treatments-xml/data/7A/32/76/7A327654505B9F46F5E3055CB572FDC7.xml
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<document id="42F923D3B22E1F552F54EE0EA7043F8F" ID-DOI="10.11646/zootaxa.3931.4.3" ID-GBIF-Dataset="d954dbb2-b586-4b02-984c-0bb8f3709ab0" ID-ISSN="1175-5326" ID-Zenodo-Dep="234953" ID-ZooBank="918B6F76-B5CA-4FC6-9F32-F499DDE7C420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461011379296" checkinUser="plazi" docAuthor="Deschodt, Christian M., Davis, Adrian L. V. &amp; Scholtz, Clarke H." docDate="2015" docId="7A327654505B9F46F5E3055CB572FDC7" docLanguage="en" docName="zt03931p527.pdf" docOrigin="Zootaxa 3931 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scarabaeus (Scarabaeolus) planipennis Davis and Deschodt" docType="treatment" docVersion="7" lastPageNumber="518" masterDocId="860B0E2C50509F4BF5740364B06DFFB4" masterDocTitle="Status changes, new synonymies, key and descriptions of seven new species in the subgenus Scarabaeus (Scarabaeolus) Balthasar 1965 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="527" masterPageNumber="505" pageNumber="516" updateTime="1698567509984" updateUser="plazi">
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<mods:title id="51F55420B1F04A8EF3728983526C341F">Status changes, new synonymies, key and descriptions of seven new species in the subgenus Scarabaeus (Scarabaeolus) Balthasar 1965 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="321CFE374ABD184093D972D8CD80DEE0">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="D9C13FDFFA35CFE55042D6497B5D0133">Davis, Adrian L. V.</mods:namePart>
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<mods:namePart id="91069017F917CD5E52030A33E64E8F74">Scholtz, Clarke H.</mods:namePart>
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<mods:date id="2AA7D7EB5AF93CE715BE49B8AFE945CE">2015</mods:date>
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<mods:number id="229A208A647F7B937898316CF5D464C4">3931</mods:number>
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<treatment id="7A327654505B9F46F5E3055CB572FDC7" ID-DOI="http://doi.org/10.5281/zenodo.6108506" ID-GBIF-Taxon="119600192" ID-Zenodo-Dep="6108506" LSID="urn:lsid:plazi:treatment:7A327654505B9F46F5E3055CB572FDC7" httpUri="http://treatment.plazi.org/id/7A327654505B9F46F5E3055CB572FDC7" lastPageId="13" lastPageNumber="518" pageId="11" pageNumber="516">
<subSubSection id="BA8194C9505B9F40F5E3055CB140F9C0" pageId="11" pageNumber="516" type="nomenclature">
<paragraph id="F224C742505B9F40F5E3055CB319F9E6" blockId="11.[151,884,1592,1652]" box="[151,884,1592,1618]" pageId="11" pageNumber="516">
<heading id="A96C702E505B9F40F5E3055CB319F9E6" bold="true" box="[151,884,1592,1618]" fontSize="11" level="1" pageId="11" pageNumber="516" reason="1">
<taxonomicName id="359BBCC1505B9F40F5E3055CB319F9E6" ID-CoL="4V2DF" authority="Davis and Deschodt" authorityName="Davis and Deschodt" box="[151,884,1592,1618]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="516" phylum="Arthropoda" rank="species" species="planipennis" subGenus="Scarabaeolus">
<emphasis id="C0EF1B50505B9F40F5E3055CB319F9E6" bold="true" box="[151,884,1592,1618]" pageId="11" pageNumber="516">
<emphasis id="C0EF1B50505B9F40F5E3055CB218F9E6" bold="true" box="[151,629,1592,1618]" italics="true" pageId="11" pageNumber="516">Scarabaeus (Scarabaeolus) planipennis</emphasis>
Davis and Deschodt
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="F224C742505B9F40F5E3053FB140F9C0" blockId="11.[151,884,1592,1652]" box="[151,301,1627,1652]" pageId="11" pageNumber="516">Figure 4, 6, 8</paragraph>
</subSubSection>
<subSubSection id="BA8194C9505B9F40F5E305F8B2C1F949" pageId="11" pageNumber="516" type="diagnosis">
<paragraph id="F224C742505B9F40F5E305F8B2C1F949" blockId="11.[151,1437,1692,2005]" pageId="11" pageNumber="516">
<emphasis id="C0EF1B50505B9F40F5E305F8B17CF901" bold="true" box="[151,273,1692,1717]" pageId="11" pageNumber="516">Diagnosis.</emphasis>
<taxonomicName id="359BBCC1505B9F40F46805F9B289F901" box="[284,740,1693,1717]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="516" phylum="Arthropoda" rank="species" species="planipennis" subGenus="Scarabaeolus">
<emphasis id="C0EF1B50505B9F40F46805F9B289F901" box="[284,740,1693,1717]" italics="true" pageId="11" pageNumber="516">Scarabaeus (Scarabaeolus) planipennis</emphasis>
</taxonomicName>
is quite distinct from other species within its geographical range. It can be separated from
<taxonomicName id="359BBCC1505B9F40F48B05A4B4ECF96D" authority="Boheman, 1860" authorityName="Boheman" authorityYear="1860" box="[511,1153,1728,1753]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="516" phylum="Arthropoda" rank="species" species="flavicornis" subGenus="Scarabaeolus">
<emphasis id="C0EF1B50505B9F40F48B05A4B3D8F96C" box="[511,949,1728,1752]" italics="true" pageId="11" pageNumber="516">Scarabaeus (Scarabaeolus) flavicornis</emphasis>
(
<bibRefCitation id="960ABAB3505B9F40F6B005A4B414F96D" author="Boheman" box="[964,1145,1728,1753]" pageId="11" pageNumber="516" refString="Boheman, C. H. (1860) Coleoptera Caffraria annis 1838 - 1845 a J. A. Wahlberg collecta. Ofversigt Kongliga Svenska Vetenskapsakademiens Forhandlingar, 17 (3), 107 - 120." type="journal article" year="1860">Boheman, 1860</bibRefCitation>
)
</taxonomicName>
(also with flat elytra) by its muted greenish appearance and smaller size.
</paragraph>
</subSubSection>
<subSubSection id="BA8194C9505B9F47F5B3046CB1B9FD1C" lastPageId="12" lastPageNumber="517" pageId="11" pageNumber="516" type="description">
<paragraph id="F224C742505B9F40F5B3046CB236F8DC" blockId="11.[151,1437,1692,2005]" pageId="11" pageNumber="516">
<emphasis id="C0EF1B50505B9F40F5B3046CB134F895" bold="true" box="[199,345,1800,1825]" pageId="11" pageNumber="516">Description.</emphasis>
<typeStatus id="2D2079E0505B9F40F42B046CB1A1F895" box="[351,460,1800,1825]" pageId="11" pageNumber="516" type="holotype">Holotype</typeStatus>
: Body length
<quantity id="35636AA7505B9F40F712046DB2A0F894" box="[614,717,1801,1825]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="11" pageNumber="516" unit="mm" value="12.8">12.8 mm</quantity>
, green. Pubescence is dark brown to black. Antennae yellow. Head: Frons punctate, each puncture with associated short black setae. Wide puncture-free mid-longitudinal area. Clypeus punctate with short black setae.
</paragraph>
<paragraph id="F224C742505B9F40F5B30411B525F861" blockId="11.[151,1437,1692,2005]" pageId="11" pageNumber="516">Pronotum: Dark green and punctate with sparse short black setae. Spaces between punctures smooth comprising one to two puncture diameters. No longitudinal smooth area. Lateral margins curved, jagged anteriorly, with one side of jagged edge almost 10 times longer than the other gradually becoming toothed posteriorly.</paragraph>
<paragraph id="F224C742505C9F47F5B303F3B1F1FF43" blockId="12.[151,1436,151,680]" pageId="12" pageNumber="517">Elytra: Dark green, lateral sides slightly curved slightly wider medially than at base and apex. Striae shallow with clear regular small punctures separated by about one puncture diameter. Interstriae very flat, matte, with very sparse small punctures.</paragraph>
<paragraph id="F224C742505C9F47F5B30260B2E6FEA8" blockId="12.[151,1436,151,680]" box="[199,651,260,285]" pageId="12" pageNumber="517">Pygidium: Shallowly emarginate, matte.</paragraph>
<paragraph id="F224C742505C9F47F5B3024CB542FED0" blockId="12.[151,1436,151,680]" pageId="12" pageNumber="517">
Metasternum: No fovea between the mesocoxae.
<collectingCountry id="8A8C87D2505C9F47F7990243B377FEF4" box="[749,794,295,320]" name="India" pageId="12" pageNumber="517">End</collectingCountry>
of metasternal elevation rounded almost right angled and steeply sloping toward mesosternum. Area behind mesocoxae matte with small widely spaced punctures.
</paragraph>
<paragraph id="F224C742505C9F47F5B3020BB3FBFE3C" blockId="12.[151,1436,151,680]" box="[199,918,367,392]" pageId="12" pageNumber="517">Abdominal sternites: Matte with short, fine black setae laterally.</paragraph>
<paragraph id="F224C742505C9F47F5B302F1B368FD31" blockId="12.[151,1436,151,680]" pageId="12" pageNumber="517">Legs: Anterior femora swollen basally and shallowly punctate ventrally with black setae. Outside margin of anterior tibiae proximally crenulate merging into small saw-like teeth with four outward pointing teeth distally, each separted by nine to ten saw-like denticles. Inside margin finely crenulate, almost straight. Meso- and metafemora slender with irregular median row of punctures and fine black setae. Inside margin of mesotibiae smooth with row of black setae, outside margin with two brushes of black setae in distal third. Vestigial second mesotibial spur absent. Metatibiae curving slightly inward with black setae. All tarsomeres of metatarsi with setae, last tarsomere longest with claws almost half its length.</paragraph>
<paragraph id="F224C742505C9F47F5B301EBB1B9FD1C" blockId="12.[151,1436,151,680]" box="[199,468,655,680]" pageId="12" pageNumber="517">Aedeagus: see Figure 6.</paragraph>
</subSubSection>
<caption id="A6E497CA505C9F47F5E304FEB2EDF864" httpUri="https://zenodo.org/record/234957/files/figure.png" pageId="12" pageNumber="517" targetBox="[170,1435,733,1905]" targetPageId="12">
<paragraph id="F224C742505C9F47F5E304FEB2EDF864" blockId="12.[151,1436,1946,2000]" pageId="12" pageNumber="517">
<emphasis id="C0EF1B50505C9F47F5E304FEB17EF804" bold="true" box="[151,275,1946,1968]" pageId="12" pageNumber="517">FIGURE 7.</emphasis>
Map showing the collecting localities of specimens of (⋅)
<taxonomicName id="359BBCC1505C9F47F61A04FFB457F804" box="[878,1082,1946,1968]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="canaliculatus" subGenus="Scarabaeolus">
<emphasis id="C0EF1B50505C9F47F61A04FFB457F804" box="[878,1082,1946,1968]" italics="true" pageId="12" pageNumber="517">S. (S.) canaliculatus</emphasis>
</taxonomicName>
, (•)
<taxonomicName id="359BBCC1505C9F47F10504FFB490F804" box="[1137,1277,1946,1968]" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="fritschi" subGenus="Scarabaeolus">
<emphasis id="C0EF1B50505C9F47F10504FFB490F804" box="[1137,1277,1946,1968]" italics="true" pageId="12" pageNumber="517">S. (S.) fritschi</emphasis>
</taxonomicName>
and (˔)
<taxonomicName id="359BBCC1505C9F47F02A04FFB0B1F864" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="517" phylum="Arthropoda" rank="species" species="reichei" subGenus="Scarabaeolus">
<emphasis id="C0EF1B50505C9F47F02A04FFB0B1F864" italics="true" pageId="12" pageNumber="517">S. (S.) reichei</emphasis>
</taxonomicName>
in SANC, TMSA and UPSA collections.
</paragraph>
</caption>
<subSubSection id="BA8194C9505D9F46F5B303F3B31CFF04" box="[199,881,151,176]" pageId="13" pageNumber="518" type="etymology">
<paragraph id="F224C742505D9F46F5B303F3B31CFF04" blockId="13.[151,1436,151,627]" box="[199,881,151,176]" pageId="13" pageNumber="518">
<emphasis id="C0EF1B50505D9F46F5B303F3B123FF04" bold="true" box="[199,334,151,176]" pageId="13" pageNumber="518">Etymology.</emphasis>
The species name is derived from the flat elytra.
</paragraph>
</subSubSection>
<subSubSection id="BA8194C9505D9F46F5B303DEB572FDC7" pageId="13" pageNumber="518" type="materials_examined">
<paragraph id="F224C742505D9F46F5B303DEB572FDC7" blockId="13.[151,1436,151,627]" pageId="13" pageNumber="518">
<emphasis id="C0EF1B50505D9F46F5B303DEB1D9FF67" bold="true" box="[199,436,186,211]" pageId="13" pageNumber="518">Material examined.</emphasis>
<typeStatus id="2D2079E0505D9F46F4CB03DEB241FF67" box="[447,556,186,211]" pageId="13" pageNumber="518" type="holotype">Holotype</typeStatus>
: ♂, Maputo Elephant Reserve,
<collectingCountry id="8A8C87D2505D9F46F6D403DEB452FF67" box="[928,1087,186,211]" name="Mozambique" pageId="13" pageNumber="518">Mozambique</collectingCountry>
,[1518]
<date id="8625E182505D9F46F1EA03DEB54EFF67" box="[1182,1315,186,211]" pageId="13" pageNumber="518" value="2006-11-26">
xi.2006,
<geoCoordinate id="97AFA185505D9F46F07103DEB54EFF67" box="[1285,1315,186,211]" direction="south" orientation="latitude" pageId="13" pageNumber="518" precision="1" value="-26.585222">26</geoCoordinate>
</date>
º3506.8”S
<geoCoordinate id="97AFA185505D9F46F5E303B9B15FFF42" box="[151,306,221,246]" direction="east" orientation="longitude" pageId="13" pageNumber="518" precision="1" value="32.779194">32º4645.1”E</geoCoordinate>
, sand, grassland, Escobar and Deschodt,
<collectionCode id="948A5F87505D9F46F60403BAB3ACFF42" box="[880,961,222,246]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="13" pageNumber="518">TMSA</collectionCode>
.
<typeStatus id="2D2079E0505D9F46F6AB03BAB43CFF42" box="[991,1105,222,246]" pageId="13" pageNumber="518" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="E49D0CCB505D9F46F11A03B9B576FF42" box="[1134,1307,221,246]" pageId="13" pageNumber="518" type="generic">15 specimens</specimenCount>
, unsexed,
<collectingCountry id="8A8C87D2505D9F46F5E30264B131FEAC" box="[151,348,256,280]" name="Mozambique" pageId="13" pageNumber="518">MOZAMBIQUE</collectingCountry>
: Maputo Elephant Reserve,
<date id="8625E182505D9F46F7D2039BB327FEAC" box="[678,842,255,280]" pageId="13" pageNumber="518" value="2007-11-10" valueMax="2007-11-16" valueMin="2007-11-10">1016.xi.2007</date>
,
<geoCoordinate id="97AFA185505D9F46F62D039BB3DEFEAC" box="[857,947,255,280]" direction="south" orientation="latitude" pageId="13" pageNumber="518" precision="925" value="-26.5">26º30S</geoCoordinate>
<geoCoordinate id="97AFA185505D9F46F6CA039BB474FEAC" box="[958,1049,255,280]" direction="east" orientation="longitude" pageId="13" pageNumber="518" precision="925" value="32.85">32º51E</geoCoordinate>
Pitfall Trap, W. Strumpher &amp; C. Deschodt, 2
<collectionCode id="948A5F87505D9F46F45A0247B115FE8F" box="[302,376,291,315]" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="13" pageNumber="518">UPSA</collectionCode>
, 5
<collectionCode id="948A5F87505D9F46F4D60247B19EFE8F" box="[418,499,291,315]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="13" pageNumber="518">TMSA</collectionCode>
, 5
<collectionCode id="948A5F87505D9F46F7690247B219FE8E" box="[541,628,291,314]" pageId="13" pageNumber="518">BMNH</collectionCode>
, 3
<collectionCode id="948A5F87505D9F46F7EA0247B29DFE8F" box="[670,752,291,315]" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="13" pageNumber="518">SAMC</collectionCode>
,
<specimenCount id="E49D0CCB505D9F46F6750247B3E8FE8F" box="[769,901,290,315]" pageId="13" pageNumber="518" type="generic">1 specimen</specimenCount>
<collectingCountry id="8A8C87D2505D9F46F6E50247B40EFE8F" box="[913,1123,291,315]" name="South Africa" pageId="13" pageNumber="518">SOUTH AFRICA</collectingCountry>
, KZN Pr. Tembe,
<geoCoordinate id="97AFA185505D9F46F0370246B5F1FE8F" box="[1347,1436,290,315]" direction="south" orientation="latitude" pageId="13" pageNumber="518" precision="925" value="-26.95">26º57S</geoCoordinate>
<geoCoordinate id="97AFA185505D9F46F5E30222B098FEEA" box="[151,245,326,350]" direction="east" orientation="longitude" pageId="13" pageNumber="518" precision="925" value="32.4">32º24E</geoCoordinate>
,
<date id="8625E182505D9F46F4710221B11DFEEA" box="[261,368,325,350]" pageId="13" pageNumber="518" value="2004-04-07">7.iv.2004</date>
, J. v.d. Merwe,
<collectionCode id="948A5F87505D9F46F7430222B2ECFEEA" box="[567,641,326,350]" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="13" pageNumber="518">UPSA</collectionCode>
,
<specimenCount id="E49D0CCB505D9F46F7E60222B34CFEEA" box="[658,801,325,350]" pageId="13" pageNumber="518" type="generic">9 specimens</specimenCount>
Thembe Elephant Park, KZN,
<collectingCountry id="8A8C87D2505D9F46F1E30221B55FFEEA" box="[1175,1330,325,350]" name="South Africa" pageId="13" pageNumber="518">South Africa</collectingCountry>
,
<geoCoordinate id="97AFA185505D9F46F0370222B5F1FEEA" box="[1347,1436,326,350]" direction="south" orientation="latitude" pageId="13" pageNumber="518" precision="925" value="-27.016666">27º01S</geoCoordinate>
32º24E,
<date id="8625E182505D9F46F58A0203B1F9FE34" box="[254,404,359,384]" pageId="13" pageNumber="518" value="1996-01-24" valueMax="1996-01-29" valueMin="1996-01-24">2429.i.1996</date>
, B.J.Van Rensburg, 2
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, 7
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,
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<collectingCountry id="8A8C87D2505D9F46F1630203B4D9FE34" box="[1047,1204,359,384]" name="South Africa" pageId="13" pageNumber="518">South Africa</collectingCountry>
: Kwazulu, Thembe Elephant Park, SE 2632 CD Ndumu,
<date id="8625E182505D9F46F73902EFB2DFFE17" box="[589,690,395,419]" pageId="13" pageNumber="518" value="1990-10-07">7.x.1990</date>
, T. Beyers,
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,
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<collectingCountry id="8A8C87D2505D9F46F15102EFB495FE17" box="[1061,1272,395,419]" name="South Africa" pageId="13" pageNumber="518">SOUTH AFRICA</collectingCountry>
: KZN, Tembe Elephant Park, Sihangwane Area, 27[º].02[]S 32[º].25[]E
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,
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R.Stals,
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,
<specimenCount id="E49D0CCB505D9F46F06502CAB5F5FE72" box="[1297,1432,429,454]" pageId="13" pageNumber="518" type="generic">1 specimen</specimenCount>
, 2705 [Muzi, N.
<collectingCountry id="8A8C87D2505D9F46F43802ABB1E6FE5C" box="[332,395,463,488]" name="South Africa" pageId="13" pageNumber="518">Natal</collectingCountry>
, (26º52
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),
<date id="8625E182505D9F46F71102ABB2D0FE5C" box="[613,701,463,488]" pageId="13" pageNumber="518" value="1980-10" valueMax="1980-11-31" valueMin="1980-10-00">xxi.80</date>
, G. Vale.],
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,
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,
<collectionCode id="948A5F87505D9F46F16B02B4B404FE53" box="[1055,1129,464,487]" pageId="13" pageNumber="518">MUZI</collectionCode>
, TONGALAND,
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(
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W), (26º52
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),
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, H.H. Aschenborn,
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,
<specimenCount id="E49D0CCB505D9F46F6210297B3B7FDBF" box="[853,986,498,523]" pageId="13" pageNumber="518" type="generic">1 specimen</specimenCount>
, S. Afr: KrugerNat. PK, Punda Milia, sand, 22[º].38[] S31[º].02[] E,
<date id="8625E182505D9F46F76A0172B2E9FD9A" box="[542,644,534,558]" pageId="13" pageNumber="518" value="1994-02-03">3.2.1994</date>
; E-Y: 2974, groundtraps, 8 days, leg. Endrödy-Younga, groundtraps with faeces bait,
<collectionCode id="948A5F87505D9F46F427015CB1C9FDE4" box="[339,420,568,592]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="13" pageNumber="518">TMSA</collectionCode>
,
<specimenCount id="E49D0CCB505D9F46F4DA015CB254FDE4" box="[430,569,567,592]" pageId="13" pageNumber="518" type="generic">8 specimens</specimenCount>
, S. Afr: KrugerNat. PK,Nyandu sands, 22[º].41[] S31[º].22[] E,
<date id="8625E182505D9F46F046015CB5F5FDE4" box="[1330,1432,568,592]" pageId="13" pageNumber="518" value="1994-02-09">9.2.1994</date>
; E-Y: 2989, groundtraps, 4 days, leg. Endrödy-Younga, groundtraps with faeces bait, 6
<collectionCode id="948A5F87505D9F46F12B013FB4C2FDC7" box="[1119,1199,603,627]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="13" pageNumber="518">TMSA</collectionCode>
, 2
<collectionCode id="948A5F87505D9F46F1A4013FB576FDC7" box="[1232,1307,603,627]" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="13" pageNumber="518">UPSA</collectionCode>
.
</paragraph>
</subSubSection>
</treatment>
</document>