240 lines
33 KiB
XML
240 lines
33 KiB
XML
<document id="587F1C4F94D0968CF475E49150FA610F" ID-CLB-Dataset="28347" ID-DOI="10.11646/zootaxa.4567.3.1" ID-GBIF-Dataset="47813581-d217-4a19-8848-a3e9cf2ddaf4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2598945" ID-ZooBank="0AEFAF80-B001-4A18-88AC-5B6A189F6DCD" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553062614686" checkinUser="plazi" docAuthor="Staples, David A." docDate="2019" docId="03895C3329184F0CFF01F8EFFD06F93C" docLanguage="en" docName="zootaxa.4567.3.1.pdf" docOrigin="Zootaxa 4567 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nymphon gravidus Staples 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="412" masterDocId="FFB0244B29114F07FF96FFB2FF80FFFD" masterDocTitle="Pycnogonids (Arthropoda, Pycnogonida) from the Southwest Indian Ridge" masterLastPageNumber="449" masterPageNumber="401" pageNumber="410" updateTime="1698701998649" updateUser="ExternalLinkService">
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<mods:title id="3CE97F3ACD15512EFD442E9AE900F908">Pycnogonids (Arthropoda, Pycnogonida) from the Southwest Indian Ridge</mods:title>
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<mods:name id="62A6CB01CF79FA0F674483B828B099D0" type="personal">
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<mods:namePart id="60E5BBE1540A1C6326D9430163C6C508">Staples, David A.</mods:namePart>
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<mods:title id="497DEFD1A65B874EE3D953C3AA7734EB">Zootaxa</mods:title>
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<mods:part id="203FB77A0A3BF7E36E2211C52B1B9D19">
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<mods:date id="2B5E40B2E0D505E0EE1CAE9A8F5B2ED2">2019</mods:date>
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<mods:number id="041FCEC4969EED62A552ED9EADA204F8">2019-03-18</mods:number>
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<mods:number id="531B1124F1423F12669B2598EF9F4FB1">4567</mods:number>
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<mods:identifier id="5DBC7EEEECF23CFCF8396B9A428436D5" type="DOI">10.11646/zootaxa.4567.3.1</mods:identifier>
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<treatment id="03895C3329184F0CFF01F8EFFD06F93C" ID-DOI="http://doi.org/10.5281/zenodo.5944878" ID-GBIF-Taxon="154435127" ID-Zenodo-Dep="5944878" LSID="urn:lsid:plazi:treatment:03895C3329184F0CFF01F8EFFD06F93C" httpUri="http://treatment.plazi.org/id/03895C3329184F0CFF01F8EFFD06F93C" lastPageId="11" lastPageNumber="412" pageId="9" pageNumber="410">
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<subSubSection id="C33ABEAE29184F0EFF01F8EFFE62F88A" box="[151,482,1885,1911]" pageId="9" pageNumber="410" type="nomenclature">
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<paragraph id="8B9FED2529184F0EFF01F8EFFE62F88A" blockId="9.[151,482,1885,1945]" box="[151,482,1885,1911]" pageId="9" pageNumber="410">
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<heading id="D0D75A4929184F0EFF01F8EFFE62F88A" bold="true" box="[151,482,1885,1911]" fontSize="11" level="1" pageId="9" pageNumber="410" reason="1">
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<emphasis id="B954313729184F0EFF01F8EFFE62F88A" bold="true" box="[151,482,1885,1911]" pageId="9" pageNumber="410">
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<taxonomicName id="4C2096A629184F0EFF01F8EFFEFDF88A" ID-CoL="83477369-a17b-48dd-9bbf-8821323f7d06" authority="Staples, 2019" authorityName="Staples" authorityYear="2019" box="[151,381,1885,1911]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="gravidus" status="sp. nov.">
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<emphasis id="B954313729184F0EFF01F8EFFEFDF88A" bold="true" box="[151,381,1885,1911]" italics="true" pageId="9" pageNumber="410">Nymphon gravidus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2678C4C29184F0EFE12F8EFFE62F88A" box="[388,482,1885,1911]" pageId="9" pageNumber="410" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33ABEAE29184F0EFF01F832FE32F865" box="[151,434,1920,1945]" pageId="9" pageNumber="410" type="description">
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<paragraph id="8B9FED2529184F0EFF01F832FE32F865" blockId="9.[151,482,1885,1945]" box="[151,434,1920,1945]" pageId="9" pageNumber="410">
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<figureCitation id="131BF1A029184F0EFF01F832FF74F864" box="[151,244,1920,1945]" captionStart="FIGURE 5" captionStartId="10.[151,250,1951,1973]" captionTargetBox="[203,1358,343,1929]" captionTargetId="figure@10.[150,1436,265,1930]" captionTargetPageId="10" captionText="FIGURE 5. Nymphon gravidus sp. nov., holotype, female. A, trunk, dorsal view; B, trunk, lateral view; C, left oviger; D, oviger claw; E, right chela; F, third leg; G, tarsus and propodus of leg 4; H, left palp; I, proboscis, ventral." httpUri="https://zenodo.org/record/2598955/files/figure.png" pageId="9" pageNumber="410">Figure 5</figureCitation>
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A–I;
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<figureCitation id="131BF1A029184F0EFEB9F832FEFBF865" box="[303,379,1920,1945]" captionStart="PLATE 2" captionStartId="41.[151,234,1740,1762]" captionTargetBox="[151,1436,193,1718]" captionTargetId="figure@41.[151,1436,193,1718]" captionTargetPageId="41" captionText="PLATE 2. Nymphon gravidus sp. nov., holotype, female. A, trunk lateral; B, trunk dorsal; C, dorsolateral view showing oocytes in trunk and limbs; D, chelae. Tanystylum lamonti sp. nov., holotype, male. E, trunk lateral; F, trunk ventral showing eggs. Paratype, female. G, trunk dorsal; H, trunk lateral." httpUri="https://zenodo.org/record/2598985/files/figure.png" pageId="9" pageNumber="410">Plate 2</figureCitation>
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A–D
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</paragraph>
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</subSubSection>
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<subSubSection id="C33ABEAE29184F0DFF01F87AFECFFF29" lastPageId="10" lastPageNumber="411" pageId="9" pageNumber="410" type="materials_examined">
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<paragraph id="8B9FED2529184F0EFF01F87AFA1CF81D" blockId="9.[151,1436,1992,2017]" box="[151,1436,1992,2017]" pageId="9" pageNumber="410">
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<emphasis id="B954313729184F0EFF01F87AFE01F81C" bold="true" box="[151,385,1992,2017]" pageId="9" pageNumber="410">Material examined.</emphasis>
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<materialsCitation id="3B48E77829184F0EFE1CF87AFA1CF81D" ID-GBIF-Occurrence="2013824385" box="[394,1436,1992,2017]" collectionCode="NHMUK" location="Southwest Indian Ocean" municipality="Middle of What Seamount" pageId="9" pageNumber="410" specimenCode="NHMUK 2018.11" specimenCount="1" typeStatus="holotype">
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<typeStatus id="549B538729184F0EFE1CF87AFE75F81C" box="[394,501,1992,2017]" pageId="9" pageNumber="410" type="holotype">Holotype</typeStatus>
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, female (
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<specimenCode id="DB86455E29184F0EFDCBF87AFCAFF81D" box="[605,815,1992,2016]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="410" type="Museum">NHMUK 2018.11</specimenCode>
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),
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<location id="8EFFBBFE29184F0EFCD2F87AFBDAF81D" LSID="urn:lsid:plazi:treatment:03895C3329184F0CFF01F8EFFD06F93C:8EFFBBFE29184F0EFCD2F87AFBDAF81D" box="[836,1114,1992,2017]" municipality="Middle of What Seamount" name="Southwest Indian Ocean" pageId="9" pageNumber="410">Southwest Indian Ocean</location>
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,
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<collectingMunicipality id="6BFB775F29184F0EFBF1F87AFA18F81D" box="[1127,1432,1992,2017]" pageId="9" pageNumber="410">Middle of What Seamount</collectingMunicipality>
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,
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</materialsCitation>
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</paragraph>
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<paragraph id="8B9FED25291B4F0DFF01FF25FECFFF29" blockId="10.[151,1436,151,213]" pageId="10" pageNumber="411">
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37˚ 57. 91'S, 50˚ 24.42'E, ROV,
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<quantity id="4CD840C0291B4F0DFDBEFF25FD04FF52" box="[552,644,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.135" pageId="10" pageNumber="411" unit="m" value="1135.0">1135 m</quantity>
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, specimen JC066-3415, parent 2584, amongst sediment sample,
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<date id="FF9ECBE5291B4F0DFA18FF2AFECBFF29" pageId="10" pageNumber="411" value="2011-12-01">1 December 2011</date>
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.
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</paragraph>
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</subSubSection>
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<caption id="DF5FBDAD291B4F0DFF01F82DFB43F82A" httpUri="https://zenodo.org/record/2598955/files/figure.png" pageId="10" pageNumber="411" startId="10.[151,250,1951,1973]" targetBox="[203,1358,343,1929]" targetPageId="10">
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<paragraph id="8B9FED25291B4F0DFF01F82DFB43F82A" blockId="10.[151,1435,1951,2007]" pageId="10" pageNumber="411">
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<emphasis id="B9543137291B4F0DFF01F82DFE95F848" bold="true" box="[151,277,1951,1973]" pageId="10" pageNumber="411">FIGURE 5.</emphasis>
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<taxonomicName id="4C2096A6291B4F0DFE88F82DFE60F848" authority="Staples, 2019" authorityName="Staples" authorityYear="2019" box="[286,480,1951,1973]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="gravidus" status="sp. nov.">
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<emphasis id="B9543137291B4F0DFE88F82DFE60F848" box="[286,480,1951,1973]" italics="true" pageId="10" pageNumber="411">Nymphon gravidus</emphasis>
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</taxonomicName>
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<emphasis id="B9543137291B4F0DFE7CF82DFDBDF848" bold="true" box="[490,573,1951,1973]" pageId="10" pageNumber="411">
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<taxonomicNameLabel id="A2678C4C291B4F0DFE7CF82DFDBDF848" box="[490,573,1951,1973]" pageId="10" pageNumber="411" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, holotype, female. A, trunk, dorsal view; B, trunk, lateral view; C, left oviger; D, oviger claw; E, right chela; F, third leg; G, tarsus and propodus of leg 4; H, left palp; I, proboscis, ventral.
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</paragraph>
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</caption>
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<subSubSection id="C33ABEAE291A4F0CFF51FF25FB4CFBA3" pageId="11" pageNumber="412" type="description">
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<paragraph id="8B9FED25291A4F0CFF51FF25FEFCFE98" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">
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<emphasis id="B9543137291A4F0CFF51FF25FED9FF4D" bold="true" box="[199,345,151,176]" pageId="11" pageNumber="412">Description.</emphasis>
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Female
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<typeStatus id="549B5387291A4F0CFE2FFF25FD9FFF4D" box="[441,543,151,176]" pageId="11" pageNumber="412" type="holotype">holotype</typeStatus>
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. Trunk (
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<figureCitation id="131BF1A0291A4F0CFDEAFF25FD76FF52" box="[636,758,151,176]" captionStart="FIGURE 5" captionStartId="10.[151,250,1951,1973]" captionTargetBox="[203,1358,343,1929]" captionTargetId="figure@10.[150,1436,265,1930]" captionTargetPageId="10" captionText="FIGURE 5. Nymphon gravidus sp. nov., holotype, female. A, trunk, dorsal view; B, trunk, lateral view; C, left oviger; D, oviger claw; E, right chela; F, third leg; G, tarsus and propodus of leg 4; H, left palp; I, proboscis, ventral." httpUri="https://zenodo.org/record/2598955/files/figure.png" pageId="11" pageNumber="412">Fig. 5A, B</figureCitation>
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.
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<figureCitation id="131BF1A0291A4F0CFC94FF25FC05FF4D" box="[770,901,151,176]" captionStart="PLATE 2" captionStartId="41.[151,234,1740,1762]" captionTargetBox="[151,1436,193,1718]" captionTargetId="figure@41.[151,1436,193,1718]" captionTargetPageId="41" captionText="PLATE 2. Nymphon gravidus sp. nov., holotype, female. A, trunk lateral; B, trunk dorsal; C, dorsolateral view showing oocytes in trunk and limbs; D, chelae. Tanystylum lamonti sp. nov., holotype, male. E, trunk lateral; F, trunk ventral showing eggs. Paratype, female. G, trunk dorsal; H, trunk lateral." httpUri="https://zenodo.org/record/2598985/files/figure.png" pageId="11" pageNumber="412">Plate 2A, C</figureCitation>
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) smooth, segmented, ovigers attached to lateral extensions of the neck well-forward of first lateral processes, neck short, strongly constricted immediately forward of oviger implants; lateral processes longer than width at bases, processes 1 and 2 separated by 1.3 times basal width, distance between remaining processes progressively decreasing. Ocular tubercle placed midway between first lateral processes and bases of ovigers, two tiny dorsal papillae; four eyes well developed but unpigmented in preserved specimen.
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</paragraph>
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<paragraph id="8B9FED25291A4F0CFF51FEDDFD33FE75" blockId="11.[151,1437,151,1730]" box="[199,691,367,392]" pageId="11" pageNumber="412">
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Proboscis tubular, distally swollen (
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<figureCitation id="131BF1A0291A4F0CFDC1FEDDFD25FE75" box="[599,677,367,392]" captionStart="FIGURE 5" captionStartId="10.[151,250,1951,1973]" captionTargetBox="[203,1358,343,1929]" captionTargetId="figure@10.[150,1436,265,1930]" captionTargetPageId="10" captionText="FIGURE 5. Nymphon gravidus sp. nov., holotype, female. A, trunk, dorsal view; B, trunk, lateral view; C, left oviger; D, oviger claw; E, right chela; F, third leg; G, tarsus and propodus of leg 4; H, left palp; I, proboscis, ventral." httpUri="https://zenodo.org/record/2598955/files/figure.png" pageId="11" pageNumber="412">
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Fig.
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<date id="FF9ECBE5291A4F0CFD1DFEDDFD25FE75" box="[651,677,367,392]" pageId="11" pageNumber="412">5I</date>
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</figureCitation>
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).
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</paragraph>
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<paragraph id="8B9FED25291A4F0CFF51FE26FF5EFE2D" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">Abdomen with obscure suture line near base, slightly swollen dorsally at base, cleft distally, length about twice width.</paragraph>
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<paragraph id="8B9FED25291A4F0CFF51FE6EFCABFDE5" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">
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Chelifore scape slightly longer than proboscis, chela fingers much longer than palm, moveable finger with 25 teeth, immoveable finger with 21 teeth (
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<figureCitation id="131BF1A0291A4F0CFDCEFE4DFD2FFDE5" box="[600,687,511,536]" captionStart="FIGURE 5" captionStartId="10.[151,250,1951,1973]" captionTargetBox="[203,1358,343,1929]" captionTargetId="figure@10.[150,1436,265,1930]" captionTargetPageId="10" captionText="FIGURE 5. Nymphon gravidus sp. nov., holotype, female. A, trunk, dorsal view; B, trunk, lateral view; C, left oviger; D, oviger claw; E, right chela; F, third leg; G, tarsus and propodus of leg 4; H, left palp; I, proboscis, ventral." httpUri="https://zenodo.org/record/2598955/files/figure.png" pageId="11" pageNumber="412">Fig. 5E</figureCitation>
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.
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<figureCitation id="131BF1A0291A4F0CFD2FFE4DFC9FFDE5" box="[697,799,511,536]" captionStart="PLATE 2" captionStartId="41.[151,234,1740,1762]" captionTargetBox="[151,1436,193,1718]" captionTargetId="figure@41.[151,1436,193,1718]" captionTargetPageId="41" captionText="PLATE 2. Nymphon gravidus sp. nov., holotype, female. A, trunk lateral; B, trunk dorsal; C, dorsolateral view showing oocytes in trunk and limbs; D, chelae. Tanystylum lamonti sp. nov., holotype, male. E, trunk lateral; F, trunk ventral showing eggs. Paratype, female. G, trunk dorsal; H, trunk lateral." httpUri="https://zenodo.org/record/2598985/files/figure.png" pageId="11" pageNumber="412">Plate 2D</figureCitation>
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).
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</paragraph>
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<paragraph id="8B9FED25291A4F0CFF51FD96FD33FD9D" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">
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Palp (
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<figureCitation id="131BF1A0291A4F0CFE9FFD96FEE1FDC0" box="[265,353,548,573]" captionStart="FIGURE 5" captionStartId="10.[151,250,1951,1973]" captionTargetBox="[203,1358,343,1929]" captionTargetId="figure@10.[150,1436,265,1930]" captionTargetPageId="10" captionText="FIGURE 5. Nymphon gravidus sp. nov., holotype, female. A, trunk, dorsal view; B, trunk, lateral view; C, left oviger; D, oviger claw; E, right chela; F, third leg; G, tarsus and propodus of leg 4; H, left palp; I, proboscis, ventral." httpUri="https://zenodo.org/record/2598955/files/figure.png" pageId="11" pageNumber="412">Fig. 5H</figureCitation>
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) five-segmented, segment 2 longest, segment 4 longer than segment 5, setae on distal segments fine, long, mostly longer than width of segment.
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</paragraph>
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<paragraph id="8B9FED25291A4F0CFF51FDDEFD8DFD0D" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">
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Oviger (
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<figureCitation id="131BF1A0291A4F0CFEB1FDDEFE01FD78" box="[295,385,620,645]" captionStart="FIGURE 5" captionStartId="10.[151,250,1951,1973]" captionTargetBox="[203,1358,343,1929]" captionTargetId="figure@10.[150,1436,265,1930]" captionTargetPageId="10" captionText="FIGURE 5. Nymphon gravidus sp. nov., holotype, female. A, trunk, dorsal view; B, trunk, lateral view; C, left oviger; D, oviger claw; E, right chela; F, third leg; G, tarsus and propodus of leg 4; H, left palp; I, proboscis, ventral." httpUri="https://zenodo.org/record/2598955/files/figure.png" pageId="11" pageNumber="412">Fig. 5C</figureCitation>
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) ten-segmented, segment 5 longest, segment 4 with prominent gland opening on posterior surface, claw with four or five teeth on inner margin, length of claw about equal to segment 10, compound spines segments 7 to 10 well developed, pointed, with three pair of upwardly-curved lateral teeth, mid-pair longest, compound spine formula 8:6:5:6.
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</paragraph>
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<paragraph id="8B9FED25291A4F0CFF51FD4EFB33FC7D" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">
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Third leg (
|
||
<figureCitation id="131BF1A0291A4F0CFED4FD4EFE17FCE8" box="[322,407,764,789]" captionStart="FIGURE 5" captionStartId="10.[151,250,1951,1973]" captionTargetBox="[203,1358,343,1929]" captionTargetId="figure@10.[150,1436,265,1930]" captionTargetPageId="10" captionText="FIGURE 5. Nymphon gravidus sp. nov., holotype, female. A, trunk, dorsal view; B, trunk, lateral view; C, left oviger; D, oviger claw; E, right chela; F, third leg; G, tarsus and propodus of leg 4; H, left palp; I, proboscis, ventral." httpUri="https://zenodo.org/record/2598955/files/figure.png" pageId="11" pageNumber="412">Fig. 5F</figureCitation>
|
||
) with scattered fine setae, coxa 2 almost three times length of coxa 1, femur shorter than tibia 1, proximal two-thirds strongly swollen with oocytes; tibia 2 longest, almost 1.5 times length of tibia1, distal seta about three times width of segment, tarsus less than half propodus length, sole spines 10 to 20 varying between legs, heel absent, auxiliary claws absent. Gonopore large, ventrodistal surface coxa 2, all legs.
|
||
</paragraph>
|
||
<paragraph id="8B9FED25291A4F0CFF51FC22FB4CFBA3" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">
|
||
Measurements of
|
||
<typeStatus id="549B5387291A4F0CFE18FC3CFE70FC5A" box="[398,496,910,935]" pageId="11" pageNumber="412" type="holotype">holotype</typeStatus>
|
||
(mm). Length trunk (frontal margin of cephalic segment to tip of 4
|
||
<superScript id="7C55406D291A4F0CFB4FFC39FB65FC64" attach="left" box="[1241,1253,907,921]" fontSize="6" pageId="11" pageNumber="412">th</superScript>
|
||
lateral process), 2.432; length cephalon, 1.224; width across 2
|
||
<superScript id="7C55406D291A4F0CFD0FFC00FD29FC3D" attach="left" box="[665,681,946,960]" fontSize="6" pageId="11" pageNumber="412">nd</superScript>
|
||
lateral processes, 0.864; proboscis length (ventral), 1.080; greatest diameter proboscis, 0.400; length abdomen (lateral) 0.376. Palp: seg. 1, 0.112; seg. 2, 0.576; seg. 3, 0.424; seg. 4, 0.240; seg. 5, 0.216. Oviger: seg. 1, 0.136; seg. 2, 0.104; seg. 3, 0.232; seg. 4, 0.680; seg. 5, 0.800; seg. 6, 0.304. seg. 7. 0.240; seg. 8, 0.280; seg. 9, 0.160; seg. 10, 0.160; claw, 0.168. Third leg: coxa 1, 0.224; coxa 2, 0.560; coxa 3, 0.264; femur, 1.624; tibia 1, 2.040; tibia 2, 0.928; tarsus, 0.256; propodus, 0.496; claw, 0.200.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ABEAE291A4F0CFF51FBDBFE4AFB5B" pageId="11" pageNumber="412" type="etymology">
|
||
<paragraph id="8B9FED25291A4F0CFF51FBDBFE4AFB5B" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">
|
||
<emphasis id="B9543137291A4F0CFF51FBDBFECEFB7F" bold="true" box="[199,334,1129,1154]" pageId="11" pageNumber="412">Etymology.</emphasis>
|
||
Gravid, from the Latin
|
||
<taxonomicName id="4C2096A6291A4F0CFDCCFBDBFD3CFB7F" authorityName="Staples" authorityYear="2019" box="[602,700,1129,1154]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="gravidus">
|
||
<emphasis id="B9543137291A4F0CFDCCFBDBFD3CFB7F" box="[602,700,1129,1154]" italics="true" pageId="11" pageNumber="412">gravidus</emphasis>
|
||
</taxonomicName>
|
||
-pregnant, references the extraordinary number of oocytes carried in the legs and body cavity.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33ABEAE291A4F0CFF51FB03FD06F93C" pageId="11" pageNumber="412" type="discussion">
|
||
<paragraph id="8B9FED25291A4F0CFF51FB03FB4CFA3B" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">
|
||
<emphasis id="B9543137291A4F0CFF51FB03FEBBFB37" bold="true" box="[199,315,1201,1226]" pageId="11" pageNumber="412">Remarks.</emphasis>
|
||
There are four uniunguiculate species of
|
||
<taxonomicName id="4C2096A6291A4F0CFC9EFB03FCF3FB37" authorityName="J. C. Fabricius" authorityYear="1794" box="[776,883,1201,1226]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9543137291A4F0CFC9EFB03FCF3FB37" box="[776,883,1201,1226]" italics="true" pageId="11" pageNumber="412">Nymphon</emphasis>
|
||
</taxonomicName>
|
||
that have the oviger implantation well forward of the first lateral processes and well-developed eyes on an ocular tubercle placed between the ovigers and the first lateral processes. These species are
|
||
<taxonomicName id="4C2096A6291A4F0CFDAFFB48FD44FAEF" authority="Staples, 2019" authorityName="Staples" authorityYear="2019" box="[569,708,1273,1298]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="dentatum" status="sp. nov.">
|
||
<emphasis id="B9543137291A4F0CFDAFFB48FD44FAEF" box="[569,708,1273,1298]" italics="true" pageId="11" pageNumber="412">N. dentatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9543137291A4F0CFD59FB4BFCACFAEF" bold="true" box="[719,812,1273,1298]" pageId="11" pageNumber="412">
|
||
<taxonomicNameLabel id="A2678C4C291A4F0CFD59FB4BFCACFAEF" box="[719,812,1273,1298]" pageId="11" pageNumber="412" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<taxonomicName id="4C2096A6291A4F0CFCA0FB48FC2AFAEC" authority="Staples, 2019" authorityName="Staples" authorityYear="2019" box="[822,938,1274,1297]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="rogersi" status="sp. nov.">
|
||
<emphasis id="B9543137291A4F0CFCA0FB48FC2AFAEC" box="[822,938,1274,1297]" italics="true" pageId="11" pageNumber="412">N. rogersi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2678C4C291A4F0CFC22FB4BFB90FAEC" box="[948,1040,1273,1298]" pageId="11" pageNumber="412" rank="species">
|
||
<emphasis id="B9543137291A4F0CFC22FB4BFB8AFAEF" bold="true" box="[948,1034,1273,1298]" pageId="11" pageNumber="412">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C2096A6291A4F0CFBB4FB48FAC5FAEF" authority="Hoek, 1881" authorityName="Hoek" authorityYear="1881" box="[1058,1349,1273,1298]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="longicoxa">
|
||
<emphasis id="B9543137291A4F0CFBB4FB48FB35FAEF" box="[1058,1205,1273,1298]" italics="true" pageId="11" pageNumber="412">N. longicoxa</emphasis>
|
||
<bibRefCitation id="EFB190D4291A4F0CFB29FB4BFAC5FAEF" author="Hoek, P. P. C." box="[1215,1349,1273,1298]" pageId="11" pageNumber="412" pagination="1 - 167" refId="ref27925" refString="Hoek, P. P. C. (1881) Report on the Pycnogonida, dredged by HMS Challenger 1873 - 76. Reports of the Scientific Results of the Exploring Foyage of HMS Challenger, 3 (10), 1 - 167. https: // doi. org / 10.5962 / bhl. title. 13132" type="journal article" year="1881">Hoek, 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2096A6291A4F0CFA14FB48FE05FACB" authority="Fage, 1942" authorityName="Fage" authorityYear="1942" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="prolatum">
|
||
<emphasis id="B9543137291A4F0CFA14FB48FF7DFACB" italics="true" pageId="11" pageNumber="412">N. prolatum</emphasis>
|
||
<bibRefCitation id="EFB190D4291A4F0CFE92FAACFE05FACB" author="Fage, L." box="[260,389,1310,1334]" pageId="11" pageNumber="412" pagination="75 - 90" refId="ref27532" refString="Fage, L. (1942) Pycnogonides de la cote occidentale d'Afrique. Archives de Zoologie Experimentale et Generale, 82 (Notes & Revue, 2), 75 - 90." type="journal article" year="1942">Fage, 1942</bibRefCitation>
|
||
</taxonomicName>
|
||
. This species it is most like
|
||
<taxonomicName id="4C2096A6291A4F0CFD52FAADFCB6FAC8" authorityName="Staples" authorityYear="2019" box="[708,822,1310,1334]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="rogersi">
|
||
<emphasis id="B9543137291A4F0CFD52FAADFCB6FAC8" box="[708,822,1310,1334]" italics="true" pageId="11" pageNumber="412">N. rogersi</emphasis>
|
||
</taxonomicName>
|
||
but from which it differs most notably in the shape of the neck region, by having much more elongate palp segments (particularly the fourth segment) and in the shape of the chela fingers. From
|
||
<taxonomicName id="4C2096A6291A4F0CFE33FAD5FDAEFA83" authorityName="Staples" authorityYear="2019" box="[421,558,1381,1406]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="dentatum">
|
||
<emphasis id="B9543137291A4F0CFE33FAD5FDAEFA83" box="[421,558,1381,1406]" italics="true" pageId="11" pageNumber="412">N. dentatum</emphasis>
|
||
</taxonomicName>
|
||
it differs conspicuously in the length of the neck. From the other two species that make up this group this species is most like
|
||
<taxonomicName id="4C2096A6291A4F0CFD2CFA38FCC9FA5F" authorityName="Hoek" authorityYear="1881" box="[698,841,1417,1442]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="longicoxa">
|
||
<emphasis id="B9543137291A4F0CFD2CFA38FCC9FA5F" box="[698,841,1417,1442]" italics="true" pageId="11" pageNumber="412">N. longicoxa</emphasis>
|
||
</taxonomicName>
|
||
but from which it differs in the much shorter second coxa.
|
||
<taxonomicName id="4C2096A6291A4F0CFF4CFA1FFE32FA3B" authorityName="Fage" authorityYear="1942" box="[218,434,1453,1478]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="prolatum">
|
||
<emphasis id="B9543137291A4F0CFF4CFA1FFE32FA3B" box="[218,434,1453,1478]" italics="true" pageId="11" pageNumber="412">Nymphon prolatum</emphasis>
|
||
</taxonomicName>
|
||
is distinguished by its long ocular tubercle and much longer abdomen.
|
||
</paragraph>
|
||
<paragraph id="8B9FED25291A4F0CFF51FA63FD06F93C" blockId="11.[151,1437,151,1730]" pageId="11" pageNumber="412">
|
||
Considering the low fecundity of some species which may carry as few as one oocyte in each femur (
|
||
<bibRefCitation id="EFB190D4291A4F0CFAD4FA63FEF2F9F3" author="Bamber, R. N. & Thurston, M. H." pageId="11" pageNumber="412" pagination="837 - 861" refId="ref26489" refString="Bamber, R. N. & Thurston, M. H. (1993) Deep water pycnogonids of the Cape Verde slope. Journal of the Marine Biological Association of the United Kingdom, 73, 837 - 861. https: // doi. org / 10.1017 / S 0025315400034767" type="journal article" year="1993">Bamber and Thurston 1993</bibRefCitation>
|
||
), the estimated many hundreds of oocytes carried by this female is remarkable. The oocytes have a diameter of about 0.040 µm and are tightly packed into the femora and tibiae and in lesser numbers, in almost the entire body cavity. They are present in the coxae, trunk, abdomen scapes and one oviger. Those in oviger segment 5 are lined up in a single row (
|
||
<figureCitation id="131BF1A0291A4F0CFDC3F9D3FD3BF987" box="[597,699,1633,1658]" captionStart="PLATE 2" captionStartId="41.[151,234,1740,1762]" captionTargetBox="[151,1436,193,1718]" captionTargetId="figure@41.[151,1436,193,1718]" captionTargetPageId="41" captionText="PLATE 2. Nymphon gravidus sp. nov., holotype, female. A, trunk lateral; B, trunk dorsal; C, dorsolateral view showing oocytes in trunk and limbs; D, chelae. Tanystylum lamonti sp. nov., holotype, male. E, trunk lateral; F, trunk ventral showing eggs. Paratype, female. G, trunk dorsal; H, trunk lateral." httpUri="https://zenodo.org/record/2598985/files/figure.png" pageId="11" pageNumber="412">Plate 2C</figureCitation>
|
||
). The presence of oocytes throughout the body cavity is common in the genera
|
||
<taxonomicName id="4C2096A6291A4F0CFED2F937FCABF963" authority="(Bamber & Thurston, 1993)" authorityName="Bamber & Thurston" authorityYear="1993" baseAuthorityName="Bamber & Thurston" baseAuthorityYear="1993" box="[324,811,1669,1694]" class="Pycnogonida" family="Austrodecidae" genus="Austrodecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9543137291A4F0CFED2F937FE4DF963" box="[324,461,1669,1694]" italics="true" pageId="11" pageNumber="412">Austrodecus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB190D4291A4F0CFE75F937FCA3F963" author="Bamber, R. N. & Thurston, M. H." box="[483,803,1669,1694]" pageId="11" pageNumber="412" pagination="837 - 861" refId="ref26489" refString="Bamber, R. N. & Thurston, M. H. (1993) Deep water pycnogonids of the Cape Verde slope. Journal of the Marine Biological Association of the United Kingdom, 73, 837 - 861. https: // doi. org / 10.1017 / S 0025315400034767" type="journal article" year="1993">Bamber & Thurston, 1993</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2096A6291A4F0CFCE6F937FB9BF963" authorityName="Costa" authorityYear="1861" box="[880,1051,1669,1694]" class="Pycnogonida" family="Rhynchothoracidae" genus="Rhynchothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9543137291A4F0CFCE6F937FB9BF963" box="[880,1051,1669,1694]" italics="true" pageId="11" pageNumber="412">Rhynchothorax</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<taxonomicName id="4C2096A6291A4F0CFBF3F935FB3BF960" authority="Staples, 2019" authorityName="Staples" authorityYear="2019" box="[1125,1211,1670,1694]" class="Pycnogonida" family="Rhynchothoracidae" genus="Rhynchothorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="swir" status="sp. nov.">
|
||
<emphasis id="B9543137291A4F0CFBF3F935FB3BF960" box="[1125,1211,1670,1694]" italics="true" pageId="11" pageNumber="412">R. swir</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2678C4C291A4F0CFB5CF937FAA8F963" box="[1226,1320,1669,1694]" pageId="11" pageNumber="412" rank="species">
|
||
<emphasis id="B9543137291A4F0CFB5CF937FAA3F963" bold="true" box="[1226,1315,1669,1694]" pageId="11" pageNumber="412">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
) but this occurrence is unique in the genus
|
||
<taxonomicName id="4C2096A6291A4F0CFD82F91BFDFFF93F" authorityName="J. C. Fabricius" authorityYear="1794" box="[532,639,1705,1730]" class="Pycnogonida" family="Nymphonidae" genus="Nymphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pantopoda" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9543137291A4F0CFD82F91BFDFFF93F" box="[532,639,1705,1730]" italics="true" pageId="11" pageNumber="412">Nymphon</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |