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<document id="3EC49E9EF95F7540E5991EF972435E1A" ID-ISSN="2201-4349" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629580321750" checkinUser="felipe" docAuthor="Cairns, S. D." docDate="2004" docId="903C803187465968FC22BD0AFEFCFC0D" docLanguage="en" docName="RecAustMus.56.3.259-329.pdf" docOrigin="Records of the Australian Museum 56" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Sphenotrochus cuneolus Cairns 2004, n.sp." docType="treatment" docVersion="2" lastPageNumber="300" masterDocId="6C05F849876E5941FFF1BE50FF96FF9F" masterDocTitle="The Azooxanthellate Scleractinia (Coelenterata: Anthozoa) of Australia" masterLastPageNumber="329" masterPageNumber="259" pageNumber="299" updateTime="1699080756492" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="46A3E73607768D5F20BB6248406907FF">The Azooxanthellate Scleractinia (Coelenterata: Anthozoa) of Australia</mods:title>
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<mods:namePart id="181F0936A135C6E526AB5063200B7DD4">Cairns, S. D.</mods:namePart>
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<mods:title id="DC911F653547A535CF396B628AA29D11">Records of the Australian Museum</mods:title>
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<mods:date id="0625C69441CD599C255DCEA8284AD606">2004</mods:date>
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<taxonomicName id="DF954AA487465969FC22BD0AFB66FCED" ID-CoL="86533" authority="Cairns, 2004" authorityName="Cairns" authorityYear="2004" box="[979,1264,858,882]" class="Anthozoa" family="Turbinoliidae" genus="Sphenotrochus" kingdom="Animalia" order="Scleractinia" pageId="40" pageNumber="299" phylum="Cnidaria" rank="species" species="cuneolus" status="sp. nov.">
<emphasis id="2AE1ED3587465969FC22BD0AFB66FCED" bold="true" box="[979,1264,858,882]" italics="true" pageId="40" pageNumber="299">Sphenotrochus cuneolus</emphasis>
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</emphasis>
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<subSubSection id="508F62AC87465969FBB3BDD3FB56FC05" box="[1090,1216,899,922]" pageId="40" pageNumber="299" type="description">
<paragraph id="182A312787465969FBB3BDD3FB56FC05" blockId="40.[1090,1216,899,922]" box="[1090,1216,899,922]" pageId="40" pageNumber="299">
<figureCitation id="80AE2DA287465969FBB3BDD3FB56FC05" box="[1090,1216,899,922]" captionStart="Fig" captionStartId="42.[166,202,1801,1822]" captionTargetBox="[166,1467,151,1776]" captionTargetId="figure-6@42.[166,1467,151,1776]" captionTargetPageId="42" captionText="Fig. 9. (AF), Sphenotrochus cuneolus (A, E, paratype, USNM 1008849, GCD=1.34 mm; BD, F, holotype, AM G16746, GCD=1.43 mm): (A, B), calicular and lateral views, ×36, ×19, respectively; (C), detail of base showing origin of C3, ×30.5; (D, F), enlargement of costae showing alternate placement of pores in intercostal regions, ×115, ×75 respectively; (F), oblique calicular view, ×35. (G, H), Sphenotrochus excavatus, holotype, Macleay Museum, oblique calicular and lateral views, ×8.3, ×5.6, respectively (GCD=5.8 mm). (IL), Trematotrochus hedleyi, USNM 82014, CD=3.6 mm: (IK), progressive enlargements of theca showing costae and porous intercostal regions, ×10.1, ×17.7, ×40, respectively; (L), view from within calice of septa and costal porosity, ×40. (MN), Monomyces rubrum: (M), AM G16298, lateral view showing contiguous basal rootlets, ×3.3 (GCD=5.0 mm); (N), USNM 1009341, lateral view of larger specimen showing lateral rootlets, ×2.8 (GCD=11.5 mm)." pageId="40" pageNumber="299">Figs. 9AF</figureCitation>
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<taxonomicName id="DF954AA487465969FCB6BDFCFBC6FC5E" ID-CoL="4YY5R" authorityName="Durham &amp; Barnard" authorityYear="1952" box="[839,1104,940,961]" class="Anthozoa" family="Turbinoliidae" genus="Sphenotrochus" kingdom="Animalia" order="Scleractinia" pageId="40" pageNumber="299" phylum="Cnidaria" rank="species" species="hancocki">
<emphasis id="2AE1ED3587465969FCB6BDFCFBC6FC5E" box="[839,1104,940,961]" italics="true" pageId="40" pageNumber="299">Sphenotrochus hancocki</emphasis>
</taxonomicName>
.
<treatmentCitation id="9934173687465969FB95BDFCFAC2FC5E" author="Cairns, S" box="[1124,1364,940,961]" page="38" pageId="40" pageNumber="299" year="1989">
<bibRefCitation id="7C044CD687465969FB95BDFCFAC2FC5E" author="Cairns, S" box="[1124,1364,940,961]" pageId="40" pageNumber="299" refId="ref82160" refString="Cairns, S. D., 1989 a. A revision of the ahermatypic Scleractinia of the Philippine Islands and adjacent waters, Part 1: Fungiacyathidae, Micrabaciidae, Turbinoliinae, Guyniidae, and Flabellidae. Smithsonian Contributions to Zoology 486: 136 pp., 42 pls., 3 figs." type="book" year="1989">Cairns, 1989a: 3839</bibRefCitation>
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(in part:
<emphasis id="2AE1ED3587465969FC94BD97FC54FC43" box="[869,962,967,988]" italics="true" pageId="40" pageNumber="299">Albatross</emphasis>
-5145).
</treatmentCitationGroup>
</paragraph>
<paragraph id="182A312787465969FCB6BDBCFB45FB09" blockId="40.[839,1467,1004,1174]" pageId="40" pageNumber="299">
<emphasis id="2AE1ED3587465969FCB6BDBCFC6BFB9B" bold="true" box="[839,1021,1004,1028]" pageId="40" pageNumber="299">Records/Types</emphasis>
.
<materialsCitation id="A8FD3B7A87465969FBE0BDBDFC36FBBE" accessionNumber="G16746" collectionCode="AM" pageId="40" pageNumber="299" specimenCount="1" typeStatus="holotype">
<typeStatus id="C72E8F8587465969FBE0BDBDFB14FB9B" box="[1041,1154,1005,1028]" pageId="40" pageNumber="299" type="holotype">Holotype</typeStatus>
: Franklin 03/99/D11,
<collectionCode id="7E84A9E287465969FA7FBDBDFA2CFB9B" box="[1422,1466,1005,1028]" country="Australia" name="Australian Museum" pageId="40" pageNumber="299" type="Museum">AM</collectionCode>
G16746
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.
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<typeStatus id="C72E8F8587465969FC59BA5AFB85FBBE" box="[936,1043,1034,1057]" pageId="40" pageNumber="299" type="paratype">Paratypes</typeStatus>
:
<collectorName id="B56054F187465969FBEDBA5AFBE0FBBE" box="[1052,1142,1034,1057]" pageId="40" pageNumber="299">Franklin</collectorName>
03/99/D8, 1,
<specimenCode id="4833995C87465969FAF5BA5AFA21FBBE" box="[1284,1463,1034,1057]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="40" pageNumber="299" type="Museum">USNM 1008848</specimenCode>
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;
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<collectorName id="B56054F187465969FCB6BA77FC37FBA1" box="[839,929,1063,1086]" pageId="40" pageNumber="299">Franklin</collectorName>
03/99/D10, 6 (including
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),
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;
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<collectorName id="B56054F187465969FCB6BA15FC08FBC3" box="[839,926,1093,1116]" pageId="40" pageNumber="299">Franklin</collectorName>
03/99/D11, 6: 5,
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<materialsCitation id="A8FD3B7A87465969FB09BA15FBB8FBE6" collectionCode="USNM" collectorName="Alb" pageId="40" pageNumber="299" specimenCode="USNM 81896" specimenCount="1" typeStatus="paratype">
and 1, ZMUZ;
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<collectorName id="B56054F187465969FA7FBA15FA24FBC3" box="[1422,1458,1093,1116]" pageId="40" pageNumber="299">Alb</collectorName>
</emphasis>
- 5145, 1,
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.
<materialsCitation id="A8FD3B7A87465969FBC5BA32FB45FB09" elevation="342" latitude="-20.2415" location="Marion Plateau" longLatPrecision="12" longitude="151.79216" pageId="40" pageNumber="299" specimenCount="342" stateProvince="Queensland" typeStatus="paratype">
Type Locality:
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(
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,
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),
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.
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</paragraph>
</subSubSection>
<subSubSection id="508F62AC87465968FCB6BAF7FD92FE04" lastPageId="41" lastPageNumber="300" pageId="40" pageNumber="299" type="description">
<paragraph id="182A312787465969FCB6BAF7FB6BF848" blockId="40.[839,1467,1191,2007]" pageId="40" pageNumber="299">
<emphasis id="2AE1ED3587465969FCB6BAF7FC5AFB20" bold="true" box="[839,972,1191,1215]" pageId="40" pageNumber="299">Description</emphasis>
. Corallum cuneiform, with planar thecal faces and rounded thecal edges and base, the edges being roughly parallel. Largest specimen (
<typeStatus id="C72E8F8587465969FB82BAB2FB43FB66" box="[1139,1237,1250,1273]" pageId="40" pageNumber="299" type="holotype">holotype</typeStatus>
) only 1.43×
<quantity id="DF6D9CC287465969FAA8BAB2FA2DFB66" box="[1369,1467,1250,1273]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="40" pageNumber="299" unit="mm" value="1.02">1.02 mm</quantity>
in CD and
<quantity id="DF6D9CC287465969FC33BB50FBB2FA88" box="[962,1060,1280,1303]" metricMagnitude="-3" metricUnit="m" metricValue="2.78" pageId="40" pageNumber="299" unit="mm" value="2.78">2.78 mm</quantity>
in height, but judging from the high frequency of juvenile forms of other turbinoliids taken at this site, this may also represent a sub-adult size. Calice elliptical, the GCD:LCD about 1.4. Costal arrangement distinctive and consistent (
<figureCitation id="80AE2DA287465969FB9FBB25FB7FFA13" box="[1134,1257,1397,1420]" captionStart="Fig" captionStartId="42.[166,202,1801,1822]" captionTargetBox="[166,1467,151,1776]" captionTargetId="figure-6@42.[166,1467,151,1776]" captionTargetPageId="42" captionText="Fig. 9. (AF), Sphenotrochus cuneolus (A, E, paratype, USNM 1008849, GCD=1.34 mm; BD, F, holotype, AM G16746, GCD=1.43 mm): (A, B), calicular and lateral views, ×36, ×19, respectively; (C), detail of base showing origin of C3, ×30.5; (D, F), enlargement of costae showing alternate placement of pores in intercostal regions, ×115, ×75 respectively; (F), oblique calicular view, ×35. (G, H), Sphenotrochus excavatus, holotype, Macleay Museum, oblique calicular and lateral views, ×8.3, ×5.6, respectively (GCD=5.8 mm). (IL), Trematotrochus hedleyi, USNM 82014, CD=3.6 mm: (IK), progressive enlargements of theca showing costae and porous intercostal regions, ×10.1, ×17.7, ×40, respectively; (L), view from within calice of septa and costal porosity, ×40. (MN), Monomyces rubrum: (M), AM G16298, lateral view showing contiguous basal rootlets, ×3.3 (GCD=5.0 mm); (N), USNM 1009341, lateral view of larger specimen showing lateral rootlets, ×2.8 (GCD=11.5 mm)." pageId="40" pageNumber="299">Figs. 9B,C</figureCitation>
). The 12 C12 are continuous, all reaching the base of the corallum. The medial C2, which bisect the lateral faces, and the principal C1, which are on the corallum edges, are straight, whereas the 4 C1 and C2 on each face between the medial C2 and principal C1 curve slightly inward near the base of the corallum and then abruptly outward, producing a moderate sinuosity. C3 also continuous but do not extend entire distance to the base, their length progressively increasing away from the medial C2: those C3 adjacent to the medial C2 extend about 60% of the distance to the base, the next pair of C3 toward the principal C1 being slightly longer (about 65% of the distance to the base), and those C3 directly adjacent to the principal C1 extend about 75% to the base. Costae uniform in width and ridge-like,
<quantity id="DF6D9CC287465969FAF1B940FA08F8B8" box="[1280,1438,1808,1831]" metricMagnitude="-5" metricUnit="m" metricValue="6.5" metricValueMax="7" metricValueMin="6.0" pageId="40" pageNumber="299" unit="mm" value="0.065" valueMax="0.07" valueMin="0.06">0.060.07 mm</quantity>
in width, and separated by broad intercostal spaces about
<quantity id="DF6D9CC287465969FA7DB97DFCE7F8FE" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="40" pageNumber="299" unit="mm" value="0.09">0.09 mm</quantity>
in width. A series of small pores, each 1011 µm in diameter, occurs along both edges of each intercostal region (
<figureCitation id="80AE2DA287465969FCBEB9D5FC32F803" box="[847,932,1925,1948]" captionStart="Fig" captionStartId="42.[166,202,1801,1822]" captionTargetBox="[166,1467,151,1776]" captionTargetId="figure-6@42.[166,1467,151,1776]" captionTargetPageId="42" captionText="Fig. 9. (AF), Sphenotrochus cuneolus (A, E, paratype, USNM 1008849, GCD=1.34 mm; BD, F, holotype, AM G16746, GCD=1.43 mm): (A, B), calicular and lateral views, ×36, ×19, respectively; (C), detail of base showing origin of C3, ×30.5; (D, F), enlargement of costae showing alternate placement of pores in intercostal regions, ×115, ×75 respectively; (F), oblique calicular view, ×35. (G, H), Sphenotrochus excavatus, holotype, Macleay Museum, oblique calicular and lateral views, ×8.3, ×5.6, respectively (GCD=5.8 mm). (IL), Trematotrochus hedleyi, USNM 82014, CD=3.6 mm: (IK), progressive enlargements of theca showing costae and porous intercostal regions, ×10.1, ×17.7, ×40, respectively; (L), view from within calice of septa and costal porosity, ×40. (MN), Monomyces rubrum: (M), AM G16298, lateral view showing contiguous basal rootlets, ×3.3 (GCD=5.0 mm); (N), USNM 1009341, lateral view of larger specimen showing lateral rootlets, ×2.8 (GCD=11.5 mm)." pageId="40" pageNumber="299">Fig. 9D</figureCitation>
), each pore separated by 3540 µm from one other and alternating in position from those on the opposite side of the intercostal space. Corallum white.
</paragraph>
<paragraph id="182A312787475968FF66BEC9FD92FE04" blockId="41.[121,749,153,411]" pageId="41" pageNumber="300">
<collectingRegion id="DA51FFC587475968FF66BEC9FF40FF2F" box="[151,214,153,176]" country="Spain" name="Ceuta" pageId="41" pageNumber="300">Septa</collectingRegion>
hexamerally arranged in 3 complete cycles, resulting in 24 septa (S12&gt;S3). All specimens reported dead when collected and thus not optimally preserved, but S12 equal in exsertness and width, having quite sinuous axial edges, the lower edges of which fuse to the columella in the case of the 6 S
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the lateral corallum position. S3 considerably smaller and have straight axial edges. Fossa shallow, containing a robust, lamellar columella, the upper edge rising above the calicular edge.
</paragraph>
</subSubSection>
<subSubSection id="508F62AC87475968FF88BFFCFE9BFC99" pageId="41" pageNumber="300" type="discussion">
<paragraph id="182A312787475968FF88BFFCFE9BFC99" blockId="41.[121,749,428,774]" pageId="41" pageNumber="300">
<emphasis id="2AE1ED3587475968FF88BFFCFF74FE5B" bold="true" box="[121,226,428,452]" pageId="41" pageNumber="300">Remarks</emphasis>
. Among the nine Recent species in the nominate subgenus of
<taxonomicName id="DF954AA487475968FF0ABF9AFE00FE7E" box="[251,406,458,481]" class="Anthozoa" family="Turbinoliidae" genus="Sphenotrochus" kingdom="Animalia" order="Scleractinia" pageId="41" pageNumber="300" phylum="Cnidaria" rank="genus">
<emphasis id="2AE1ED3587475968FF0ABF9AFE00FE7E" box="[251,406,458,481]" italics="true" pageId="41" pageNumber="300">Sphenotrochus</emphasis>
</taxonomicName>
, defined as those species bearing costae that are continuous (not fragmented into numerous short carinae, see Cairns, 1997),
<taxonomicName id="DF954AA487475968FE22BC55FDDAFD83" authorityName="Cairns" authorityYear="2004" box="[467,588,517,540]" class="Anthozoa" family="Turbinoliidae" genus="Sphenotrochus" kingdom="Animalia" order="Scleractinia" pageId="41" pageNumber="300" phylum="Cnidaria" rank="species" species="cuneolus">
<emphasis id="2AE1ED3587475968FE22BC55FDDAFD83" box="[467,588,517,540]" italics="true" pageId="41" pageNumber="300">S. cuneolus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="DF954AA487475968FF64BC72FDA6FDA6" authority="Durham &amp; Barnard, 1952" authorityName="Durham &amp; Barnard" authorityYear="1952" box="[149,560,546,569]" class="Anthozoa" family="Turbinoliidae" genus="Sphenotrochus" kingdom="Animalia" order="Scleractinia" pageId="41" pageNumber="300" phylum="Cnidaria" rank="species" species="hancocki">
<emphasis id="2AE1ED3587475968FF64BC72FE87FDA6" box="[149,273,546,569]" italics="true" pageId="41" pageNumber="300">S. hancocki</emphasis>
Durham &amp; Barnard, 1952
</taxonomicName>
, a species known from throughout the tropical Pacific at
<quantity id="DF6D9CC287475968FDD3BC6FFD19FDC9" box="[546,655,575,598]" metricMagnitude="2" metricUnit="m" metricValue="1.46" metricValueMax="2.7399999999999998" metricValueMin="0.18" pageId="41" pageNumber="300" unit="m" value="146.0" valueMax="274.0" valueMin="18.0">18274 m</quantity>
(
<bibRefCitation id="7C044CD687475968FD6FBC6FFF29FDEB" author="Cairns, S" pageId="41" pageNumber="300" refId="ref82160" refString="Cairns, S. D., 1989 a. A revision of the ahermatypic Scleractinia of the Philippine Islands and adjacent waters, Part 1: Fungiacyathidae, Micrabaciidae, Turbinoliinae, Guyniidae, and Flabellidae. Smithsonian Contributions to Zoology 486: 136 pp., 42 pls., 3 figs." type="book" year="1989">Cairns, 1989a</bibRefCitation>
). These two species are similar in size and septal and columellar morphology, but
<taxonomicName id="DF954AA487475968FE58BC2AFDB4FD0E" authorityName="Cairns" authorityYear="2004" box="[425,546,634,657]" class="Anthozoa" family="Turbinoliidae" genus="Sphenotrochus" kingdom="Animalia" order="Scleractinia" pageId="41" pageNumber="300" phylum="Cnidaria" rank="species" species="cuneolus">
<emphasis id="2AE1ED3587475968FE58BC2AFDB4FD0E" box="[425,546,634,657]" italics="true" pageId="41" pageNumber="300">S. cuneolus</emphasis>
</taxonomicName>
differs in its costal arrangement and in having sinuous axial edges of its S12.
<emphasis id="2AE1ED3587475968FF88BCE5FE25FD53" box="[121,435,693,716]" italics="true" pageId="41" pageNumber="300">
SSphenotrochus
<taxonomicName id="DF954AA487475968FEB6BCE5FE25FD53" authorityName="Durham &amp; Barnard" authorityYear="1952" box="[327,435,693,716]" class="Anthozoa" family="Turbinoliidae" genus="Sphenotrochus" kingdom="Animalia" order="Scleractinia" pageId="41" pageNumber="300" phylum="Cnidaria" rank="species" species="hancocki">hancocki</taxonomicName>
</emphasis>
not only has a different arrangement of costae but also a fragmentation of its costae near the base.
</paragraph>
</subSubSection>
<subSubSection id="508F62AC87475968FF88BD47FDBBFCD3" pageId="41" pageNumber="300" type="etymology">
<paragraph id="182A312787475968FF88BD47FDBBFCD3" blockId="41.[121,748,791,844]" pageId="41" pageNumber="300">
<emphasis id="2AE1ED3587475968FF88BD47FF63FCB0" bold="true" box="[121,245,791,815]" pageId="41" pageNumber="300">Etymology</emphasis>
.
<taxonomicName id="DF954AA487475968FEF0BD48FE33FCB0" authority=", Latin" authorityName="Latin" box="[257,421,792,815]" class="Anthozoa" family="Turbinoliidae" genus="Sphenotrochus" kingdom="Animalia" order="Scleractinia" pageId="41" pageNumber="300" phylum="Cnidaria" rank="species" species="cuneolus">
<emphasis id="2AE1ED3587475968FEF0BD48FEF7FCB0" box="[257,353,792,815]" italics="true" pageId="41" pageNumber="300">cuneolus</emphasis>
, Latin
</taxonomicName>
for “small wedge”, an allusion to the small size of this cuneiform coral.
</paragraph>
</subSubSection>
<subSubSection id="508F62AC87475968FF88BD0DFEFCFC0D" pageId="41" pageNumber="300" type="distribution">
<paragraph id="182A312787475968FF88BD0DFEFCFC0D" blockId="41.[121,748,861,914]" pageId="41" pageNumber="300">
<emphasis id="2AE1ED3587475968FF88BD0DFF68FCEA" bold="true" box="[121,254,861,885]" pageId="41" pageNumber="300">Distribution</emphasis>
. Marion Plateau, Queensland; SuluArchipelago,
<collectingCountry id="608271B787475968FF88BD2BFF67FC0D" box="[121,241,891,914]" name="Philippines" pageId="41" pageNumber="300">Philippines</collectingCountry>
;
<quantity id="DF6D9CC287475968FF0BBD2BFEF1FC0D" box="[250,359,891,914]" metricMagnitude="2" metricUnit="m" metricValue="1.92" metricValueMax="3.42" metricValueMin="0.42" pageId="41" pageNumber="300" unit="m" value="192.0" valueMax="342.0" valueMin="42.0">
42
<specimenCount id="0E93FAAE87475968FED3BD2BFEF1FC0D" box="[290,359,891,914]" pageId="41" pageNumber="300" type="generic">342 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>