187 lines
16 KiB
XML
187 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. & Tishechkin, Alexey K." docDate="2013" docId="DA7F1E87735AC018BB612721F12EDCB8" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus lama Mazur 1988" docType="treatment" docVersion="3" lastPageNumber="324" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="321" updateTime="1668155382655" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Caterino, Michael S.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2013</mods:date>
|
||
<mods:detail type="volume">
|
||
<mods:number>271</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>401</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-GBIF-Taxon="152040868" LSID="urn:lsid:plazi:treatment:DA7F1E87735AC018BB612721F12EDCB8" httpUri="http://treatment.plazi.org/id/DA7F1E87735AC018BB612721F12EDCB8" lastPageId="323" lastPageNumber="324" pageId="320" pageNumber="321">
|
||
<subSubSection pageId="320" pageNumber="321" type="nomenclature">
|
||
<paragraph pageId="320" pageNumber="321">
|
||
<taxonomicName LSID="http://species-id.net/wiki/Operclipygus_lama" authority="Mazur, 1988" authorityName="Mazur" authorityYear="1988" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus lama" order="Coleoptera" pageId="320" pageNumber="321" phylum="Arthropoda" rank="species" species="lama">Operclipygus lama Mazur, 1988</taxonomicName>
|
||
Figs 85
|
||
<normalizedToken originalValue="F–H87A–EMap">F-H87A-EMap</normalizedToken>
|
||
31
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="320" pageNumber="321" type="reference_group">
|
||
<paragraph pageId="320" pageNumber="321">
|
||
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus lama" order="Coleoptera" pageId="320" pageNumber="321" phylum="Arthropoda" rank="species" species="lama">Operclipygus lama</taxonomicName>
|
||
Mazur, 1988: 295.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="320" pageNumber="321" type="type locality">
|
||
<paragraph pageId="320" pageNumber="321">Type locality.</paragraph>
|
||
<paragraph pageId="320" pageNumber="321">
|
||
ECUADOR: Napo: Puerto
|
||
<normalizedToken originalValue="Misahuallí">Misahualli</normalizedToken>
|
||
[
|
||
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-1.0333333">1°02'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-77.666664">77°40'W</geoCoordinate>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="320" pageNumber="321" type="type material">
|
||
<paragraph pageId="320" pageNumber="321">Type material.</paragraph>
|
||
<paragraph pageId="320" pageNumber="321">Holotype male: Ecuador, Napo, Misahualli, II/19/1983; not examined (LUND).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="321" lastPageNumber="322" pageId="320" pageNumber="321" type="other material">
|
||
<paragraph pageId="320" pageNumber="321">Other material.</paragraph>
|
||
<paragraph lastPageId="321" lastPageNumber="322" pageId="320" pageNumber="321">
|
||
BOLIVIA: Cochabamba: 6: Est. Biol. Valle Sajta, Univ. San Simon, 67.5km E Villa Tunari,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-17.105278">17°06'19"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-64.7825">64°46'57"W</geoCoordinate>
|
||
, 300m, 9-13.ii.1999, FIT, lowland rain forest, F. Genier (CMNC), 2: 7-9.ii.1999, FIT, lowland rain forest, F. Genier (CMNC); La Paz: 1: Guanay, Uyapi,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-15.416667">15°25'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-67.76667">67°46'W</geoCoordinate>
|
||
, 20.x.1999, rotting logs, G. Carrasco (CHND). ECUADOR: Orellana: 1:
|
||
<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
|
||
Res. Stn.,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-0.67444444">00°40'28"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-76.64722">76°38'50"W</geoCoordinate>
|
||
, 215m, 5-10.ix.1999, FIT, Primary forest, E.G. Riley (TAMU); same data except as noted: 2: 20-29.vi.1999, FIT, C.E. Carlton & A.K. Tishechkin (LSAM), 1: 28-5.vii.1999 (LSAM), 3: 22-28.vi.1999 (LSAM), 2: 12-20.vii.1999 (LSAM), 2: 26.vii-4.viii.1999 (LSAM), 1: 26.vii-4.viii.1999 (LSAM), 3: 5-12.vii.1999 (LSAM), 1: 23-30.vi.1999 (LSAM), 1: 5-11.vii.1999 (LSAM), 2: 28.
|
||
<normalizedToken originalValue="vi–">vi-</normalizedToken>
|
||
5.vii.1999 (LSAM), 1: 14-23.vii.1999 (LSAM), 1: 28.
|
||
<normalizedToken originalValue="vi–">vi-</normalizedToken>
|
||
5.vii.1999 (LSAM), 1: 11-18.vii.1999 (LSAM); 1: Yuturi Lodge, Rio Napo,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-0.5983333">0°35'54"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-76.03833">76°2'18"W</geoCoordinate>
|
||
, 270m, 20-21.iii.1999, FIT, R. Brooks & D. Brzoska (SEMC); 16: Tiputini Biodiversity Station,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="5" value="-0.6376">0.6376°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="5" value="-76.1499">76.1499°W</geoCoordinate>
|
||
, 4-9.vi.2011, FIT, M.S. Caterino & A.K. Tishechkin, DNA Extract MSC-2174 (SBMNH, USFQ), 4: 5-25.ix.2000, D.J. Inward & K.A. Jackson (BMNH); 8: Parque Nac.
|
||
<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
|
||
, Via Maxus at Puente
|
||
<normalizedToken originalValue="Piraña">Pirana</normalizedToken>
|
||
,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.65833336">0°39.5'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.433334">76°26'W</geoCoordinate>
|
||
, 14-20.vii.2007, FIT, A.K. Tishechkin (AKTC), 1: 20-24.vii.2008, FIT, A.K. Tishechkin (AKTC); 2: Payamino Research Station,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="1" value="-0.49333608">0°29'36.01"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-77.29143">77°17'29.15"W</geoCoordinate>
|
||
, 300m, 30.vii-12.viii.2007, FIT, tropical rainforest, CPDT Gillett (BMNH);
|
||
<normalizedToken originalValue="Sucumbíos">Sucumbios</normalizedToken>
|
||
: 1: Sacha Lodge,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-0.47055554">0°28'14"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-76.45972">76°27'35"W</geoCoordinate>
|
||
, 270m, 21-24.iii.1999, FIT, R. Brooks (SEMC). PERU: Ayacucho: 1: La Mar, Santa Rosa, 640m, 8-15.ix.1976, R. Gordon (USNM); Loreto: 2: 1.5km N Teniente Lopez,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-2.5943334">2°35.66'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-76.11533">76°06.92'W</geoCoordinate>
|
||
, 210-240m, 20.vii.1993, FIT, R. Leschen (SEMC); 9: 1.5km N Teniente Lopez,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="9" value="-2.5943334">2°35.66'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="9" value="-76.11533">76°06.92'W</geoCoordinate>
|
||
, 210-240m, 18.vii.1993, FIT, R. Leschen (SEMC); Madre de Dios: 6: 24.x.1982, rotten palm flowers, L. Watrous & G. Mazurek (FMNH); 1: Manu National
|
||
<pageBreakToken pageId="321" pageNumber="322" start="start">Park</pageBreakToken>
|
||
, Zona res., Rio Manu, Cocha Juarez, trail nr. Manu Lodge, 18-24.ix.1991, FIT, A. Hartman (FMNH); 6: Manu National Park, Cocha Salvador,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.003612">12°0'13"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-71.52667">71°31'36"W</geoCoordinate>
|
||
, 310m, 20-21.x.2000, FIT, R. Brooks (SEMC); 1: Manu National Park, Pantiacolla Lodge, Alto Madre de Dios River,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-12.656111">12°39'22"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-71.23194">71°13'55"W</geoCoordinate>
|
||
, 400m, 23-26.x.2000, FIT, R. Brooks (SEMC); 9: Manu National Park, Cocha Cashu Bio. Sta.,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-11.895833">11°53'45"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="15" value="-71.40667">71°24'24"W</geoCoordinate>
|
||
, 350m, 17-19.x.2000, FIT, R. Brooks (SEMC); 1: Manu National Park, Rio Alto Madre de Dios, Pantiacolla Lodge,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-12.655">12°39.3'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-71.23167">71°13.9'W</geoCoordinate>
|
||
, 14-19.xi.2007, FIT, D. Brzoska (SEMC); 1: Amazonas Lodge, N Atalaya,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="92" value="-12.87">12°52.2'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="92" value="-71.37666">71°22.6'W</geoCoordinate>
|
||
, 480m, 10-13.xi.2007, FIT, D. Brzoska (SEMC); 1: Tambopata, Reserva Cuzco Amazonico, 15km NE Pto. Maldonado,
|
||
<geoCoordinate direction="south" orientation="latitude" precision="925" value="-12.55">12°33'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate direction="west" orientation="longitude" precision="925" value="-69.05">69°03'W</geoCoordinate>
|
||
, 200m, 16.vii.1989,
|
||
<taxonomicName class="Magnoliopsida" family="Combretaceae" genus="Buchenavia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Buchenavia" order="Myrtales" pageId="321" pageNumber="322" phylum="Tracheophyta" rank="genus">Buchenavia</taxonomicName>
|
||
fruit fall, J. Ashe & R. Leschen (SEMC), 8: 24.vi.1989, FIT (SEMC), 22: 22.vi.1989, FIT (SEMC), 3: 20.vi.1989, FIT (SEMC), 6: 16.vii.1989, FIT (SEMC), 2: 19.vii.1989, FIT (SEMC), 5: 13.vii.1989, FIT (SEMC), 5: 13.vi.1989, FIT (SEMC), 1: 30.vi.1989, FIT (CHSM), 3: 28.vi.1989, FIT (SEMC), 4: 26.vi.1989, FIT (SEMC), 7: 17.vi.1989, FIT (SEMC), 5: 15.vi.1989, FIT (SEMC), 1: 16.vi.1989, Agaricales (SEMC), 1: 27.vi.1989, flat ascomycete (SEMC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="323" lastPageNumber="324" pageId="321" pageNumber="322" type="description">
|
||
<paragraph pageId="321" pageNumber="322">Diagnostic description.</paragraph>
|
||
<paragraph lastPageId="323" lastPageNumber="324" pageId="321" pageNumber="322">
|
||
Length: 1.93-2.18 mm, width: 1.93-2.00 mm; body piceous, rounded, convex; frons weakly depressed, frontal stria complete, continuous with complete supraorbital stria; pronotum with lateral submarginal stria complete, continued anteriorly by anterior marginal stria, which is then interrupted for the distance of the outwardly arcuate anterior pronotal emargination; anterior submarginal
|
||
<pageBreakToken pageId="322" pageNumber="323" start="start">pronotal</pageBreakToken>
|
||
stria detached, barely recurved posterad at apices, median pronotal gland openings just beyond the ends of this stria; pronotal disk with few or no lateral punctures and lacking prescutellar impression; elytron with two complete epipleural striae, outer subhumeral stria entire, sinuate at middle, inner subhumeral stria absent, striae 1-3 complete, 4th stria present in apical half, 5th stria present in apical third, sutural stria present in apical three-fourths; prosternal keel projecting into mesoventral emargination, prosternal carinal striae connected in wide anterior arch, about one-fifth short of presternal suture; prosternal lobe short, marginal stria continued laterally by series of punctures; mesoventral marginal stria usually interrupted; mesometaventral stria arched forward in middle, metaventral disk lacking punctures; 1st abdominal ventrite
|
||
<pageBreakToken pageId="323" pageNumber="324" start="start">with</pageBreakToken>
|
||
two striae on each side, outer stria frequently interrupted; abdominal ventrites 3 and 4 with single series of very large punctures at sides; propygidium with fine, sparse ground punctation and a dense group of deep punctures slightly removed from all margins; pygidium with ground punctation denser than that of propygidium with coarser punctures concentrated along basal margin, more sparsely scattered elsewhere; marginal pygidial stria complete, finely crentulate. Male genitalia (Figs 87
|
||
<normalizedToken originalValue="A–E">A-E</normalizedToken>
|
||
): accessory sclerites absent; S8 with apical guides strongly developed from base to apex; S9 desclerotized along midline, with deep apical emargination and small, separate apical flanges; tegmen with basomedial process divided into two lateral processes; median lobe short, broad.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="323" pageNumber="324" type="remarks">
|
||
<paragraph pageId="323" pageNumber="324">Remarks.</paragraph>
|
||
<paragraph pageId="323" pageNumber="324">This is a rather isolated species, with the divided medioventral processes of the aedeagus (Fig. 87E) found in no other species. Externally, it is most easily recognized by the large deep punctures on ventrites 3 and 4 (Fig. 85G), and the small but deep punctures on the propygidium (Fig. 85H).</paragraph>
|
||
<paragraph pageId="323" pageNumber="324">
|
||
Although we did not study the type specimen, we have seen numerous specimens determined by the
|
||
<normalizedToken originalValue="species’">species'</normalizedToken>
|
||
author, all agreeing in characters with the original description, and to this widespread, more or less invariable species.
|
||
</paragraph>
|
||
<caption pageId="323" pageNumber="324">
|
||
<paragraph pageId="323" pageNumber="324">
|
||
Figure 87. Male genitalia of
|
||
<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus lama" order="Coleoptera" pageId="323" pageNumber="324" phylum="Arthropoda" rank="species" species="lama">Operclipygus lama</taxonomicName>
|
||
. A T8 B S8 C T9 & T10 D S9 E Aedeagus, dorsal and lateral views.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |