239 lines
20 KiB
XML
239 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.228.3401" ID-GBIF-Dataset="14392d6c-4f18-4c00-863b-4d0c709409a5" ID-GBIF-Taxon="152037318" ID-Pensoft-Pub="1313-2970-228-77" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-228-77" ModsDocOrigin="ZooKeys 228" ModsDocTitle="Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines" checkinTime="1451248671826" checkinUser="pensoft" docAuthor="Zhao, Danyang & Tian, Mingyi" docDate="2012" docId="5A9D0E601DCC69CDF67C63886E0DCC13" docLanguage="en" docName="ZooKeys 228: 77-100" docOrigin="ZooKeys 228" docSource="http://dx.doi.org/10.3897/zookeys.228.3401" docTitle="Macrocheilus Hope 1838" docType="treatment" docVersion="2" lastPageId="2" lastPageNumber="78" masterDocId="FFD6FFE3FF8DFFD04741FF923F7CFFAC" masterDocTitle="Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines" masterLastPageNumber="100" masterPageNumber="77" pageId="0" pageNumber="77" updateTime="1643546355349" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic notes on the Macrocheilus Hope (Coleoptera, Carabidae, Helluonini) from Oriental Region, with description of one new species from the Philippines</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhao, Danyang</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tian, Mingyi</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>228</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>77</mods:start>
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<mods:end>100</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.228.3401</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.228.3401</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-228-77</mods:identifier>
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<mods:identifier type="ZooBank">76973A68-EEE6-4C43-ACE8-1573BB6964C3</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152037318" LSID="urn:lsid:plazi:treatment:5A9D0E601DCC69CDF67C63886E0DCC13" httpUri="http://treatment.plazi.org/id/5A9D0E601DCC69CDF67C63886E0DCC13" lastPageId="2" lastPageNumber="78" pageId="0" pageNumber="77">
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<subSubSection pageId="0" pageNumber="77" type="nomenclature">
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<paragraph pageId="0" pageNumber="77">
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Genus
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<taxonomicName LSID="http://species-id.net/wiki/Macrocheilus" authority="Hope, 1838" authorityName="Hope" authorityYear="1838" class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus" order="Coleoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Macrocheilus Hope, 1838</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="77" type="reference_group">
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<paragraph pageId="0" pageNumber="77">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus" order="Coleoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Macrocheilus</taxonomicName>
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<bibRefCitation pageId="0" pageNumber="77">Hope 1838</bibRefCitation>
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: 166. Type species:
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus bensoni" order="Coleoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="bensoni">Macrocheilus bensoni</taxonomicName>
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Hope, 1838.
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<bibRefCitation pageId="0" pageNumber="77">Jeannel 1949</bibRefCitation>
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: 1041.
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<bibRefCitation author="Zhao, DY" journalOrPublisher="Polish Journal of Entomology," pageId="14" pageNumber="91" pagination="3 - 23" title="A review of the genus Macrocheilus Hope (Coleoptera: Carabidae: Helluonini) of China, with descriptions of seven new species." volume="29" year="2010">Zhao and Tian 2010</bibRefCitation>
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:4.
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</paragraph>
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<paragraph pageId="0" pageNumber="77">
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<taxonomicName authority="Reiche, 1843" authorityName="Reiche" authorityYear="1843" class="Insecta" family="Carabidae" genus="Acanthogenius" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acanthogenius" order="Coleoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="genus">Acanthogenius</taxonomicName>
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<bibRefCitation pageId="0" pageNumber="77">Reiche 1842</bibRefCitation>
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: 334. Type species:
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<taxonomicName class="Insecta" family="Carabidae" genus="Helluo" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helluo impictus" order="Coleoptera" pageId="0" pageNumber="77" phylum="Arthropoda" rank="species" species="impictus">Helluo impictus</taxonomicName>
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Wiedemann, 1823.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="78" type="generic diagnosis">
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<paragraph pageId="1" pageNumber="78">
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<pageBreakToken pageId="1" pageNumber="78" start="start">Generic</pageBreakToken>
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diagnosis.
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</paragraph>
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<paragraph pageId="1" pageNumber="78">Medium sized (length from 8.0 to 17.0 mm), elongate, whole body coarsely punctuate and pubescent, except for labrum and middle region of ventral side of head. Head with two supraorbital setae on each side; ligula fairly wide, deeply impressed beneath at sides of base; mentum deeply emarginated, with a long and slender tooth; palpi varied in form; labrum extraordinarily or well developed; mandibles dorsally covered by labrum or not covered; antennae stout and flat, densely pubescent from antennomere 5. Pronotum truncate-cordiform or quadrate, basal margin more or less produced backwards in middle; lateral margin with two setae, one just before middle, and the other at hind angle. Elytra with deep striae and setiferous pores; intervals slightly or rather convex, interval 8 usually wide. Metepisterna elongate and very narrow in all species. Tarsomere 4 emarginate. Wings fully developed. No externally visible sexual dimorphic characters present. Median lobe of aedeagus varied in form. Parameres of aedeagus quite similar, elongate, the left one larger than the right one.</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="78" type="differences with other genera of helluonini in the oriental region">
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<paragraph pageId="1" pageNumber="78">
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Differences with other genera of
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<taxonomicName lsidName="" pageId="1" pageNumber="78" rank="tribe" tribe="Helluonini">Helluonini</taxonomicName>
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in the Oriental Region.
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</paragraph>
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<paragraph pageId="1" pageNumber="78">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Macrocheilus</taxonomicName>
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spp., with larger body size, simple 4th tarsomeres, and non-securiform labial palpomere are distinctly different from
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<taxonomicName class="Insecta" family="Carabidae" genus="Colfax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Colfax" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Colfax</taxonomicName>
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spp. (smaller body size, securiform labial palpomere) and
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<taxonomicName class="Insecta" family="Carabidae" genus="Creagris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Creagris" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Creagris</taxonomicName>
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spp. (bilobed 4th tarsomere; smaller body size). Functional wings and the occurrence of a long spiniform median tooth of mentum of
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Macrocheilus</taxonomicName>
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distinguish them from
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<taxonomicName class="Insecta" family="Carabidae" genus="Omphra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omphra" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Omphra</taxonomicName>
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spp. (which are brachypterous and have a short median tooth of the mentum).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="78" type="generic range">
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<paragraph pageId="1" pageNumber="78">Generic range.</paragraph>
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<paragraph pageId="1" pageNumber="78">Oriental Region (China, Vietnam, Laos, Cambodia, Myanmar, India, Sri Lanka, the Philippines, Malaysia and Indonesia), Palaearctic Region (Pakistan, Syria, Asia Minor), Ethiopian Region (Ethiopia, Tanzania, Uganda, Senegal, Guinea, Guinea-Bissau, Sierra Leone, Togo, Nigeria, Central Africa, Cameroon, Gabon, Congo, Zimbabwe, Angola, South Africa, Madagascar).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="79" pageId="1" pageNumber="78" type="key to species of oriental macrocheilus">
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<paragraph pageId="1" pageNumber="78">
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Key to species of Oriental
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="genus">Macrocheilus</taxonomicName>
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</paragraph>
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<paragraph lastPageId="2" lastPageNumber="79" pageId="1" pageNumber="78">
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<table lastPageId="2" lastPageNumber="79" pageId="1" pageNumber="78">
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 9Fig. 1</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus bicolor" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="bicolor">Macrocheilus bicolor</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 10Fig. 2</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus impictus" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="impictus">Macrocheilus impictus</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 3Fig. 11</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus immanis" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="immanis">Macrocheilus immanis</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 46Fig. 12</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus niger" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="niger">Macrocheilus niger</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 47</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus asteriscus" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="asteriscus">Macrocheilus asteriscus</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus vitalisi" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="vitalisi">Macrocheilus vitalisi</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 21</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 5</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus binotatus" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="binotatus">Macrocheilus binotatus</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus macromaculatus" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="macromaculatus">Macrocheilus macromaculatus</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus gigas" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="gigas">Macrocheilus gigas</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 51</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus parvimaculatus" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="parvimaculatus">Macrocheilus parvimaculatus</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 52Fig. 6Fig. 22</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus tripustulatus" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="tripustulatus">Macrocheilus tripustulatus</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 53</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus chaudoiri" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="chaudoiri">Macrocheilus chaudoiri</taxonomicName>
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</td>
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</tr>
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<tr pageId="1" pageNumber="78">
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">Fig. 7</td>
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<td colspan="1" pageId="1" pageNumber="78" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus nigrotibialis" order="Coleoptera" pageId="1" pageNumber="78" phylum="Arthropoda" rank="species" species="nigrotibialis">Macrocheilus nigrotibialis</taxonomicName>
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</td>
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</tr>
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<tr pageId="2" pageNumber="79">
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus bensoni" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="bensoni">
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<pageBreakToken pageId="2" pageNumber="79" start="start">Macrocheilus</pageBreakToken>
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bensoni
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</taxonomicName>
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</td>
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</tr>
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<tr pageId="2" pageNumber="79">
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">Fig. 24Fig. 8</td>
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus deuvie" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="deuvie">Macrocheilus deuvie</taxonomicName>
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</td>
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</tr>
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<tr pageId="2" pageNumber="79">
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus fuscipennis" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="fuscipennis">Macrocheilus fuscipennis</taxonomicName>
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</td>
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</tr>
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<tr pageId="2" pageNumber="79">
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<bibRefCitation author="Zhao, DY" journalOrPublisher="Polish Journal of Entomology," pageId="14" pageNumber="91" pagination="3 - 23" title="A review of the genus Macrocheilus Hope (Coleoptera: Carabidae: Helluonini) of China, with descriptions of seven new species." volume="29" year="2010">Zhao and Tian 2010</bibRefCitation>
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</td>
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus solidipalpis" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="solidipalpis">Macrocheilus solidipalpis</taxonomicName>
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</td>
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</tr>
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<tr pageId="2" pageNumber="79">
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<bibRefCitation author="Zhao, DY" journalOrPublisher="Polish Journal of Entomology," pageId="14" pageNumber="91" pagination="3 - 23" title="A review of the genus Macrocheilus Hope (Coleoptera: Carabidae: Helluonini) of China, with descriptions of seven new species." volume="29" year="2010">Zhao and Tian 2010</bibRefCitation>
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</td>
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus cheni" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="cheni">Macrocheilus cheni</taxonomicName>
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</td>
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</tr>
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<tr pageId="2" pageNumber="79">
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<bibRefCitation author="Zhao, DY" journalOrPublisher="Polish Journal of Entomology," pageId="14" pageNumber="91" pagination="3 - 23" title="A review of the genus Macrocheilus Hope (Coleoptera: Carabidae: Helluonini) of China, with descriptions of seven new species." volume="29" year="2010">Zhao and Tian 2010</bibRefCitation>
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</td>
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus quadratus" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="quadratus">Macrocheilus quadratus</taxonomicName>
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</td>
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</tr>
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<tr pageId="2" pageNumber="79">
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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<bibRefCitation author="Zhao, DY" journalOrPublisher="Polish Journal of Entomology," pageId="14" pageNumber="91" pagination="3 - 23" title="A review of the genus Macrocheilus Hope (Coleoptera: Carabidae: Helluonini) of China, with descriptions of seven new species." volume="29" year="2010">Zhao and Tian 2010</bibRefCitation>
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</td>
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<td colspan="1" pageId="2" pageNumber="79" rowspan="1">
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|
<taxonomicName class="Insecta" family="Carabidae" genus="Macrocheilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Macrocheilus sinuatilabris" order="Coleoptera" pageId="2" pageNumber="79" phylum="Arthropoda" rank="species" species="sinuatilabris">Macrocheilus sinuatilabris</taxonomicName>
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</td>
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</tr>
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|
</table>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |