170 lines
19 KiB
XML
170 lines
19 KiB
XML
<document id="459FFCB92A3A891B632FF30807D796B1" ID-DOI="10.5281/zenodo.4525461" ID-ISSN="1638-9387" ID-Zenodo-Dep="4525461" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612878885009" checkinUser="felipe" docAuthor="Mah, Christopher L." docDate="2006" docId="038D664EFF90FFB31542AA97FF3D6151" docLanguage="en" docName="Zoosystema.28.4.917-954.pdf" docOrigin="Zoosystema 28 (4)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Floriaster maya Downey 1980" docType="treatment" docVersion="3" lastPageNumber="927" masterDocId="FFB41E36FF99FFB9164AAE59FFD46376" masterDocTitle="Phylogeny and biogeography of the deep-sea goniasterid Circeaster (Echinodermata, Asteroidea, Goniasteridae) including descriptions of six new species" masterLastPageNumber="954" masterPageNumber="917" pageNumber="926" updateTime="1698868963781" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="C4F13816BA04404504ECA14F523E67D0">
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<mods:title id="6AC8DDCD556AD91F45D8F7F45E950D0A">Phylogeny and biogeography of the deep-sea goniasterid Circeaster (Echinodermata, Asteroidea, Goniasteridae) including descriptions of six new species</mods:title>
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<mods:name id="D02A7B1722664A741AF3060BE1A8B1C1" type="personal">
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<mods:roleTerm id="2C07AB69CDB1C929033B675F827BD437">Author</mods:roleTerm>
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<mods:namePart id="302D2012317DB10482F42C43595B68A9">Mah, Christopher L.</mods:namePart>
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<mods:affiliation id="0525774D05012FA9E4FE4F4F6AE1917F">Department of Geology, University of Illinois at Urbana-Champaign, 245 NHB, 1301 W. Green, Urbana, Illinois 61801 (USA) Present address: Department of Invertebrate Zoology, National Museum of Natural History, Mail Stop MRC 163, P. O. Box 37012, Washington, DC, 20013 - 7012 (USA) mahch @ si. edu</mods:affiliation>
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<mods:nameIdentifier id="B9BAD35003B408E60E1CFC34A0F95092" type="email">mahch@si.edu</mods:nameIdentifier>
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<mods:typeOfResource id="45B4937C8A6DB7CAC20D8616971437AA">text</mods:typeOfResource>
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<mods:titleInfo id="177483365C88E787C39894E51040A2B5">
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<mods:title id="192CCE2EFB6E8852DB3D43F9BE50CC16">Zoosystema</mods:title>
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</mods:titleInfo>
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<mods:part id="06D7BB73A039E0578768B201EAF660BB">
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<mods:date id="86769E4CE815266BA45E30792CB77490">2006</mods:date>
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<mods:detail id="7C573232EDE57145728F374D0A308127" type="volume">
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<mods:number id="BF5B54CCCEB222E1795AEF6292A06050">28</mods:number>
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<mods:number id="DC34F1A24F3E992613B587697C9BC1B9">4</mods:number>
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<mods:classification id="7AF9F580AD279E123292D589C6A10822">journal article</mods:classification>
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<mods:identifier id="BBCE69048B8464BA455C500FAB1AA5F6" type="DOI">10.5281/zenodo.4525461</mods:identifier>
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<mods:identifier id="CF74A78C66FA1E0BDD7BD81C9ECB2055" type="ISSN">1638-9387</mods:identifier>
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<treatment id="038D664EFF90FFB31542AA97FF3D6151" LSID="urn:lsid:plazi:treatment:038D664EFF90FFB31542AA97FF3D6151" httpUri="http://treatment.plazi.org/id/038D664EFF90FFB31542AA97FF3D6151" lastPageId="10" lastPageNumber="927" pageId="9" pageNumber="926">
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<subSubSection id="C33E84D3FF90FFB01542AA97FB82679E" box="[776,1110,1230,1256]" pageId="9" pageNumber="926" type="nomenclature">
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<paragraph id="8B9BD758FF90FFB01542AA97FB82679E" blockId="9.[776,1110,1230,1288]" box="[776,1110,1230,1256]" pageId="9" pageNumber="926">
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<heading id="D0D36034FF90FFB01542AA97FB82679E" box="[776,1110,1230,1256]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="926" reason="2">
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<taxonomicName id="4C24ACDBFF90FFB01542AA97FB82679E" ID-CoL="6J9Y6" authority="Downey, 1980" authorityName="Downey" authorityYear="1980" box="[776,1110,1230,1256]" class="Asteroidea" family="Goniasteridae" genus="Floriaster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="926" phylum="Echinodermata" rank="species" species="maya">
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<emphasis id="B9500B4AFF90FFB01542AA97FC65679E" bold="true" box="[776,945,1230,1256]" italics="true" pageId="9" pageNumber="926">Floriaster maya</emphasis>
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<bibRefCitation id="EFB5AAA9FF90FFB015FDAA97FB82679E" author="DOWNEY M. E." box="[951,1110,1230,1256]" pageId="9" pageNumber="926" pagination="346 - 349" refId="ref23352" refString="DOWNEY M. E. 1980. - Floriaster maya, new genus and species of the family Goniasteridae. Proceedings of the Biological Society of Washington 93 (2): 346 - 349." type="journal article" year="1980">Downey, 1980</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E84D3FF90FFB0153AAAB7FC3A667E" box="[880,1006,1262,1288]" pageId="9" pageNumber="926" type="description">
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<paragraph id="8B9BD758FF90FFB0153AAAB7FC3A667E" blockId="9.[776,1110,1230,1288]" box="[880,1006,1262,1288]" pageId="9" pageNumber="926">
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(
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<figureCitation id="131FCBDDFF90FFB01532AAB7FC32667E" box="[888,998,1262,1288]" captionStart="FIG" captionStartId="8.[113,124,1634,1651]" captionTargetBox="[119,1193,217,1578]" captionTargetId="figure-18@8.[116,665,754,1305]" captionTargetPageId="8" captionText="FIG. 2. — A-D, Lydiaster johannae (MNHN EcAs 11724); A, abactinal surface; B, pedicellariae; C, abactinal arm surface; D, actinal surface; E, F, Floriaster maya (NMNH E38662); E, abactinal surface; F, actinal surface; G, H, Cladaster analogus (NMNH E38569); G, abactinal surface; H, actinal surface. Scale bars: A, C-E, G, 1.0 cm; B, F, H, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.4525465" httpUri="https://zenodo.org/record/4525465/files/figure.png" pageId="9" pageNumber="926">Fig. 2E, F</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E84D3FF90FFB014EFAB7AFB786623" pageId="9" pageNumber="926" type="reference_group">
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<paragraph id="8B9BD758FF90FFB014EFAB7AFB786623" blockId="9.[675,1210,1315,1461]" pageId="9" pageNumber="926">
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<treatmentCitationGroup id="AB34F076FF90FFB014EFAB7AFB786623" pageId="9" pageNumber="926">
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<treatmentCitation id="0A85F149FF90FFB014EFAB7AFC25664C" author="DOWNEY M. E." box="[677,1009,1315,1338]" page="105" pageId="9" pageNumber="926" year="1980">
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<taxonomicName id="4C24ACDBFF90FFB014EFAB7AFC25664C" ID-CoL="6J9Y6" authority="Downey, 1980: 105" authorityName="Downey" authorityPageNumber="105" authorityYear="1980" box="[677,1009,1315,1338]" class="Asteroidea" family="Goniasteridae" genus="Floriaster" kingdom="Animalia" order="Valvatida" pageId="9" pageNumber="926" phylum="Echinodermata" rank="species" species="maya">
|
||
<emphasis id="B9500B4AFF90FFB014EFAB7AFCFB664C" box="[677,815,1315,1338]" italics="true" pageId="9" pageNumber="926">Floriaster maya</emphasis>
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<bibRefCitation id="EFB5AAA9FF90FFB0157EAB7AFC25664C" author="DOWNEY M. E." box="[820,1009,1315,1338]" pageId="9" pageNumber="926" pagination="346 - 349" refId="ref23352" refString="DOWNEY M. E. 1980. - Floriaster maya, new genus and species of the family Goniasteridae. Proceedings of the Biological Society of Washington 93 (2): 346 - 349." type="journal article" year="1980">Downey, 1980: 105</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
, figs 1, 2. —
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<treatmentCitation id="0A85F149FF90FFB01221AB7AFCB76623" author="CLARK A. M. & DOWNEY M. E." page="244" pageId="9" pageNumber="926" year="1992">
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<bibRefCitation id="EFB5AAA9FF90FFB01221AB7AFCB76623" author="CLARK A. M. & DOWNEY M. E." pageId="9" pageNumber="926" refId="ref23167" refString="CLARK A. M. & DOWNEY M. E. 1992. - Starfishes of the Atlantic. Chapman and Hall, London, 794 p." type="book" year="1992">Clark & Downey 1992: 244</bibRefCitation>
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</treatmentCitation>
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||
, pl. 56G-H. —
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<treatmentCitation id="0A85F149FF90FFB01249AB67FB7C6623" author="CLARK A. M." box="[1027,1192,1342,1365]" page="253" pageId="9" pageNumber="926" year="1993">
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<bibRefCitation id="EFB5AAA9FF90FFB01249AB67FB7C6623" author="CLARK A. M." box="[1027,1192,1342,1365]" pageId="9" pageNumber="926" pagination="187 - 366" refId="ref23139" refString="CLARK A. M. 1993. - An index of names of recent Asteroidea Part 2: Valvatida. Echinoderm Studies 4: 187 - 366." type="journal article" year="1993">Clark 1993: 253</bibRefCitation>
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||
</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C33E84D3FF90FFB014EFAB30FBAD652A" pageId="9" pageNumber="926" type="materials_examined">
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<paragraph id="8B9BD758FF90FFB014EFAB30FB6E66C3" blockId="9.[675,1210,1315,1461]" pageId="9" pageNumber="926">
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<materialsCitation id="3B4CDD05FF90FFB014EFAB30FB6E66C3" accessionNumber="E18324" collectingDate="1970-07-06" collectionCode="R, NMNH" elevation="979" latitude="20.75" location="Channel" longLatPrecision="1266" longitude="-86.45" pageId="9" pageNumber="926" specimenCount="1" stateProvince="Yucatan" typeStatus="holotype">
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<typeStatus id="549F69FAFF90FFB014EFAB30FCD96609" box="[677,781,1385,1408]" pageId="9" pageNumber="926" type="holotype">HOLOTYPE</typeStatus>
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||
. —
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<emphasis id="B9500B4AFF90FFB0157DAB33FC1E66F7" bold="true" box="[823,970,1386,1409]" pageId="9" pageNumber="926">Caribbean Sea.</emphasis>
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||
Southern end of
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||
<collectingRegion id="49E019BAFF90FFB01227AB30FB6E66F6" box="[1133,1210,1385,1408]" country="Mexico" name="Yucatan" pageId="9" pageNumber="926">Yucatan</collectingRegion>
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<location id="8EFB8183FF90FFB014E9ABDDFD2D66ED" LSID="urn:lsid:plazi:treatment:038D664EFF90FFB31542AA97FF3D6151:8EFB8183FF90FFB014E9ABDDFD2D66ED" box="[675,761,1412,1435]" latitude="20.75" longLatPrecision="1266" longitude="-86.45" name="Channel" pageId="9" pageNumber="926" stateProvince="Yucatan">Channel</location>
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,
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<geoCoordinate id="EE10B19FFF90FFB0154BABDDFC8366ED" box="[769,855,1412,1435]" degrees="20" direction="north" minutes="45" orientation="latitude" pageId="9" pageNumber="926" precision="925" value="20.75">20°45’N</geoCoordinate>
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,
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<geoCoordinate id="EE10B19FFF90FFB0152AABDDFC6366ED" box="[864,951,1412,1435]" degrees="86" direction="west" minutes="27" orientation="longitude" pageId="9" pageNumber="926" precision="925" value="-86.45">86°27’W</geoCoordinate>
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,
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<quantity id="4CDC7ABDFF90FFB0158AABDDFBEF66ED" box="[960,1083,1412,1435]" metricMagnitude="2" metricUnit="m" metricValue="9.785" metricValueMax="10.24" metricValueMin="9.33" pageId="9" pageNumber="926" unit="m" value="978.5" valueMax="1024.0" valueMin="933.0">
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<elevation id="0009306BFF90FFB0158AABDDFBEF66ED" box="[960,1083,1412,1435]" metricMagnitude="2" metricUnit="m" metricValue="9.785" metricValueMax="10.24" metricValueMin="9.33" pageId="9" pageNumber="926" unit="m" value="978.5" valueMax="1024.0" valueMin="933.0">933-1024 m</elevation>
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</quantity>
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,
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<date id="FF9AF198FF90FFB0120EABDDFB6066ED" box="[1092,1204,1412,1435]" pageId="9" pageNumber="926" value="1970-07-06">
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<collectingDate id="EFDE0870FF90FFB0120EABDDFB6066ED" box="[1092,1204,1412,1435]" pageId="9" pageNumber="926" value="1970-07-06">6.VII.1970</collectingDate>
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||
</date>
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||
, 1 dry spec.,
|
||
<collectionCode id="ED354F9DFF90FFB01553ABC7FCFD66C3" box="[793,809,1438,1461]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="926">R</collectionCode>
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||
= 4.0 cm, r = 2.0 cm (
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||
<collectionCode id="ED354F9DFF90FFB0124EABC7FB8C66C3" box="[1028,1112,1438,1461]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="926" type="Museum">NMNH</collectionCode>
|
||
<accessionNumber id="94774ABBFF90FFB01214ABC7FB7B66C3" box="[1118,1199,1438,1461]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/E18324" pageId="9" pageNumber="926" type="EnaNcbi">E18324</accessionNumber>
|
||
).
|
||
</materialsCitation>
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||
</paragraph>
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<paragraph id="8B9BD758FF90FFB014EFAB92FBAD652A" blockId="9.[675,1210,1482,1628]" pageId="9" pageNumber="926">
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<materialsCitation id="3B4CDD05FF90FFB014EFAB92FB5C6560" collectingDate="1980-09-21" collectionCode="OTHER, MATERIAL, EXAMINED, R" country="Bahamas" elevation="1088" latitude="28.1" location="Blake Plateau" longLatPrecision="1233" longitude="-77.13333" municipality="North Atlantic Ocean" pageId="9" pageNumber="926" specimenCount="1">
|
||
<collectionCode id="ED354F9DFF90FFB014EFAB92FC786696" box="[677,940,1483,1507]" pageId="9" pageNumber="926">OTHER MATERIAL EXAMINED</collectionCode>
|
||
. —
|
||
<emphasis id="B9500B4AFF90FFB0159FAB93FBE76697" bold="true" box="[981,1075,1482,1505]" pageId="9" pageNumber="926">
|
||
<collectingCountry id="F33397C8FF90FFB0159FAB93FBFA6697" box="[981,1070,1482,1505]" name="Bahamas" pageId="9" pageNumber="926">Bahamas</collectingCountry>
|
||
.
|
||
</emphasis>
|
||
<location id="8EFB8183FF90FFB0127DAB93FB636697" LSID="urn:lsid:plazi:treatment:038D664EFF90FFB31542AA97FF3D6151:8EFB8183FF90FFB0127DAB93FB636697" box="[1079,1207,1482,1505]" country="Bahamas" latitude="28.1" longLatPrecision="1233" longitude="-77.13333" municipality="North Atlantic Ocean" name="Blake Plateau" pageId="9" pageNumber="926">Blake Plateau</location>
|
||
,
|
||
<collectingMunicipality id="6BFF4D22FF90FFB014E9ABBCFCA4668A" box="[675,880,1509,1532]" pageId="9" pageNumber="926">North Atlantic Ocean</collectingMunicipality>
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,
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<geoCoordinate id="EE10B19FFF90FFB0153DABBCFC1E668A" box="[887,970,1509,1532]" degrees="28" direction="north" minutes="06" orientation="latitude" pageId="9" pageNumber="926" precision="925" value="28.1">28°06’N</geoCoordinate>
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||
,
|
||
<geoCoordinate id="EE10B19FFF90FFB01598ABBCFBF3668A" box="[978,1063,1509,1532]" degrees="77" direction="west" minutes="08" orientation="longitude" pageId="9" pageNumber="926" precision="925" value="-77.13333">77°08’W</geoCoordinate>
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||
,
|
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<quantity id="4CDC7ABDFF90FFB01265ABBCFB60668A" box="[1071,1204,1509,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.0879999999999999" metricValueMax="1.153" metricValueMin="1.023" pageId="9" pageNumber="926" unit="m" value="1088.0" valueMax="1153.0" valueMin="1023.0">
|
||
<elevation id="0009306BFF90FFB01265ABBCFB60668A" box="[1071,1204,1509,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.0879999999999999" metricValueMax="1.153" metricValueMin="1.023" pageId="9" pageNumber="926" unit="m" value="1088.0" valueMax="1153.0" valueMin="1023.0">1023-1153 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FF9AF198FF90FFB014E9ABA6FCC26560" box="[675,790,1535,1558]" pageId="9" pageNumber="926" value="1980-09-21">
|
||
<collectingDate id="EFDE0870FF90FFB014E9ABA6FCC26560" box="[675,790,1535,1558]" pageId="9" pageNumber="926" value="1980-09-21">21.IX.1980</collectingDate>
|
||
</date>
|
||
, 2 dry specs,
|
||
<collectionCode id="ED354F9DFF90FFB015D7ABA6FC796560" box="[925,941,1535,1558]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="926">R</collectionCode>
|
||
=
|
||
<quantity id="4CDC7ABDFF90FFB01583ABA6FBC56560" box="[969,1041,1535,1558]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="9" pageNumber="926" unit="cm" value="3.9">3.9 cm</quantity>
|
||
, r = 2.0 cm
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B4CDD05FF90FFB012D9ABA6FBA1652A" accessionNumber="E38662" collectionCode="R, NMNH" country="Bahamas" elevation="1043" pageId="9" pageNumber="926" specimenCount="1">
|
||
<collectionCode id="ED354F9DFF90FFB012D9ABA6FB776560" box="[1171,1187,1535,1558]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="926">R</collectionCode>
|
||
=
|
||
<quantity id="4CDC7ABDFF90FFB014EFA843FD386547" box="[677,748,1562,1585]" metricMagnitude="-2" metricUnit="m" metricValue="5.1" pageId="9" pageNumber="926" unit="cm" value="5.1">5.1 cm</quantity>
|
||
, r =
|
||
<quantity id="4CDC7ABDFF90FFB0155DA843FC8F6547" box="[791,859,1562,1585]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="926" unit="cm" value="2.1">2.1 cm</quantity>
|
||
(
|
||
<collectionCode id="ED354F9DFF90FFB01523A843FC696547" box="[873,957,1562,1585]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="926" type="Museum">NMNH</collectionCode>
|
||
E38662). DISTRIBUTION. — Caribbean Sea.
|
||
<quantity id="4CDC7ABDFF90FFB015BDA81CFBA1652A" box="[1015,1141,1605,1628]" metricMagnitude="3" metricUnit="m" metricValue="1.043" metricValueMax="1.153" metricValueMin="0.933" pageId="9" pageNumber="926" unit="m" value="1043.0" valueMax="1153.0" valueMin="933.0">
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<elevation id="0009306BFF90FFB015BDA81CFBA1652A" box="[1015,1141,1605,1628]" metricMagnitude="3" metricUnit="m" metricValue="1.043" metricValueMax="1.153" metricValueMin="0.933" pageId="9" pageNumber="926" unit="m" value="1043.0" valueMax="1153.0" valueMin="933.0">933-1153 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.
|
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</paragraph>
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</subSubSection>
|
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<subSubSection id="C33E84D3FF90FFB314EFA828FF3D6151" lastPageId="10" lastPageNumber="927" pageId="9" pageNumber="926" type="diagnosis">
|
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<paragraph id="8B9BD758FF90FFB014EFA828FB6F65D5" blockId="9.[677,1211,1649,1699]" pageId="9" pageNumber="926">DIAGNOSIS. — R/r ≈ 2.0-2.5. Abactinal plates round to polygonal in shape small, irregularly distributed</paragraph>
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<paragraph id="8B9BD758FF93FFB3163BAE81FDC862B4" blockId="10.[111,648,216,450]" pageId="10" pageNumber="927">
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relative to
|
||
<taxonomicName id="4C24ACDBFF93FFB3169EAE81FEF86399" authorityName="Verrill" authorityYear="1899" box="[212,300,216,239]" class="Asteroidea" family="Goniasteridae" genus="Cladaster" kingdom="Animalia" order="Valvatida" pageId="10" pageNumber="927" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B9500B4AFF93FFB3169EAE81FEF86399" box="[212,300,216,239]" italics="true" pageId="10" pageNumber="927">Cladaster</emphasis>
|
||
</taxonomicName>
|
||
. Enlarged, round tubercles covering abactinal, actinal surfaces. Abactinal, actinal plates with coarse, angular peripheral granules. Marginal plates with tubercles and coarse, nubbin-like granules. Approximately 30 marginal plates present from interradius to interradius. Interradial arcs linear. Paddle-like pedicellariae on adambulacral plates; subambulacral spine, enlarged, blunt, abradial to pedicellariae. Furrow spines five to seven, compressed, angular in cross-section.
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||
</paragraph>
|
||
<paragraph id="8B9BD758FF93FFB3163BAF85FF3D6151" blockId="10.[113,646,475,551]" pageId="10" pageNumber="927">
|
||
APOMORPHY LIST. — Nodes 24 to
|
||
<taxonomicName id="4C24ACDBFF93FFB3178EAF82FD806284" authorityName="Downey" authorityYear="1980" box="[452,596,475,498]" class="Asteroidea" family="Goniasteridae" genus="Floriaster" kingdom="Animalia" order="Valvatida" pageId="10" pageNumber="927" phylum="Echinodermata" rank="species" species="maya">
|
||
<emphasis id="B9500B4AFF93FFB3178EAF82FD806284" box="[452,596,475,498]" italics="true" pageId="10" pageNumber="927">Floriaster maya</emphasis>
|
||
</taxonomicName>
|
||
: 1.5, single tubercle present; 1.8, tubercles present on distal arm surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |