treatments-xml/data/D8/9D/F8/D89DF85F64A751AC96A34B873FC92FF4.xml

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<mods:title id="8A94CB0FE53A02BF61CEA1BA1CAB7591">Reclassification of Cybistrinae Sharp, 1880 in the Neotropical Region (Coleoptera, Adephaga, Dytiscidae), with description of new taxa</mods:title>
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<mods:namePart id="329D290E8AC7A489714CDAD1AB0DB45B">Miller, Kelly B.</mods:namePart>
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<mods:affiliation id="FDD9DF20B0237288A655CBE2E7F6C212">Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131 - 0001, USA</mods:affiliation>
<mods:nameIdentifier id="359837E14267B28E649E8A2F7F61B3F3" type="email">kbmiller@unm.edu</mods:nameIdentifier>
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<mods:namePart id="24D73B342C8D8DB2D54AD577F983BC44">Michat, Mariano C.</mods:namePart>
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<mods:affiliation id="9FADC04EBD36C6C0526D0C102E69C026">Instituto de Biodiversidad y Biologia Experimental y Aplicada, CONICET-Universidad de Buenos Aires, Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="53265458B9108EE71A9FCA0468D3B261">Ferreira Jr, Nelson</mods:namePart>
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<mods:affiliation id="5EA0527A75AEE050E6BBEDAB3B4FE70B">Laboratorio de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Rio de Janeiro, RJ, Brasi</mods:affiliation>
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<mods:date id="B619ED9A7538EDC3F029712016F098DB">2024</mods:date>
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<paragraph id="73D69492DAD61192F76003F4C4477599" pageId="0" pageNumber="125">
<taxonomicName id="C637DD35FC15D46EB4B7A12DAB6C57A0" LSID="https://zoobank.org/3F2BEED6-5AA7-4394-9309-788F3ACFACD1" authority="Miller &amp; Michat &amp; Ferreira Jr, 2024" authorityName="Miller &amp; Michat &amp; Ferreira Jr" authorityYear="2024" family="Dytiscidae" genus="Nilssondytes" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nilssondytes diversus" order="Coleoptera" pageId="0" pageNumber="125" rank="species" species="diversus" status="sp. nov.">Nilssondytes diversus</taxonomicName>
<taxonomicNameLabel id="46D1731DCDAC4ADA001862763A015BB0" pageId="0" pageNumber="125">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="D764FAF1672CE0B9FF8213FBE89A8005" pageId="0" pageNumber="125" type="description">
<paragraph id="585CA1F7A9779330EA913CC02E474DB8" pageId="0" pageNumber="125">
<figureCitation id="490D9519B846E0468AD952E5E1257193" captionStart="Figures 9–14" captionStartId="F2" captionText="Figures 9 - 14. Neotropical Cybistrinae species, habitus 9 Cybister festae 10 Cybister puncticollis 11 Nilssondytes diversus 12 Megadytes parvus 13 Megadytes latus 14 species &quot; IR 57 &quot; (Ribera et al. 2008). Scale bar: 10.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures9-14" httpUri="https://binary.pensoft.net/fig/963584" pageId="0" pageNumber="125">Figs 11</figureCitation>
<figureCitation id="B5160B295315A83F44ED4D7D67601F32" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">, 19</figureCitation>
<figureCitation id="915EC3DD38968733D49CADB9B645CFD1" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">, 20</figureCitation>
<figureCitation id="3D70B0FA2DC48FD434E2F4CF803CCDC6" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">, 36-38</figureCitation>
<figureCitation id="3B39EB21148A08CACEEF3433ACD9DD2D" captionStart="Figures 51–57" captionStartId="F5" captionText="Figures 51 - 57. Neotropical Cybistrinae species, male sternite IX, ventral aspect 51 Cybister festae 52 Cybister puncticollis 53 Nilssondytes diversus 54 Megadytes latus 55 Megadytes parvus 56 Paramegadytes glaucus 57 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures51-57" httpUri="https://binary.pensoft.net/fig/963587" pageId="0" pageNumber="125">, 53</figureCitation>
<figureCitation id="1ECEADE0150922D85FE2620E5398ACCB" captionStart="Figures 58–61" captionStartId="F6" captionText="Figures 58 - 61. Neotropical Cybistrinae species, female genitalia including left gonocoxosternite, ventral aspect, except Fig. 60 without internal genitalia 58 Cybister festae 59 Cybister puncticollis 60 Megadytes latus 61 Nilssondytes diversus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures58-61" httpUri="https://binary.pensoft.net/fig/963588" pageId="0" pageNumber="125">, 61</figureCitation>
<figureCitation id="6EFD97AFDB9953907E218C4DAF4A184D" captionStart="Figure 73" captionStartId="F10" captionText="Figure 73. Nilssondytes diversus, holotype specimen. Scale bar: 5.0 mm." figureDoi="10.3897/zookeys.1188.110081.figure73" pageId="0" pageNumber="125">, 73</figureCitation>
<figureCitation id="66C12A9F70798E006C509ADECA2361E5" captionStart="Figure 74" captionStartId="F11" captionText="Figure 74. Distribution of Nilssondytes diversus in northern South America." figureDoi="10.3897/zookeys.1188.110081.figure74" httpUri="https://binary.pensoft.net/fig/963592" pageId="0" pageNumber="125">, 74</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C5C78BE5362D95F5CA6626BEEE9C8097" pageId="0" pageNumber="125" type="type locality">
<paragraph id="A49169916BDFD5CA9F6D28DC7D9B3FB0" pageId="0" pageNumber="125">Type locality.</paragraph>
<paragraph id="8EF1BF0F295971E728E1E910A6943349" pageId="0" pageNumber="125">
Venezuela, Amazonas State, roadside pond ca. 7 km S Samariapo 
<geoCoordinate id="37A8E37BE6D2F51A6D15ACFF60DB7F24" degrees="5" direction="north" minutes="10.900" orientation="latitude" precision="1" value="5.181667">5°10.900'N</geoCoordinate>
, 
<geoCoordinate id="ADE662C5073E7430012E2EEEEC3F1857" degrees="67" direction="west" minutes="46.078" orientation="longitude" precision="1" value="-67.76797">67°46.078'W</geoCoordinate>
, 95 m elev.
</paragraph>
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<subSubSection id="A82B2491E7C66E53DD96C8CB7F66B1B7" pageId="0" pageNumber="125" type="diagnosis">
<paragraph id="06AFFB8D6BABA7FAD2AAB4273C4B5684" pageId="0" pageNumber="125">Diagnosis.</paragraph>
<paragraph id="D941D7EC08F10B97D73AC4B0F5E985CE" pageId="0" pageNumber="125">
This is the only species in the genus and is characterized by its diagnostic combination (see above). Typically, species-level features include the shape of the male median lobe which is unique. In ventral aspect the apex is abruptly constricted with the apex narrowly truncate with laterally pointed processes (Fig. 
<figureCitation id="6CE9F8999B46EAA5F3E0B45823781C7B" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">37</figureCitation>
). In lateral aspect, the median lobe is moderately evenly curved on the dorsal margin, lobe apically abruptly narrowed with the apex elongate and slender, apically narrowly rounded (Fig. 
<figureCitation id="E47F8A923FA81FF065EFE90B734839BC" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">36</figureCitation>
).
</paragraph>
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<subSubSection id="C9E6AEC6847B6C8FCB9E135FA927A19A" pageId="0" pageNumber="125" type="description">
<paragraph id="BCBB508AE8C560D44DD74C1DFD42D7D2" pageId="0" pageNumber="125">Description.</paragraph>
<paragraph id="296DA87A225F81B620C5302EDE91000A" pageId="0" pageNumber="125">
<emphasis id="920528F30B383E44F69A2F90A7E167A2" bold="true" pageId="0" pageNumber="125">
<emphasis id="18245A6D45EA541C4A19615ACB001CBF" italics="true" pageId="0" pageNumber="125">Measurements</emphasis>
.
</emphasis>
TL = 16.7-19.4 mm, GW = 9.6-10.7 mm, PW = 7.0-8.1 mm, HW = 4.2-4.7 mm, EW = 2.7-2.9 mm, TL/GW = 1.7-1.8, HW/EW = 1.6-1.7, WC/WV = 3.1-3.2. Body shape suboval, slightly expanded posteriorly, widest at ~ 3/5 of length; lateral margins evenly, continuously curved between pronotum and elytron. Depressed and somewhat flattened in lateral aspect (Figs 
<figureCitation id="A5F5DF53A24D7511B8C8B40A31BF1D40" captionStart="Figures 9–14" captionStartId="F2" captionText="Figures 9 - 14. Neotropical Cybistrinae species, habitus 9 Cybister festae 10 Cybister puncticollis 11 Nilssondytes diversus 12 Megadytes parvus 13 Megadytes latus 14 species &quot; IR 57 &quot; (Ribera et al. 2008). Scale bar: 10.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures9-14" httpUri="https://binary.pensoft.net/fig/963584" pageId="0" pageNumber="125">11</figureCitation>
, 
<figureCitation id="4A1370440ABD5B5DFF003E8D18AC77BB" captionStart="Figure 73" captionStartId="F10" captionText="Figure 73. Nilssondytes diversus, holotype specimen. Scale bar: 5.0 mm." figureDoi="10.3897/zookeys.1188.110081.figure73" pageId="0" pageNumber="125">73</figureCitation>
).
</paragraph>
<paragraph id="18A722B94B3CFD178A1ECBC973CB2FC5" pageId="0" pageNumber="125">
<emphasis id="2A49B58553B33ACBFF95779601AAD608" bold="true" italics="true" pageId="0" pageNumber="125">Coloration</emphasis>
(Figs 
<figureCitation id="E4AC57D2C8308AEE3527C60685D09A61" captionStart="Figures 9–14" captionStartId="F2" captionText="Figures 9 - 14. Neotropical Cybistrinae species, habitus 9 Cybister festae 10 Cybister puncticollis 11 Nilssondytes diversus 12 Megadytes parvus 13 Megadytes latus 14 species &quot; IR 57 &quot; (Ribera et al. 2008). Scale bar: 10.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures9-14" httpUri="https://binary.pensoft.net/fig/963584" pageId="0" pageNumber="125">11</figureCitation>
, 
<figureCitation id="557DAA38412E88712CFBA77DF714D862" captionStart="Figure 73" captionStartId="F10" captionText="Figure 73. Nilssondytes diversus, holotype specimen. Scale bar: 5.0 mm." figureDoi="10.3897/zookeys.1188.110081.figure73" pageId="0" pageNumber="125">73</figureCitation>
). Head dark green, anterior clypeal margin yellow, more so laterally, testaceous near eyes. Pronotum dark green with broad lateral yellow margin, posteriorly interrupted and green in one of the four examined specimens, in other specimen yellow extending to posterior angle. Elytron dark green with broad lateral yellow band, separated narrowly from lateral margin, slightly expanded near apex. Ventral surfaces largely black, testaceous on head, basal leg segments and elytral epipleuron.
</paragraph>
<paragraph id="BAE87F850B3E53852C1CC66A32ACE198" pageId="0" pageNumber="125">
<emphasis id="19F8840C2DD5AE0A0EDAA6C3E1027F0B" bold="true" pageId="0" pageNumber="125">
<emphasis id="B5807B9925613AF20831C6080672B7A3" italics="true" pageId="0" pageNumber="125">Sculpture and structure</emphasis>
.
</emphasis>
Head broad, frontoclypeal lines elongate, straight, strongly oblique; anterior clypeal margin broadly, shallowly, and evenly concave; dorsal surface evenly covered with fine microsculpture and micropunctures. Pronotum with lateral margins evenly and broadly curved; surface similar to surface of head in microsculpture and micropunctation. Elytral lateral margin evenly and slightly curved for most of length, apically broadly curved; surface of elytron similar to surface of head in microsculpture and micropunctation. Prosternal process apically rounded, ventral surface distinctly sulcate, apex robust, acutely pointed. Metaventral wing broad, slightly less than 1/3 width of lateral portion of metacoxa (WC/WV = 3.1-3.2); surface smooth, without sculpturing. Lateral portion of metacoxa large, broad, surface smooth, without sculpturing; metacoxal lines short, extending less than half distance across metacoxa. Abdominal ventrites smooth, unsculptured.
</paragraph>
<paragraph id="284A0D718D8E9983A1569EC28CA9B327" pageId="0" pageNumber="125">
<emphasis id="020BFDEB5A39B01E5428D24636F6FA6B" bold="true" pageId="0" pageNumber="125">
<emphasis id="DFE90C523AA88F9F43673EFB8442A667" italics="true" pageId="0" pageNumber="125">Male genitalia</emphasis>
.
</emphasis>
Male median lobe in lateral aspect shallowly curved, apically abruptly narrowed, apex narrowed, slightly curved, apically narrowly rounded, broad medially (Fig. 
<figureCitation id="8F8B24C9E2E493D14DD29694E5A7BF45" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">36</figureCitation>
). In dorsal aspect broad throughout most of length, apically abruptly narrowed, apex laterally produced, submedially with broad, elongate lobes on each side, ventral sclerite short, apically sharp, acuminate, extending to 3/5 length of median lobe, apex sharply pointed (Fig. 
<figureCitation id="EA01F768DF36C88B7B913CFADCCE63BA" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">37</figureCitation>
). Lateral lobe broad in basal half, apically distinctly narrowed, apex narrowly rounded, with series of elongate setae along more than apical half of dorsal margin of lateral lobe (Fig. 
<figureCitation id="B9F34EDC4624267D8F84D5EE098F5023" captionStart="Figures 30–50" captionStartId="F4" captionText="Figures 30 - 50. Neotropical Cybistrinae species, male genitalia 30 - 32 Cybister festae: 30 median lobe, right lateral aspect 31 median lobe, ventral aspect 32 right lateral lobe, right lateral aspect 33 - 35 Cybister puncticollis: 33 median lobe, right lateral aspect 34 median lobe, ventral aspect 35 right lateral lobe, right lateral aspect 36 - 38 Nilssondytes diversus: 36 median lobe, right lateral aspect 37 median lobe, ventral aspect 38 right lateral lobe, right lateral aspect 39 - 43 Megadytes parvus: 39 median lobe, right lateral aspect 40 median lobe, ventral aspect 41 right lateral lobe, right lateral aspect 42 apex of median lobe, right lateral aspect 43 apex of median lobe, ventral aspect 44 - 47 Megadytes latus: 44 median lobe, right lateral aspect 45 median lobe, ventral aspect 46 right lateral lobe, right lateral aspect 47 apex of median lobe, ventral aspect 48 - 50 species &quot; IR 57 &quot; (Ribera et al. 2008): 48 median lobe, right lateral aspect 49 median lobe, ventral aspect 50 right lateral lobe, right lateral aspect." figureDoi="10.3897/zookeys.1188.110081.figures30-50" httpUri="https://binary.pensoft.net/fig/963586" pageId="0" pageNumber="125">38</figureCitation>
).
</paragraph>
<paragraph id="2EEEB227157FEDBB5868EC80EA7C043A" pageId="0" pageNumber="125">
<emphasis id="15C4F660474B3F7E37E68CF918561576" bold="true" pageId="0" pageNumber="125">
<emphasis id="4C486685B3A71BC837F57ECC108C3345" italics="true" pageId="0" pageNumber="125">Female genitalia</emphasis>
.
</emphasis>
With a single genital opening, vagina elongate, slender, with enlarged, bulbous region at base of common oviduct; spermatheca short, curved, at apex of enlarged region, with soft tissue region on each side of enlarged region (Fig. 
<figureCitation id="ACDB10B8A4E812CAF13009AC374901CB" captionStart="Figures 58–61" captionStartId="F6" captionText="Figures 58 - 61. Neotropical Cybistrinae species, female genitalia including left gonocoxosternite, ventral aspect, except Fig. 60 without internal genitalia 58 Cybister festae 59 Cybister puncticollis 60 Megadytes latus 61 Nilssondytes diversus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures58-61" httpUri="https://binary.pensoft.net/fig/963588" pageId="0" pageNumber="125">61</figureCitation>
); gonocoxae together broad, apically broadly pointed (Fig. 
<figureCitation id="FCA599929B9F4D3E7703D5E895B736D4" captionStart="Figures 58–61" captionStartId="F6" captionText="Figures 58 - 61. Neotropical Cybistrinae species, female genitalia including left gonocoxosternite, ventral aspect, except Fig. 60 without internal genitalia 58 Cybister festae 59 Cybister puncticollis 60 Megadytes latus 61 Nilssondytes diversus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures58-61" httpUri="https://binary.pensoft.net/fig/963588" pageId="0" pageNumber="125">61</figureCitation>
); gonocoxosternite broad, with elongate anterolateral lobe, with medial margin sublinear, without conspicuous setae (Fig. 
<figureCitation id="3D5BAF4A20390A1D78C98590FD695415" captionStart="Figures 58–61" captionStartId="F6" captionText="Figures 58 - 61. Neotropical Cybistrinae species, female genitalia including left gonocoxosternite, ventral aspect, except Fig. 60 without internal genitalia 58 Cybister festae 59 Cybister puncticollis 60 Megadytes latus 61 Nilssondytes diversus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures58-61" httpUri="https://binary.pensoft.net/fig/963588" pageId="0" pageNumber="125">61</figureCitation>
).
</paragraph>
<paragraph id="21F5D7E5A85F535BC94E191CA623F0FE" pageId="0" pageNumber="125">
<emphasis id="4BBBDEBA8664769288430F940F75C5AB" bold="true" pageId="0" pageNumber="125">
<emphasis id="638196E09155C1C63B957765EBCC8FB5" italics="true" pageId="0" pageNumber="125">Sexual dimorphism</emphasis>
.
</emphasis>
Males have a characteristic broad protarsal palette with ventral adhesive setae. Males also have mesotarsomeres with clumps of posteroventral setae. Females lack pro- and mesotarsal expansions or adhesive setae. Both males and females have two metatarsal claws with the posterior shorter than the anterior, but females have the posterior somewhat more curved than in males (Figs 
<figureCitation id="18FE0174B4701FB240C3D2741F0178FF" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">19</figureCitation>
, 
<figureCitation id="909627BDDD2A72382092C50A6680DFB2" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">20</figureCitation>
). Females have distinctive microsculpture on the surface of the elytron anteriorly in the form of a field of short striae which is absent in males.
</paragraph>
<paragraph id="CE08228CCF03F4F679CA04B6158F3E07" pageId="0" pageNumber="125">
<emphasis id="38CBD97F3CC1851669D5E1406C56779A" bold="true" pageId="0" pageNumber="125">
<emphasis id="6D1BFDEE26E666C494620F3B74A60157" italics="true" pageId="0" pageNumber="125">Variation</emphasis>
.
</emphasis>
Five specimens were examined. One specimen has the lateral pronotal yellow band extending to the posterior margin of the pronotum, the others have a narrow dark green separation from the posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="2AF1A6ABE77604082D8010E6FBAA3050" pageId="0" pageNumber="125" type="distribution">
<paragraph id="5A76BAE7834C2BD3B3AFB75F33F67AF3" pageId="0" pageNumber="125">Distribution.</paragraph>
<paragraph id="CBA40BD9D1AE8B7C59F0BB51C7A49F84" pageId="0" pageNumber="125">
This species is known from few localities in Venezuela along the northwestern margins of the Guiana Shield craton (Fig. 
<figureCitation id="8D1D9600CAD9780F7BE159B3A67A71D1" captionStart="Figure 74" captionStartId="F11" captionText="Figure 74. Distribution of Nilssondytes diversus in northern South America." figureDoi="10.3897/zookeys.1188.110081.figure74" httpUri="https://binary.pensoft.net/fig/963592" pageId="0" pageNumber="125">74</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7483E419D6A0A01DD841362FA8388970" pageId="0" pageNumber="125" type="natural history">
<paragraph id="B8BDF7294475E9ECB894824B2F9D2915" pageId="0" pageNumber="125">Natural history.</paragraph>
<paragraph id="4D07682B2E6495429EB68D7B56B17EC7" pageId="0" pageNumber="125">The only natural history information available from labels is &quot;roadside pond,&quot; &quot;river margin,&quot; and &quot;rock outcropping.&quot;</paragraph>
</subSubSection>
<subSubSection id="01B7B143E8DBB3298B866F9AD16ACBFB" pageId="0" pageNumber="125" type="etymology">
<paragraph id="4E05AD8FC94B2952740D74899FC01BE1" pageId="0" pageNumber="125">Etymology.</paragraph>
<paragraph id="0C5667EF7DF4C21304BD4CE7DCB8ED75" pageId="0" pageNumber="125">
The species is named from the Latin 
<taxonomicName id="781D58C074EAF02D450931564C627086" authorityName="Miller &amp; Michat &amp; Ferreira Jr" authorityYear="2024" family="Dytiscidae" genus="Nilssondytes" kingdom="Animalia" lsidName="Nilssondytes diversus" order="Coleoptera" pageId="0" pageNumber="125" rank="species" species="diversus">
<emphasis id="60CD426169753D1564B1CD22740C9E0F" italics="true" pageId="0" pageNumber="125">Nilssondytes diversus</emphasis>
</taxonomicName>
, meaning 
<normalizedToken id="DAC5F04FD9187850183AA9FB9286CA2B" originalValue="“different,”">&quot;different,&quot;</normalizedToken>
in recognition of the different lengths of the metatarsal claws in both males and females (Figs 
<figureCitation id="57AB86EFB093836C0A4D2FAB35D32F6C" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">19</figureCitation>
, 
<figureCitation id="7A4D7705357F4E64D1F009B6EE7CEE94" captionStart="Figures 15–29" captionStartId="F3" captionText="Figures 15 - 29. Neotropical Cybistrinae species, metatarsal claws and tarsomere VI of males and females 15, 16 Cybister festae 17, 18 Cybister puncticollis 19, 20 Nilssondytes diversus 21, 22 Megadytes latus 23, 24 Megadytes parvus 25 species &quot; IR 57 &quot; (Ribera et al. 2008) 26, 27 Paramegadytes glaucus 28, 29 Metaxydytes fraternus. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1188.110081.figures15-29" httpUri="https://binary.pensoft.net/fig/963585" pageId="0" pageNumber="125">20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EFF8C07DF6ECB2D7E1B81EA06CC4B932" pageId="0" pageNumber="125" type="materials_examined">
<paragraph id="8BCD265DE37ADE28D04842996BAC88A2" pageId="0" pageNumber="125">Material examined.</paragraph>
<paragraph id="9239AEACF86156F770F8C1F1089E735C" pageId="0" pageNumber="125">
<materialsCitation id="B3166D65E763CCA3C96F509CF90FF53E" collectingDate="2009-01-15" collectorName="S. Samariapo, Short" country="VENEZUELA" elevation="95" latitude="5.181667" location="Amazonas State" longLatPrecision="1" longitude="-67.76797" specimenCount="1" stateProvince="Amazonas State" typeStatus="Holotype">
<emphasis id="704AAD42DB9F9FF396F6BAA256373FFE" bold="true" italics="true" pageId="0" pageNumber="125">
<typeStatus id="31CD2DBC152C2E5B3D5A22C392BE3287" type="holotype">Holotype</typeStatus>
</emphasis>
, male labeled, &quot; 
<collectingCountry id="BE15753A8BC9465507FD9F7A063A4BFF" name="Venezuela">VENEZUELA</collectingCountry>
: 
<collectingRegion id="832919E178029EE06BE786000857F823" country="Venezuela" name="Amazonas">Amazonas State</collectingRegion>
<geoCoordinate id="DA4A43DB45EF1A00774C4D4F5C3197E3" degrees="5" direction="north" minutes="10.900" orientation="latitude" precision="1" value="5.181667">5°10.900'N</geoCoordinate>
, 
<geoCoordinate id="3FACE740CA2F1F5EF1D40B1D36A4D3AD" degrees="67" direction="west" minutes="46.078" orientation="longitude" precision="1" value="-67.76797">67°46.078'W</geoCoordinate>
, 
<quantity id="AF1AAC56712DD854AFAFD2C79AB98E86" metricMagnitude="1" metricUnit="m" metricValue="9.5" unit="m" value="95.0">
<elevation id="A1FF4F8C636FEE92376855C61A2AAF31" metricMagnitude="1" metricUnit="m" metricValue="9.5" unit="m" value="95.0">95 m</elevation>
</quantity>
ca. 
<quantity id="A7CA18967268564B4C79C636E9A4B20B" metricMagnitude="3" metricUnit="m" metricValue="7.0" unit="km" value="7.0">7 km</quantity>
<collectorName id="8A2A23E276777F59BAB4C16F89F53949">S. Samariapo</collectorName>
<collectingDate id="DA38719932F0A466E612C0AB1F53BDB3" value="2009-01-15">15.i.2009</collectingDate>
; leg. 
<collectorName id="E3CB0530D9AC789C6BA5EB22650BCF80">Short</collectorName>
, Miller, 
<normalizedToken id="2A695F51FCDB3C4627D67FD4E93C6A63" originalValue="García">Garcia</normalizedToken>
, Camacho, Joly VZ09-0115-02X: roadside pond/ SM0846115 KUNHM-ENT [barcode label]/ 
<typeStatus id="E325CB59133B33B90D014CCFD7FB24CF" type="holotype">HOLOTYPE</typeStatus>
: 
<taxonomicName id="6A65BD628BF1F280206C0EDEB1A2F2A0" authorityName="Miller &amp; Michat &amp; Ferreira Jr" authorityYear="2024" family="Dytiscidae" genus="Nilssondytes" kingdom="Animalia" lsidName="Nilssondytes diversus" order="Coleoptera" pageId="0" pageNumber="125" rank="species" species="diversus">
<emphasis id="8CF0A7ADFD469A05874744C2AB786AD5" italics="true" pageId="0" pageNumber="125">Nilssondytes diversus</emphasis>
</taxonomicName>
Miller, Michat and Ferreira-Jr., 2023 [red label with double black line border].&quot;
</materialsCitation>
<materialsCitation id="D80D0AE8DB2D95D91EB483E4B8B924D5" collectingDate="1900-04-20" collectorName="E. A. Klages." country="Venezuela" location="Caura River" municipality="Suapure" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="Paratypes">
<emphasis id="152D331C153C24F18C134E570CCA5650" bold="true" italics="true" pageId="0" pageNumber="125">
<typeStatus id="7A8546B1B8FF97B0C343D0960F52EB3D" type="paratype">Paratypes</typeStatus>
</emphasis>
, 
<specimenCount id="3D4235E46E0C8A459299A03A578C23C8" count="1" type="male">1 male</specimenCount>
labeled, &quot; 
<collectingMunicipality id="515EB2D086988A697746D25B19E461E7">Suapure</collectingMunicipality>
VENEZ. 
<location id="D4721530399D183FD3876AA11CB57303" LSID="urn:lsid:plazi:treatment:D89DF85F64A751AC96A34B873FC92FF4:D4721530399D183FD3876AA11CB57303" country="Venezuela" municipality="Suapure" name="Caura River" stateProvince="Amazonas">Caura River</location>
<collectingDate id="537B783676451EA70B392655B8B57219" value="1900-04-20">4.20.1900</collectingDate>
[handwritten] 
<collectorName id="5DB1AC16E7A1541532D7E454AEE02AEB">E.A. Klages.</collectorName>
&quot;
</materialsCitation>
, 
<materialsCitation id="B0B6334A3953BADE6F36792F08EAFA1D" collectingDate="2008-08-05" collectorName="A. Short, M. Garcia" country="VENEZUELA" elevation="134" latitude="7.689889" location="14 km E Rio Aro" longLatPrecision="1" longitude="-64.03223" specimenCount="1" specimenCount-female="1" stateProvince="Bolivar State" typeStatus="paratype">
<specimenCount id="7FABE4EAB2843CAA0B840C08EE4A1AC8" count="1" type="female">1 female</specimenCount>
labeled &quot; 
<collectingCountry id="5BD813AA422D89B302DF9CE6B6DE25AE" name="Venezuela">VENEZUELA</collectingCountry>
: 
<collectingRegion id="DE96CB0C84F36AC29506832673CF7850" country="Venezuela" name="Bolivar">Bolivar State</collectingRegion>
<geoCoordinate id="9DFE0C7C508324212323125B28CD6F78" degrees="7" direction="north" minutes="41" orientation="latitude" precision="1" seconds="23.6" value="7.689889">7°41'23.6&quot;N</geoCoordinate>
, 
<geoCoordinate id="663E635D3F525745367C16EFDAB777E9" degrees="64" direction="west" minutes="1" orientation="longitude" precision="1" seconds="56.0" value="-64.03223">64°1'56.0&quot;W</geoCoordinate>
, 
<quantity id="D2E8F0A55B4C6D0422412BF9FB312D12" metricMagnitude="2" metricUnit="m" metricValue="1.34" unit="m" value="134.0">
<elevation id="92FAFE34D96B73034608DC109C1C6C4D" metricMagnitude="2" metricUnit="m" metricValue="1.34" unit="m" value="134.0">134 m</elevation>
</quantity>
ca. 
<location id="2D129BFFF413312062C7AACC21C05567" LSID="urn:lsid:plazi:treatment:D89DF85F64A751AC96A34B873FC92FF4:2D129BFFF413312062C7AACC21C05567" country="VENEZUELA" latitude="7.689889" longLatPrecision="1" longitude="-64.03223" name="14 km E Rio Aro" stateProvince="Bolivar State">
<quantity id="87F25082B0832A228818A6E2034E67F7" metricMagnitude="4" metricUnit="m" metricValue="1.4" unit="km" value="14.0">14 km</quantity>
E Rio Aro
</location>
; 
<collectingDate id="6C4A453BED2C7092D08A9A27B6BC3027" value="2008-08-05">5.viii.2008</collectingDate>
leg. 
<collectorName id="C5093A2B807F6963B9F2F1626BD14CC0">A. Short</collectorName>
$ 
<collectorName id="C304F325FA166B0ABA34508584EC6056">
M. 
<normalizedToken id="A13C72B2846DBE4190A9A7EE6F8B7F60" originalValue="García">Garcia</normalizedToken>
</collectorName>
AS-08-073; rock outcropping/ SM0829328 KUNMH-ENT [barcode label],&quot;
</materialsCitation>
<materialsCitation id="3B8093D9F590492A531A293E79E4A479" collectingDate="2009-01-10" collectorName="Short, Miller, Joly, Garcia, Camacho" country="VENEZUELA" elevation="52" latitude="8.103766" location="Rio Aguaro" longLatPrecision="1" longitude="-66.43713" municipality="San Nicolasito Field Station" specimenCount="1" specimenCount-female="1" stateProvince="Guarico State" typeStatus="paratype">
<specimenCount id="C0AC8C718E6D80CBC2105C010D082FC1" count="1" type="female">1 female</specimenCount>
labeled &quot; 
<collectingCountry id="1C28CA8B72CD1824D3937F793099A05D" name="Venezuela">VENEZUELA</collectingCountry>
: 
<collectingRegion id="74927FCAA447C1BDBE47E78B01EED83D" country="Venezuela" name="Guarico">
<normalizedToken id="CD101A1589449428ABDB1B47EC94651B" originalValue="Guárico">Guarico</normalizedToken>
State
</collectingRegion>
<geoCoordinate id="90EF1C60A4DA8AA32DB530FF5F6D6F73" degrees="8" direction="north" minutes="6.226" orientation="latitude" precision="1" value="8.103766">8°6.226'N</geoCoordinate>
, 
<geoCoordinate id="46248839BA4136B7D0CC224288CC6E44" degrees="66" direction="west" minutes="26.228" orientation="longitude" precision="1" value="-66.43713">66°26.228'W</geoCoordinate>
, 
<quantity id="DFACA04948049B2597F1AE5D2BA77DA0" metricMagnitude="1" metricUnit="m" metricValue="5.2" unit="m" value="52.0">
<elevation id="C9DE3211E713CF6BF941D931A4D43E32" metricMagnitude="1" metricUnit="m" metricValue="5.2" unit="m" value="52.0">52 m</elevation>
</quantity>
UCV 
<collectingMunicipality id="0CAFD6AA424F5722273E7991453919B4">San Nicolasito Field Station</collectingMunicipality>
: 
<location id="ED1AE7E808B71E693120B4D838865B8F" LSID="urn:lsid:plazi:treatment:D89DF85F64A751AC96A34B873FC92FF4:ED1AE7E808B71E693120B4D838865B8F" country="VENEZUELA" latitude="8.103766" longLatPrecision="1" longitude="-66.43713" municipality="San Nicolasito Field Station" name="Rio Aguaro" stateProvince="Guarico State">Rio Aguaro</location>
; 
<collectingDate id="7BEE867A1F4465B095EA82169EE8F992" value="2009-01-10">10.i.2009</collectingDate>
leg. 
<collectorName id="BDAFEEBF3BEE600159551D54BA8E10F7">Short</collectorName>
, 
<collectorName id="EA90ECA95769D2999026D884A7635942">Miller</collectorName>
, 
<collectorName id="8EC1CCFAE4BF9426812EB099CF94B314">Joly</collectorName>
, 
<normalizedToken id="454645C0E0736A1B604B09EBF802BE6C" originalValue="García">
<collectorName id="73BC11A2E8E6905021B58AB72448D6A8">Garcia</collectorName>
</normalizedToken>
, 
<collectorName id="7F2EF366FF461C0BD5196069A002FABF">Camacho</collectorName>
; VZ09-0110-01A/ SEMC0852602 KUNHM-ENT,&quot;
</materialsCitation>
<materialsCitation id="F5EE2AAB726E1DAE39E600154CBDCB1C" collectingDate="2009-01-12" collectorName="Short Miller VZ, All" country="VENEZUELA" latitude="6.58694" location="Rio Caripito" longLatPrecision="1" longitude="-67.02912" specimenCount="1" specimenCount-male="1" stateProvince="Bolivar State" typeStatus="paratypes">
<specimenCount id="5359E9D33BEBB1C8BA38E3993AFBD569" count="1" type="male">1 male</specimenCount>
labeled &quot; 
<collectingCountry id="59DEF7FF8D876E685DB7EF44C44B6C89" name="Venezuela">VENEZUELA</collectingCountry>
: 
<collectingRegion id="AC59614D6759BC074FCEADFDFFB4FFC8" country="Venezuela" name="Bolivar">Bolivar State</collectingRegion>
<geoCoordinate id="E45C37826FC5D61F0EC8AF0A92164D6D" degrees="6.58694" direction="north" orientation="latitude" precision="1" value="6.58694">6.58694°N</geoCoordinate>
; 
<geoCoordinate id="99AE54F636788BB97984D0F108CCD4DF" degrees="67.02912" direction="west" orientation="longitude" precision="1" value="-67.02912">67.02912°W</geoCoordinate>
<location id="51BBD622E2602A104A7C63EE8EEB9E63" LSID="urn:lsid:plazi:treatment:D89DF85F64A751AC96A34B873FC92FF4:51BBD622E2602A104A7C63EE8EEB9E63" country="VENEZUELA" latitude="6.58694" longLatPrecision="1" longitude="-67.02912" name="Rio Caripito" stateProvince="Bolivar State">Rio Caripito</location>
<collectingDate id="684EC9C820DEB5EA7F51362EF6AD352D" value="2009-01-12">12.i.2009</collectingDate>
; leg. 
<collectorName id="4D052C458602519CCB8E271FE44A8BFD">Short Miller VZ</collectorName>
09-0112-02A: river margin/ SM0844405 KUNHM-ENT [barcode label].&quot; 
<collectorName id="167385BFEBCB9A3855A9906ED19101E2">All</collectorName>
<typeStatus id="274B7CDC202678FD3DDA25CE022303A6" type="paratype">paratypes</typeStatus>
with, 
<normalizedToken id="5E88148E98CA57B30F1751C050C0EC83" originalValue="“…">&quot;...</normalizedToken>
<typeStatus id="AFA248059DD06765D3EBAD51F4B1F1A3" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="8F9F7032A491A4C6FB2F804318AC9F22" authorityName="Miller &amp; Michat &amp; Ferreira Jr" authorityYear="2024" family="Dytiscidae" genus="Nilssondytes" kingdom="Animalia" lsidName="Nilssondytes diversus" order="Coleoptera" pageId="0" pageNumber="125" rank="species" species="diversus">
<emphasis id="3238A63B2D4AF9BA711965E801DC09C2" italics="true" pageId="0" pageNumber="125">Nilssondytes diversus</emphasis>
</taxonomicName>
Miller, Michat and Ferreira-Jr., 2023 [blue label with black line border].&quot;
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>