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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1171.104142" ID-Pensoft-Pub="1313-2970-1171-1" ID-Pensoft-UUID="7559C2D42DE85144AEFFC98A16172F97" ID-ZooBank="267D0D4559CA4A18A08034768E652607" ModsDocID="1313-2970-1171-1" checkinTime="1689969829564" checkinUser="pensoft" docAuthor="Short, Andrew Edward Z. &amp; Giron, Jennifer C." docDate="2023" docId="59C78BC226FB55249D864CFFEF6979A5" docLanguage="en" docName="ZooKeys 1171: 1-112" docOrigin="ZooKeys 1171" docPubDate="2023-07-20" docSource="http://dx.doi.org/10.3897/zookeys.1171.104142" docTitle="Novochares pallipes" docType="treatment" docVersion="1" id="7559C2D42DE85144AEFFC98A16172F97" lastPageNumber="1" masterDocId="7559C2D42DE85144AEFFC98A16172F97" masterDocTitle="Revision of the Neotropical water scavenger beetle genus Novochares Giron &amp; Short (Coleoptera, Hydrophilidae, Acidocerinae)" masterLastPageNumber="112" masterPageNumber="1" pageNumber="1" updateTime="1689969829564" updateUser="pensoft">
<mods:mods id="BF8EDA5017228A2F16B8A2E4EA3B8B95" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ADF49D9EDCB444763D649420BF1EA6CA">
<mods:title id="A87A62F84D467E0442E21F987DCD5CE2">Revision of the Neotropical water scavenger beetle genus Novochares Giron &amp; Short (Coleoptera, Hydrophilidae, Acidocerinae)</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="B7A771A578E6FFFB9DEC5DDD4F364B76">Short, Andrew Edward Z.</mods:namePart>
<mods:nameIdentifier id="D8F5EABEA02BCD2AA584C72184A3A870" type="ORCID">https://orcid.org/0000-0002-7467-7116</mods:nameIdentifier>
<mods:affiliation id="74DCD4B1EBEE3D0984440A10695CF1D2">Department of Entomology &amp; Nematology, University of Florida, Gainesville, FL, 32611, USA</mods:affiliation>
<mods:nameIdentifier id="4D33769FD4574DD620C8BAFE2A2FF21A" type="email">aezshort@ku.edu</mods:nameIdentifier>
</mods:name>
<mods:name id="DB583041B371EF6BD7A767DB59131E5C" type="personal">
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<mods:roleTerm id="336DD39A33BD43A726B5EF0009584968">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="06A990B7900C33D5273CD711DE9C0625">Giron, Jennifer C.</mods:namePart>
<mods:nameIdentifier id="BF00825425DE22C7FBE332FDEE811ECB" type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
<mods:affiliation id="8F9A47B0214E061EF5F4F56C6DBD15BA">Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, TX 79409, USA</mods:affiliation>
</mods:name>
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<mods:title id="7978C37C4806B0547582280C5775D7B6">ZooKeys</mods:title>
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<mods:date id="3C8045227CFE68227E1915D5F5ECE338">2023</mods:date>
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<mods:number id="33C71C2C40A7F7671C9A0FDAD153AB62">2023-07-20</mods:number>
</mods:detail>
<mods:detail id="FF565D0BE8BF998D937961ED34A038C7" type="volume">
<mods:number id="18422C5F2D77AA7823BD4B032822C6F5">1171</mods:number>
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</mods:location>
<mods:classification id="764C366D05C6A5B017B385AC3E1628AF">journal article</mods:classification>
<mods:identifier id="DEC3EEEDB905272735F7DC561025F075" type="DOI">http://dx.doi.org/10.3897/zookeys.1171.104142</mods:identifier>
<mods:identifier id="500B7A052E8C7D4915A953267ED1A308" type="Pensoft-Pub">1313-2970-1171-1</mods:identifier>
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<treatment id="59C78BC226FB55249D864CFFEF6979A5" LSID="urn:lsid:plazi:treatment:59C78BC226FB55249D864CFFEF6979A5" httpUri="http://treatment.plazi.org/id/59C78BC226FB55249D864CFFEF6979A5" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="56993BB15ACF44418388FFCA14BCFAED" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="B47A6DACC6AAAEC19612170A76E37D7E" pageId="0" pageNumber="1">
<taxonomicName id="7B778190ED6F2EC6FE9A33A9886B5A82" ID-CoL="8Y6QX" LSID="59C78BC2-26FB-5524-9D86-4CFFEF6979A5" authority="(Brulle, 1841)" baseAuthorityName="Brulle" baseAuthorityYear="1841" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares pallipes" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pallipes">
Novochares pallipes (
<normalizedToken id="36FABC84EF040CEBD3CC7D0F7BDA94CD" originalValue="Brullé">Brulle</normalizedToken>
, 1841)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="47EC2E4F6B022C50B09E8E891E4F00AE" pageId="0" pageNumber="1" type="description">
<paragraph id="E58CBA0A58FD4021D8097FBDCCE1CEAA" pageId="0" pageNumber="1">
<figureCitation id="6F6EC45B994DF407E4F68F33CFF5AF2C" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Aedeagi and habitus of the Novochares abbreviatus species group A-E N. oculatus: A-C aedeagus, Mexico D, E paralectotype from Guatemala D aedeagus E habitus F-K miscellaneous forms of aedeagi in the Novochares abbreviatus species group F, G Argentina H, I Venezuela J, K Bolivia L, M, R N. pallipes N, O N. baca A, D, F, H, J, L, N dorsal view B, M ventral view C, G, I, K, O lateral view R oblique view." figureDoi="10.3897/zookeys.1171.104142.figure9" httpUri="https://binary.pensoft.net/fig/881176" pageId="0" pageNumber="1">Figs 9L-R</figureCitation>
<figureCitation id="707C72FE3322D24841BF6FEF4F6046BB" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution of Novochares abbreviatus species group A N. abbreviatus: examined specimens (red) and literature / unconfirmed records (blue) B N. oculatus: examined specimens (red) and literature / unconfirmed records (blue) C N. pallipes (red), N. latus (yellow), N. baca (blue), N. pilatus (green)." figureDoi="10.3897/zookeys.1171.104142.figure10" httpUri="https://binary.pensoft.net/fig/881177" pageId="0" pageNumber="1">, 10C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="827B0F104DFFB85E58E2475A11015DA8" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="DE85D8D1208495A6A39EAAD53A9297AA" pageId="0" pageNumber="1">
<taxonomicName id="06E99888E091ECFBB3163DFE7E86074B" authorityName="E.Blanchard" authorityYear="1841" class="Insecta" family="Hydrophilidae" genus="Hydrophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydrophilus (Philhydrus) pallipes" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pallipes" subGenus="Philhydrus">Hydrophilus (Philhydrus) pallipes</taxonomicName>
<normalizedToken id="1D86F6884BFD43142CCC51CF3EAB6B92" originalValue="Brullé">Brulle</normalizedToken>
, 1841: 58 - Uruguay, Montevideo.
</paragraph>
<paragraph id="4323BDB73B8947E961CBA62C581890DD" pageId="0" pageNumber="1">
<taxonomicName id="385705F3FB064BAC3453A11812C86225" class="Insecta" family="Hydrophilidae" genus="Philhydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philhydrus pallipes" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pallipes">Philhydrus pallipes</taxonomicName>
(
<normalizedToken id="14DE048A5EC4929956BA0658E03C00B7" originalValue="Brullé">Brulle</normalizedToken>
, 1841);
<bibRefCitation id="AAFE14044503B50C49916FDD6000D239" DOI="https://doi.org/10.5962/bhl.title.67686" author="Lacordaire, T" journalOrPublisher="Librairie encyclopedique de Roret, Paris" pageId="0" pageNumber="1" refId="B29" refString="Lacordaire, T, 1854. Histoire naturelle des insectes: Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome Premier. Contenant les familles des Cicindeletes, Carabiques, Dytiscides, Gyrinides et Palpicornes. Librairie encyclopedique de Roret, Paris, DOI: https://doi.org/10.5962/bhl.title.67686" title="Histoire naturelle des insectes: Genera des Coleopteres ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome Premier. Contenant les familles des Cicindeletes, Carabiques, Dytiscides, Gyrinides et Palpicornes." url="https://doi.org/10.5962/bhl.title.67686" year="1854">Lacordaire 1854</bibRefCitation>
: 457.
</paragraph>
<paragraph id="F5FC4B8D0914E635E8AB280DFB8CA6AA" pageId="0" pageNumber="1">
<taxonomicName id="035A80B9E7914A06CD329E5FD2E8D8A1" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helochares pallipes" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pallipes">Helochares pallipes</taxonomicName>
(
<normalizedToken id="D014F5DAC57876571835002720DF16F9" originalValue="Brullé">Brulle</normalizedToken>
, 1841);
<bibRefCitation id="CC3BA61355E8FD8C15E07991224A822B" author="Bedel, L" journalOrPublisher="Annals &amp; Magazine of Natural History" pageId="0" pageNumber="1" publicationUrl="https://biodiversitylibrary.org/page/8996339" refId="B2" refString="Bedel, L, 1881. (Synonymie de quelques Hydrophilidae et Sphaeridiidae exotiques decrits par Brulle). Annales de la Societe Entomologique de France 6(1): XCIV-XCV. https://biodiversitylibrary.org/page/8996339" title="(Synonymie de quelques Hydrophilidae et Sphaeridiidae exotiques decrits par Brulle). Annales de la Societe Entomologique de France 6 (1): XCIV-XCV." url="https://biodiversitylibrary.org/page/8996339" year="1881">Bedel 1881</bibRefCitation>
: XCIV.
</paragraph>
<paragraph id="4AF5861CAFE48F4E3727AC6CDA5B9DFB" pageId="0" pageNumber="1">
<taxonomicName id="4F4021A8A3FE4C9FDA772C93E50A2EFB" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helochares pallipes" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pallipes">Helochares (s. str.) pallipes</taxonomicName>
(
<normalizedToken id="97A908CBF39D4AFE31146DD3FFBE0F93" originalValue="Brullé">Brulle</normalizedToken>
, 1841);
<bibRefCitation id="1F4F48A2469020E8027BF5474A052B9D" author="Fernandez, LA" journalOrPublisher="Limnobios" pageId="0" pageNumber="1" pagination="439 - 449" refId="B17" refString="Fernandez, LA, 1983. Helochares (Sindolus) talarum sp. nov., y redescripcion de Helochares (Helochares) pallipes (Brulle), con descripcion de los estados preimaginales (Coleoptera: Hydrophilidae). Limnobios 2: 439 - 449" title="Helochares (Sindolus) talarum sp. nov., y redescripcion de Helochares (Helochares) pallipes (Brulle), con descripcion de los estados preimaginales (Coleoptera: Hydrophilidae)." volume="2" year="1983">
<normalizedToken id="1A5FAAEB14BC2899730E2C1CF163FE7A" originalValue="Fernández">Fernandez</normalizedToken>
1983
</bibRefCitation>
: 444;
<bibRefCitation id="629ABD6041A686D5FA46AF95E5DEC92D" author="Fernandez, LA" journalOrPublisher="Revista de la Sociedad Entomologica Argentina" pageId="0" pageNumber="1" pagination="145 - 151" publicationUrl="https://www.biotaxa.org/RSEA/article/view/36600" refId="B18" refString="Fernandez, LA, 1989. Notas sobre el genero Helochares. II (Coleoptera: Hydrophilidae). Descripcion de dos especies nuevas neotropicales. Clave para determinar las especies argentinas y de areas vecinas. Revista de la Sociedad Entomologica Argentina 45: 145 - 151, https://www.biotaxa.org/RSEA/article/view/36600" title="Notas sobre el genero Helochares. II (Coleoptera: Hydrophilidae). Descripcion de dos especies nuevas neotropicales. Clave para determinar las especies argentinas y de areas vecinas." url="https://www.biotaxa.org/RSEA/article/view/36600" volume="45" year="1989">
<normalizedToken id="307F750387FBDC386FCAC04271315EA2" originalValue="Fernández">Fernandez</normalizedToken>
1989
</bibRefCitation>
: 148 [in key].
</paragraph>
<paragraph id="DF8897FC52077C77B224A3FFBE060023" pageId="0" pageNumber="1">
<taxonomicName id="4CE03E556FCA8E7DABBB1C6F3F8587F1" baseAuthorityName="Brulle" baseAuthorityYear="1841" class="Insecta" family="Hydrophilidae" genus="Novochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Novochares pallipes" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pallipes">Novochares pallipes</taxonomicName>
(
<normalizedToken id="00C2CCEC8A607706C31960D5F4088FBC" originalValue="Brullé">Brulle</normalizedToken>
, 1841);
<bibRefCitation id="FF455F12035213ACCE1FC7E166AA0593" DOI="https://doi.org/10.3897/zookeys.1045.63810" author="Giron, JC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 236" refId="B23" refString="Giron, JC, Short, AEZ, 2021. The Acidocerinae (Coleoptera, Hydrophilidae): Taxonomy, classification, and catalog of species. ZooKeys 1045: 1 - 236, DOI: https://doi.org/10.3897/zookeys.1045.63810" title="The Acidocerinae (Coleoptera, Hydrophilidae): Taxonomy, classification, and catalog of species." url="https://doi.org/10.3897/zookeys.1045.63810" volume="1045" year="2021">
<normalizedToken id="FF354798F09F72885D463A7401AA3540" originalValue="Girón">Giron</normalizedToken>
and Short 2021
</bibRefCitation>
: 205.
</paragraph>
</subSubSection>
<subSubSection id="6256335739E1846CD3C7B742B32F5488" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="29511DA48CE6571FF007BBC1FB855080" pageId="0" pageNumber="1">Material examined</paragraph>
<paragraph id="0F77F2D0F44BD856ED7382BF08CA5088" pageId="0" pageNumber="1">
<emphasis id="8E2F25A08DB600B1AD43E5A6DC2439F7" bold="true" pageId="0" pageNumber="1">
(
<specimenCount id="F5F072985D92E30D8C021CF4F9895ED6" type="generic">14 exs.</specimenCount>
).
<materialsCitation id="3C62E272D3D551565CDF281954F1E0AC" collectingDate="1953-12" collectingDateMax="1962-03-10" collectingDateMin="1953-12" country="Argentina" location="Martinez" specimenCount="1" stateProvince="Buenos Aires">
<collectingCountry id="5284C6CF2ED9853784504E79D2C792EF" name="Argentina">Argentina</collectingCountry>
:
<collectingRegion id="435A56925D41DB1C7705A07765F16F51" country="Argentina" name="Ciudad Autonoma de Buenos Aires">Buenos Aires</collectingRegion>
:
<location id="C0ABFE90132ADB753041CFB6E9B9F4AE" LSID="urn:lsid:plazi:treatment:59C78BC226FB55249D864CFFEF6979A5:C0ABFE90132ADB753041CFB6E9B9F4AE" country="Argentina" name="Martinez" stateProvince="Buenos Aires">Martinez</location>
,
<collectingDate id="3ED4EB874CE7BCC746CB6EF601D7E998" value="1953-12">xii.1953</collectingDate>
(2, CAS); San Isidro,
<collectingDate id="829713946DD84C7044BB0B53E18D8BC3" value="1955-12">xii.1955</collectingDate>
(6, CAS, SEMC); Martinez,
<collectingDate id="5FB5EFB0D9BC3522162FDF6E43DE175B" value="1957-12">xii.1957</collectingDate>
(2, USNM); Choya, &quot;Sta. del Estero&quot;,
<collectingDate id="38AB7A719257868796738C942832796F" value="1962-03-10">10.iii.1962</collectingDate>
(1, CAS)
</materialsCitation>
</emphasis>
.
<materialsCitation id="B6B853705A1F579477A5C3C218A607A5" collectingDate="1979-12-10" collectorName="C. M. &amp; S. Flint, Flint, Jr." country="Argentina" location="Rio Parana Ibicuy" specimenCount="1" stateProvince="Entre Rios">
<emphasis id="527ECF6EC06E136F7787D3EB0F9B28A8" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="02AC63705D2B15F4E671A72F07AF6E79" country="Argentina" name="Entre Rios">
Entre
<normalizedToken id="4426925811CCC54C122F83C9A3AF1B44" originalValue="Ríos">Rios</normalizedToken>
</collectingRegion>
</emphasis>
:
<location id="B63950B7AB135AF340199E82CF911DF9" LSID="urn:lsid:plazi:treatment:59C78BC226FB55249D864CFFEF6979A5:B63950B7AB135AF340199E82CF911DF9" country="Argentina" name="Rio Parana Ibicuy" stateProvince="Entre Rios">
<normalizedToken id="7057B119D9F3929C7F8E8ABC5504A753" originalValue="Río">Rio</normalizedToken>
<normalizedToken id="EBB869D7B74180E6CCC5E8AB0FEADA8A" originalValue="Paraná">Parana</normalizedToken>
Ibicuy
</location>
, Pto. Ibicuy,
<collectingDate id="403569332EC93A52DB2B8CC9B6434029" value="1979-12-10">10.xii.1979</collectingDate>
, leg.
<collectorName id="67B4A5A1A1CEFF57CBFDAC11E6367281">
C.M. &amp; S.
<collectorName id="C7CE7480C4BD3B7111BE4EEBC6368341">Flint, Jr.</collectorName>
</collectorName>
(3, USNM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5A982FFF8F293460D41E9882FA953741" pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph id="C22CD026D109882282AE8D7E7BC60688" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph id="BD4492F9461E2BC7F51534C4F990C10F" pageId="0" pageNumber="1">
The strongly curved parameres and the extremely large distinctive fork of the dorsal plate of the median lobe (Fig.
<figureCitation id="A2D5E9B9DF1C1339E508D918812CFEA7" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Aedeagi and habitus of the Novochares abbreviatus species group A-E N. oculatus: A-C aedeagus, Mexico D, E paralectotype from Guatemala D aedeagus E habitus F-K miscellaneous forms of aedeagi in the Novochares abbreviatus species group F, G Argentina H, I Venezuela J, K Bolivia L, M, R N. pallipes N, O N. baca A, D, F, H, J, L, N dorsal view B, M ventral view C, G, I, K, O lateral view R oblique view." figureDoi="10.3897/zookeys.1171.104142.figure9" httpUri="https://binary.pensoft.net/fig/881176" pageId="0" pageNumber="1">9L-R</figureCitation>
) serve to separate this species fairly easily from other members of the species group.
</paragraph>
</subSubSection>
<subSubSection id="314F0EAE1B37A8193AE04ED7EC211971" pageId="0" pageNumber="1" type="description">
<paragraph id="68D7A39B1247D4DE80D5282C9ACC3C85" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="6FF3508D9C9C026E8B08B4A5A210B26B" pageId="0" pageNumber="1">
Body length 7.2-7.8 mm.
<emphasis id="F8D808200D2F7083D23B705C4169BFF3" bold="true" italics="true" pageId="0" pageNumber="1">Coloration</emphasis>
: Dorsal surfaces pale brown to orange-brown, sometimes with weakly paler (orange) clypeus and margins of pronotum and elytra.
<emphasis id="C2E638EC02C054AFB1198C6AC6583C75" bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
: Maxillary palps nearly 1.3
<normalizedToken id="9800E3D5A83F6B2DB3E00B7F08E0AA20" originalValue="×">x</normalizedToken>
width of head, uniformly yellow to orange in color.
<emphasis id="155D40A8F975CBAA73F1EAA2C787EBD8" bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
: Elytra without rows of serial punctures, faint rows (one dorsal and two or three lateral) of scarce and weakly marked systematic punctures barely noticeable. Prosternum medially broadly convex. Posterior elevation of mesoventrite broadly elevated, with low medial longitudinal ridge extending anteriorly.
<emphasis id="44ADFAFBA18AD613E0E8F1CCF1B764E8" bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
: Apical emargination of fifth ventrite moderately deep and relatively narrow.
<emphasis id="84EE2BC46328105499D4C5A105EEC740" bold="true" italics="true" pageId="0" pageNumber="1">Aedeagus</emphasis>
: (Fig.
<figureCitation id="92441E9F89802400F06653C65713CD50" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Aedeagi and habitus of the Novochares abbreviatus species group A-E N. oculatus: A-C aedeagus, Mexico D, E paralectotype from Guatemala D aedeagus E habitus F-K miscellaneous forms of aedeagi in the Novochares abbreviatus species group F, G Argentina H, I Venezuela J, K Bolivia L, M, R N. pallipes N, O N. baca A, D, F, H, J, L, N dorsal view B, M ventral view C, G, I, K, O lateral view R oblique view." figureDoi="10.3897/zookeys.1171.104142.figure9" httpUri="https://binary.pensoft.net/fig/881176" pageId="0" pageNumber="1">9L-R</figureCitation>
) Apical region of each paramere medially and dorsally curved; outer margin of apical region of paramere rounded, latero-ventrally weakly pointed; dorsal plate of median lobe with long and relatively broad neck, widened at apical region; neck with paired longitudinal, laminar elevations along second 1/2; distal arms laminate, dorso-ventrally oriented, 1/2 as long as neck of dorsal plate of median lobe; ventral plate of median lobe narrow and triangular, moderately sclerotized, reaching to beyond base of fork of dorsal plate of median lobe.
</paragraph>
</subSubSection>
<subSubSection id="03AC2645D5B55CDB765E25361BFB5AB0" pageId="0" pageNumber="1" type="distribution">
<paragraph id="852267A6AFDD860F4F60D24A34CB9574" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="E1C6094492A8BA8D19F07C1C33F6D6FB" pageId="0" pageNumber="1">
Argentina, Uruguay (Fig.
<figureCitation id="6F599E7AD5AB8305A5E618A4F87C333F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution of Novochares abbreviatus species group A N. abbreviatus: examined specimens (red) and literature / unconfirmed records (blue) B N. oculatus: examined specimens (red) and literature / unconfirmed records (blue) C N. pallipes (red), N. latus (yellow), N. baca (blue), N. pilatus (green)." figureDoi="10.3897/zookeys.1171.104142.figure10" httpUri="https://binary.pensoft.net/fig/881177" pageId="0" pageNumber="1">10C</figureCitation>
). Prior records from Brazil and Paraguay were erroneously reported and are removed from the known distribution of this species.
</paragraph>
</subSubSection>
<subSubSection id="CA7507ED6CC1088147608CF53EF2A3E8" pageId="0" pageNumber="1" type="remarks">
<paragraph id="9EFA2C92BE6DD9F581DF7C3DA99459DF" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="B003449825267B63DAA087E2E7F1F41A" pageId="0" pageNumber="1">
There has been some confusion about the distribution of this species in the literature. We have only been able to confirm the presence of this species in Argentina and Uruguay. Literature reports of this species occurring in Mato Grosso do Sul (
<normalizedToken id="376BCFF2C2DF9B044B427C5A597037EF" originalValue="Corumbá">Corumba</normalizedToken>
) and Paraguay (
<normalizedToken id="8DFEED9CE1AAA66D25FAB5DA4CE590EA" originalValue="Río">Rio</normalizedToken>
Alto
<normalizedToken id="C027E01B4B0954C8644C667BABD2ABE2" originalValue="Paraná">Parana</normalizedToken>
) appear to be derived from the original description of
<taxonomicName id="A41187C882AE6770C66719786DF2B0AA" authorityName="Orchymont" authorityYear="1939" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helochares parhedrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="parhedrus">Helochares (s. str.) parhedrus</taxonomicName>
. This species was briefly synonymized with
<taxonomicName id="0B0FF1B22F2D3B3B4D02BF1210677CE4" lsidName="N. pallipes" pageId="0" pageNumber="1" rank="species" species="pallipes">
<emphasis id="56E01833CD2EE46DF3088B0D806DBB2A" italics="true" pageId="0" pageNumber="1">N. pallipes</emphasis>
</taxonomicName>
but later discovered to actually be a synonym of
<taxonomicName id="497457A0FB550B7E1F3BE41A7BC3E14F" lsidName="H. atratus" pageId="0" pageNumber="1" rank="species" species="atratus">
<emphasis id="5B612CA30D3D5428CB8947DA11ECEC40" italics="true" pageId="0" pageNumber="1">H. atratus</emphasis>
</taxonomicName>
(
<normalizedToken id="9BA4C8F5BBF5A8E514D0CF205C7E46EF" originalValue="Fernández">Fernandez</normalizedToken>
, 1982). However, because for a short time the distributional records of
<taxonomicName id="F4FE57E451A840C351E3BC9E312B54E7" lsidName="N. pallipes" pageId="0" pageNumber="1" rank="species" species="pallipes">
<emphasis id="F63381118EC3B8EFD5C0FAF4D27B1295" italics="true" pageId="0" pageNumber="1">N. pallipes</emphasis>
</taxonomicName>
and
<taxonomicName id="BDA6271F7C8DE521277FE04E7E10C874" lsidName="H. parhedrus" pageId="0" pageNumber="1" rank="species" species="parhedrus">
<emphasis id="95D0415BE6E95758AA17ABFD423B0C3E" italics="true" pageId="0" pageNumber="1">H. parhedrus</emphasis>
</taxonomicName>
were fused, this muddied the literature.
<bibRefCitation id="006B6B104683095BE5012C1A8074E9FC" author="Fernandez, LA" journalOrPublisher="Limnobios" pageId="0" pageNumber="1" pagination="439 - 449" refId="B17" refString="Fernandez, LA, 1983. Helochares (Sindolus) talarum sp. nov., y redescripcion de Helochares (Helochares) pallipes (Brulle), con descripcion de los estados preimaginales (Coleoptera: Hydrophilidae). Limnobios 2: 439 - 449" title="Helochares (Sindolus) talarum sp. nov., y redescripcion de Helochares (Helochares) pallipes (Brulle), con descripcion de los estados preimaginales (Coleoptera: Hydrophilidae)." volume="2" year="1983">
<normalizedToken id="78AE248842C910B81BEE39F8A57907E9" originalValue="Fernández">Fernandez</normalizedToken>
(1983)
</bibRefCitation>
appears to have accidentally repeated the fused distribution which then carried into later papers. The record of this species from the state of Minas Gerais, Brazil derives from
<bibRefCitation id="6BDB03C9E138B8C764C8FDC58DCF0AA2" author="d'Orchymont, A" journalOrPublisher="Bulletin et Annales de la Societe entomologique de Belgique" pageId="0" pageNumber="1" pagination="201 - 248" publicationUrl="https://babel.hathitrust.org/cgi/pt?id=uc1.b3307256&amp;view=1up&amp;seq=207" refId="B8" refString="d'Orchymont, A, 1926. Contribution a l'etude des Hydrophilides VI. Bulletin et Annales de la Societe entomologique de Belgique 66: 201 - 248, https://babel.hathitrust.org/cgi/pt?id=uc1.b3307256&amp;view=1up&amp;seq=207" title="Contribution a l'etude des Hydrophilides VI." url="https://babel.hathitrust.org/cgi/pt?id=uc1.b3307256&amp;view=1up&amp;seq=207" volume="66" year="1926">
<normalizedToken id="4AE4362A8220A53614B31A1DFEE392C5" originalValue="d’Orchymont">d'Orchymont</normalizedToken>
(1926)
</bibRefCitation>
. However, later
<bibRefCitation id="FFC0DA911FDABCF0639BEE26E1486FEF" author="d'Orchymont, A" journalOrPublisher="Revista de la Sociedad Entomologica Argentina" pageId="0" pageNumber="1" pagination="253 - 264" refId="B10" refString="d'Orchymont, A, 1939. Notes sur quelques Palpicornia de la Republique Argentine. Revista de la Sociedad Entomologica Argentina 10: 253 - 264" title="Notes sur quelques Palpicornia de la Republique Argentine." volume="10" year="1939">
<normalizedToken id="4A75E96046DB5749432A40AFA6391ECC" originalValue="d’Orchymont">d'Orchymont</normalizedToken>
(1939)
</bibRefCitation>
noted that these records were erroneous and referred them to
<taxonomicName id="76C5804A497ACA7B5D79E3E36796DB7F" lsidName="H. parhedrus" pageId="0" pageNumber="1" rank="species" species="parhedrus">
<emphasis id="F462BB43855D523693C20C7AB9D0D34E" italics="true" pageId="0" pageNumber="1">H. parhedrus</emphasis>
</taxonomicName>
, which later was synonymized with
<taxonomicName id="4C0B0B7BC336FCE7207A8255764864A7" lsidName="N. atratus" pageId="0" pageNumber="1" rank="species" species="atratus">
<emphasis id="EC9F533583892E61904D9EA5B747E25B" italics="true" pageId="0" pageNumber="1">N. atratus</emphasis>
</taxonomicName>
. We are not aware of any verified Brazilian records of
<taxonomicName id="72AF743E0805E798B1760CC505725BCA" lsidName="H. pallipes" pageId="0" pageNumber="1" rank="species" species="pallipes">
<emphasis id="9C9933EC301DDCD950F0CB20194116DF" italics="true" pageId="0" pageNumber="1">H. pallipes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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