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<document id="4070A601B1E89A90C3D30BA1E007F013" ID-DOI="10.5281/zenodo.5329313" ID-ISSN="0374-1036" ID-Zenodo-Dep="5329313" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630162999482" checkinUser="felipe" docAuthor="Hisamatsu, Sadatomo" docDate="2011" docId="0388D148EA1F872BFE1FFCAFFE49604E" docLanguage="en" docName="ActaEntMusNatPra.2011.51.2.551-585.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 51 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Sibirhelus corpulentus" docType="treatment" docVersion="3" lastPageNumber="574" masterDocId="FFB1A930EA0A873CFF97FFF6FFB1633A" masterDocTitle="A review of the Japanese Kateretidae fauna (Coleoptera: Cucujoidea)" masterLastPageNumber="585" masterPageNumber="551" pageNumber="572" updateTime="1699169862046" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DAB7FEA2DDFA43FA0EFEE49A31658DA4">A review of the Japanese Kateretidae fauna (Coleoptera: Cucujoidea)</mods:title>
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<mods:namePart id="AB878087D552AA1D7E4FAEAF2F5B768A">Hisamatsu, Sadatomo</mods:namePart>
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<mods:title id="D91660F6374BE5592E182EBED7C84F43">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="E7CB5CF6ACBF0CAF57CE0DECFB14A577">2011</mods:date>
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<subSubSection id="C33B33D5EA1F8729FE1FFCAFFC9B604B" box="[392,810,857,882]" pageId="21" pageNumber="572" type="nomenclature">
<paragraph id="8B9E605EEA1F8729FE1FFCAFFC9B604B" blockId="21.[392,810,857,882]" box="[392,810,857,882]" pageId="21" pageNumber="572">
<heading id="D0D6D732EA1F8729FE1FFCAFFC9B604B" bold="true" box="[392,810,857,882]" centered="true" fontSize="8" level="2" pageId="21" pageNumber="572" reason="3">
<taxonomicName id="4C211BDDEA1F8729FE1FFCAFFC9B604B" authority="(Reitter, 1900)" baseAuthorityName="Reitter" baseAuthorityYear="1900" box="[392,810,857,882]" class="Insecta" family="Kateretidae" genus="Sibirhelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="572" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B955BC4CEA1F8729FE1FFCAFFC9B604B" bold="true" box="[392,810,857,882]" pageId="21" pageNumber="572">
<emphasis id="B955BC4CEA1F8729FE1FFCAFFD306048" bold="true" box="[392,641,857,882]" italics="true" pageId="21" pageNumber="572">Sibirhelus corpulentus</emphasis>
(
<bibRefCitation id="EFB01DAFEA1F8729FD18FCAFFC92604B" author="REITTER E." box="[655,803,857,881]" pageId="21" pageNumber="572" pagination="225 - 232" refId="ref20598" refString="REITTER E. 1900: Beschreibung und Abbildung von neun Coleopteren der palaearctischen Fauna. Wiener Entomologische Zeitung 19: 225 - 232, pl. 2." type="journal article" year="1900">Reitter, 1900</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33B33D5EA1F8729FDB5FC8EFD2160B1" box="[546,656,888,907]" pageId="21" pageNumber="572" type="description">
<paragraph id="8B9E605EEA1F8729FDB5FC8EFD2160B1" blockId="21.[113,1090,888,1143]" box="[546,656,888,907]" pageId="21" pageNumber="572">
(
<figureCitation id="131A7CDBEA1F8729FDBEFC8EFDC460B1" box="[553,629,888,907]" captionStart="Fig" captionStartId="3.[113,144,1488,1507]" captionTargetBox="[232,958,532,1461]" captionTargetId="figure-117@3.[220,982,514,1467]" captionTargetPageId="3" captionText="Fig. 2. Habitus and labels of Kateretidae spp. AB Heterhelus (Heterhelus) morio (Reitter, 1878): A lectotype, male; B ditto, labels; C Heterhelus (Boreades) solani (Heer, 1841), male (Hokkaido); D Sibirhelus corpulentus (Reitter, 1900), male (Iwate Pref.); E Brachypterus urticae (Fabricius, 1792), male (Hokkaido); FG Brachypterolus shimoyamai Hisamatsu, 1985: F holotype, male; G ditto, labels. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5329319" httpUri="https://zenodo.org/record/5329319/files/figure.png" pageId="21" pageNumber="572">Figs. 2D</figureCitation>
,
<figureCitation id="131A7CDBEA1F8729FDE9FC8EFD3860B1" box="[638,649,888,907]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="21" pageNumber="572">9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33B33D5EA1F8729FFE6FC6DFDEE60C7" pageId="21" pageNumber="572" type="reference_group">
<paragraph id="8B9E605EEA1F8729FFE6FC6DFCDC6097" blockId="21.[113,1090,888,1143]" box="[113,877,921,942]" pageId="21" pageNumber="572">
<treatmentCitationGroup id="AB314770EA1F8729FFE6FC6DFCDC6097" box="[113,877,921,942]" pageId="21" pageNumber="572">
<treatmentCitation id="0A80464FEA1F8729FFE6FC6DFE6B6097" author="REITTER E." box="[113,474,922,941]" page="231" pageId="21" pageNumber="572" year="1900">
<taxonomicName id="4C211BDDEA1F8729FFE6FC6DFE6B6097" authority="Reitter, 1900: 231" authorityName="Reitter" authorityPageNumber="231" authorityYear="1900" box="[113,474,922,941]" class="Insecta" family="Kateretidae" genus="Heterhelus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="572" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B955BC4CEA1F8729FFE6FC6DFE886097" box="[113,313,923,941]" italics="true" pageId="21" pageNumber="572">Heterhelus corpulentus</emphasis>
<bibRefCitation id="EFB01DAFEA1F8729FEA9FC6CFE6B6097" author="REITTER E." box="[318,474,922,941]" pageId="21" pageNumber="572" pagination="225 - 232" refId="ref20598" refString="REITTER E. 1900: Beschreibung und Abbildung von neun Coleopteren der palaearctischen Fauna. Wiener Entomologische Zeitung 19: 225 - 232, pl. 2." type="journal article" year="1900">Reitter, 1900: 231</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, tab. II, fig. 7.
<emphasis id="B955BC4CEA1F8729FDC0FC6CFD636097" bold="true" box="[599,722,921,941]" pageId="21" pageNumber="572">
<typeStatus id="549ADEFCEA1F8729FDC0FC6CFD356097" box="[599,644,922,941]" pageId="21" pageNumber="572">Type</typeStatus>
locality:
</emphasis>
[
<collectingCountry id="F33620CEEA1F8729FD4AFC6CFCAC6097" box="[733,797,922,941]" name="Russia" pageId="21" pageNumber="572">Russia</collectingCountry>
:]
<collectingRegion id="49E5AEBCEA1F8729FCBDFC6CFCD86097" box="[810,873,922,941]" country="Russia" name="Irkutsk" pageId="21" pageNumber="572">Irkutsk</collectingRegion>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9E605EEA1F8729FFE6FC40FDDC60F2" blockId="21.[113,1090,888,1143]" box="[113,621,949,968]" pageId="21" pageNumber="572">
<treatmentCitationGroup id="AB314770EA1F8729FFE6FC40FDDC60F2" box="[113,621,949,968]" pageId="21" pageNumber="572">
<treatmentCitation id="0A80464FEA1F8729FFE6FC40FDB060F2" author="GROUVELLE A." box="[113,513,949,968]" page=")" pageId="21" pageNumber="572" year="1913">
<taxonomicName id="4C211BDDEA1F8729FFE6FC40FDB060F2" authority=": GROUVELLE (1913: 15)" authorityName="GROUVELLE" authorityPageNumber="15" authorityYear="1913" box="[113,513,949,968]" class="Insecta" family="Kateretidae" genus="Heterhelus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="572" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B955BC4CEA1F8729FFE6FC40FE8860F2" box="[113,313,950,968]" italics="true" pageId="21" pageNumber="572">Heterhelus corpulentus</emphasis>
:
<bibRefCitation id="EFB01DAFEA1F8729FED3FC43FDB060F2" author="GROUVELLE A." box="[324,513,949,968]" pageId="21" pageNumber="572" refId="ref18397" refString="GROUVELLE A. 1913: Byturidae, Nitidulidae. In: JUNK W. &amp; SCHENKLING S. (eds.): Coleopterorum Catalogus. Vol. 56. W. Junk, Berlin, 223 pp." type="book" year="1913">GROUVELLE (1913: 15)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[catalogue].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9E605EEA1F8729FFE6FC27FDEE60C7" blockId="21.[113,1090,888,1143]" pageId="21" pageNumber="572">
<treatmentCitationGroup id="AB314770EA1F8729FFE6FC27FDEE60C7" pageId="21" pageNumber="572">
<treatmentCitation id="0A80464FEA1F8729FFE6FC27FDB260D9" author="KIREJTSHUK A. G." box="[113,515,976,995]" page=")" pageId="21" pageNumber="572" year="1989">
<taxonomicName id="4C211BDDEA1F8729FFE6FC27FDB260D9" authority=": KIREJTSHUK (1989: 147)" authorityName="KIREJTSHUK" authorityPageNumber="147" authorityYear="1989" box="[113,515,976,995]" class="Insecta" family="Kateretidae" genus="Sibirhelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="572" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B955BC4CEA1F8729FFE6FC27FE8060D9" box="[113,305,977,995]" italics="true" pageId="21" pageNumber="572">Sibirhelus corpulentus</emphasis>
:
<bibRefCitation id="EFB01DAFEA1F8729FEABFC26FDB260D9" author="KIREJTSHUK A. G." box="[316,515,976,995]" pageId="21" pageNumber="572" pagination="145 - 149" refId="ref19515" refString="KIREJTSHUK A. G. 1989: Novye palearkticheskie rod i vidy sem. Kateretidae (Coleoptera) i zamechaniya po sinonimii. (New Palaearctic genus and species of the family Kateretidae (Coleoptera) and notes on the synonymy). Zoologicheskii Zhurnal 68: 145 - 149 (in Russian, English title and summary)." type="journal article" year="1989">KIREJTSHUK (1989: 147)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[redescription];
<treatmentCitation id="0A80464FEA1F8729FD07FC26FCE960D9" author="KIREJTSHUK A. G." box="[656,856,976,995]" page="214" pageId="21" pageNumber="572" year="1992">
<bibRefCitation id="EFB01DAFEA1F8729FD07FC26FCE960D9" author="KIREJTSHUK A. G." box="[656,856,976,995]" pageId="21" pageNumber="572" pagination="210 - 216" refId="ref19575" refString="KIREJTSHUK A. G. 1992: Family Kateretidae Erichson, 1846. Pp. 210 - 216. In: LEHR P. A. (ed.): Opredelitel' nasekomykh Dal'nego Vostoka SSSR. [Identification key to the insects of the Far East of the USSR. Volume 3. Part 2.]. Nauka Pren, St. Petersburg, Russia, 704 pp (in Russian)." type="book chapter" year="1992">KIREJTSHUK (1992: 214)</bibRefCitation>
</treatmentCitation>
[in key];
<treatmentCitation id="0A80464FEA1F8729FC3DFC26FF1C60C7" author="HISAMATSU S." page="14" pageId="21" pageNumber="572" year="2003">
<bibRefCitation id="EFB01DAFEA1F8729FC3DFC26FF1C60C7" author="HISAMATSU S." pageId="21" pageNumber="572" pagination="14 - 15" refId="ref18926" refString="HISAMATSU S. 2003: Two nitidulid genera new to Japan (Coleoptera). Shikoku Tyuho 37: 14 - 15 (in Japanese, with English title)." type="journal article" year="2003">HISAMATSU (2003: 14)</bibRefCitation>
</treatmentCitation>
[note];
<treatmentCitation id="0A80464FEA1F8729FF67FC1DFE4260C7" author="JELINEK J. &amp; AUDISIO P." box="[240,499,1002,1022]" page="458" pageId="21" pageNumber="572" year="2007">
<bibRefCitation id="EFB01DAFEA1F8729FF67FC1DFE4260C7" author="JELINEK J. &amp; AUDISIO P." box="[240,499,1002,1022]" pageId="21" pageNumber="572" pagination="459 - 491" refId="ref19280" refString="JELINEK J. &amp; AUDISIO P. 2007: Family Nitidulidae. Pp. 459 - 491. In: LOBL I. &amp; SMETANA A. (eds): Catalogue of Palaearctic Coleoptera. Vol. 4: Elateroidea - Derodontoidea - Bostrichoidea - Lymexyloidea - Cleroidea - Cucujoidea. Apollo Books, Stenstrup, 935 pp." type="book chapter" year="2007">JELÍNEK &amp; AUDISIO (2007: 458)</bibRefCitation>
</treatmentCitation>
[catalogue].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33B33D5EA1F8729FFE6FBE5FDA9674D" pageId="21" pageNumber="572" type="materials_examined">
<paragraph id="8B9E605EEA1F8729FFE6FBE5FBF06778" blockId="21.[113,1090,888,1143]" pageId="21" pageNumber="572">
<emphasis id="B955BC4CEA1F8729FFE6FBE5FE93671D" bold="true" box="[113,290,1043,1063]" pageId="21" pageNumber="572">Material examined.</emphasis>
<materialsCitation id="3B496A03EA1F8729FEB1FBE5FF206778" collectingDate="2007-05-09" collectorName="Y. Takahashi" country="Japan" location="IWATE" pageId="21" pageNumber="572" specimenCount="1" specimenCount-female="1" stateProvince="Iwate">
<emphasis id="B955BC4CEA1F8729FEB1FBE5FE01671D" bold="true" box="[294,432,1043,1063]" pageId="21" pageNumber="572">
<collectingCountry id="F33620CEEA1F8729FEB1FBE5FEDF671D" box="[294,366,1043,1063]" name="Japan" pageId="21" pageNumber="572">JAPAN</collectingCountry>
:
<collectingRegion id="49E5AEBCEA1F8729FEE2FBE2FE18671C" box="[373,425,1044,1063]" country="Japan" name="Iwate" pageId="21" pageNumber="572">IWATE</collectingRegion>
:
</emphasis>
1J
<specimenCount id="9D27ABD7EA1F8729FE41FBE2FE45671D" box="[470,500,1038,1063]" pageId="21" pageNumber="572" type="female">1 ♀</specimenCount>
, Kouyanotutumi, Kanegasaki-chô, Iwasa-gun,
<date id="FF9F469EEA1F8729FC14FBE2FC7F671D" box="[899,974,1044,1063]" pageId="21" pageNumber="572" value="2007-05-09">
<collectingDate id="EFDBBF76EA1F8729FC14FBE2FC7F671D" box="[899,974,1044,1063]" pageId="21" pageNumber="572" value="2007-05-09">9.v.2007</collectingDate>
</date>
,
<collectorName id="26D40588EA1F8729FC42FBE2FBF1671D" box="[981,1088,1044,1063]" pageId="21" pageNumber="572">Y.Takahashi</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B496A03EA1F8729FF03FBD9FB8A6778" box="[148,1083,1065,1091]" collectingDate="1992-05-02" collectorName="H. Ohkawa" country="Japan" location="TOCHIGI" pageId="21" pageNumber="572" specimenCount="4" specimenCount-female="4" stateProvince="Tochigi">
<emphasis id="B955BC4CEA1F8729FF03FBD9FF506778" bold="true" box="[148,225,1071,1090]" pageId="21" pageNumber="572">
<collectingRegion id="49E5AEBCEA1F8729FF03FBD9FF6B677A" box="[148,218,1071,1090]" country="Japan" name="Tochigi" pageId="21" pageNumber="572">TOCHIGI</collectingRegion>
:
</emphasis>
2JJ
<specimenCount id="9D27ABD7EA1F8729FE81FBD9FEF06778" box="[278,321,1065,1090]" pageId="21" pageNumber="572" type="female">4♀♀</specimenCount>
, Watarase-yûsuichi,Fujioka-chô,
<date id="FF9F469EEA1F8729FDCBFBD9FD176778" box="[604,678,1071,1090]" pageId="21" pageNumber="572" value="1992-05-02">
<collectingDate id="EFDBBF76EA1F8729FDCBFBD9FD176778" box="[604,678,1071,1090]" pageId="21" pageNumber="572" value="1992-05-02">2.v.1992</collectingDate>
</date>
,
<collectorName id="26D40588EA1F8729FD3AFBD9FCBA6778" box="[685,779,1071,1090]" pageId="21" pageNumber="572">H.Ohkawa</collectorName>
leg., by sweeping of
<taxonomicName id="4C211BDDEA1F8729FC20FBC6FB8A6778" box="[951,1083,1072,1090]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="21" pageNumber="572" phylum="Tracheophyta" rank="species" species="dispalata">
<emphasis id="B955BC4CEA1F8729FC20FBC6FB8A6778" box="[951,1083,1072,1090]" italics="true" pageId="21" pageNumber="572">Carex dispalata</emphasis>
</taxonomicName>
</materialsCitation>
;
</paragraph>
<paragraph id="8B9E605EEA1F8729FFE6FBBFFBF36766" blockId="21.[113,1090,888,1143]" box="[113,1090,1091,1117]" pageId="21" pageNumber="572">
10 JJ
<specimenCount id="9D27ABD7EA1F8729FF39FBBFFF6A6766" box="[174,219,1091,1116]" pageId="21" pageNumber="572" type="female">2 ♀♀</specimenCount>
, Watarase-yûsuichi, Fujioka-chô,
<date id="FF9F469EEA1F8729FE6AFBBFFDCF6766" box="[509,638,1097,1116]" pageId="21" pageNumber="572" value="2010-04-24" valueMax="2010-04-25" valueMin="2010-04-24">24.25.iv.2010</date>
, S-T. Hisamatsu leg.; 4JJ
<specimenCount id="9D27ABD7EA1F8729FCF0FBBFFC226766" box="[871,915,1091,1116]" pageId="21" pageNumber="572" type="female">5♀♀</specimenCount>
, Watarase-yûsuichi,
</paragraph>
<paragraph id="8B9E605EEA1F8729FFE6FB92FDA9674D" blockId="21.[113,1090,888,1143]" box="[113,536,1124,1143]" pageId="21" pageNumber="572">
Fujioka-chô,
<date id="FF9F469EEA1F8729FF74FB92FE8E674D" box="[227,319,1124,1143]" pageId="21" pageNumber="572" value="2010-04-24">24.iv.2010</date>
,
<collectionCode id="ED30F89BEA1F8729FED0FB92FEE4674D" box="[327,341,1124,1143]" name="Tavera, Department of Geology and Geophysics" pageId="21" pageNumber="572">T</collectionCode>
. Kurihara leg. (
<collectionCode id="ED30F89BEA1F8729FE4EFB92FDBC674D" box="[473,525,1124,1143]" pageId="21" pageNumber="572">EUM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33B33D5EA1F8729FFE6FB79FF77663D" pageId="21" pageNumber="572" type="diagnosis">
<paragraph id="8B9E605EEA1F8729FFE6FB79FF77663D" blockId="21.[113,1089,1167,1639]" pageId="21" pageNumber="572">
<emphasis id="B955BC4CEA1F8729FFE6FB79FF52679D" bold="true" box="[113,227,1167,1191]" pageId="21" pageNumber="572">Diagnosis.</emphasis>
Body coloration fully luteous to dark brown (
<figureCitation id="131A7CDBEA1F8729FD2FFB79FCBA6792" box="[696,779,1167,1192]" captionStart="Fig" captionStartId="3.[113,144,1488,1507]" captionTargetBox="[232,958,532,1461]" captionTargetId="figure-117@3.[220,982,514,1467]" captionTargetPageId="3" captionText="Fig. 2. Habitus and labels of Kateretidae spp. AB Heterhelus (Heterhelus) morio (Reitter, 1878): A lectotype, male; B ditto, labels; C Heterhelus (Boreades) solani (Heer, 1841), male (Hokkaido); D Sibirhelus corpulentus (Reitter, 1900), male (Iwate Pref.); E Brachypterus urticae (Fabricius, 1792), male (Hokkaido); FG Brachypterolus shimoyamai Hisamatsu, 1985: F holotype, male; G ditto, labels. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5329319" httpUri="https://zenodo.org/record/5329319/files/figure.png" pageId="21" pageNumber="572">Fig. 2D</figureCitation>
); antennae almost moniliform (
<figureCitation id="131A7CDBEA1F8729FFEEFB59FF7F67F2" box="[121,206,1199,1224]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="21" pageNumber="572">Fig. 9G</figureCitation>
). Medial tuft of setae on abdominal sternites III (
<figureCitation id="131A7CDBEA1F8729FD5AFB59FC9167F2" box="[717,800,1199,1224]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="21" pageNumber="572">Fig. 9K</figureCitation>
) in Japanese specimens, on abdominal sternites IIIV in Russian specimens. Male protibiae (
<figureCitation id="131A7CDBEA1F8729FCBBFB39FCC467D2" box="[812,885,1231,1256]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="21" pageNumber="572">Fig. 9I</figureCitation>
) strongly medially arcuate.
</paragraph>
</subSubSection>
<subSubSection id="C33B33D5EA1F872BFFE6FAF9FF686009" lastPageId="23" lastPageNumber="574" pageId="21" pageNumber="572" type="description">
<paragraph id="8B9E605EEA1F8729FFE6FAF9FE46661D" blockId="21.[113,1089,1167,1639]" box="[113,503,1295,1319]" pageId="21" pageNumber="572">
<emphasis id="B955BC4CEA1F8729FFE6FAF9FEA6661D" bold="true" box="[113,279,1295,1319]" pageId="21" pageNumber="572">Redescription.</emphasis>
Length
<quantity id="4CD9CDBBEA1F8729FEF8FAF9FE42661D" box="[367,499,1295,1319]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.6" metricValueMin="2.7" pageId="21" pageNumber="572" unit="mm" value="3.15" valueMax="3.6" valueMin="2.7">2.73.6 mm</quantity>
.
</paragraph>
<paragraph id="8B9E605EEA1F8729FF07FAD9FCF9665D" blockId="21.[113,1089,1167,1639]" pageId="21" pageNumber="572">
Male. Body (
<figureCitation id="131A7CDBEA1F8729FE89FAD9FEC36672" box="[286,370,1327,1352]" captionStart="Fig" captionStartId="3.[113,144,1488,1507]" captionTargetBox="[232,958,532,1461]" captionTargetId="figure-117@3.[220,982,514,1467]" captionTargetPageId="3" captionText="Fig. 2. Habitus and labels of Kateretidae spp. AB Heterhelus (Heterhelus) morio (Reitter, 1878): A lectotype, male; B ditto, labels; C Heterhelus (Boreades) solani (Heer, 1841), male (Hokkaido); D Sibirhelus corpulentus (Reitter, 1900), male (Iwate Pref.); E Brachypterus urticae (Fabricius, 1792), male (Hokkaido); FG Brachypterolus shimoyamai Hisamatsu, 1985: F holotype, male; G ditto, labels. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5329319" httpUri="https://zenodo.org/record/5329319/files/figure.png" pageId="21" pageNumber="572">Fig. 2D</figureCitation>
) elongate oval, strongly convex, feebly shining; dorsal disc with a long, yellowish setae. Coloration fully luteous or rarely dark brown.
</paragraph>
<paragraph id="8B9E605EEA1F8729FF07FA99FCA56512" blockId="21.[113,1089,1167,1639]" pageId="21" pageNumber="572">
Head densely punctate, each puncture larger than eye-facet, separated by &lt;1 diameter; interspaces finely reticulate or smooth. Frontoclypeal suture incomplete, distinctly visible. Front margin of clypeus with medial arcuate emargination. Labrum broadly notched at middle. Mandibles slightly bent inward. Antennae (
<figureCitation id="131A7CDBEA1F8729FD0CFA39FD4166D2" box="[667,752,1487,1512]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="21" pageNumber="572">Fig. 9G</figureCitation>
) 1.16 times longer than HW (n = 2), appearing moniliform, without club; approximate ratio of each segment (n = 1) is 2.00: 1.14: 1.43: 1.14: 1.14: 1.00: 1.00: 1.00: 1.14: 1.29: 2.14.
</paragraph>
<paragraph id="8B9E605EEA1F872AFF07F9C6FEB36269" blockId="21.[113,1089,1167,1639]" lastBlockId="22.[151,1127,219,691]" lastPageId="22" lastPageNumber="573" pageId="21" pageNumber="572">Pronotum strongly convex, 1.35 times as wide as long (n =1); lateral margins narrowly explanate, arcuately rounded, widest at mid-length; anterior corner not prominent; posterior corner obtusely angulate; anterior margin nearly straight or with slight curvature, clearly bordered; basal margin gently arcuated, slightly sinuated at mid-width, distinctly bordered; punctures on disc larger than those on head, separated by &lt;1 diameter; interspaces finely reticulate.</paragraph>
<paragraph id="8B9E605EEA1C872AFF22FEADFE3D62C9" blockId="22.[151,1127,219,691]" pageId="22" pageNumber="573">Elytra conjointly 1.03 times as long as wide (n = 1), 1.75 times as long as pronotum (n = 1), subparallel-sided; punctures on disc larger than those on pronotum, extremely dense, separated by &lt;1 diameter; interspaces smooth. Abdominal tergite VI partially obscured by elytra. Abdominal tergite VII fully exposed, apex arcuately emarginate. Abdominal tergite VIII externally visible.</paragraph>
<paragraph id="8B9E605EEA1C872BFF22FE0AFEE76249" blockId="22.[151,1127,219,691]" lastBlockId="23.[113,1089,219,884]" lastPageId="23" lastPageNumber="574" pageId="22" pageNumber="573">
Prosternum (excluding prosternal process) rather long, 0.56 times as long as mesoventrite (n = 1), 0.40 times as long as metaventrite (n = 1); prosternal process slightly converging towards apex. Mesoventrite depressed below the level of the metaventrite, apical margin between inter-mesocoxa straight. Metaventrite convex, strongly shining, metathoracic discrimen in basal 1/2; disc with sparse punctures, separated by 1 or 2 diameter(s) at the middle; punctures becoming denser laterally. Inter-mesocoxal distance separated by 1.67 times width of inter-procoxal distance. Inter-metacoxal distance separated by 2.50 times width of interprocoxal distance. Abdominal sternites III (
<figureCitation id="131A7CDBEA1D872BFDD4FF0DFD29622E" box="[579,664,251,276]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="23" pageNumber="574">Fig. 9K</figureCitation>
) with medial tuft of setae; approximate ratio of length of abdominal sternites IIIVII (n = 1) is 3.00: 1.00: 1.00: 1.90: 3.00. Legs slender and long, protibiae (
<figureCitation id="131A7CDBEA1D872BFE0AFECDFE57626E" box="[413,486,315,340]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="23" pageNumber="574">Fig. 9I</figureCitation>
) strongly medially arcuate; claws (
<figureCitation id="131A7CDBEA1D872BFCC9FECDFC01626E" box="[862,944,315,340]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="23" pageNumber="574">Fig. 9E</figureCitation>
) more or less expanded at the base.
</paragraph>
<caption id="DF5E30D6EA1C872AFF01F9FDFE8B6554" ID-DOI="http://doi.org/10.5281/zenodo.5329341" ID-Zenodo-Dep="5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="22" pageNumber="573" startId="22.[150,181,1547,1566]" targetBox="[297,982,717,1504]" targetPageId="22">
<paragraph id="8B9E605EEA1C872AFF01F9FDFE8B6554" blockId="22.[150,1127,1547,1646]" pageId="22" pageNumber="573">
Fig. 9.
<taxonomicName id="4C211BDDEA1C872AFF46F9FAFDA16524" authority="(Reitter, 1900)" baseAuthorityName="Reitter" baseAuthorityYear="1900" box="[209,528,1547,1566]" class="Insecta" family="Kateretidae" genus="Sibirhelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="573" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B955BC4CEA1C872AFF46F9FAFE216524" box="[209,400,1548,1566]" italics="true" pageId="22" pageNumber="573">Sibirhelus corpulentus</emphasis>
(
<bibRefCitation id="EFB01DAFEA1C872AFE0EF9FDFDBA6524" author="REITTER E." box="[409,523,1547,1566]" pageId="22" pageNumber="573" pagination="225 - 232" refId="ref20598" refString="REITTER E. 1900: Beschreibung und Abbildung von neun Coleopteren der palaearctischen Fauna. Wiener Entomologische Zeitung 19: 225 - 232, pl. 2." type="journal article" year="1900">Reitter, 1900</bibRefCitation>
)
</taxonomicName>
.A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi,
</paragraph>
</caption>
<paragraph id="8B9E605EEA1D872BFF07FE8DFEE862E9" blockId="23.[113,1089,219,884]" pageId="23" pageNumber="574">
Male genitalia sclerotized; tegmen (
<figureCitation id="131A7CDBEA1D872BFD87FE8DFDD562AE" box="[528,612,379,404]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="23" pageNumber="574">Fig. 9A</figureCitation>
) with parameres asymmetrical, bearing long setae; interparameral lobe broadly rounded; median lobe (
<figureCitation id="131A7CDBEA1D872BFD44FE6DFC95628E" box="[723,804,411,436]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="23" pageNumber="574">Fig. 9C</figureCitation>
) abruptly bent inward near apex in lateral aspect.
</paragraph>
<paragraph id="8B9E605EEA1D872BFF07FE2DFE796129" blockId="23.[113,1089,219,884]" pageId="23" pageNumber="574">
Female. Apical margin of abdominal tergite VII rounded. Ovipositor (
<figureCitation id="131A7CDBEA1D872BFC05FE2DFC5962CE" box="[914,1000,475,500]" captionStart="Fig" captionStartId="22.[150,181,1547,1566]" captionTargetBox="[297,982,717,1504]" captionTargetId="figure-184@22.[290,987,714,1518]" captionTargetPageId="22" captionText="Fig. 9. Sibirhelus corpulentus (Reitter, 1900).A tegmen (ventral view); B median lobe (ventral view); C median lobe (lateral view); D ovipositor; E tarsal claw of right foreleg; F left maxillary palpus; G male right antenna; H female right protibia; I male right protibia; J pronotum; K abdominal sternites. AK = Tochigi Pref., Watarase-yûsuichi," figureDoi="http://doi.org/10.5281/zenodo.5329341" httpUri="https://zenodo.org/record/5329341/files/figure.png" pageId="23" pageNumber="574">Fig. 9D</figureCitation>
) deeply bifid, with extremely short styli.
</paragraph>
<paragraph id="8B9E605EEA1D872BFFE6FDEDFF7C61E9" blockId="23.[113,1089,219,884]" pageId="23" pageNumber="574">
<emphasis id="B955BC4CEA1D872BFFE6FDEDFF5E6109" bold="true" box="[113,239,539,563]" pageId="23" pageNumber="574">Variability.</emphasis>
Russian specimens have medial tuft of setae on abdominal sternites IIIV (Kirejtshuk, pers. comm.), but Japanese specimens have these tufts on abdominal sternite III only. The tarsal claws were described as subdentatis in the original description (
<bibRefCitation id="EFB01DAFEA1D872BFC36FDAAFB84614E" author="REITTER E." box="[929,1077,604,628]" pageId="23" pageNumber="574" pagination="225 - 232" refId="ref20598" refString="REITTER E. 1900: Beschreibung und Abbildung von neun Coleopteren der palaearctischen Fauna. Wiener Entomologische Zeitung 19: 225 - 232, pl. 2." type="journal article" year="1900">REITTER 1900</bibRefCitation>
), and as dentate in
<bibRefCitation id="EFB01DAFEA1D872BFEAFFD8AFE4E61AE" author="KIREJTSHUK A. G." box="[312,511,636,660]" pageId="23" pageNumber="574" pagination="210 - 216" refId="ref19575" refString="KIREJTSHUK A. G. 1992: Family Kateretidae Erichson, 1846. Pp. 210 - 216. In: LEHR P. A. (ed.): Opredelitel' nasekomykh Dal'nego Vostoka SSSR. [Identification key to the insects of the Far East of the USSR. Volume 3. Part 2.]. Nauka Pren, St. Petersburg, Russia, 704 pp (in Russian)." type="book chapter" year="1992">KIREJTSHUK (1992)</bibRefCitation>
. However, in the Japanese specimens examined so far, the base of the tarsal claws is more or less expanded at the base with no apparent tooth or denticle.
</paragraph>
<paragraph id="8B9E605EEA1D872BFFE6FD2DFF686009" blockId="23.[113,1089,219,884]" pageId="23" pageNumber="574">
<emphasis id="B955BC4CEA1D872BFFE6FD2DFF5E61C9" bold="true" box="[113,239,731,755]" pageId="23" pageNumber="574">Bionomics.</emphasis>
Twelve specimens of
<taxonomicName id="4C211BDDEA1D872BFE41FD2BFDDC61CE" baseAuthorityName="Reitter" baseAuthorityYear="1900" box="[470,621,733,756]" class="Insecta" family="Kateretidae" genus="Sibirhelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="574" phylum="Arthropoda" rank="species" species="corpulentus">
<emphasis id="B955BC4CEA1D872BFE41FD2BFDDC61CE" box="[470,621,733,756]" italics="true" pageId="23" pageNumber="574">S. corpulentus</emphasis>
</taxonomicName>
were collected from flowers of
<taxonomicName id="4C211BDDEA1D872BFC2EFD2BFEC9602E" authority="Boott" authorityName="Boott" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="23" pageNumber="574" phylum="Tracheophyta" rank="species" species="dispalata">
<emphasis id="B955BC4CEA1D872BFC2EFD2BFF2B602E" italics="true" pageId="23" pageNumber="574">Carex dispalata</emphasis>
Boott (Cyperaceae)
</taxonomicName>
, in Watarase-yûsuichi,
<collectingRegion id="49E5AEBCEA1D872BFDE1FD0DFCB26029" box="[630,771,763,787]" country="Japan" name="Tochigi" pageId="23" pageNumber="574">Tochigi Pref.</collectingRegion>
No additional information is available.
</paragraph>
</subSubSection>
<subSubSection id="C33B33D5EA1D872BFFE6FCCDFE49604E" pageId="23" pageNumber="574" type="distribution">
<paragraph id="8B9E605EEA1D872BFFE6FCCDFE49604E" blockId="23.[113,1089,219,884]" pageId="23" pageNumber="574">
<emphasis id="B955BC4CEA1D872BFFE6FCCDFEB56069" bold="true" box="[113,260,827,851]" pageId="23" pageNumber="574">Distribution.</emphasis>
<collectingCountry id="F33620CEEA1D872BFE9AFCCAFEFB6069" box="[269,330,828,851]" name="Japan" pageId="23" pageNumber="574">Japan</collectingCountry>
(Honshû) first recorded by
<bibRefCitation id="EFB01DAFEA1D872BFD0FFCCAFCEC606E" author="HISAMATSU S." box="[664,861,828,852]" pageId="23" pageNumber="574" pagination="14 - 15" refId="ref18926" refString="HISAMATSU S. 2003: Two nitidulid genera new to Japan (Coleoptera). Shikoku Tyuho 37: 14 - 15 (in Japanese, with English title)." type="journal article" year="2003">HISAMATSU (2003)</bibRefCitation>
;
<collectingCountry id="F33620CEEA1D872BFCFCFCCDFC036069" box="[875,946,827,851]" name="Russia" pageId="23" pageNumber="574">Russia</collectingCountry>
(
<collectingRegion id="49E5AEBCEA1D872BFC54FCCDFBA16069" box="[963,1040,827,851]" country="Russia" name="Irkutsk" pageId="23" pageNumber="574">Irkutsk</collectingRegion>
and
<collectingRegion id="49E5AEBCEA1D872BFFE6FCADFEAF604E" box="[113,286,859,884]" country="Russia" name="Primorskiy" pageId="23" pageNumber="574">Primorsky Kray</collectingRegion>
) (
<bibRefCitation id="EFB01DAFEA1D872BFEA3FCAAFE5D604E" author="KIREJTSHUK A. G." box="[308,492,860,884]" pageId="23" pageNumber="574" pagination="210 - 216" refId="ref19575" refString="KIREJTSHUK A. G. 1992: Family Kateretidae Erichson, 1846. Pp. 210 - 216. In: LEHR P. A. (ed.): Opredelitel' nasekomykh Dal'nego Vostoka SSSR. [Identification key to the insects of the Far East of the USSR. Volume 3. Part 2.]. Nauka Pren, St. Petersburg, Russia, 704 pp (in Russian)." type="book chapter" year="1992">KIREJTSHUK 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>