88AD-455AF27C71AB" docUuidSource="ZooBank" docVersion="6" lastPageNumber="45" masterDocId="FFB48875FFE58569FFB62A512B12180B" masterDocTitle="To the Pterophoridae fauna of Paraguay (Lepidoptera). Part 3" masterLastPageNumber="45" masterPageNumber="38" pageNumber="41" updateTime="1699525149320" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5AE1AC9D7B4F77EE871734CE35A4E4CF">To the Pterophoridae fauna of Paraguay (Lepidoptera). Part 3</mods:title>
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<mods:namePart id="16D14F805B69338762AEDA87031ED569">Ustjuzhanin, Petr Ya.</mods:namePart>
<mods:nameIdentifier id="EDE11C901ED49372536A1CF47FDA88C4" type="ORCID">0000-0002-5222-2241</mods:nameIdentifier>
<mods:affiliation id="EAB555764E092DDEF987532131C82F4B">Altai State University, Lenina 61, Barnaul, RU- 656049, Russia. &amp; Biological Institute, Tomsk State University, Lenina 36, Tomsk, RU - 634050, Russia. &amp; Corresponding author. E-mail: petrust @ mail. ru, https: // orcid. org / 0000 - 0002 - 5222 - 2241</mods:affiliation>
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<mods:namePart id="1A7C24EECE741025580560A42AFFBE74">Kovtunovich, Vasily N.</mods:namePart>
<mods:nameIdentifier id="6EF0496B5C4A8F5F76BDB93F1B583B3C" type="ORCID">0000-0001-5091-4263</mods:nameIdentifier>
<mods:affiliation id="3F19B9345C77A84678D0092075949068">Moscow, Russia, E-mail: vasko- 69 @ mail. ru, https: // orcid. org / 0000 - 0001 - 5091 - 4263</mods:affiliation>
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<mods:title id="C28E821380C4340D5A245EC71F97730C">Ecologica Montenegrina</mods:title>
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<mods:date id="827EB141639C7185C888BFF1E50B0C1B">2022</mods:date>
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<mods:number id="29F6E1B9514F25C70D0375D9292E7EAE">2022-11-30</mods:number>
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<mods:number id="D9E4B686D204A49BAD1256C4776649BF">59</mods:number>
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<treatment id="038DF00DFFE6856EFF212F1C2E5A19C1" ID-DOI="http://doi.org/10.5281/zenodo.8044326" ID-GBIF-Taxon="209695466" ID-Zenodo-Dep="8044326" LSID="urn:lsid:zoobank.org:act:E742FEC4-E5BB-49F9-88AD-455AF27C71AB" httpUri="http://treatment.plazi.org/id/038DF00DFFE6856EFF212F1C2E5A19C1" lastPageId="7" lastPageNumber="45" pageId="3" pageNumber="41">
<subSubSection id="C33E1290FFE6856AFF212F1C2F931D8E" pageId="3" pageNumber="41" type="nomenclature">
<paragraph id="8B9B411BFFE6856AFF212F1C28181D6C" blockId="3.[151,1153,1357,1447]" box="[151,778,1357,1383]" pageId="3" pageNumber="41">
<heading id="D0D3F677FFE6856AFF212F1C28181D6C" box="[151,778,1357,1383]" fontSize="11" level="2" pageId="3" pageNumber="41" reason="2">
<emphasis id="B9509D09FFE6856AFF212F1C28181D6C" bold="true" box="[151,778,1357,1383]" pageId="3" pageNumber="41">
<taxonomicName id="4C243A98FFE6856AFF212F1C29A91D6C" ID-CoL="B7QGZ" authority="Ustjuzhanin &amp; Kovtunovich" authorityName="Ustjuzhanin &amp; Kovtunovich" authorityYear="2022" box="[151,699,1357,1383]" class="Insecta" family="Pterophoridae" genus="Emmelina" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="41" phylum="Arthropoda" rank="species" species="vera" status="sp. nov.">
<emphasis id="B9509D09FFE6856AFF212F1C2A421D6C" bold="true" box="[151,336,1357,1383]" italics="true" pageId="3" pageNumber="41">Emmelina vera</emphasis>
Ustjuzhanin &amp; Kovtunovich
</taxonomicName>
,
<taxonomicNameLabel id="A2632072FFE6856AFD7E2F1C28181D6C" box="[712,778,1357,1383]" pageId="3" pageNumber="41" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B411BFFE6856AFF212F3C2F931D8E" blockId="3.[151,1153,1357,1447]" box="[151,1153,1389,1413]" pageId="3" pageNumber="41">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="FF827BCEFFE6856AFDC42F3C2F931D8E" box="[626,1153,1389,1413]" pageId="3" pageNumber="41">E742FEC4-E5BB-49F9-88AD-455AF27C71AB</uuid>
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<subSubSection id="C33E1290FFE6856AFF212FDD2BEA1DAC" box="[151,248,1420,1447]" pageId="3" pageNumber="41" type="description">
<paragraph id="8B9B411BFFE6856AFF212FDD2BEA1DAC" blockId="3.[151,1153,1357,1447]" box="[151,248,1420,1447]" pageId="3" pageNumber="41">
<figureCitation id="131F5D9EFFE6856AFF212FDD2BEA1DAC" box="[151,248,1420,1447]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="5.[150,224,759,783]" captionStartId-1="5.[150,224,2011,2035]" captionStartId-2="6.[931,1005,1941,1965]" captionTargetBox-0="[164,1415,199,713]" captionTargetBox-1="[174,1387,1039,1928]" captionTargetBox-2="[265,763,189,2055]" captionTargetId-0="figure-21@5.[151,1434,189,719]" captionTargetId-1="figure-70@5.[151,1436,965,1993]" captionTargetId-2="figure-7@6.[265,763,189,2055]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Figure 4. Emmelina vera Ustjuzhanin &amp; Kovtunovich, sp. nov. Adult, male, holotype." captionText-1="Figure 5. Emmelina vera Ustjuzhanin &amp; Kovtunovich, sp. nov. Male genitalia, holotype (ZISP gen.pr. Nr. 1988)." captionText-2="Figure 6. Emmelina vera Ustjuzhanin &amp; Kovtunovich, sp. nov. Female genitalia, paratype (ZISP gen.pr. Nr. 1989)." figureDoi-0="http://doi.org/10.5281/zenodo.8044296" figureDoi-1="http://doi.org/10.5281/zenodo.8044302" figureDoi-2="http://doi.org/10.5281/zenodo.8044306" httpUri-0="https://zenodo.org/record/8044296/files/figure.png" httpUri-1="https://zenodo.org/record/8044302/files/figure.png" httpUri-2="https://zenodo.org/record/8044306/files/figure.png" pageId="3" pageNumber="41">Figs 4-6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33E1290FFE6856AFF212F812E601E5B" pageId="3" pageNumber="41" type="materials_examined">
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<emphasis id="B9509D09FFE6856AFF212F812A5C1DE1" bold="true" box="[151,334,1488,1514]" pageId="3" pageNumber="41">Type material</emphasis>
:
<materialsCitation id="3B4C4B46FFE6856AFED52F8128611E06" ID-GBIF-Occurrence="4132920305" collectingDate="2011-11-10" collectingDateMax="2011-11-14" collectingDateMin="2011-11-10" collectionCode="ZISP" collectorName="U. Drechsel" country="Paraguay" latitude="-25.65" location="Tacuara Renda" longLatPrecision="1245" longitude="-56.916668" pageId="3" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Cordillera" typeStatus="holotype">
<emphasis id="B9509D09FFE6856AFED52F812AC71DE1" bold="true" box="[355,469,1488,1514]" pageId="3" pageNumber="41">
<typeStatus id="549FFFB9FFE6856AFED52F812AC71DE1" box="[355,469,1488,1514]" pageId="3" pageNumber="41" type="holotype">Holotype</typeStatus>
</emphasis>
,
<specimenCount id="9D228A92FFE6856AFE5F2F802AED1DE0" box="[489,511,1489,1515]" pageId="3" pageNumber="41" type="male">♂</specimenCount>
(
<collectionCode id="ED35D9DEFFE6856AFDA32F8129471DE0" box="[533,597,1488,1515]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="41">ZISP</collectionCode>
, gen.pr. Nr. 1988),
<emphasis id="B9509D09FFE6856AFCE42F812F101DE1" bold="true" box="[850,1026,1488,1514]" pageId="3" pageNumber="41">
<collectingCountry id="F333018BFFE6856AFCE42F8128EE1DE1" box="[850,1020,1488,1514]" name="Paraguay" pageId="3" pageNumber="41">PARAGUAY</collectingCountry>
,
</emphasis>
<collectingRegion id="49E08FF9FFE6856AFBB82F802FDF1DE0" box="[1038,1229,1488,1515]" country="Paraguay" name="Cordillera" pageId="3" pageNumber="41">Dep. Cordillera</collectingRegion>
,
<location id="8EFB17C0FFE6856AFB692F802E8A1DE0" LSID="urn:lsid:plazi:treatment:038DF00DFFE6856EFF212F1C2E5A19C1:8EFB17C0FFE6856AFB692F802E8A1DE0" box="[1247,1432,1488,1515]" country="Paraguay" latitude="-25.65" longLatPrecision="1245" longitude="-56.916668" name="Tacuara Renda" pageId="3" pageNumber="41" stateProvince="Cordillera">Tacuara Renda</location>
,
<geoCoordinate id="EE1027DCFFE6856AFF212FA32A191E06" box="[151,267,1522,1549]" degrees="25" direction="south" minutes="39" orientation="latitude" pageId="3" pageNumber="41" precision="925" value="-25.65">25°39’ S</geoCoordinate>
,
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,
<date id="FF9A67DBFFE6856AFE0E2FA329951E06" box="[440,647,1522,1549]" pageId="3" pageNumber="41" value="2011-11-10" valueMax="2011-11-14" valueMin="2011-11-10">
<collectingDate id="EFDE9E33FFE6856AFE0E2FA329951E06" box="[440,647,1522,1549]" pageId="3" pageNumber="41" value="2011-11-10" valueMax="2011-11-14" valueMin="2011-11-10">10–14. xi. 2011</collectingDate>
</date>
,
<collectorName id="26D124CDFFE6856AFD2B2FA228211E06" box="[669,819,1522,1549]" pageId="3" pageNumber="41">U. Drechsel</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B4C4B46FFE6856AFC3F2FA32E731E25" ID-GBIF-Occurrence="4132920302" collectingDate="2011-06-18" collectionCode="ZISP" collectorName="U. Drechsel" country="Paraguay" latitude="-24.916666" location="Puerto Barbero" longLatPrecision="1249" longitude="-57.3" pageId="3" pageNumber="41" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B9509D09FFE6856AFC3F2FA32F151E07" bold="true" box="[905,1031,1522,1548]" pageId="3" pageNumber="41">
<typeStatus id="549FFFB9FFE6856AFC3F2FA32F151E07" box="[905,1031,1522,1548]" pageId="3" pageNumber="41" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D228A92FFE6856AFB972FA32F571E06" box="[1057,1093,1522,1549]" pageId="3" pageNumber="41" type="female">1♀</specimenCount>
(
<collectionCode id="ED35D9DEFFE6856AFBE92FA32F8D1E06" box="[1119,1183,1522,1549]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="3" pageNumber="41">ZISP</collectionCode>
, gen.pr. Nr. 1989),
<emphasis id="B9509D09FFE6856AFF212C452A551E25" bold="true" box="[151,327,1556,1582]" pageId="3" pageNumber="41">
<collectingCountry id="F333018BFFE6856AFF212C452A531E25" box="[151,321,1556,1582]" name="Paraguay" pageId="3" pageNumber="41">PARAGUAY</collectingCountry>
,
</emphasis>
Dep. Pte. Hayes,
<location id="8EFB17C0FFE6856AFD902C4529CD1E25" LSID="urn:lsid:plazi:treatment:038DF00DFFE6856EFF212F1C2E5A19C1:8EFB17C0FFE6856AFD902C4529CD1E25" box="[550,735,1555,1582]" country="Paraguay" latitude="-24.916666" longLatPrecision="1249" longitude="-57.3" name="Puerto Barbero" pageId="3" pageNumber="41">Puerto Barbero</location>
,
<geoCoordinate id="EE1027DCFFE6856AFD582C4228491E25" box="[750,859,1555,1582]" degrees="24" direction="south" minutes="55" orientation="latitude" pageId="3" pageNumber="41" precision="925" value="-24.916666">24°55’ S</geoCoordinate>
,
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,
<date id="FF9A67DBFFE6856AFC422C422F981E25" box="[1012,1162,1555,1582]" pageId="3" pageNumber="41" value="2011-06-18">
<collectingDate id="EFDE9E33FFE6856AFC422C422F981E25" box="[1012,1162,1555,1582]" pageId="3" pageNumber="41" value="2011-06-18">18. vi. 2011</collectingDate>
</date>
,
<collectorName id="26D124CDFFE6856AFB2F2C452E3B1E25" box="[1177,1321,1555,1582]" pageId="3" pageNumber="41">U. Drechsel</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B4C4B46FFE6856AFAC62C422E601E5B" ID-GBIF-Occurrence="4132920318" collectingDate="2015-05-13" collectingDateMax="2015-05-17" collectingDateMin="2015-05-13" collectorName="U. Drechsel" country="Paraguay" latitude="-21.5" location="Laguna Jacare" longLatPrecision="1264" longitude="-57.966667" pageId="3" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Alto Paraguay" typeStatus="paratype">
<specimenCount id="9D228A92FFE6856AFAC62C422E841E25" box="[1392,1430,1555,1582]" pageId="3" pageNumber="41" type="male">1♂</specimenCount>
,
<emphasis id="B9509D09FFE6856AFF212C672A551E5B" bold="true" box="[151,327,1590,1616]" pageId="3" pageNumber="41">
<collectingCountry id="F333018BFFE6856AFF212C672A531E5B" box="[151,321,1590,1616]" name="Paraguay" pageId="3" pageNumber="41">PARAGUAY</collectingCountry>
,
</emphasis>
<collectingRegion id="49E08FF9FFE6856AFEF82C67292B1E5B" box="[334,569,1589,1616]" country="Paraguay" name="Alto Paraguay" pageId="3" pageNumber="41">Dep. Alto Paraguay</collectingRegion>
,
<location id="8EFB17C0FFE6856AFDF22C6729E31E5B" LSID="urn:lsid:plazi:treatment:038DF00DFFE6856EFF212F1C2E5A19C1:8EFB17C0FFE6856AFDF22C6729E31E5B" box="[580,753,1590,1616]" country="Paraguay" latitude="-21.5" longLatPrecision="1264" longitude="-57.966667" name="Laguna Jacare" pageId="3" pageNumber="41" stateProvince="Alto Paraguay">Laguna Jacare</location>
,
<geoCoordinate id="EE1027DCFFE6856AFD4A2C6428771E5B" box="[764,869,1589,1616]" degrees="21" direction="south" minutes="30" orientation="latitude" pageId="3" pageNumber="41" precision="925" value="-21.5">21°30’ S</geoCoordinate>
,
<geoCoordinate id="EE1027DCFFE6856AFCC42C6428FA1E5B" box="[882,1000,1589,1616]" degrees="57" direction="west" minutes="58" orientation="longitude" pageId="3" pageNumber="41" precision="925" value="-57.966667">57°58’ W</geoCoordinate>
,
<date id="FF9A67DBFFE6856AFC422C642FBA1E5B" box="[1012,1192,1589,1616]" pageId="3" pageNumber="41" value="2015-05-13" valueMax="2015-05-17" valueMin="2015-05-13">
<collectingDate id="EFDE9E33FFE6856AFC422C642FBA1E5B" box="[1012,1192,1589,1616]" pageId="3" pageNumber="41" value="2015-05-13" valueMax="2015-05-17" valueMin="2015-05-13">13–17. v. 2015</collectingDate>
</date>
,
<collectorName id="26D124CDFFE6856AFB052C672E2D1E5B" box="[1203,1343,1589,1616]" pageId="3" pageNumber="41">U. Drechsel</collectorName>
leg.
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</paragraph>
</subSubSection>
<subSubSection id="C33E1290FFE6856CFF402C0929601BB3" lastPageId="5" lastPageNumber="43" pageId="3" pageNumber="41" type="description">
<paragraph id="8B9B411BFFE6856AFF402C09286C1F37" blockId="3.[151,1437,1488,2021]" pageId="3" pageNumber="41">
<emphasis id="B9509D09FFE6856AFF402C092ADB1E79" bold="true" box="[246,457,1624,1650]" pageId="3" pageNumber="41">DESCRIPTION.</emphasis>
External characters. Head, thorax and tegulae with grey-brown scales. Collar on head brown. Labial palpi thin, straight, slightly longer than eye diameter in length. Antennae thin, lightbrown, scape slightly thickened. Wingspan
<quantity id="4CDCECFEFFE6856AFD142CCA280C1EBD" box="[674,798,1691,1718]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="3" pageNumber="41" unit="mm" value="21.5" valueMax="23.0" valueMin="20.0">20-23 mm</quantity>
(
<typeStatus id="549FFFB9FFE6856AFC992CCA288A1EBD" box="[815,920,1691,1718]" pageId="3" pageNumber="41" type="holotype">holotype</typeStatus>
<quantity id="4CDCECFEFFE6856AFC172CCA28E81EBD" box="[929,1018,1691,1718]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="41" unit="mm" value="23.0">23 mm</quantity>
). Fore wings grey-brown sputtered with dark-brown scales. Elongated dark-brown spot along costal margin above cleft. Small dark-brown spot on outer margin of first lobe. Both lobes noticeably darkened apically and framed with dark-brown scales. Big blurred brownish spot in front of cleft. Fringe inside cleft dark-grey. Hind wings unicolorous, noticeably lighter colored than fore wings, with light-grey fringe on lobes.
</paragraph>
<paragraph id="8B9B411BFFE6856AFF402D152A951FEE" blockId="3.[151,1437,1488,2021]" pageId="3" pageNumber="41">
<emphasis id="B9509D09FFE6856AFF402D152AA01F55" bold="true" box="[246,434,1860,1886]" pageId="3" pageNumber="41">Male genitalia.</emphasis>
Valves asymmetric. Saccular process on left valve long, apically arched, tip hook-like, curved, acute. Saccular process on right valve shaped as wide fold located in medium part of valve. Uncus simple, thin, slightly curved. Anellus arms asymmetric, left arm basally wide then narrowing, right arm wide in all length, slightly shorter than left arm. Aedeagus thin, slightly curved basally, slightly shorter than valve in length.
</paragraph>
<caption id="DF5B1193FFE1856DFF202D812BCF100D" ID-DOI="http://doi.org/10.5281/zenodo.8044294" ID-Zenodo-Dep="8044294" httpUri="https://zenodo.org/record/8044294/files/figure.png" pageId="4" pageNumber="42" startId="4.[150,226,2000,2024]" targetBox="[466,1201,189,1981]" targetPageId="4" targetType="figure">
<paragraph id="8B9B411BFFE1856DFF202D812BCF100D" blockId="4.[150,1437,1999,2054]" pageId="4" pageNumber="42">
<emphasis id="B9509D09FFE1856DFF202D812BEB1FE3" bold="true" box="[150,249,2000,2024]" pageId="4" pageNumber="42">Figure 3</emphasis>
.
<taxonomicName id="4C243A98FFE1856DFEBC2D8128551FEC" authority="Ustjuzhanin &amp; Kovtunovich" authorityName="Ustjuzhanin &amp; Kovtunovich" authorityYear="2022" box="[266,839,1999,2024]" class="Insecta" family="Pterophoridae" genus="Michaelophorus" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="42" phylum="Arthropoda" rank="species" species="maria" status="sp. nov.">
<emphasis id="B9509D09FFE1856DFEBC2D8129181FE3" bold="true" box="[266,522,2000,2024]" italics="true" pageId="4" pageNumber="42">Michaelophorus maria</emphasis>
Ustjuzhanin &amp; Kovtunovich
</taxonomicName>
<emphasis id="B9509D09FFE1856DFCE72D8128B81FE3" bold="true" box="[849,938,2000,2024]" pageId="4" pageNumber="42">
<taxonomicNameLabel id="A2632072FFE1856DFCE72D8128B81FE3" box="[849,938,2000,2024]" pageId="4" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female genitalia, paratype (ZISP gen.pr. Nr. 1990).
</paragraph>
</caption>
<caption id="DF5B1193FFE0856CFF2028A62F2B1B04" ID-DOI="http://doi.org/10.5281/zenodo.8044296" ID-Zenodo-Dep="8044296" box="[150,1081,759,783]" httpUri="https://zenodo.org/record/8044296/files/figure.png" pageId="5" pageNumber="43" startId="5.[150,224,759,783]" targetBox="[164,1415,199,713]" targetPageId="5" targetType="figure">
<paragraph id="8B9B411BFFE0856CFF2028A62F2B1B04" blockId="5.[150,1081,759,783]" box="[150,1081,759,783]" pageId="5" pageNumber="43">
<emphasis id="B9509D09FFE0856CFF2028A62BE11B04" bold="true" box="[150,243,759,783]" pageId="5" pageNumber="43">Figure 4</emphasis>
.
<taxonomicName id="4C243A98FFE0856CFEB628A629F21B04" authority="Ustjuzhanin &amp; Kovtunovich" authorityName="Ustjuzhanin &amp; Kovtunovich" authorityYear="2022" box="[256,736,759,783]" class="Insecta" family="Pterophoridae" genus="Emmelina" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="vera" status="sp. nov.">
<emphasis id="B9509D09FFE0856CFEB628A62ABA1B04" bold="true" box="[256,424,759,783]" italics="true" pageId="5" pageNumber="43">Emmelina vera</emphasis>
Ustjuzhanin &amp; Kovtunovich
</taxonomicName>
,
<emphasis id="B9509D09FFE0856CFD5D28A628501B04" bold="true" box="[747,834,759,783]" pageId="5" pageNumber="43">
<taxonomicNameLabel id="A2632072FFE0856CFD5D28A628501B04" box="[747,834,759,783]" pageId="5" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Adult, male, holotype.
</paragraph>
</caption>
<paragraph id="8B9B411BFFE0856CFF21290B29601BB3" blockId="5.[151,1436,857,952]" pageId="5" pageNumber="43">
<emphasis id="B9509D09FFE0856CFF21290B2A661B7F" bold="true" box="[151,372,858,884]" pageId="5" pageNumber="43">Female genitalia.</emphasis>
Papillae anales wide, triangle, apically acute. Posterior apophyses straight. Anterior apophyses absent. Antrum short, tubulate. Ductus bursae long, thin, membranous. Bursa copulatrix small, round, poorly sclerotized, without signa.
</paragraph>
</subSubSection>
<caption id="DF5B1193FFE0856CFF202D8A2E491FF9" ID-DOI="http://doi.org/10.5281/zenodo.8044302" ID-Zenodo-Dep="8044302" box="[150,1371,2011,2035]" httpUri="https://zenodo.org/record/8044302/files/figure.png" pageId="5" pageNumber="43" startId="5.[150,224,2011,2035]" targetBox="[174,1387,1039,1928]" targetPageId="5" targetType="figure">
<paragraph id="8B9B411BFFE0856CFF202D8A2E491FF9" blockId="5.[150,1371,2011,2035]" box="[150,1371,2011,2035]" pageId="5" pageNumber="43">
<emphasis id="B9509D09FFE0856CFF202D8A2BE11FF8" bold="true" box="[150,243,2011,2035]" pageId="5" pageNumber="43">Figure 5</emphasis>
.
<taxonomicName id="4C243A98FFE0856CFEB62D8A29F21FF8" authority="Ustjuzhanin &amp; Kovtunovich" authorityName="Ustjuzhanin &amp; Kovtunovich" authorityYear="2022" box="[256,736,2011,2035]" class="Insecta" family="Pterophoridae" genus="Emmelina" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="43" phylum="Arthropoda" rank="species" species="vera" status="sp. nov.">
<emphasis id="B9509D09FFE0856CFEB62D8A2ABA1FF8" bold="true" box="[256,424,2011,2035]" italics="true" pageId="5" pageNumber="43">Emmelina vera</emphasis>
Ustjuzhanin &amp; Kovtunovich
</taxonomicName>
,
<emphasis id="B9509D09FFE0856CFD5D2D8A28501FF8" bold="true" box="[747,834,2011,2035]" pageId="5" pageNumber="43">
<taxonomicNameLabel id="A2632072FFE0856CFD5D2D8A28501FF8" box="[747,834,2011,2035]" pageId="5" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male genitalia, holotype (ZISP gen.pr. Nr. 1988).
</paragraph>
</caption>
<caption id="DF5B1193FFE3856FFC152DC42E1C1FE2" ID-DOI="http://doi.org/10.5281/zenodo.8044306" ID-Zenodo-Dep="8044306" httpUri="https://zenodo.org/record/8044306/files/figure.png" pageId="6" pageNumber="44" startId="6.[931,1005,1941,1965]" targetBox="[265,763,189,2055]" targetPageId="6" targetType="figure">
<paragraph id="8B9B411BFFE3856FFC152DC42E1C1FE2" blockId="6.[931,1407,1940,2026]" pageId="6" pageNumber="44">
<emphasis id="B9509D09FFE3856FFC152DC42F1A1FA6" bold="true" box="[931,1032,1941,1965]" pageId="6" pageNumber="44">Figure 6</emphasis>
.
<taxonomicName id="4C243A98FFE3856FFBAB2DC42F211FC0" authority="Ustjuzhanin &amp; Kovtunovich" authorityName="Ustjuzhanin &amp; Kovtunovich" authorityYear="2022" class="Insecta" family="Pterophoridae" genus="Emmelina" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="44" phylum="Arthropoda" rank="species" species="vera" status="sp. nov.">
<emphasis id="B9509D09FFE3856FFBAB2DC42FDF1FA6" bold="true" box="[1053,1229,1941,1965]" italics="true" pageId="6" pageNumber="44">Emmelina vera</emphasis>
Ustjuzhanin &amp; Kovtunovich
</taxonomicName>
,
<emphasis id="B9509D09FFE3856FFBFF2DE22FB81FC0" bold="true" box="[1097,1194,1971,1995]" pageId="6" pageNumber="44">
<taxonomicNameLabel id="A2632072FFE3856FFBFF2DE22FB81FC0" box="[1097,1194,1971,1995]" pageId="6" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Female genitalia, paratype (ZISP gen.pr. Nr. 1989).
</paragraph>
</caption>
<subSubSection id="C33E1290FFE2856EFF402A9529A5198C" pageId="7" pageNumber="45" type="diagnosis">
<paragraph id="8B9B411BFFE2856EFF402A952F96196E" blockId="7.[151,1436,195,458]" pageId="7" pageNumber="45">
<emphasis id="B9509D09FFE2856EFF402A952A6418D5" bold="true" box="[246,374,196,222]" pageId="7" pageNumber="45">Diagnosis.</emphasis>
In the male genitalia, in the curved saccular process on the left valve, the species resembles
<taxonomicName id="4C243A98FFE2856EFEA12AB4283A18F4" authority="(Barnes &amp; Lindsey, 1921)" baseAuthorityName="Barnes &amp; Lindsey" baseAuthorityYear="1921" box="[279,808,229,256]" class="Insecta" family="Pterophoridae" genus="Emmelina" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="buscki">
<emphasis id="B9509D09FFE2856EFEA12AB42AF1190B" box="[279,483,229,256]" italics="true" pageId="7" pageNumber="45">Emmelina buscki</emphasis>
(Barnes &amp; Lindsey, 1921
<emphasis id="B9509D09FFE2856EFCA82AB4283A18F4" bold="true" box="[798,808,229,255]" pageId="7" pageNumber="45">)</emphasis>
</taxonomicName>
<emphasis id="B9509D09FFE2856EFC9E2AB4283D18F4" bold="true" box="[808,815,229,255]" pageId="7" pageNumber="45">,</emphasis>
but in
<taxonomicName id="4C243A98FFE2856EFC3D2AB728EB190B" baseAuthorityName="Barnes &amp; Lindsey" baseAuthorityYear="1921" box="[907,1017,229,256]" class="Insecta" family="Pterophoridae" genus="Emmelina" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="buscki">
<emphasis id="B9509D09FFE2856EFC3D2AB728EB190B" box="[907,1017,229,256]" italics="true" pageId="7" pageNumber="45">E. buscki</emphasis>
</taxonomicName>
this process is curved towards the lover margin of the valve, while in the new species it is curved towards the upper margin. Additionally, the new species differs well from
<taxonomicName id="4C243A98FFE2856EFDBD2B7B29681948" baseAuthorityName="Barnes &amp; Lindsey" baseAuthorityYear="1921" box="[523,634,296,323]" class="Insecta" family="Pterophoridae" genus="Emmelina" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="buscki">
<emphasis id="B9509D09FFE2856EFDBD2B7B29681948" box="[523,634,296,323]" italics="true" pageId="7" pageNumber="45">E. buscki</emphasis>
</taxonomicName>
in the special shape of the anellus arms and the aedeagus. In the female genitalia, the new species differs from
<taxonomicName id="4C243A98FFE2856EFD002B1A2833196E" baseAuthorityName="Barnes &amp; Lindsey" baseAuthorityYear="1921" box="[694,801,330,357]" class="Insecta" family="Pterophoridae" genus="Emmelina" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="45" phylum="Arthropoda" rank="species" species="buscki">
<emphasis id="B9509D09FFE2856EFD002B1A2833196E" box="[694,801,330,357]" italics="true" pageId="7" pageNumber="45">E. buscki</emphasis>
</taxonomicName>
in the round bursa copulatrix.
</paragraph>
<paragraph id="8B9B411BFFE2856EFF402B3D29A5198C" blockId="7.[151,1436,195,458]" box="[246,695,364,391]" pageId="7" pageNumber="45">
<emphasis id="B9509D09FFE2856EFF402B3D2AB1198D" bold="true" box="[246,419,364,390]" pageId="7" pageNumber="45">Flight period:</emphasis>
May, June, November.
</paragraph>
</subSubSection>
<subSubSection id="C33E1290FFE2856EFF402BDF290619A2" box="[246,532,398,425]" pageId="7" pageNumber="45" type="distribution">
<paragraph id="8B9B411BFFE2856EFF402BDF290619A2" blockId="7.[151,1436,195,458]" box="[246,532,398,425]" pageId="7" pageNumber="45">
<emphasis id="B9509D09FFE2856EFF402BDF2A8A19A3" bold="true" box="[246,408,398,424]" pageId="7" pageNumber="45">Distribution:</emphasis>
<collectingCountry id="F333018BFFE2856EFE162BDE291F19A2" box="[416,525,399,425]" name="Paraguay" pageId="7" pageNumber="45">Paraguay</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E1290FFE2856EFF402BE12E5A19C1" box="[246,1352,431,458]" pageId="7" pageNumber="45" type="etymology">
<paragraph id="8B9B411BFFE2856EFF402BE12E5A19C1" blockId="7.[151,1436,195,458]" box="[246,1352,431,458]" pageId="7" pageNumber="45">
<emphasis id="B9509D09FFE2856EFF402BE12A9419C1" bold="true" box="[246,390,432,458]" pageId="7" pageNumber="45">Etymology:</emphasis>
The species is named after Vera Aleynikova (Medvedeva) (
<collectingRegion id="49E08FF9FFE2856EFBFE2BFE2FCE19C1" box="[1096,1244,431,458]" country="Russia" name="Novosibirsk" pageId="7" pageNumber="45">Novosibirsk</collectingRegion>
,
<collectingCountry id="F333018BFFE2856EFB5E2BFE2E2E19C1" box="[1256,1340,431,458]" name="Russia" pageId="7" pageNumber="45">Russia</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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