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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.403.7145" ID-GBIF-Dataset="a6fda06f-575a-4ae4-8d40-ae927a01f959" ID-PMC="PMC4023238" ID-Pensoft-Pub="1313-2970-403-15" ID-PubMed="24843265" ID-ZBK="328E01EFBF324352AD7DBE989A3D716B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-403-15" ModsDocOrigin="ZooKeys 403" ModsDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" checkinTime="1451245966222" checkinUser="pensoft" docAuthor="Yoshida, Takahiro &amp; Hirowatari, Toshiya" docDate="2014" docId="3543C1FAF43E59D90AB81E1A52505865" docLanguage="en" docName="ZooKeys 403: 15-45" docOrigin="ZooKeys 403" docSource="http://dx.doi.org/10.3897/zookeys.403.7145" docTitle="Psammoecus bipunctatus Fabricius 1792" docType="treatment" docVersion="3" lastPageNumber="23" masterDocId="F8057B36FF80C436875CFFBFFFE6FF8F" masterDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="45" masterPageNumber="15" pageNumber="21" updateTime="1668158244412" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yoshida, Takahiro</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hirowatari, Toshiya</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>403</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>15</mods:start>
<mods:end>45</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.403.7145</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.403.7145</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-403-15</mods:identifier>
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<mods:identifier type="ZooBank">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
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<treatment ID-GBIF-Taxon="152052620" LSID="urn:lsid:plazi:treatment:3543C1FAF43E59D90AB81E1A52505865" httpUri="http://treatment.plazi.org/id/3543C1FAF43E59D90AB81E1A52505865" lastPageId="8" lastPageNumber="23" pageId="6" pageNumber="21">
<subSubSection pageId="6" pageNumber="21" type="nomenclature">
<paragraph pageId="6" pageNumber="21">
<taxonomicName LSID="http://species-id.net/wiki/Psammoecus_bipunctatus" authority="Fabricius, 1792" authorityName="Fabricius" authorityYear="1792" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus bipunctatus" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="bipunctatus">Psammoecus bipunctatus (Fabricius, 1792)</taxonomicName>
Figs 1B, 4and 13
<normalizedToken originalValue="DF">D-F</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="21" type="reference_group">
<paragraph pageId="6" pageNumber="21">
<taxonomicName class="Insecta" family="Anthicidae" genus="Notoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Notoxus bipunctatus" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="bipunctatus">Notoxus bipunctatus</taxonomicName>
Fabricius, 1792: 212. Type locality: Germania Dom.
</paragraph>
<paragraph pageId="6" pageNumber="21">
<taxonomicName class="Insecta" family="Ptiliidae" genus="Latridius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Latridius bipunctatus" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="bipunctatus">Latridius bipunctatus</taxonomicName>
: Herbst 1793: 10.
</paragraph>
<paragraph pageId="6" pageNumber="21">
<taxonomicName class="Insecta" family="Anthicidae" genus="Anthicus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anthicus bipunctatus" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="bipunctatus">Anthicus bipunctatus</taxonomicName>
Fabricius, 1801: 291.
</paragraph>
<paragraph pageId="6" pageNumber="21">
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus bipunctatus" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="bipunctatus">Psammoecus bipunctatus</taxonomicName>
:
<bibRefCitation author="Latreille, PA" editor="Cuvier, G" journalOrPublisher="Deterville, Libraire, Paris" pageId="27" pageNumber="42" title="Le Regne Animal distribue d'apres son Organization, pour server de base a l'histoire naturelle des animaux et d'introduction a l'anatomie comparee" year="1829">Latreille 1829</bibRefCitation>
: 135. -
<bibRefCitation author="Schaufuss, C" journalOrPublisher="Band I. E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart" pageId="28" pageNumber="43" title="Calwer's Kaeferbuch Einfuehrung in die Kenntnis der Kaefer Europas." year="1916">Schaufuss 1916</bibRefCitation>
: 457, fig. 4 in pl. 14. -
<bibRefCitation author="Hetschko, A" journalOrPublisher="W. Junk, Berlin" pageId="27" pageNumber="42" pagination="1 - 90" title="Coleopterorum Catalogus Pars 109" year="1930">Hetschko 1930</bibRefCitation>
: 81. (catalogue) -
<bibRefCitation author="Halstead, DGH" editor="Loebl, I" journalOrPublisher="Apollo Books, Stenstrup" pageId="27" pageNumber="42" pagination="496 - 501" title="Silvanidae." volumeTitle="Catalogue of Palaearctic Coleoptera" year="2007">Halstead et al. 2007</bibRefCitation>
-
<bibRefCitation author="Hirano, Y" journalOrPublisher="Kanagawa-chuho" pageId="27" pageNumber="42" pagination="57 - 83" title="Notes on Japanese Silvanidae (Nihonsan hosohiratamushi-ka ni tsuite)." volume="168" year="2009">Hirano 2009</bibRefCitation>
: 63, 64, 82, fig. 3. -
<bibRefCitation author="Hirano, Y" journalOrPublisher="Roppon-Ashi Entomological Books, Tokyo" pageId="27" pageNumber="42" title="Cucujoidea of Japan Vol. 2 Silvanidae, Byturidae, Biphyllidae." year="2010">Hirano 2010</bibRefCitation>
: 11, 12.
</paragraph>
<paragraph pageId="6" pageNumber="21">
<taxonomicName class="Insecta" genus="Psammaechus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Psammaechus" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="genus">Psammaechus</taxonomicName>
[sic.] bipunctatus:
<bibRefCitation author="Smith, F" journalOrPublisher="List of the Coleopterous Insects in the Collection of the British Museum" pageId="28" pageNumber="43" pagination="1 - 25" title="Catalogue of the Cucujidae, &amp; c." volume="1" year="1851">Smith 1851</bibRefCitation>
: 15. (catalogue)
</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="21" type="diagnosis">
<paragraph pageId="6" pageNumber="21">Diagnosis.</paragraph>
<paragraph pageId="6" pageNumber="21">
This species is similar to
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
and
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus boreas" order="Coleoptera" pageId="6" pageNumber="21" phylum="Arthropoda" rank="species" species="boreas">Psammoecus boreas</taxonomicName>
sp. n., but can be distinguished from the former by the shorter teeth on the lateral margins of the pronotum and from the latter by the shorter body length and the shorter antennae.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="22" pageId="6" pageNumber="21" type="description">
<paragraph pageId="6" pageNumber="21">Description.</paragraph>
<paragraph pageId="6" pageNumber="21">Body length. 2.25-2.83 mm (n=9).</paragraph>
<paragraph pageId="6" pageNumber="21">Coloration (Fig. 1B). Head dark brown, gradually darker toward anterior. Pronotum yellowish-brown to reddish-brown. Elytra yellowish-brown with round dark maculae on posterior 1/3, elytral suture dark in posterior half. Elytra of lighter colored specimens with reduced dark maculae and dark areas of elytral suture. Antennae yellowish-brown basally, 9th, 10th and posterior end of 8th antennomeres blackish-brown, 11th (apex) yellowish-brown.</paragraph>
<paragraph pageId="6" pageNumber="21">Head (Fig. 4A, B, C). Triangular, HW/HL 1.55-2.06; IE/HL 1.03-1.33. Dorsal surface with strong dense punctures, temples and posterior ventral surface also with irregularly dense punctures. Temples well enlarged, immediately narrowing thereafter. Eyes prominent, relatively small. Antennae 1.09-1.20 mm, comparatively thick, short; covered with medium length erect pubescence on each antennomere; approximate ratio of one of examined specimens as follows: 2.6: 1.1: 1.2: 1.1: 1.2: 1.2: 1.1: 1.0: 1.1: 1.0: 1.8 (Fig. 4A).</paragraph>
<paragraph lastPageId="7" lastPageNumber="22" pageId="6" pageNumber="21">
Pronotum (Fig. 4B, C). Roundly subquadrate, PW/PL 1.16-1.24. Dorsal surface with strong moderately dense punctuation, denser than that on ventral surface; some punctures on ventral surface in rows. Pubescence composed of many short setae on dorsal surface and several medium length setae on each lateral margin, distance between medium length setae
<pageBreakToken pageId="7" pageNumber="22" start="start">approximately</pageBreakToken>
regular. Each anterior angle with several very small teeth, each lateral margin with four similarly sized small teeth, the distance between them irregular and variable; each posterior angle with a small tooth, similar in size to those on lateral margins.
</paragraph>
<paragraph pageId="7" pageNumber="22">Elytra (Fig. 4E). Elongate, EW/BL 0.36-0.47. Rows of punctures almost same width as interstices. Pubescence composed of numerous comparatively short setae, without long setae.</paragraph>
<paragraph pageId="7" pageNumber="22">9th abdominal sternite (Fig. 13D). Strut Y-shaped, cut at anterior 1/3, diverging in posterior 1/4, branches relatively wide, ends of each branch pointed and curved inwards. Lateral sclerites elongate, membranous.</paragraph>
<paragraph pageId="7" pageNumber="22">Aedeagus (Figs 13E and F). Parameres club-shaped, wide and thick at bases, each apical portion with a long seta, stick-shaped portions with sparse punctuation and several short setae, lateral portions and inner margins of bases with successions of dense punctures, inner margin of bases with numerous short setae. Phallobase enlarged toward posterior margin from posterior 1/3, anterior margin roundly incised, anterior portion extended. Penis flat, wide, narrowed gradually toward apex, apical portion pointed, apical portion of dorsal part with sparse punctures.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="23" type="specimens examined">
<paragraph pageId="8" pageNumber="23">
<pageBreakToken pageId="8" pageNumber="23" start="start">Specimens</pageBreakToken>
examined.
</paragraph>
<paragraph pageId="8" pageNumber="23">
JAPAN: [Hokkaido Pref.] 1 male, Takkobu-numa Lake, Kushiro City, 1
<normalizedToken originalValue="IX">-IX-</normalizedToken>
2006, T. Yoshida leg. (ELKU); 1 ex., Iwabokki, Kushiro City, 25
<normalizedToken originalValue="VIII">-VIII-</normalizedToken>
1990, M. Sakai leg. (EUMJ); 6 exs., same locality, 26
<normalizedToken originalValue="VIII">-VIII-</normalizedToken>
1990, M. Sakai leg. (EUMJ); 1 ex., same locality, 29
<normalizedToken originalValue="VIII">-VIII-</normalizedToken>
1990, M. Sakai leg. (EUMJ).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="23" type="distribution">
<paragraph pageId="8" pageNumber="23">Distribution.</paragraph>
<paragraph pageId="8" pageNumber="23">JAPAN: Hokkaido; Africa; Europe; Russia; Turkestan.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="23" type="biological notes">
<paragraph pageId="8" pageNumber="23">Biological notes.</paragraph>
<paragraph pageId="8" pageNumber="23">
This species is known to inhabit marshland.
<bibRefCitation author="Warne, AC" journalOrPublisher="Nature in Cambridgeshire" pageId="28" pageNumber="43" pagination="24 - 26" title="The insects of Thriplow meadows." url="http://www.natureincambridgeshire.org.uk/volumes/nature-in-cambs-vol-06-1963.pdf" volume="6" year="1963">Warne (1963)</bibRefCitation>
reported that large numbers of this species were found on
<taxonomicName class="Liliopsida" family="Cyperaceae" genus="Carex" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Carex" order="Poales" pageId="8" pageNumber="23" phylum="Tracheophyta" rank="genus">Carex</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="23" type="remarks">
<paragraph pageId="8" pageNumber="23">Remarks.</paragraph>
<paragraph pageId="8" pageNumber="23">
The habitat and distribution of this species are unusual in the genus
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="8" pageNumber="23" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
. This species inhabits marshland and is distributed in comparatively high latitudes where species richness of
<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="8" pageNumber="23" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
is poor. Coloration is variable. In the past, four aberrations and two varieties (e.g.
<bibRefCitation author="Lucas, PH" journalOrPublisher="Guerin-Meneville" pageId="27" pageNumber="42" pagination="145 - 147" title="Note sur quelques nouvelles especes d'insectes de la famille des Trachelides qui habitent les possessions francaises du nord de l'Afrique. Revue Zoologique, par la Societe Cuvierienne; Association Universelle pour l'Avancement de la Zoologie, de l'Anatomie Comparee et de la Palaeontologie; Journal Mensuel. Publie sous la Direction de M. F. - E." volume="7" year="1843">Lucas 1843</bibRefCitation>
;
<bibRefCitation author="Gerhardt, J" journalOrPublisher="Jahresheft des Vereins fuer schlesische Insektenkunde zu Breslau" pageId="26" pageNumber="41" pagination="5 - 6" title="Neuheiten der schlesischen Koleopterenfauna aus dem Jahre 1911." volume="5" year="1912">Gerhardt 1912</bibRefCitation>
) were described.
</paragraph>
</subSubSection>
</treatment>
</document>