treatments-xml/data/78/20/06/7820060EFFD2C23EACA0FEB7B7CFFF69.xml

<document id="E4181A2363D75ED3A73094C88EF0D107" ID-DOI="10.11646/zootaxa.3717.4.6" ID-GBIF-Dataset="95ca01fa-ce8c-4423-9643-5a3e186d75f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="247606" ID-ZooBank="1E052509-825D-4527-8F24-1F1C7781C47B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460685103033" checkinUser="plazi" docAuthor="Hoskin, Conrad J. &amp; Couper, Patrick" docDate="2013" docId="7820060EFFD2C23EACA0FEB7B7CFFF69" docLanguage="en" docName="zt03717p558.pdf" docOrigin="Zootaxa 3717 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Saltuarius eximius Hoskin &amp; Couper, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="553" masterDocId="84197E76FFD0C235AC37FD18B351FFBC" masterDocTitle="A spectacular new leaf-tailed gecko (Carphodactylidae: Saltuarius) from the Melville Range, north-east Australia" masterLastPageNumber="558" masterPageNumber="543" pageNumber="545" updateTime="1698351021033" updateUser="plazi">
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<mods:title id="3BE2B352FE324DB1155D4AFB6D05816E">A spectacular new leaf-tailed gecko (Carphodactylidae: Saltuarius) from the Melville Range, north-east Australia</mods:title>
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<mods:namePart id="D70EAF605F1F31E057DBEAB8B08B2EE7">Hoskin, Conrad J.</mods:namePart>
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<mods:namePart id="9F3FB55F6E2228076BB072C38CC87123">Couper, Patrick</mods:namePart>
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<mods:date id="B66AD6FD92EBD025A302028AC1CB8B93">2013</mods:date>
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<paragraph id="F036B718FFD2C237ACA0FEB7B28EFC75" blockId="2.[151,520,943,1039]" box="[151,479,943,969]" pageId="2" pageNumber="545">
<heading id="AB7E0074FFD2C237ACA0FEB7B28EFC75" bold="true" box="[151,479,943,969]" fontSize="11" level="1" pageId="2" pageNumber="545" reason="1">
<emphasis id="C2FD6B0AFFD2C237ACA0FEB7B28EFC75" bold="true" box="[151,479,943,969]" pageId="2" pageNumber="545">
<taxonomicName id="3789CC9BFFD2C237ACA0FEB7B22BFC75" box="[151,378,943,969]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="545" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD2C237ACA0FEB7B22BFC75" bold="true" box="[151,378,943,969]" italics="true" pageId="2" pageNumber="545">Saltuarius eximius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9CED671FFD2C237ADB6FEB7B28EFC75" box="[385,479,943,969]" pageId="2" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="F036B718FFD2C237ACA0FECBB2EEFBB3" blockId="2.[151,520,943,1039]" pageId="2" pageNumber="545">
Cape Melville Leaf-tailed Gecko (
<figureCitation id="68B2AB9DFFD2C237ACA8FEEEB3B4FBB3" box="[159,229,1014,1039]" captionStart="FIGURE 1" captionStartId="4.[151,250,1989,2011]" captionTargetBox="[188,1355,193,1942]" captionTargetId="figure@4.[188,1356,193,1947]" captionTargetPageId="4" captionText="FIGURE 1. Saltuarius eximius sp. nov. in life: (A, B) original tail (QMJ 92377), (C) regenerated tail (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247607/files/figure.png" pageId="2" pageNumber="545">Figs 1</figureCitation>
, 
<figureCitation id="68B2AB9DFFD2C237ACC4FEEFB252FBB3" box="[243,259,1015,1039]" captionStart="FIGURE 2" captionStartId="5.[151,250,873,895]" captionTargetBox="[167,1435,193,835]" captionTargetId="figure@5.[167,1452,193,836]" captionTargetPageId="5" captionText="FIGURE 2. Saltuarius eximius sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/247608/files/figure.png" pageId="2" pageNumber="545">2</figureCitation>
, 
<figureCitation id="68B2AB9DFFD2C237AD38FEEFB24EFBB3" box="[271,287,1015,1039]" captionStart="FIGURE 3" captionStartId="5.[151,250,1873,1895]" captionTargetBox="[151,1436,945,1852]" captionTargetId="figure@5.[151,1436,945,1852]" captionTargetPageId="5" captionText="FIGURE 3. Original tail of S. eximius sp. nov. (QMJ 92377): (A) dorsal, (B) ventral (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247609/files/figure.png" pageId="2" pageNumber="545">3</figureCitation>
, 
<figureCitation id="68B2AB9DFFD2C237AD1CFEEFB26BFBB3" box="[299,314,1015,1039]" captionStart="FIGURE 4" captionStartId="7.[151,250,1968,1990]" captionTargetBox="[227,1337,193,1929]" captionTargetId="figure@7.[227,1339,193,1932]" captionTargetPageId="7" captionText="FIGURE 4. Comparison of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Jeff Wright)." httpUri="https://zenodo.org/record/247610/files/figure.png" pageId="2" pageNumber="545">4</figureCitation>
A, 5A, 6, 7)
</paragraph>
</subSubSection>
<subSubSection id="B893E493FFD2C237ACA0F926B1AFFB23" pageId="2" pageNumber="545" type="materials_examined">
<paragraph id="F036B718FFD2C237ACA0F926B1AFFB23" blockId="2.[151,1437,1086,1981]" pageId="2" pageNumber="545">
<emphasis id="C2FD6B0AFFD2C237ACA0F926B2ADFBEB" bold="true" box="[151,508,1086,1111]" pageId="2" pageNumber="545">
Material examined. 
<typeStatus id="2F3209BAFFD2C237ADB1F926B2A6FBEB" box="[390,503,1086,1111]" pageId="2" pageNumber="545" type="holotype">Holotype</typeStatus>
:
</emphasis>
QMJ92377, female, Melville Range (
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, 
<geoCoordinate id="95BDD1DFFFD2C237A801F927B79CFBEA" box="[1078,1229,1087,1111]" direction="east" orientation="longitude" pageId="2" pageNumber="545" precision="15" value="144.49112">144°29'28&quot; E</geoCoordinate>
, elevation 
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), Cape Melville National Park, north-east Queensland, C. J. Hoskin, 
<date id="843791D8FFD2C237AFB1F97BB765FBC7" box="[902,1076,1122,1147]" pageId="2" pageNumber="545" value="2013-03-20">20 March 2013</date>
. 
<emphasis id="C2FD6B0AFFD2C237A808F97AB791FBC7" bold="true" box="[1087,1216,1122,1147]" pageId="2" pageNumber="545">
<typeStatus id="2F3209BAFFD2C237A808F97AB7EAFBC7" box="[1087,1211,1122,1147]" pageId="2" pageNumber="545" type="paratype">Paratypes</typeStatus>
:
</emphasis>
QMJ92378 (male), QMJ92379 (female), collection details as for 
<typeStatus id="2F3209BAFFD2C237AEA3F99EB1ABFB23" box="[660,762,1158,1183]" pageId="2" pageNumber="545" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="B893E493FFD2C237ACF0F9B2B7CDFA5F" pageId="2" pageNumber="545" type="diagnosis">
<paragraph id="F036B718FFD2C237ACF0F9B2B7CDFA5F" blockId="2.[151,1437,1086,1981]" pageId="2" pageNumber="545">
<emphasis id="C2FD6B0AFFD2C237ACF0F9B2B210FB7F" bold="true" box="[199,321,1194,1219]" pageId="2" pageNumber="545">Diagnosis.</emphasis>
<taxonomicName id="3789CC9BFFD2C237AD7FF9B3B146FB7E" box="[328,535,1195,1219]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="545" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD2C237AD7FF9B3B146FB7E" box="[328,535,1195,1219]" italics="true" pageId="2" pageNumber="545">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD2C237AE29F9B2B129FB7F" bold="true" box="[542,632,1194,1219]" pageId="2" pageNumber="545">
<taxonomicNameLabel id="D9CED671FFD2C237AE29F9B2B129FB7F" box="[542,632,1194,1219]" pageId="2" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is highly distinct in its long and slender form, short head and very large eyes (
<figureCitation id="68B2AB9DFFD2C237ACEEF9D6B217FB5B" box="[217,326,1230,1255]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="4.[151,250,1989,2011]" captionStartId-1="5.[151,250,873,895]" captionStartId-2="5.[151,250,1873,1895]" captionStartId-3="7.[151,250,1968,1990]" captionStartId-4="8.[151,250,1965,1987]" captionTargetBox-0="[188,1355,193,1942]" captionTargetBox-1="[167,1435,193,835]" captionTargetBox-2="[151,1436,945,1852]" captionTargetBox-3="[227,1337,193,1929]" captionTargetBox-4="[382,1209,193,1932]" captionTargetId-0="figure@4.[188,1356,193,1947]" captionTargetId-1="figure@5.[167,1452,193,836]" captionTargetId-2="figure@5.[151,1436,945,1852]" captionTargetId-3="figure@7.[227,1339,193,1932]" captionTargetId-4="figure@8.[381,1209,193,1932]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Saltuarius eximius sp. nov. in life: (A, B) original tail (QMJ 92377), (C) regenerated tail (photos: Conrad Hoskin)." captionText-1="FIGURE 2. Saltuarius eximius sp. nov. in life (photo: Conrad Hoskin)." captionText-2="FIGURE 3. Original tail of S. eximius sp. nov. (QMJ 92377): (A) dorsal, (B) ventral (photos: Conrad Hoskin)." captionText-3="FIGURE 4. Comparison of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Jeff Wright)." captionText-4="FIGURE 5. Comparison of heads of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Steve Wilson)." httpUri-0="https://zenodo.org/record/247607/files/figure.png" httpUri-1="https://zenodo.org/record/247608/files/figure.png" httpUri-2="https://zenodo.org/record/247609/files/figure.png" httpUri-3="https://zenodo.org/record/247610/files/figure.png" httpUri-4="https://zenodo.org/record/247611/files/figure.png" pageId="2" pageNumber="545">Figs. 1–5</figureCitation>
). 
<taxonomicName id="3789CC9BFFD2C237AD6BF9D6B17CFB5A" box="[348,557,1230,1254]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="545" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD2C237AD6BF9D6B17CFB5A" box="[348,557,1230,1254]" italics="true" pageId="2" pageNumber="545">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD2C237AE02F9D6B1C1FB5B" bold="true" box="[565,656,1230,1255]" pageId="2" pageNumber="545">
<taxonomicNameLabel id="D9CED671FFD2C237AE02F9D6B1C1FB5B" box="[565,656,1230,1255]" pageId="2" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is readily distinguished from all its congeners by the following nonoverlapping (or largely non-overlapping) body measurements (as % SVL): its more elongate and narrow body form (AG = 49–50% 
<emphasis id="C2FD6B0AFFD2C237AD7DF80FB230FA92" box="[330,353,1303,1326]" italics="true" pageId="2" pageNumber="545">vs</emphasis>
40–51% combined range for all other 
<taxonomicName id="3789CC9BFFD2C237AF38F80EB0D3FA92" box="[783,898,1302,1326]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="545" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD2C237AF38F80EB0D3FA92" box="[783,898,1302,1326]" italics="true" pageId="2" pageNumber="545">Saltuarius</emphasis>
</taxonomicName>
species; midBW = 11–12% 
<emphasis id="C2FD6B0AFFD2C237A8F6F80FB789FA92" box="[1217,1240,1303,1326]" italics="true" pageId="2" pageNumber="545">vs</emphasis>
13–21%); longer limbs (L1 = 55–57% 
<emphasis id="C2FD6B0AFFD2C237ADA2F823B2FDFAEE" box="[405,428,1339,1362]" italics="true" pageId="2" pageNumber="545">vs</emphasis>
42–52%, L2 = 63–64% 
<emphasis id="C2FD6B0AFFD2C237AEE5F823B1B8FAEE" box="[722,745,1339,1362]" italics="true" pageId="2" pageNumber="545">vs</emphasis>
52–63%); narrower neck (NW = 6.3–6.5% 
<emphasis id="C2FD6B0AFFD2C237A8D8F823B657FAEE" box="[1263,1286,1339,1362]" italics="true" pageId="2" pageNumber="545">vs</emphasis>
7.5–10.7%); narrower head (HW = 18–19% 
<emphasis id="C2FD6B0AFFD2C237ADCEF847B141FACA" box="[505,528,1375,1398]" italics="true" pageId="2" pageNumber="545">vs</emphasis>
19–24%); shorter head (HL = 24–25% 
<emphasis id="C2FD6B0AFFD2C237AFF9F847B0B4FACA" box="[974,997,1375,1398]" italics="true" pageId="2" pageNumber="545">vs</emphasis>
25–30%); shorter temporal region (EE = 6.0–7.8% 
<emphasis id="C2FD6B0AFFD2C237AD28F89BB267FA26" box="[287,310,1411,1434]" italics="true" pageId="2" pageNumber="545">vs</emphasis>
7.4–10.3%); and larger eye diameter (ED = 7.8–8.6% 
<emphasis id="C2FD6B0AFFD2C237AFAFF89BB0FEFA26" box="[920,943,1411,1434]" italics="true" pageId="2" pageNumber="545">vs</emphasis>
5.2–7.3%) (Table 1). 
<taxonomicName id="3789CC9BFFD2C237A893F89BB622FA26" box="[1188,1395,1411,1435]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="545" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD2C237A893F89BB622FA26" box="[1188,1395,1411,1435]" italics="true" pageId="2" pageNumber="545">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD2C237A94DF89AB399FA03" bold="true" pageId="2" pageNumber="545">
<taxonomicNameLabel id="D9CED671FFD2C237A94DF89AB399FA03" pageId="2" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also has an unpatterned grey eye versus the patterned eyes of all congeners. 
<taxonomicName id="3789CC9BFFD2C237A804F8BEB652FA02" box="[1075,1283,1446,1470]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="545" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD2C237A804F8BEB652FA02" box="[1075,1283,1446,1470]" italics="true" pageId="2" pageNumber="545">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD2C237A93BF8BEB636FA03" bold="true" box="[1292,1383,1446,1471]" pageId="2" pageNumber="545">
<taxonomicNameLabel id="D9CED671FFD2C237A93BF8BEB636FA03" box="[1292,1383,1446,1471]" pageId="2" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also has a distinct tail that is elaborately frilled along the margin with a short, fine attenuated tip.
</paragraph>
</subSubSection>
<subSubSection id="B893E493FFD2C237ACF0F8F6B63AF997" pageId="2" pageNumber="545" type="etymology">
<paragraph id="F036B718FFD2C237ACF0F8F6B63AF997" blockId="2.[151,1437,1086,1981]" pageId="2" pageNumber="545">
<emphasis id="C2FD6B0AFFD2C237ACF0F8F6B21FF9BB" bold="true" box="[199,334,1518,1543]" pageId="2" pageNumber="545">Etymology.</emphasis>
<taxonomicName id="3789CC9BFFD2C237AD6DF8F6B2EAF9BB" box="[346,443,1518,1543]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="557" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">Eximius</taxonomicName>
; from the Latin meaning exceptional, extraordinary, exquisite. In recognition of the particularly fine form and distinctiveness of this species. The species epithet is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="B893E493FFD2C23FACF0FB2EB795F980" lastPageId="10" lastPageNumber="553" pageId="2" pageNumber="545" type="description">
<paragraph id="F036B718FFD2C237ACF0FB2EB638F959" blockId="2.[151,1437,1086,1981]" pageId="2" pageNumber="545">
<emphasis id="C2FD6B0AFFD2C237ACF0FB2EB1B2F9F3" bold="true" box="[199,739,1590,1615]" pageId="2" pageNumber="545">
Measurements and scale counts of 
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.
</emphasis>
SVL = 
<quantity id="37711AFDFFD2C237AF73FB2EB0E9F9F2" box="[836,952,1590,1615]" metricMagnitude="-1" metricUnit="m" metricValue="1.064" pageId="2" pageNumber="545" unit="mm" value="106.4">106.4 mm</quantity>
, AG = 
<quantity id="37711AFDFFD2C237A827FB2EB726F9F2" box="[1040,1143,1590,1615]" metricMagnitude="-2" metricUnit="m" metricValue="5.35" pageId="2" pageNumber="545" unit="mm" value="53.5">53.5 mm</quantity>
, midBW = 
<quantity id="37711AFDFFD2C237A8C8FB2EB635F9F2" box="[1279,1380,1590,1615]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="2" pageNumber="545" unit="mm" value="11.5">11.5 mm</quantity>
, Tail (original) = 
<quantity id="37711AFDFFD2C237AD2DFB42B22CF9CE" box="[282,381,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="6.83" pageId="2" pageNumber="545" unit="mm" value="68.3">68.3 mm</quantity>
, Tip = 
<quantity id="37711AFDFFD2C237ADFEFB43B17DF9CE" box="[457,556,1627,1651]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="2" pageNumber="545" unit="mm" value="17.2">17.2 mm</quantity>
, TailW = 33.9, L1 = 
<quantity id="37711AFDFFD2C237AF38FB42B022F9CE" box="[783,883,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="6.07" pageId="2" pageNumber="545" unit="mm" value="60.7">60.7 mm</quantity>
, L2 = 
<quantity id="37711AFDFFD2C237AF8EFB42B74DF9CE" box="[953,1052,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="6.859999999999999" pageId="2" pageNumber="545" unit="mm" value="68.6">68.6 mm</quantity>
, FLL = 
<quantity id="37711AFDFFD2C237A843FB42B786F9CE" box="[1140,1239,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="2" pageNumber="545" unit="mm" value="26.3">26.3 mm</quantity>
, HLL = 
<quantity id="37711AFDFFD2C237A903FB42B6C6F9CE" box="[1332,1431,1626,1651]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" pageId="2" pageNumber="545" unit="mm" value="28.6">28.6 mm</quantity>
, HL = 
<quantity id="37711AFDFFD2C237ACEEFB66B26CF92A" box="[217,317,1662,1687]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="2" pageNumber="545" unit="mm" value="25.8">25.8 mm</quantity>
, HW = 
<quantity id="37711AFDFFD2C237ADA3FB67B2A8F92A" box="[404,505,1663,1687]" metricMagnitude="-2" metricUnit="m" metricValue="1.97" pageId="2" pageNumber="545" unit="mm" value="19.7">19.7 mm</quantity>
, snout = 
<quantity id="37711AFDFFD2C237AE6AFB67B191F92A" box="[605,704,1663,1687]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="2" pageNumber="545" unit="mm" value="11.8">11.8 mm</quantity>
, EE = 
<quantity id="37711AFDFFD2C237AF3CFB66B030F92A" box="[779,865,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="545" unit="mm" value="6.4">6.4 mm</quantity>
, ED = 9.2, NL = 
<quantity id="37711AFDFFD2C237A815FB67B7D4F92A" box="[1058,1157,1663,1687]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" pageId="2" pageNumber="545" unit="mm" value="20.8">20.8 mm</quantity>
, NW = 
<quantity id="37711AFDFFD2C237A8EAFB66B662F92A" box="[1245,1331,1662,1687]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="545" unit="mm" value="6.7">6.7 mm</quantity>
, enlarged lamellae 4th finger = 19, enlarged lamellae 4th toe = 18, lamellae to junction 3rd and 4th finger = 19, lamellae to junction 3rd and 4th toe = 21 (includes 3 smaller scales at basal end), supralabials = 15 (1st in series fused), infralabials = 12.
</paragraph>
<paragraph id="F036B718FFD2C236ACF0FBE8B435FC91" blockId="2.[151,1437,1086,1981]" lastBlockId="3.[125,1919,653,913]" lastPageId="3" lastPageNumber="546" pageId="2" pageNumber="545">
<emphasis id="C2FD6B0AFFD2C237ACF0FBE8B141F8B5" bold="true" box="[199,528,1776,1801]" pageId="2" pageNumber="545">
Description of 
<typeStatus id="2F3209BAFFD2C237ADBDFBE8B2ECF8B5" box="[394,445,1776,1801]" pageId="2" pageNumber="545">type</typeStatus>
series
</emphasis>
(
<figureCitation id="68B2AB9DFFD2C237AE1EFBE8B129F8B5" box="[553,632,1776,1801]" captionStart="FIGURE 6" captionStartId="9.[151,250,1972,1994]" captionTargetBox="[192,1410,193,1930]" captionTargetId="figure@9.[192,1434,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. Type series of S. eximius sp. nov.. Specimens (left-right): QMJ 92378 (paratype, male), QMJ 92377 (holotype, female), QMJ 92379 (paratype, female). (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247612/files/figure.png" pageId="2" pageNumber="545">Fig. 6</figureCitation>
): Data presented as range followed by mean in brackets. 
<emphasis id="C2FD6B0AFFD2C237A960FBE8B212F891" bold="true" pageId="2" pageNumber="545">Adult measurements</emphasis>
(mm): SVL 106.4–118.3 (112.4); AG 53.5–58.3 (55.6); midBW 11.5–14.4 (13.2); Tail (original) 68.3 (only 
<typeStatus id="2F3209BAFFD2C237AD24FA20B224F8ED" box="[275,373,1848,1873]" pageId="2" pageNumber="545" type="holotype">holotype</typeStatus>
has an original tail); Tip (original) 17. 2; TailL 53.1–68.3 (59.7); TailW 33.9–41.2 (37.3); L1 60.7–65.3 (62.5); L2 68.6–74.5 (71.2); FLL 26.3–30.5 (28.3); HLL 28.6–31.5 (29.9); HL 25.8–29.1 (27.5); HW 19.7–22.3 (21.2); snout 11.8–13.2 (12.5); NW 6.7–7.6 (7.2); NL 20.8–22.4 (21.8); EE 6.4–9.3 (7.8); ED 8.8–9.5 (9.1). 
<emphasis id="C2FD6B0AFFD2C237ACEDFABCB2ECF801" bold="true" box="[218,445,1956,1981]" pageId="2" pageNumber="545">Adult proportions:</emphasis>
see Table 1. 
<emphasis id="C2FD6B0AFFD2C237AE79FABCB1C5F801" bold="true" box="[590,660,1956,1981]" italics="true" pageId="2" pageNumber="545">Head:</emphasis>
Short, deep, triangular, distinct from neck; head depth 48–53% of head 
<emphasis id="C2FD6B0AFFD3C236AC4AFF96B3A0FD1B" bold="true" box="[125,241,654,679]" pageId="3" pageNumber="546">TABLE 1.</emphasis>
Morphological comparison of 
<taxonomicName id="3789CC9BFFD3C236AE54FF95B187FD1B" box="[611,726,653,679]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD3C236AE54FF95B187FD1B" box="[611,726,653,679]" italics="true" pageId="3" pageNumber="546">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236AEEAFF96B06EFD1B" bold="true" box="[733,831,654,679]" pageId="3" pageNumber="546">
<taxonomicNameLabel id="D9CED671FFD3C236AEEAFF96B06EFD1B" box="[733,831,654,679]" pageId="3" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with other 
<taxonomicName id="3789CC9BFFD3C236AFFCFF95B714FD1B" box="[971,1093,653,679]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD3C236AFFCFF95B714FD1B" box="[971,1093,653,679]" italics="true" pageId="3" pageNumber="546">Saltuarius</emphasis>
</taxonomicName>
species and 
<taxonomicName id="3789CC9BFFD3C236A8E9FF95B6CAFD1B" box="[1246,1435,653,679]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="occultus">
<emphasis id="C2FD6B0AFFD3C236A8E9FF95B6CAFD1B" box="[1246,1435,653,679]" italics="true" pageId="3" pageNumber="546">Orraya occultus</emphasis>
</taxonomicName>
. Table shows data for SVL (mm) then length of traits as a proportion (%) of SVL. Mean is presented above range. Ranges for traits with proportions over 10% are rounded to nearest percent. For 
<taxonomicName id="3789CC9BFFD3C236AD20FFC8B2DBFD56" box="[279,394,720,746]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD3C236AD20FFC8B2DBFD56" box="[279,394,720,746]" italics="true" pageId="3" pageNumber="546">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236ADA6FFC9B2A2FD56" bold="true" box="[401,499,721,746]" pageId="3" pageNumber="546">
<taxonomicNameLabel id="D9CED671FFD3C236ADA6FFC9B2A2FD56" box="[401,499,721,746]" pageId="3" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="3789CC9BFFD3C236AE1AFFC8B1FDFD56" box="[557,684,720,746]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="occultus">
<emphasis id="C2FD6B0AFFD3C236AE1AFFC8B1FDFD56" box="[557,684,720,746]" italics="true" pageId="3" pageNumber="546">
<collectingCountry id="889EF788FFD3C236AE1AFFC8B110FD56" box="[557,577,720,746]" name="Iceland" pageId="3" pageNumber="546">O</collectingCountry>
. occultus
</emphasis>
</taxonomicName>
, only a single specimen had an original tail, while for 
<taxonomicName id="3789CC9BFFD3C236A904FFC8B6CCFD56" box="[1331,1437,720,746]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="kateae">
<emphasis id="C2FD6B0AFFD3C236A904FFC8B6CCFD56" box="[1331,1437,720,746]" italics="true" pageId="3" pageNumber="546">S. kateae</emphasis>
</taxonomicName>
, no specimens with original tails were available. Morphology codes are: SVL: snout to vent length, Tail: original tail length, Tip: original tail tip length, AG: axilla to groin length, midBW: mid–body width, L1: forelimb length, L2: hindlimb length, HL: head length, HW: head width, snout: snout length, NW: neck width, NL: neck length,
</paragraph>
<paragraph id="F036B718FFD3C236AC96FE2DB1F2FC2D" blockId="3.[125,1919,653,913]" pageId="3" pageNumber="546">
: eye to ear distance, ED: eye diameter. For more details see Methods. For 
<taxonomicName id="3789CC9BFFD3C236A82EFE2CB7DCFCF2" box="[1049,1165,820,846]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD3C236A82EFE2CB7DCFCF2" box="[1049,1165,820,846]" italics="true" pageId="3" pageNumber="546">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236A8A3FE2DB7A7FCF2" bold="true" box="[1172,1270,821,846]" pageId="3" pageNumber="546">
<taxonomicNameLabel id="D9CED671FFD3C236A8A3FE2DB7A7FCF2" box="[1172,1270,821,846]" pageId="3" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the data comes from the three specimens comprising the 
<typeStatus id="2F3209BAFFD3C236AC9DFE4EB3B1FCD3" box="[170,224,854,879]" pageId="3" pageNumber="546">type</typeStatus>
series. For the other species, the data is a summary of measurements taken from the specimens listed in the Appendix and data from Couper 
<emphasis id="C2FD6B0AFFD3C236ACECFE6FB244FC2D" box="[219,277,887,913]" italics="true" pageId="3" pageNumber="546">et al.</emphasis>
(2008) and Couper 
<emphasis id="C2FD6B0AFFD3C236AE3FFE6FB113FC2D" box="[520,578,887,913]" italics="true" pageId="3" pageNumber="546">et al.</emphasis>
(1993).
</paragraph>
<paragraph id="F036B718FFD3C236ADB4FEA4B6E1FC4A" blockId="3.[145,1934,956,1551]" pageId="3" pageNumber="546">SVL Tail/SVLTip/Tail AG/SVLmidBW/ L1/SVL L2/SVL HL/SVL HW/SVL snout/ NW/SVL NL/SVL EE/SVL ED/SVL SVL SVL</paragraph>
<paragraph id="F036B718FFD3C233ACA1FEE7B00CFB31" blockId="3.[145,1934,956,1551]" lastBlockId="6.[151,1437,151,2029]" lastPageId="6" lastPageNumber="549" pageId="3" pageNumber="546">
<taxonomicName id="3789CC9BFFD3C236ACA1FEE7B3A1FBA5" box="[150,240,1023,1049]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="eximius">
<emphasis id="C2FD6B0AFFD3C236ACA1FEE7B3A1FBA5" box="[150,240,1023,1049]" italics="true" pageId="3" pageNumber="546">eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236ACC0F918B423FBA5" bold="true" box="[247,1906,1024,1049]" pageId="3" pageNumber="546">sp nov. 112 64.1 25.2 49.5 11.7 55.6 63.4 24.5 18.8 11.1 6.4 19.4 6.9 8.1</emphasis>
106–118 49– 
<date id="843791D8FFD3C236AEC0F939B047FB86" box="[759,790,1057,1082]" pageId="3" pageNumber="546" value="1950-12-11">50</date>
11–12 55–57 63–64 24– 
<date id="843791D8FFD3C236A889F939B78DFB86" box="[1214,1244,1057,1082]" pageId="3" pageNumber="546" value="1925-11-18" valueMax="1925-11-19" valueMin="1925-11-18">25</date>
18–19 11–11 6.3–6.5 19–20 6.0–7.8 7.8–8.6 
<taxonomicName id="3789CC9BFFD3C236ACA1F95AB3AEFBE0" box="[150,255,1090,1116]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="cornutus">
<emphasis id="C2FD6B0AFFD3C236ACA1F95AB3AEFBE0" box="[150,255,1090,1116]" italics="true" pageId="3" pageNumber="546">cornutus</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236ADB6F95AB423FBE7" bold="true" box="[385,1906,1090,1115]" pageId="3" pageNumber="546">126 63.2 35.8 45.5 16.2 47.8 56.9 28.1 21.8 13.5 9.4 17.8 9.3 6.3</emphasis>
116–136 59–68 32–40 41–49 13–19 47–51 55–61 27–29 21–22 13–14 8.9–9.8 15–21 8.6–10.0 5.6–7.2 
<taxonomicName id="3789CC9BFFD3C236ACA1F99DB243FB23" box="[150,274,1157,1183]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="salebrosus">
<emphasis id="C2FD6B0AFFD3C236ACA1F99DB243FB23" box="[150,274,1157,1183]" italics="true" pageId="3" pageNumber="546">salebrosus</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236ADB6F99DB423FB22" bold="true" box="[385,1906,1157,1182]" pageId="3" pageNumber="546">132 63.9 25.6 44.6 17.7 47.3 55.5 26.6 22.7 12.3 9.4 19.6 9.7 5.8</emphasis>
121–144 58–72 22–30 43–47 14–21 45–51 52–61 26– 
<date id="843791D8FFD3C236A889F9BFB78DFB7C" box="[1214,1244,1191,1216]" pageId="3" pageNumber="546" value="1927-12-22" valueMax="1927-12-24" valueMin="1927-12-22">27</date>
22–24 12–13 8.8–10.7 18–21 9.3–10.0 5.2–6.5 
<taxonomicName id="3789CC9BFFD3C236ACA1F9D0B3B2FB5E" box="[150,227,1224,1250]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="swaini">
<emphasis id="C2FD6B0AFFD3C236ACA1F9D0B3B2FB5E" box="[150,227,1224,1250]" italics="true" pageId="3" pageNumber="546">swaini</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236ADB6F9D0B423FB5D" bold="true" box="[385,1906,1224,1249]" pageId="3" pageNumber="546">124 66.9 40.6 46.2 15.9 48.2 58.7 26.2 20.4 12.2 8.9 18.6 8.3 6.1</emphasis>
99–134 59–72 35–49 41–51 15–17 44–51 53–62 25– 
<date id="843791D8FFD3C236A88AF9F2B78DFABF" box="[1213,1244,1258,1283]" pageId="3" pageNumber="546" value="1929-11-19" valueMax="1929-11-22" valueMin="1929-11-19">29</date>
19–22 11–14 8.0–9.6 15–22 7.4–9.4 5.6–6.5 
<taxonomicName id="3789CC9BFFD3C236ACA1F812B3ACFA98" box="[150,253,1290,1316]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="wyberba">
<emphasis id="C2FD6B0AFFD3C236ACA1F812B3ACFA98" box="[150,253,1290,1316]" italics="true" pageId="3" pageNumber="546">wyberba</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236ADBDF813B423FA98" bold="true" box="[394,1906,1291,1316]" pageId="3" pageNumber="546">99 71.2 40.6 45.0 17.1 47.2 57.4 27.3 22.1 12.1 9.8 17.7 8.9 7.0</emphasis>
86–108 62–81 33–46 40–48 16–18 44–51 53–63 25– 
<date id="843791D8FFD3C236A88AF834B78DFAF9" box="[1213,1244,1324,1349]" pageId="3" pageNumber="546" value="1929-11-20" valueMax="1929-11-24" valueMin="1929-11-20">29</date>
20–24 11–14 8.3–10.6 15–21 7.5–10.0 6.6–7.3 
<taxonomicName id="3789CC9BFFD3C236ACA1F855B3B7FADB" box="[150,230,1357,1383]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="kateae">
<emphasis id="C2FD6B0AFFD3C236ACA1F855B3B7FADB" box="[150,230,1357,1383]" italics="true" pageId="3" pageNumber="546">kateae</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236ADB6F856B423FADB" bold="true" box="[385,1906,1358,1383]" pageId="3" pageNumber="546">104 46.8 17.8 46.3 58.0 27.2 21.3 12.2 8.3 17.4 8.3 6.4</emphasis>
97–110 45–48 17–19 45–47 55–60 27– 
<date id="843791D8FFD3C236A88AF877B78AFA34" box="[1213,1243,1391,1416]" pageId="3" pageNumber="546" value="1928-12-21" valueMax="1928-12-22" valueMin="1928-12-21">28</date>
21–22 12–13 7.5–8.7 15–20 7.5–9.2 6.3–6.6 
<taxonomicName id="3789CC9BFFD3C236ACA1F888B3BCFA16" box="[150,237,1424,1450]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="moritzi">
<emphasis id="C2FD6B0AFFD3C236ACA1F888B3BCFA16" box="[150,237,1424,1450]" italics="true" pageId="3" pageNumber="546">moritzi</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236ADBDF889B423FA16" bold="true" box="[394,1906,1425,1450]" pageId="3" pageNumber="546">98 68.9 39.3 45.1 16.7 47.4 57.7 26.9 21.2 12.2 9.1 19.4 8.4 6.8</emphasis>
88–109 63–76 29–47 40–50 15–19 42–52 52–62 25– 
<date id="843791D8FFD3C236A88AF8AAB78DFA77" box="[1213,1244,1458,1483]" pageId="3" pageNumber="546" value="1930-11-20" valueMax="1930-11-24" valueMin="1930-11-20">30</date>
20–24 11–14 8.3–10.6 16–23 7.4–10.3 6.4–7.1. 
<taxonomicName id="3789CC9BFFD3C236ACADF8CBB3ADFA51" box="[154,252,1491,1517]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="546" phylum="Chordata" rank="species" species="occultus">
<emphasis id="C2FD6B0AFFD3C236ACADF8CBB3ADFA51" box="[154,252,1491,1517]" italics="true" pageId="3" pageNumber="546">occultus</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD3C236ADB6F8CBB423FA50" bold="true" box="[385,1906,1491,1516]" pageId="3" pageNumber="546">103 48.4 21.5 43.4 15.9 47.1 59.0 25.8 17.2 13.0 6.7 24.2 6.9 6.5</emphasis>
95–108 41–45 13–18 45–49 56–61 25– 
<date id="843791D8FFD3C236A88AF8EEB78DF9B3" box="[1213,1244,1526,1551]" pageId="3" pageNumber="546" value="1927-12-16" valueMax="1927-12-19" valueMin="1927-12-16">27</date>
16–19 12–14 6.1–7.2 23–27 6.3–7.3 5.6–7.2 width (mean = 50%); covered in small granules which are intermixed with larger conical tubercles; skin of head coossified with skull; rostral much broader than tall (
<figureCitation id="68B2AB9DFFD5C230AEE6FAA0B047F86D" box="[721,790,1976,2001]" captionStart="FIGURE 2" captionStartId="5.[151,250,873,895]" captionTargetBox="[167,1435,193,835]" captionTargetId="figure@5.[167,1452,193,836]" captionTargetPageId="5" captionText="FIGURE 2. Saltuarius eximius sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/247608/files/figure.png" pageId="5" pageNumber="548">Fig. 2</figureCitation>
), fully divided by a single medial groove (
<figureCitation id="68B2AB9DFFD5C230A8CDFAA0B66EF86D" box="[1274,1343,1976,2001]" captionStart="FIGURE 2" captionStartId="5.[151,250,873,895]" captionTargetBox="[167,1435,193,835]" captionTargetId="figure@5.[167,1452,193,836]" captionTargetPageId="5" captionText="FIGURE 2. Saltuarius eximius sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/247608/files/figure.png" pageId="5" pageNumber="548">Fig. 2</figureCitation>
); rostral shield in contact with nostril (
<figureCitation id="68B2AB9DFFD5C230ADDCFAC4B160F849" box="[491,561,2012,2037]" captionStart="FIGURE 2" captionStartId="5.[151,250,873,895]" captionTargetBox="[167,1435,193,835]" captionTargetId="figure@5.[167,1452,193,836]" captionTargetPageId="5" captionText="FIGURE 2. Saltuarius eximius sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/247608/files/figure.png" pageId="5" pageNumber="548">Fig. 2</figureCitation>
); 5–6 scales bordering dorsal edge of rostral scale (mean = 5.3); ear opening elliptical, vertical, ~ ¼ as large as eye; supralabials 14–15 (mean = 14.3); infralabials 12. 
<emphasis id="C2FD6B0AFFD6C233A845FD8FB7E4FF0C" bold="true" box="[1138,1205,151,176]" italics="true" pageId="6" pageNumber="549">Neck:</emphasis>
Distinct and narrow, with large, sharp-tipped tubercules on dorsum. 
<emphasis id="C2FD6B0AFFD6C233AEA0FDA3B18BFF68" bold="true" box="[663,730,187,212]" italics="true" pageId="6" pageNumber="549">Body:</emphasis>
Narrow and elongate, covered in small granules; dorsal granules intermixed with larger conical, sharp-tipped tubercles; tubercles largest on back and upper flanks, and generally arranged as longitudinal rows (
<figureCitation id="68B2AB9DFFD6C233ADD8FC1CB165FEA0" box="[495,564,260,285]" captionStart="FIGURE 2" captionStartId="5.[151,250,873,895]" captionTargetBox="[167,1435,193,835]" captionTargetId="figure@5.[167,1452,193,836]" captionTargetPageId="5" captionText="FIGURE 2. Saltuarius eximius sp. nov. in life (photo: Conrad Hoskin)." httpUri="https://zenodo.org/record/247608/files/figure.png" pageId="6" pageNumber="549">Fig. 2</figureCitation>
); ventrolateral skin fold with a row of small, bluntly tipped tubercules; medial scales on belly considerably larger than the small outer granules that lie adjacent to the ventrolateral skin fold; a patch of enlarged, centrally placed scales is present in both the pectoral and pelvic regions; preanal pores absent in males; armpit with two pronounced skin flaps that form a pocket enclosing the axillary region (
<figureCitation id="68B2AB9DFFD6C233A86BFC77B7CFFE34" box="[1116,1182,367,392]" captionStart="FIGURE 6" captionStartId="9.[151,250,1972,1994]" captionTargetBox="[192,1410,193,1930]" captionTargetId="figure@9.[192,1434,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. Type series of S. eximius sp. nov.. Specimens (left-right): QMJ 92378 (paratype, male), QMJ 92377 (holotype, female), QMJ 92379 (paratype, female). (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247612/files/figure.png" pageId="6" pageNumber="549">Fig. 6</figureCitation>
), the lower of which is continuous with the ventrolateral fold. 
<emphasis id="C2FD6B0AFFD6C233AE7FFC8BB1C6FE10" bold="true" box="[584,663,403,428]" italics="true" pageId="6" pageNumber="549">Limbs:</emphasis>
Long and slender, covered in pronounced pointed tubercles dorsally; digits long and strongly compressed distally; enlarged subdigital lamellae 4th finger 19–21 (mean = 20); dorsal surface of fingers without enlarged conical tubercles extending along digits; enlarged subdigital lamellae 4th toe 18–20 (n = 3, mean = 19.3); dorsal surface of toes with enlarged conical tubercles on basal portion of all digits except the first digit; subdigital scales from tip of 4th finger to junction of 3rd and 4th fingers 19; subdigital scales from tip of 4th toe to junction of 3rd and 4th toes 21–22 (mean = 21.7). 
<emphasis id="C2FD6B0AFFD6C233AEDEFF4AB026FDD7" bold="true" box="[745,887,594,619]" italics="true" pageId="6" pageNumber="549">Original tail</emphasis>
(
<typeStatus id="2F3209BAFFD6C233AFBDFF4BB0BAFDD0" box="[906,1003,595,620]" pageId="6" pageNumber="549" type="holotype">holotype</typeStatus>
only; 
<figureCitation id="68B2AB9DFFD6C233A80EFF4BB72FFDD7" box="[1081,1150,595,620]" captionStart="FIGURE 3" captionStartId="5.[151,250,1873,1895]" captionTargetBox="[151,1436,945,1852]" captionTargetId="figure@5.[151,1436,945,1852]" captionTargetPageId="5" captionText="FIGURE 3. Original tail of S. eximius sp. nov. (QMJ 92377): (A) dorsal, (B) ventral (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247609/files/figure.png" pageId="6" pageNumber="549">Fig. 3</figureCitation>
): Broadly flared with an elaborate, undulating (‘frilled’) outer flange, and a short, fine attenuated tip; outer frill edged with small sharp spines; attenuated tip with very small spines on first 2/3rds of its length. 
<emphasis id="C2FD6B0AFFD6C233AF70FF85B0AFFD0A" bold="true" box="[839,1022,669,694]" italics="true" pageId="6" pageNumber="549">Regenerated tail</emphasis>
(
<typeStatus id="2F3209BAFFD6C233A83CFF86B729FD0A" box="[1035,1144,670,694]" pageId="6" pageNumber="549" type="paratype">paratypes</typeStatus>
; 
<figureCitation id="68B2AB9DFFD6C233A8B4FF85B79CFD0A" box="[1155,1229,669,694]" captionStart="FIGURE 1" captionStartId="4.[151,250,1989,2011]" captionTargetBox="[188,1355,193,1942]" captionTargetId="figure@4.[188,1356,193,1947]" captionTargetPageId="4" captionText="FIGURE 1. Saltuarius eximius sp. nov. in life: (A, B) original tail (QMJ 92377), (C) regenerated tail (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247607/files/figure.png" pageId="6" pageNumber="549">Figs. 1</figureCitation>
, 
<figureCitation id="68B2AB9DFFD6C233A8EDFF85B7BBFD0A" box="[1242,1258,669,694]" captionStart="FIGURE 6" captionStartId="9.[151,250,1972,1994]" captionTargetBox="[192,1410,193,1930]" captionTargetId="figure@9.[192,1434,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. Type series of S. eximius sp. nov.. Specimens (left-right): QMJ 92378 (paratype, male), QMJ 92377 (holotype, female), QMJ 92379 (paratype, female). (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247612/files/figure.png" pageId="6" pageNumber="549">6</figureCitation>
): Broadly flared and paddle-shaped, ‘pinched-in’ about 1/3rd of the way along its length and with a broadly rounded posterior margin. 
<emphasis id="C2FD6B0AFFD6C233ACA0FFF0B155FCBD" bold="true" box="[151,516,744,769]" pageId="6" pageNumber="549">Colour pattern in preservative</emphasis>
(
<figureCitation id="68B2AB9DFFD6C233AE23FFF0B108FCBD" box="[532,601,744,769]" captionStart="FIGURE 6" captionStartId="9.[151,250,1972,1994]" captionTargetBox="[192,1410,193,1930]" captionTargetId="figure@9.[192,1434,193,1932]" captionTargetPageId="9" captionText="FIGURE 6. Type series of S. eximius sp. nov.. Specimens (left-right): QMJ 92378 (paratype, male), QMJ 92377 (holotype, female), QMJ 92379 (paratype, female). (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247612/files/figure.png" pageId="6" pageNumber="549">Fig. 6</figureCitation>
): Dorsal base colour grey to tan, overlain with dark grey/brown markings. 
<emphasis id="C2FD6B0AFFD6C233ACA0FE14B38FFC99" bold="true" box="[151,222,780,805]" italics="true" pageId="6" pageNumber="549">Head:</emphasis>
Predominantly grey to tan; dark nuchal band in the supraoccipital region; labials pale with two broad dark bars (one at mid-snout, the other below the anterior orbit); dorsal surface of snout bridged by a series of dark chevrons; a dark, deep, V-shaped marking extending back medially between eyes. 
<emphasis id="C2FD6B0AFFD6C233A87BFE4CB7C1FCD1" bold="true" box="[1100,1168,852,877]" italics="true" pageId="6" pageNumber="549">Body:</emphasis>
Flanks with bold, dark mottling; dorsum with dark chevrons that merge with flank pattern and enclose four to five pale blotches between the pectoral and pelvic regions. 
<emphasis id="C2FD6B0AFFD6C233AE32FE84B1C4FC09" bold="true" box="[517,661,924,949]" italics="true" pageId="6" pageNumber="549">Original tail</emphasis>
(
<typeStatus id="2F3209BAFFD6C233AE91FE84B05EFC09" box="[678,783,924,949]" pageId="6" pageNumber="549" type="holotype">holotype</typeStatus>
): Alternating dark and light bands; basal region with two broad, poorly defined, pale bands and three broad dark bands; attenuated tip with two pale bands (anterior-most widest) and one dark band. 
<emphasis id="C2FD6B0AFFD6C233ADE0FEFCB1C4FC41" bold="true" box="[471,661,996,1021]" italics="true" pageId="6" pageNumber="549">Regenerated tail</emphasis>
(
<typeStatus id="2F3209BAFFD6C233AE91FEFDB048FC41" box="[678,793,997,1021]" pageId="6" pageNumber="549" type="paratype">paratypes</typeStatus>
): Pale with heavy dark grey and brown mottling. 
<emphasis id="C2FD6B0AFFD6C233A963FEFCB244FB9D" bold="true" italics="true" pageId="6" pageNumber="549">Limbs and digits:</emphasis>
Pale, with bold, well-defined, dark bands. 
<emphasis id="C2FD6B0AFFD6C233AF23F910B08CFB9D" bold="true" box="[788,989,1032,1057]" italics="true" pageId="6" pageNumber="549">Ventral surfaces:</emphasis>
<collectingCountry id="889EF788FFD6C233AFDFF910B770FB9D" box="[1000,1057,1032,1057]" name="China" pageId="6" pageNumber="549">Chin</collectingCountry>
, throat and trunk off-white with obscure brown mottling; limbs pale, peppered with dark flecks; original tail has broad dark and pale bands (dark bands are continuous with those on the dorsal surface but each is split in two; 
<figureCitation id="68B2AB9DFFD6C233AFC8F948B710FBD5" box="[1023,1089,1104,1129]" captionStart="FIGURE 3" captionStartId="5.[151,250,1873,1895]" captionTargetBox="[151,1436,945,1852]" captionTargetId="figure@5.[151,1436,945,1852]" captionTargetPageId="5" captionText="FIGURE 3. Original tail of S. eximius sp. nov. (QMJ 92377): (A) dorsal, (B) ventral (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247609/files/figure.png" pageId="6" pageNumber="549">Fig. 3</figureCitation>
); regenerated tail mottled (less pronounced than on dorsal surface), with a paler central region.
</paragraph>
<caption id="A4F6E790FFD4C231ACA0FADDB6CAF867" box="[151,1435,1989,2012]" httpUri="https://zenodo.org/record/247607/files/figure.png" pageId="4" pageNumber="547" targetBox="[188,1355,193,1942]" targetPageId="4">
<paragraph id="F036B718FFD4C231ACA0FADDB6CAF867" blockId="4.[151,1435,1989,2012]" box="[151,1435,1989,2012]" pageId="4" pageNumber="547">
<emphasis id="C2FD6B0AFFD4C231ACA0FADDB242F867" bold="true" box="[151,275,1989,2011]" pageId="4" pageNumber="547">FIGURE 1.</emphasis>
<taxonomicName id="3789CC9BFFD4C231AD2EFADDB285F860" box="[281,468,1989,2012]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="547" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD4C231AD2EFADDB285F860" box="[281,468,1989,2012]" italics="true" pageId="4" pageNumber="547">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD4C231ADECFADEB17AF867" bold="true" box="[475,555,1990,2011]" pageId="4" pageNumber="547">
<taxonomicNameLabel id="D9CED671FFD4C231ADECFADEB17AF867" box="[475,555,1990,2011]" pageId="4" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life: (A, B) original tail (QMJ92377), (C) regenerated tail (photos: Conrad Hoskin).
</paragraph>
</caption>
<caption id="A4F6E790FFD5C230ACA0FE71B03CFCC3" box="[151,877,873,895]" httpUri="https://zenodo.org/record/247608/files/figure.png" pageId="5" pageNumber="548" targetBox="[167,1435,193,835]" targetPageId="5">
<paragraph id="F036B718FFD5C230ACA0FE71B03CFCC3" blockId="5.[151,877,873,895]" box="[151,877,873,895]" pageId="5" pageNumber="548">
<emphasis id="C2FD6B0AFFD5C230ACA0FE71B242FCC2" bold="true" box="[151,275,873,895]" pageId="5" pageNumber="548">FIGURE 2.</emphasis>
<taxonomicName id="3789CC9BFFD5C230AD2EFE71B285FCC3" box="[281,468,873,895]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD5C230AD2EFE71B285FCC3" box="[281,468,873,895]" italics="true" pageId="5" pageNumber="548">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD5C230ADECFE71B17AFCC2" bold="true" box="[475,555,873,894]" pageId="5" pageNumber="548">
<taxonomicNameLabel id="D9CED671FFD5C230ADECFE71B17AFCC2" box="[475,555,873,894]" pageId="5" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in life (photo: Conrad Hoskin).
</paragraph>
</caption>
<caption id="A4F6E790FFD5C230ACA0FA49B7A4F8DB" box="[151,1269,1873,1895]" httpUri="https://zenodo.org/record/247609/files/figure.png" pageId="5" pageNumber="548" targetBox="[151,1436,945,1852]" targetPageId="5">
<paragraph id="F036B718FFD5C230ACA0FA49B7A4F8DB" blockId="5.[151,1269,1873,1895]" box="[151,1269,1873,1895]" pageId="5" pageNumber="548">
<emphasis id="C2FD6B0AFFD5C230ACA0FA49B242F8DA" bold="true" box="[151,275,1873,1895]" pageId="5" pageNumber="548">FIGURE 3.</emphasis>
Original tail of 
<taxonomicName id="3789CC9BFFD5C230AD82FA4AB14DF8DB" box="[437,540,1874,1895]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="548" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD5C230AD82FA4AB14DF8DB" box="[437,540,1874,1895]" italics="true" pageId="5" pageNumber="548">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD5C230AE14FA4AB122F8DB" bold="true" box="[547,627,1874,1895]" pageId="5" pageNumber="548">
<taxonomicNameLabel id="D9CED671FFD5C230AE14FA4AB122F8DB" box="[547,627,1874,1895]" pageId="5" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(QMJ92377): (A) dorsal, (B) ventral (photos: Conrad Hoskin).
</paragraph>
</caption>
<paragraph id="F036B718FFD6C233ACF0F980B65EFB44" blockId="6.[151,1437,151,2029]" pageId="6" pageNumber="549">
<emphasis id="C2FD6B0AFFD6C233ACF0F980B132FB0D" bold="true" box="[199,611,1176,1201]" pageId="6" pageNumber="549">Measurements of live individuals:</emphasis>
Four adults were measured in the field. Measurements of the 
<typeStatus id="2F3209BAFFD6C233A90DF980B6CDFB0D" box="[1338,1436,1176,1201]" pageId="6" pageNumber="549" type="holotype">holotype</typeStatus>
(QMJ92377) in life were: SVL 
<quantity id="37711AFDFFD6C233AE34F9A5B135FB68" box="[515,612,1213,1237]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" pageId="6" pageNumber="549" unit="mm" value="109.0">109 mm</quantity>
, WT 
<quantity id="37711AFDFFD6C233AE90F9A4B18CFB69" box="[679,733,1212,1237]" metricMagnitude="-2" metricUnit="kg" metricValue="1.6" pageId="6" pageNumber="549" unit="g" value="16.0">16 g</quantity>
. Measurements for 
<typeStatus id="2F3209BAFFD6C233AFF7F9A5B771FB69" box="[960,1056,1213,1237]" pageId="6" pageNumber="549" type="paratype">paratype</typeStatus>
QMJ92378 were: SVL 
<quantity id="37711AFDFFD6C233A901F9A5B6C6FB68" box="[1334,1431,1213,1237]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="549" unit="mm" value="120.0">120 mm</quantity>
, WT 
<quantity id="37711AFDFFD6C233ACFEF9F9B3ACFB45" box="[201,253,1249,1273]" metricMagnitude="-2" metricUnit="kg" metricValue="2.2" pageId="6" pageNumber="549" unit="g" value="22.0">22 g</quantity>
. Snout to vent lengths of two individuals that were not collected were: 
<quantity id="37711AFDFFD6C233A82DF9F9B727FB44" box="[1050,1142,1249,1273]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="6" pageNumber="549" unit="mm" value="118.0">118 mm</quantity>
and 
<quantity id="37711AFDFFD6C233A89BF9F9B65AFB44" box="[1196,1291,1249,1273]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" pageId="6" pageNumber="549" unit="mm" value="119.0">119 mm</quantity>
.
</paragraph>
<paragraph id="F036B718FFD6C233ACF0F81CB152F9A5" blockId="6.[151,1437,151,2029]" pageId="6" pageNumber="549">
<emphasis id="C2FD6B0AFFD6C233ACF0F81CB296FAA1" bold="true" box="[199,455,1284,1309]" pageId="6" pageNumber="549">Colour pattern in life</emphasis>
(
<figureCitation id="68B2AB9DFFD6C233ADE1F81CB112FAA1" box="[470,579,1284,1309]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="4.[151,250,1989,2011]" captionStartId-1="5.[151,250,873,895]" captionStartId-2="5.[151,250,1873,1895]" captionStartId-3="7.[151,250,1968,1990]" captionStartId-4="8.[151,250,1965,1987]" captionTargetBox-0="[188,1355,193,1942]" captionTargetBox-1="[167,1435,193,835]" captionTargetBox-2="[151,1436,945,1852]" captionTargetBox-3="[227,1337,193,1929]" captionTargetBox-4="[382,1209,193,1932]" captionTargetId-0="figure@4.[188,1356,193,1947]" captionTargetId-1="figure@5.[167,1452,193,836]" captionTargetId-2="figure@5.[151,1436,945,1852]" captionTargetId-3="figure@7.[227,1339,193,1932]" captionTargetId-4="figure@8.[381,1209,193,1932]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="8" captionText-0="FIGURE 1. Saltuarius eximius sp. nov. in life: (A, B) original tail (QMJ 92377), (C) regenerated tail (photos: Conrad Hoskin)." captionText-1="FIGURE 2. Saltuarius eximius sp. nov. in life (photo: Conrad Hoskin)." captionText-2="FIGURE 3. Original tail of S. eximius sp. nov. (QMJ 92377): (A) dorsal, (B) ventral (photos: Conrad Hoskin)." captionText-3="FIGURE 4. Comparison of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Jeff Wright)." captionText-4="FIGURE 5. Comparison of heads of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Steve Wilson)." httpUri-0="https://zenodo.org/record/247607/files/figure.png" httpUri-1="https://zenodo.org/record/247608/files/figure.png" httpUri-2="https://zenodo.org/record/247609/files/figure.png" httpUri-3="https://zenodo.org/record/247610/files/figure.png" httpUri-4="https://zenodo.org/record/247611/files/figure.png" pageId="6" pageNumber="549">Figs. 1–5</figureCitation>
): Dorsal colour an even mix of pale and dark markings; in some individuals this is a mix of white/pale grey and dark grey/black markings, while in others this is a mix of creamy brown and dark brown markings. Prominent markings include the following: a dark V behind the eyes; five pale bands along the back, with smaller pale markings in between; pale bands along the legs and fine pale bands on the digits; two broad, pale bands on the original tail and two white bands on the attenuated tail tip; and dark mottling on the regenerated tails. The iris is grey, tinged with yellowish green. Ventral surface is white but finely mottled with brown; underside of original tail white with soft brown bands; underside of regenerated tail pale in center but heavily mottled towards margin.
</paragraph>
<paragraph id="F036B718FFD6C23FACF0FB3CB7EBFA6C" blockId="6.[151,1437,151,2029]" lastBlockId="10.[151,1436,1211,2029]" lastPageId="10" lastPageNumber="553" pageId="6" pageNumber="549">
<emphasis id="C2FD6B0AFFD6C233ACF0FB3CB233F981" bold="true" box="[199,354,1572,1597]" pageId="6" pageNumber="549">Comparison.</emphasis>
<taxonomicName id="3789CC9BFFD6C233AD5BFB3DB111F980" box="[364,576,1573,1597]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD6C233AD5BFB3DB111F980" box="[364,576,1573,1597]" italics="true" pageId="6" pageNumber="549">Saltuarius eximius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9CED671FFD6C233AE7DFB3CB1F6F980" box="[586,679,1572,1597]" pageId="6" pageNumber="549" rank="species">
<emphasis id="C2FD6B0AFFD6C233AE7DFB3CB1F1F981" bold="true" box="[586,672,1572,1597]" pageId="6" pageNumber="549">sp. nov</emphasis>
.
</taxonomicNameLabel>
is a slender, long-bodied, long-legged, small-headed, large-eyed, large-tailed gecko which differs from all other Australian leaf-tailed geckos (
<taxonomicName id="3789CC9BFFD6C233AFCBFB51B71FF9DC" box="[1020,1102,1609,1632]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233AFCBFB51B71FF9DC" box="[1020,1102,1609,1632]" italics="true" pageId="6" pageNumber="549">Orraya</emphasis>
</taxonomicName>
, 
<taxonomicName id="3789CC9BFFD6C233A86AFB51B79BF9DD" box="[1117,1226,1609,1633]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233A86AFB51B79BF9DD" box="[1117,1226,1609,1633]" italics="true" pageId="6" pageNumber="549">Phyllurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="3789CC9BFFD6C233A935FB51B624F9DD" box="[1282,1397,1609,1633]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233A935FB51B624F9DD" box="[1282,1397,1609,1633]" italics="true" pageId="6" pageNumber="549">Saltuarius</emphasis>
</taxonomicName>
) in the following ways. It is distinguished from 
<taxonomicName id="3789CC9BFFD6C233AEAAFB75B1A1F938" box="[669,752,1645,1668]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233AEAAFB75B1A1F938" box="[669,752,1645,1668]" italics="true" pageId="6" pageNumber="549">Orraya</emphasis>
</taxonomicName>
(which it most resembles in general body form; 
<figureCitation id="68B2AB9DFFD6C233A919FB74B622F939" box="[1326,1395,1644,1669]" captionStart="FIGURE 4" captionStartId="7.[151,250,1968,1990]" captionTargetBox="[227,1337,193,1929]" captionTargetId="figure@7.[227,1339,193,1932]" captionTargetPageId="7" captionText="FIGURE 4. Comparison of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Jeff Wright)." httpUri="https://zenodo.org/record/247610/files/figure.png" pageId="6" pageNumber="549">Fig. 4</figureCitation>
) in possessing two lumbar vertebrae (
<emphasis id="C2FD6B0AFFD6C233AE2BFB89B162F914" box="[540,563,1681,1704]" italics="true" pageId="6" pageNumber="549">vs</emphasis>
three in 
<taxonomicName id="3789CC9BFFD6C233AEACFB89B1BCF914" box="[667,749,1681,1704]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233AEACFB89B1BCF914" box="[667,749,1681,1704]" italics="true" pageId="6" pageNumber="549">Orraya</emphasis>
</taxonomicName>
) (
<figureCitation id="68B2AB9DFFD6C233AF30FB88B01CF915" box="[775,845,1680,1705]" captionStart="FIGURE 7" captionStartId="10.[151,250,1114,1136]" captionTargetBox="[151,1436,193,1092]" captionTargetId="figure@10.[151,1436,193,1093]" captionTargetPageId="10" captionText="FIGURE 7. X-ray of S. eximius sp. nov. type series. Specimens (left-right): QMJ 92378 (paratype, male), QMJ 92377 (holotype, female), QMJ 92379 (paratype, female). (image: Mike Jeffery)." httpUri="https://zenodo.org/record/247613/files/figure.png" pageId="6" pageNumber="549">Fig. 7</figureCitation>
); by the shape of the cervical vertebrae (marginally elongated 
<emphasis id="C2FD6B0AFFD6C233AD3AFBADB275F970" box="[269,292,1717,1740]" italics="true" pageId="6" pageNumber="549">vs</emphasis>
distinctly elongated in 
<taxonomicName id="3789CC9BFFD6C233AE00FBADB1DBF970" box="[567,650,1717,1740]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233AE00FBADB1DBF970" box="[567,650,1717,1740]" italics="true" pageId="6" pageNumber="549">Orraya</emphasis>
</taxonomicName>
) (
<figureCitation id="68B2AB9DFFD6C233AE92FBACB1BCF971" box="[677,749,1716,1741]" captionStart="FIGURE 7" captionStartId="10.[151,250,1114,1136]" captionTargetBox="[151,1436,193,1092]" captionTargetId="figure@10.[151,1436,193,1093]" captionTargetPageId="10" captionText="FIGURE 7. X-ray of S. eximius sp. nov. type series. Specimens (left-right): QMJ 92378 (paratype, male), QMJ 92377 (holotype, female), QMJ 92379 (paratype, female). (image: Mike Jeffery)." httpUri="https://zenodo.org/record/247613/files/figure.png" pageId="6" pageNumber="549">Fig. 7</figureCitation>
); by the size of the preanal pores (male preanal pores very small 
<emphasis id="C2FD6B0AFFD6C233ACEAFBC1B3A5F94C" box="[221,244,1753,1776]" italics="true" pageId="6" pageNumber="549">vs</emphasis>
greatly enlarged in 
<taxonomicName id="3789CC9BFFD6C233ADEAFBC1B161F94C" box="[477,560,1753,1776]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233ADEAFBC1B161F94C" box="[477,560,1753,1776]" italics="true" pageId="6" pageNumber="549">Orraya</emphasis>
</taxonomicName>
); by the shape of the original tail (very large and broadly flared with a fine attenuated tip 
<emphasis id="C2FD6B0AFFD6C233AD0EFBE5B201F8A8" box="[313,336,1789,1812]" italics="true" pageId="6" pageNumber="549">vs</emphasis>
small and leaf-shaped in 
<taxonomicName id="3789CC9BFFD6C233AE40FBE5B19BF8A8" box="[631,714,1789,1812]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233AE40FBE5B19BF8A8" box="[631,714,1789,1812]" italics="true" pageId="6" pageNumber="549">Orraya</emphasis>
</taxonomicName>
); and by the spinosity of the regenerated tail (smooth-edged 
<emphasis id="C2FD6B0AFFD6C233A9B3FBE5B6CAF8A8" box="[1412,1435,1789,1812]" italics="true" pageId="6" pageNumber="549">vs</emphasis>
enlarged spines along margin in 
<taxonomicName id="3789CC9BFFD6C233AE2FFA39B13BF884" box="[536,618,1825,1848]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233AE2FFA39B13BF884" box="[536,618,1825,1848]" italics="true" pageId="6" pageNumber="549">Orraya</emphasis>
</taxonomicName>
) (
<figureCitation id="68B2AB9DFFD6C233AEB0FA38B19EF885" box="[647,719,1824,1849]" captionStart="FIGURE 1" captionStartId="4.[151,250,1989,2011]" captionTargetBox="[188,1355,193,1942]" captionTargetId="figure@4.[188,1356,193,1947]" captionTargetPageId="4" captionText="FIGURE 1. Saltuarius eximius sp. nov. in life: (A, B) original tail (QMJ 92377), (C) regenerated tail (photos: Conrad Hoskin)." httpUri="https://zenodo.org/record/247607/files/figure.png" pageId="6" pageNumber="549">Figs 1</figureCitation>
C, 4C). 
<taxonomicName id="3789CC9BFFD6C233AF04FA39B756F884" box="[819,1031,1825,1849]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD6C233AF04FA39B756F884" box="[819,1031,1825,1849]" italics="true" pageId="6" pageNumber="549">Saltuarius eximius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9CED671FFD6C233A825FA38B721F884" box="[1042,1136,1824,1849]" pageId="6" pageNumber="549" rank="species">
<emphasis id="C2FD6B0AFFD6C233A825FA38B738F885" bold="true" box="[1042,1129,1824,1849]" pageId="6" pageNumber="549">sp. nov</emphasis>
.
</taxonomicNameLabel>
is further separated from 
<taxonomicName id="3789CC9BFFD6C233ACA0FA5DB3BBF8E0" box="[151,234,1861,1884]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233ACA0FA5DB3BBF8E0" box="[151,234,1861,1884]" italics="true" pageId="6" pageNumber="549">Orraya</emphasis>
</taxonomicName>
in having a longer and narrower body, longer limbs, a shorter head and snout, a shorter neck, and larger eyes (non-overlapping differences, Table 1; 
<figureCitation id="68B2AB9DFFD6C233AEB5FA70B198F83D" box="[642,713,1896,1921]" captionStart="FIGURE 4" captionStartId="7.[151,250,1968,1990]" captionTargetBox="[227,1337,193,1929]" captionTargetId="figure@7.[227,1339,193,1932]" captionTargetPageId="7" captionText="FIGURE 4. Comparison of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Jeff Wright)." httpUri="https://zenodo.org/record/247610/files/figure.png" pageId="6" pageNumber="549">Figs 4</figureCitation>
, 
<figureCitation id="68B2AB9DFFD6C233AEE3FA70B1B5F83D" box="[724,740,1896,1921]" captionStart="FIGURE 5" captionStartId="8.[151,250,1965,1987]" captionTargetBox="[382,1209,193,1932]" captionTargetId="figure@8.[381,1209,193,1932]" captionTargetPageId="8" captionText="FIGURE 5. Comparison of heads of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Steve Wilson)." httpUri="https://zenodo.org/record/247611/files/figure.png" pageId="6" pageNumber="549">5</figureCitation>
). 
<taxonomicName id="3789CC9BFFD6C233AECEFA71B099F83C" box="[761,968,1897,1921]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD6C233AECEFA71B099F83C" box="[761,968,1897,1921]" italics="true" pageId="6" pageNumber="549">Saltuarius eximius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D9CED671FFD6C233AFF9FA70B776F83C" box="[974,1063,1896,1921]" pageId="6" pageNumber="549" rank="species">
<emphasis id="C2FD6B0AFFD6C233AFF9FA70B771F83D" bold="true" box="[974,1056,1896,1921]" pageId="6" pageNumber="549">sp. nov</emphasis>
.
</taxonomicNameLabel>
differs from 
<taxonomicName id="3789CC9BFFD6C233A88DFA71B676F83D" box="[1210,1319,1897,1921]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233A88DFA71B676F83D" box="[1210,1319,1897,1921]" italics="true" pageId="6" pageNumber="549">Phyllurus</emphasis>
</taxonomicName>
in that the rostral contacts the nostril (
<emphasis id="C2FD6B0AFFD6C233ADE6FA95B2B9F818" box="[465,488,1933,1956]" italics="true" pageId="6" pageNumber="549">vs</emphasis>
nostril not in contact with rostral in 
<taxonomicName id="3789CC9BFFD6C233AFA7FA95B0ACF819" box="[912,1021,1933,1957]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233AFA7FA95B0ACF819" box="[912,1021,1933,1957]" italics="true" pageId="6" pageNumber="549">Phyllurus</emphasis>
</taxonomicName>
), male preanal pores are present (
<emphasis id="C2FD6B0AFFD6C233A9B3FA95B6CAF818" box="[1412,1435,1933,1956]" italics="true" pageId="6" pageNumber="549">vs</emphasis>
absent in 
<taxonomicName id="3789CC9BFFD6C233AD3DFAA9B226F875" box="[266,375,1969,1993]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233AD3DFAA9B226F875" box="[266,375,1969,1993]" italics="true" pageId="6" pageNumber="549">Phyllurus</emphasis>
</taxonomicName>
), and the original tail is strongly flared with an elaborate spinose outer flange (
<emphasis id="C2FD6B0AFFD6C233A913FAA9B66AF874" box="[1316,1339,1969,1992]" italics="true" pageId="6" pageNumber="549">vs</emphasis>
without elaborate outer flange in 
<taxonomicName id="3789CC9BFFD6C233AD8BFACDB178F851" box="[444,553,2005,2029]" class="Reptilia" family="Carphodactylidae" genus="Phyllurus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233AD8BFACDB178F851" box="[444,553,2005,2029]" italics="true" pageId="6" pageNumber="549">Phyllurus</emphasis>
</taxonomicName>
). 
<taxonomicName id="3789CC9BFFD6C233AE7FFACDB046F851" box="[584,791,2005,2029]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD6C233AE7FFACDB046F851" box="[584,791,2005,2029]" italics="true" pageId="6" pageNumber="549">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD6C233AF29FACCB029F851" bold="true" box="[798,888,2004,2029]" pageId="6" pageNumber="549">
<taxonomicNameLabel id="D9CED671FFD6C233AF29FACCB029F851" box="[798,888,2004,2029]" pageId="6" pageNumber="549" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from all 
<taxonomicName id="3789CC9BFFD6C233A89DFACDB64CF851" box="[1194,1309,2005,2029]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="549" phylum="Chordata" rank="genus">
<emphasis id="C2FD6B0AFFD6C233A89DFACDB64CF851" box="[1194,1309,2005,2029]" italics="true" pageId="6" pageNumber="549">Saltuarius</emphasis>
</taxonomicName>
species in having a longer and more slender body, longer limbs, a shorter and narrower head, a shorter snout, a shorter temporal region, a narrower neck, and larger eyes (Table 1). 
<taxonomicName id="3789CC9BFFDAC23FAF5EF9F8B76DFB4B" box="[873,1084,1248,1272]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFDAC23FAF5EF9F8B76DFB4B" box="[873,1084,1248,1272]" italics="true" pageId="10" pageNumber="553">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFDAC23FA87FF9C7B7F7FB44" bold="true" box="[1096,1190,1247,1272]" pageId="10" pageNumber="553">
<taxonomicNameLabel id="D9CED671FFDAC23FA87FF9C7B7F7FB44" box="[1096,1190,1247,1272]" pageId="10" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also differs from all congeners in having an unpatterned grey eye (
<emphasis id="C2FD6B0AFFDAC23FAE94F81CB1EBFAA7" box="[675,698,1284,1307]" italics="true" pageId="10" pageNumber="553">vs</emphasis>
patterned) (
<figureCitation id="68B2AB9DFFDAC23FAF73F81BB0D8FAA0" box="[836,905,1283,1308]" captionStart="FIGURE 5" captionStartId="8.[151,250,1965,1987]" captionTargetBox="[382,1209,193,1932]" captionTargetId="figure@8.[381,1209,193,1932]" captionTargetPageId="8" captionText="FIGURE 5. Comparison of heads of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Steve Wilson)." httpUri="https://zenodo.org/record/247611/files/figure.png" pageId="10" pageNumber="553">Fig. 5</figureCitation>
). It is most closely related to 
<taxonomicName id="3789CC9BFFDAC23FA8ECF81CB60BFAA7" box="[1243,1370,1284,1307]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="cornutus">
<emphasis id="C2FD6B0AFFDAC23FA8ECF81CB60BFAA7" box="[1243,1370,1284,1307]" italics="true" pageId="10" pageNumber="553">S. cornutus</emphasis>
</taxonomicName>
(
<figureCitation id="68B2AB9DFFDAC23FA95DF81BB3F6FAFC" captionStart="FIGURE 4" captionStartId="7.[151,250,1968,1990]" captionTargetBox="[227,1337,193,1929]" captionTargetId="figure@7.[227,1339,193,1932]" captionTargetPageId="7" captionText="FIGURE 4. Comparison of: (A) S. eximius sp. nov. (QMJ 92377), (B) S. cornutus, and (C) Orraya occultus (photos: A, B, Conrad Hoskin; C, Jeff Wright)." httpUri="https://zenodo.org/record/247610/files/figure.png" pageId="10" pageNumber="553">Figs 4</figureCitation>
B, 5B) and 
<taxonomicName id="3789CC9BFFDAC23FAD1BF830B293FAFC" box="[300,450,1320,1344]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="salebrosus">
<emphasis id="C2FD6B0AFFDAC23FAD1BF830B293FAFC" box="[300,450,1320,1344]" italics="true" pageId="10" pageNumber="553">S. salebrosus</emphasis>
</taxonomicName>
, with which it shares preanal pores in males. In addition to the aforementioned traits, 
<taxonomicName id="3789CC9BFFDAC23FACA0F854B236FADF" box="[151,359,1356,1380]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFDAC23FACA0F854B236FADF" box="[151,359,1356,1380]" italics="true" pageId="10" pageNumber="553">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFDAC23FAD59F853B299FAD8" bold="true" box="[366,456,1355,1380]" pageId="10" pageNumber="553">
<taxonomicNameLabel id="D9CED671FFDAC23FAD59F853B299FAD8" box="[366,456,1355,1380]" pageId="10" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also smaller than both these species (max SVL = 
<quantity id="37711AFDFFDAC23FA82FF854B724FADF" box="[1048,1141,1356,1380]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="10" pageNumber="553" unit="mm" value="118.0">118 mm</quantity>
<emphasis id="C2FD6B0AFFDAC23FA84BF854B7C2FADF" box="[1148,1171,1356,1379]" italics="true" pageId="10" pageNumber="553">vs</emphasis>
<quantity id="37711AFDFFDAC23FA8ACF853B7A9FADF" box="[1179,1272,1355,1380]" metricMagnitude="-1" metricUnit="m" metricValue="1.3599999999999999" pageId="10" pageNumber="553" unit="mm" value="136.0">136 mm</quantity>
in 
<taxonomicName id="3789CC9BFFDAC23FA92AF854B6CDFADF" box="[1309,1436,1356,1379]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="cornutus">
<emphasis id="C2FD6B0AFFDAC23FA92AF854B6CDFADF" box="[1309,1436,1356,1379]" italics="true" pageId="10" pageNumber="553">S. cornutus</emphasis>
</taxonomicName>
and 
<quantity id="37711AFDFFDAC23FACFFF868B279FA3B" box="[200,296,1392,1416]" metricMagnitude="-1" metricUnit="m" metricValue="1.44" pageId="10" pageNumber="553" unit="mm" value="144.0">144 mm</quantity>
in 
<taxonomicName id="3789CC9BFFDAC23FAD78F868B2B5FA34" box="[335,484,1392,1416]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="salebrosus">
<emphasis id="C2FD6B0AFFDAC23FAD78F868B2B5FA34" box="[335,484,1392,1416]" italics="true" pageId="10" pageNumber="553">S. salebrosus</emphasis>
</taxonomicName>
). 
<taxonomicName id="3789CC9BFFDAC23FADC9F868B181FA3B" box="[510,720,1392,1416]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFDAC23FADC9F868B181FA3B" box="[510,720,1392,1416]" italics="true" pageId="10" pageNumber="553">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFDAC23FAEEEF877B064FA34" bold="true" box="[729,821,1391,1416]" pageId="10" pageNumber="553">
<taxonomicNameLabel id="D9CED671FFDAC23FAEEEF877B064FA34" box="[729,821,1391,1416]" pageId="10" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is further distinguished from the southern ‘ 
<taxonomicName id="3789CC9BFFDAC23FA91CF868B6C3FA3B" box="[1323,1426,1392,1415]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="swaini">
<emphasis id="C2FD6B0AFFDAC23FA91CF868B6C3FA3B" box="[1323,1426,1392,1415]" italics="true" pageId="10" pageNumber="553">S. swaini</emphasis>
</taxonomicName>
’ species group in that the males possess preanal pores (
<emphasis id="C2FD6B0AFFDAC23FAF33F88CB04AFA17" box="[772,795,1428,1451]" italics="true" pageId="10" pageNumber="553">vs</emphasis>
preanal pores absent in 
<taxonomicName id="3789CC9BFFDAC23FA805F88CB7C8FA17" box="[1074,1177,1428,1451]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="swaini">
<emphasis id="C2FD6B0AFFDAC23FA805F88CB7C8FA17" box="[1074,1177,1428,1451]" italics="true" pageId="10" pageNumber="553">S. swaini</emphasis>
</taxonomicName>
, 
<taxonomicName id="3789CC9BFFDAC23FA89FF88CB674FA10" box="[1192,1317,1428,1452]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="wyberba">
<emphasis id="C2FD6B0AFFDAC23FA89FF88CB674FA10" box="[1192,1317,1428,1452]" italics="true" pageId="10" pageNumber="553">S. wyberba</emphasis>
</taxonomicName>
, 
<taxonomicName id="3789CC9BFFDAC23FA902F88CB6CDFA10" box="[1333,1436,1428,1452]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="kateae">
<emphasis id="C2FD6B0AFFDAC23FA902F88CB6CDFA10" box="[1333,1436,1428,1452]" italics="true" pageId="10" pageNumber="553">S. kateae</emphasis>
</taxonomicName>
and 
<taxonomicName id="3789CC9BFFDAC23FACF1F8A0B260FA73" box="[198,305,1464,1487]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="moritzi">
<emphasis id="C2FD6B0AFFDAC23FACF1F8A0B260FA73" box="[198,305,1464,1487]" italics="true" pageId="10" pageNumber="553">S. moritzi</emphasis>
</taxonomicName>
). 
<taxonomicName id="3789CC9BFFDAC23FAD7EF8A0B148FA73" box="[329,537,1464,1488]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFDAC23FAD7EF8A0B148FA73" box="[329,537,1464,1488]" italics="true" pageId="10" pageNumber="553">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFDAC23FAE28F8AFB128FA6C" bold="true" box="[543,633,1463,1488]" pageId="10" pageNumber="553">
<taxonomicNameLabel id="D9CED671FFDAC23FAE28F8AFB128FA6C" box="[543,633,1463,1488]" pageId="10" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
does not co-occur with any other leaf-tailed gecko.
</paragraph>
<caption id="A4F6E790FFD7C232ACA0FAA8B28FF858" httpUri="https://zenodo.org/record/247610/files/figure.png" pageId="7" pageNumber="550" targetBox="[227,1337,193,1929]" targetPageId="7">
<paragraph id="F036B718FFD7C232ACA0FAA8B28FF858" blockId="7.[151,1436,1968,2020]" pageId="7" pageNumber="550">
<emphasis id="C2FD6B0AFFD7C232ACA0FAA8B244F879" bold="true" box="[151,277,1968,1990]" pageId="7" pageNumber="550">FIGURE 4.</emphasis>
Comparison of: (A) 
<taxonomicName id="3789CC9BFFD7C232ADC4FAA9B10DF87A" box="[499,604,1969,1990]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="550" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD7C232ADC4FAA9B10DF87A" box="[499,604,1969,1990]" italics="true" pageId="7" pageNumber="550">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD7C232AE52FAA8B1E9F879" bold="true" box="[613,696,1968,1989]" pageId="7" pageNumber="550">
<taxonomicNameLabel id="D9CED671FFD7C232AE52FAA8B1E9F879" box="[613,696,1968,1989]" pageId="7" pageNumber="550" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(QMJ92377), (B) 
<taxonomicName id="3789CC9BFFD7C232AF4AFAA9B0A0F87A" box="[893,1009,1969,1990]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="550" phylum="Chordata" rank="species" species="cornutus">
<emphasis id="C2FD6B0AFFD7C232AF4AFAA9B0A0F87A" box="[893,1009,1969,1990]" italics="true" pageId="7" pageNumber="550">S. cornutus</emphasis>
</taxonomicName>
, and (C) 
<taxonomicName id="3789CC9BFFD7C232A86EFAA9B651F87A" box="[1113,1280,1968,1990]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="550" phylum="Chordata" rank="species" species="occultus">
<emphasis id="C2FD6B0AFFD7C232A86EFAA9B651F87A" box="[1113,1280,1968,1990]" italics="true" pageId="7" pageNumber="550">Orraya occultus</emphasis>
</taxonomicName>
(photos: A, B, Conrad Hoskin; C, Jeff Wright).
</paragraph>
</caption>
<caption id="A4F6E790FFD8C23DACA0FAB5B17CF85E" httpUri="https://zenodo.org/record/247611/files/figure.png" pageId="8" pageNumber="551" targetBox="[382,1209,193,1932]" targetPageId="8">
<paragraph id="F036B718FFD8C23DACA0FAB5B17CF85E" blockId="8.[151,1437,1965,2018]" pageId="8" pageNumber="551">
<emphasis id="C2FD6B0AFFD8C23DACA0FAB5B242F87F" bold="true" box="[151,275,1965,1987]" pageId="8" pageNumber="551">FIGURE 5.</emphasis>
Comparison of heads of: (A) 
<taxonomicName id="3789CC9BFFD8C23DAE70FAB6B1FFF878" box="[583,686,1966,1988]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD8C23DAE70FAB6B1FFF878" box="[583,686,1966,1988]" italics="true" pageId="8" pageNumber="551">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD8C23DAE81FAB6B056F87F" bold="true" box="[694,775,1966,1987]" pageId="8" pageNumber="551">
<taxonomicNameLabel id="D9CED671FFD8C23DAE81FAB6B056F87F" box="[694,775,1966,1987]" pageId="8" pageNumber="551" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(QMJ92377), (B) 
<taxonomicName id="3789CC9BFFD8C23DAFF0FAB6B76BF878" box="[967,1082,1966,1988]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="cornutus">
<emphasis id="C2FD6B0AFFD8C23DAFF0FAB6B76BF878" box="[967,1082,1966,1988]" italics="true" pageId="8" pageNumber="551">S. cornutus</emphasis>
</taxonomicName>
, and (C) 
<taxonomicName id="3789CC9BFFD8C23DA8ABFAB6B610F87F" box="[1180,1345,1965,1987]" class="Reptilia" family="Carphodactylidae" genus="Orraya" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="551" phylum="Chordata" rank="species" species="occultus">
<emphasis id="C2FD6B0AFFD8C23DA8ABFAB6B610F87F" box="[1180,1345,1965,1987]" italics="true" pageId="8" pageNumber="551">Orraya occultus</emphasis>
</taxonomicName>
(photos: A, B, Conrad Hoskin; C, Steve Wilson).
</paragraph>
</caption>
<caption id="A4F6E790FFD9C23CACA0FAACB063F854" httpUri="https://zenodo.org/record/247612/files/figure.png" pageId="9" pageNumber="552" targetBox="[192,1410,193,1930]" targetPageId="9">
<paragraph id="F036B718FFD9C23CACA0FAACB063F854" blockId="9.[151,1436,1972,2024]" pageId="9" pageNumber="552">
<emphasis id="C2FD6B0AFFD9C23CACA0FAACB247F875" bold="true" box="[151,278,1972,1994]" pageId="9" pageNumber="552">FIGURE 6.</emphasis>
Type series of 
<taxonomicName id="3789CC9BFFD9C23CAD8BFAACB177F876" box="[444,550,1972,1994]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="552" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFD9C23CAD8BFAACB177F876" box="[444,550,1972,1994]" italics="true" pageId="9" pageNumber="552">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFD9C23CAE18FAACB1D3F875" bold="true" box="[559,642,1972,1993]" pageId="9" pageNumber="552">
<taxonomicNameLabel id="D9CED671FFD9C23CAE18FAACB1D3F875" box="[559,642,1972,1993]" pageId="9" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Specimens (left-right): QMJ92378 (paratype, male), QMJ92377 (holotype, female), QMJ92379 (paratype, female). (photos: Conrad Hoskin).
</paragraph>
</caption>
<caption id="A4F6E790FFDAC23FACA0F942B0D3FB33" httpUri="https://zenodo.org/record/247613/files/figure.png" pageId="10" pageNumber="553" targetBox="[151,1436,193,1092]" targetPageId="10">
<paragraph id="F036B718FFDAC23FACA0F942B0D3FB33" blockId="10.[151,1435,1114,1167]" pageId="10" pageNumber="553">
<emphasis id="C2FD6B0AFFDAC23FACA0F942B248FBCC" bold="true" box="[151,281,1114,1136]" pageId="10" pageNumber="553">FIGURE 7.</emphasis>
X-ray of 
<taxonomicName id="3789CC9BFFDAC23FADB8F943B2ADFBCC" box="[399,508,1115,1136]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFDAC23FADB8F943B2ADFBCC" box="[399,508,1115,1136]" italics="true" pageId="10" pageNumber="553">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFDAC23FAE3EF943B131FBCC" bold="true" box="[521,608,1115,1136]" pageId="10" pageNumber="553">
<taxonomicNameLabel id="D9CED671FFDAC23FAE3EF943B131FBCC" box="[521,608,1115,1136]" pageId="10" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
type series. Specimens (left-right): QMJ92378 (paratype, male), QMJ92377 (holotype, female), QMJ92379 (paratype, female). (image: Mike Jeffery).
</paragraph>
</caption>
<paragraph id="F036B718FFDAC23FACF0F8C3B795F980" blockId="10.[151,1436,1211,2029]" pageId="10" pageNumber="553">
<emphasis id="C2FD6B0AFFDAC23FACF0F8C3B264FA48" bold="true" box="[199,309,1499,1524]" pageId="10" pageNumber="553">Genetics.</emphasis>
12S/cyt- 
<emphasis id="C2FD6B0AFFDAC23FADACF8C4B2F8FA48" box="[411,425,1500,1524]" italics="true" pageId="10" pageNumber="553">b</emphasis>
mtDNA data places 
<taxonomicName id="3789CC9BFFDAC23FAEA3F8C4B054FA4F" box="[660,773,1500,1523]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFDAC23FAEA3F8C4B054FA4F" box="[660,773,1500,1523]" italics="true" pageId="10" pageNumber="553">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFDAC23FAF3AF8C3B036FA48" bold="true" box="[781,871,1499,1524]" pageId="10" pageNumber="553">
<taxonomicNameLabel id="D9CED671FFDAC23FAF3AF8C3B036FA48" box="[781,871,1499,1524]" pageId="10" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
as a divergent (9.1%) sister-species to 
<taxonomicName id="3789CC9BFFDAC23FA929F8C4B6CDFA4F" box="[1310,1436,1500,1523]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="cornutus">
<emphasis id="C2FD6B0AFFDAC23FA929F8C4B6CDFA4F" box="[1310,1436,1500,1523]" italics="true" pageId="10" pageNumber="553">S. cornutus</emphasis>
</taxonomicName>
(Hoskin, unpub. data). 12S and cyt 
<emphasis id="C2FD6B0AFFDAC23FAE16FB18B17EF9A4" box="[545,559,1536,1560]" italics="true" pageId="10" pageNumber="553">b</emphasis>
sequences for two individuals are deposited on GenBank (12S: QMJ92378 = 
<accessionNumber id="EFDA2AFBFFDAC23FACA0FB3BB240F980" box="[151,273,1571,1596]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF553907" pageId="10" pageNumber="553">KF553907</accessionNumber>
, QMJ92379 = 
<accessionNumber id="EFDA2AFBFFDAC23FAD8FFB3BB165F980" box="[440,564,1571,1596]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF553908" pageId="10" pageNumber="553">KF553908</accessionNumber>
; cyt 
<emphasis id="C2FD6B0AFFDAC23FAE56FB3CB13EF980" box="[609,623,1572,1596]" italics="true" pageId="10" pageNumber="553">b</emphasis>
: QMJ92378 = 
<accessionNumber id="EFDA2AFBFFDAC23FAF2DFB3BB0C5F980" box="[794,916,1571,1596]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF553909" pageId="10" pageNumber="553">KF553909</accessionNumber>
, QMJ92379 = 
<accessionNumber id="EFDA2AFBFFDAC23FA80CFB3BB7E9F980" box="[1083,1208,1571,1596]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF553910" pageId="10" pageNumber="553">KF553910</accessionNumber>
).
</paragraph>
</subSubSection>
<subSubSection id="B893E493FFDAC23EACF0FB5FB7CFFF69" lastPageId="11" lastPageNumber="554" pageId="10" pageNumber="553" type="distribution">
<paragraph id="F036B718FFDAC23FACF0FB5FB785F938" blockId="10.[151,1436,1211,2029]" pageId="10" pageNumber="553">
<emphasis id="C2FD6B0AFFDAC23FACF0FB5FB231F9DC" bold="true" box="[199,352,1607,1632]" pageId="10" pageNumber="553">Distribution.</emphasis>
Known only from the vicinity of the 
<typeStatus id="2F3209BAFFDAC23FAF22FB50B014F9DC" box="[789,837,1608,1632]" pageId="10" pageNumber="553">type</typeStatus>
locality in the uplands of the Melville Range, Cape Melville, north-east 
<collectingCountry id="889EF788FFDAC23FAD4CFB73B2B3F938" box="[379,482,1643,1668]" name="Australia" pageId="10" pageNumber="553">Australia</collectingCountry>
(
<figureCitation id="68B2AB9DFFDAC23FADC6FB73B164F938" box="[497,565,1643,1668]" captionStart="FIGURE 8" captionStartId="11.[151,250,1609,1631]" captionTargetBox="[268,1339,254,1591]" captionTargetId="figure@11.[268,1339,254,1592]" captionTargetPageId="11" captionText="FIGURE 8. Map of north-east Queensland, showing locality of Cape Melville, McIlwraith Range and the northern Wet Tropics, and the leaf-tailed gecko present in each of these areas. The inset shows Australia. Background image is from Google Maps (2013)." httpUri="https://zenodo.org/record/247614/files/figure.png" pageId="10" pageNumber="553">Fig. 8</figureCitation>
). The distribution falls within Cape Melville National Park.
</paragraph>
<paragraph id="F036B718FFDAC23EACF0FB97B7CFFF69" blockId="10.[151,1436,1211,2029]" lastBlockId="11.[151,1436,151,213]" lastPageId="11" lastPageNumber="554" pageId="10" pageNumber="553">
<emphasis id="C2FD6B0AFFDAC23FACF0FB97B2E5F914" bold="true" box="[199,436,1679,1704]" pageId="10" pageNumber="553">Habitat and habits.</emphasis>
Six individuals have been found, all in close proximity in an area of granite boulders covered by a rainforest canopy (
<figureCitation id="68B2AB9DFFDAC23FAE3CFBABB103F970" box="[523,594,1715,1740]" captionStart="FIGURE 9" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[150,1423,225,1902]" captionTargetId="figure@12.[139,1424,193,1904]" captionTargetPageId="12" captionText="FIGURE 9. Habitat of S. eximius sp. nov., Cape Melville Range (photos: A, Conrad Hoskin; B, Tim Laman)." httpUri="https://zenodo.org/record/247615/files/figure.png" pageId="10" pageNumber="553">Fig. 9</figureCitation>
). 
<taxonomicName id="3789CC9BFFDAC23FAE5EFBACB06AF977" box="[617,827,1716,1740]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFDAC23FAE5EFBACB06AF977" box="[617,827,1716,1740]" italics="true" pageId="10" pageNumber="553">Saltuarius eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFDAC23FAF72FBABB0F0F970" bold="true" box="[837,929,1715,1740]" pageId="10" pageNumber="553">
<taxonomicNameLabel id="D9CED671FFDAC23FAF72FBABB0F0F970" box="[837,929,1715,1740]" pageId="10" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was not found in searches of other areas of rocky rainforest in the general vicinity of the 
<typeStatus id="2F3209BAFFDAC23FAE86FBC0B1B0F94C" box="[689,737,1752,1776]" pageId="10" pageNumber="553">type</typeStatus>
locality or in a lowland area of Cape Melville to the south (
<geoCoordinate id="95BDD1DFFFDAC23FACA8FBE3B279F8A8" box="[159,296,1787,1812]" direction="south" orientation="latitude" pageId="10" pageNumber="553" precision="15" value="-14.315001">14°18'54&quot; S</geoCoordinate>
, 
<geoCoordinate id="95BDD1DFFFDAC23FAD02FBE3B29CF8A8" box="[309,461,1787,1812]" direction="east" orientation="longitude" pageId="10" pageNumber="553" precision="15" value="144.49834">144°29'54&quot; E</geoCoordinate>
). Nor was the species found in searches of exposed boulder-fields in the vicinity of the 
<typeStatus id="2F3209BAFFDAC23FACA0FA39B396F885" box="[151,199,1825,1849]" pageId="10" pageNumber="553">type</typeStatus>
locality (as seen in the background of 
<figureCitation id="68B2AB9DFFDAC23FAE42FA07B1E6F884" box="[629,695,1823,1848]" captionStart="FIGURE 9" captionStartId="12.[151,250,1953,1975]" captionTargetBox="[150,1423,225,1902]" captionTargetId="figure@12.[139,1424,193,1904]" captionTargetPageId="12" captionText="FIGURE 9. Habitat of S. eximius sp. nov., Cape Melville Range (photos: A, Conrad Hoskin; B, Tim Laman)." httpUri="https://zenodo.org/record/247615/files/figure.png" pageId="10" pageNumber="553">Fig. 9</figureCitation>
A). At the 
<typeStatus id="2F3209BAFFDAC23FAF1AFA39B00CF885" box="[813,861,1825,1849]" pageId="10" pageNumber="553">type</typeStatus>
locality, individuals were found at night on vertical surfaces of boulders or trees, between one and five meters above the ground. When first sighted the geckos were motionless and positioned head-down, with the anterior half of their body raised well off the surface and their head held parallel with the surface. In this foraging pose, 
<taxonomicName id="3789CC9BFFDAC23FAEEEFA94B018F81F" box="[729,841,1932,1955]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFDAC23FAEEEFA94B018F81F" box="[729,841,1932,1955]" italics="true" pageId="10" pageNumber="553">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFDAC23FAF67FA93B0F8F818" bold="true" box="[848,937,1931,1956]" pageId="10" pageNumber="553">
<taxonomicNameLabel id="D9CED671FFDAC23FAF67FA93B0F8F818" box="[848,937,1931,1956]" pageId="10" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is no doubt primarily a sit-and-wait predator like other leaftail geckos. Several of the geckos made an extended squeaking sound when first captured. The only other gecko species found in micro-sympatry with 
<taxonomicName id="3789CC9BFFDAC23FAED7FACCB002F850" box="[736,851,2004,2028]" class="Reptilia" family="Carphodactylidae" genus="Saltuarius" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="eximius" status="sp. nov.">
<emphasis id="C2FD6B0AFFDAC23FAED7FACCB002F850" box="[736,851,2004,2028]" italics="true" pageId="10" pageNumber="553">S. eximius</emphasis>
</taxonomicName>
<emphasis id="C2FD6B0AFFDAC23FAF6AFACCB0E8F851" bold="true" box="[861,953,2004,2029]" pageId="10" pageNumber="553">
<taxonomicNameLabel id="D9CED671FFDAC23FAF6AFACCB0E8F851" box="[861,953,2004,2029]" pageId="10" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was 
<taxonomicName id="3789CC9BFFDAC23EAFC1FACCB24DFF0C" authority="Lucas &amp; Frost 1900" authorityName="Lucas &amp; Frost" authorityYear="1900" class="Reptilia" family="Gekkonidae" genus="Cyrtodactylus" kingdom="Animalia" lastPageId="11" lastPageNumber="554" order="Squamata" pageId="10" pageNumber="553" phylum="Chordata" rank="species" species="tuberculatus">
<emphasis id="C2FD6B0AFFDAC23FAFC1FACCB679F850" box="[1014,1320,2004,2028]" italics="true" pageId="10" pageNumber="553">Cyrtodactylus tuberculatus</emphasis>
(Lucas &amp; Frost 1900)
</taxonomicName>
, which was present at higher abundance. Two frog species, 
<taxonomicName id="3789CC9BFFDBC23EAFF1FD80B63AFF0C" authority="McDonald 1997" authorityName="McDonald" authorityYear="1997" box="[966,1387,151,176]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="11" pageNumber="554" phylum="Chordata" rank="species" species="andiirrmalin">
<emphasis id="C2FD6B0AFFDBC23EAFF1FD80B7FAFF0C" box="[966,1195,152,176]" italics="true" pageId="11" pageNumber="554">Litoria andiirrmalin</emphasis>
<collectingCountry id="889EF788FFDBC23EA884FD8FB67AFF0C" box="[1203,1323,151,176]" name="Heard Island and McDonald Islands" pageId="11" pageNumber="554">McDonald</collectingCountry>
1997
</taxonomicName>
and 
<taxonomicName id="3789CC9BFFDBC23EACA0FDA4B299FF69" authority="Davies" authorityName="Davies" box="[151,456,188,213]" class="Amphibia" family="Microhylidae" genus="Cophixalus" kingdom="Animalia" order="Anura" pageId="11" pageNumber="554" phylum="Chordata" rank="species" species="zweifeli">
<emphasis id="C2FD6B0AFFDBC23EACA0FDA4B223FF68" box="[151,370,188,212]" italics="true" pageId="11" pageNumber="554">Cophixalus zweifeli</emphasis>
Davies
</taxonomicName>
&amp; 
<collectingCountry id="889EF788FFDBC23EADDBFDA4B135FF69" box="[492,612,188,213]" name="Heard Island and McDonald Islands" pageId="11" pageNumber="554">McDonald</collectingCountry>
1998, were also found foraging at night at the site.
</paragraph>
</subSubSection>
</treatment>
</document>