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<document id="4A423ADA1FC42A452410E208E224D2E2" ID-CLB-Dataset="304417" ID-DOI="10.11646/zootaxa.5519.4.1" ID-GBIF-Dataset="d2f13ee2-c472-4616-88bd-7eaf486a0e45" ID-ISSN="1175-5326" ID-Zenodo-Dep="13916138" ID-ZooBank="1434ADD6-A021-4DFB-8471-D834438F8682" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728587332582" checkinUser="plazi" docAuthor="Dupérré, Nadine &amp; Tapia, Elicio" docDate="2024" docId="8B3647425861FF99FF2866BACDA0FEA5" docLanguage="en" docName="zootaxa.5519.4.1.pdf" docOrigin="Zootaxa 5519 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5519.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paratropis arenosa Dupérré &amp; Tapia, 2024, new species" docType="treatment" docVersion="4" lastPageNumber="477" masterDocId="770F3F3A5876FF83FFBF670DCC3BFFC0" masterDocTitle="Seven new species of the enigmatic spider genus Paratropis Simon, 1889 (Mygalomorphae, Paratropididae) from Ecuador" masterLastPageNumber="486" masterPageNumber="451" pageNumber="474" updateTime="1728668132093" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E8E30749C75850D519DD4D80B69D0A7A">Seven new species of the enigmatic spider genus Paratropis Simon, 1889 (Mygalomorphae, Paratropididae) from Ecuador</mods:title>
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<mods:name id="A00261E59E0001853BA66570EDF4CDC3" type="personal">
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<mods:namePart id="97F699D62C346C482C8462E485B29CEE">Dupérré, Nadine</mods:namePart>
<mods:affiliation id="1EDE12B661CA5B14FCF25B4A2F2C37C8">Museum of Nature Hamburg, Zoology, Leibniz-Institute for the Analysis of Biodiversity Change (LIB), Center for Taxonomy and Morphology, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. &amp; Research Associate, American Museum of Natural History, New York, NY, U. S. A.</mods:affiliation>
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<mods:namePart id="4786673BE44FF7E8C770F3C32F5B42FA">Tapia, Elicio</mods:namePart>
<mods:affiliation id="1AF8C717C8BEDA1D34CDE5903F6A7161">Museum of Nature Hamburg, Zoology, Leibniz-Institute for the Analysis of Biodiversity Change (LIB), Center for Taxonomy and Morphology, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
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<mods:title id="5CD3033C1DB5CA232D46198933BA4890">Zootaxa</mods:title>
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<mods:part id="8F34B4BE127686D120A0FEB8175B4A6E">
<mods:date id="AA3556B48A990FAD7070DF64FA1C6E9F">2024</mods:date>
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<mods:number id="1591C36614CC680367A2631A5305E629">2024-10-10</mods:number>
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<mods:number id="03C60F58FDBB3C30493A9504F4DC7156">5519</mods:number>
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<mods:identifier id="37D752CA030A314D5D341C5DD2E29B72" type="DOI">10.11646/zootaxa.5519.4.1</mods:identifier>
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<treatment id="8B3647425861FF99FF2866BACDA0FEA5" ID-DOI="http://doi.org/10.5281/zenodo.13921370" ID-Zenodo-Dep="13921370" LSID="urn:lsid:plazi:treatment:8B3647425861FF99FF2866BACDA0FEA5" httpUri="http://treatment.plazi.org/id/8B3647425861FF99FF2866BACDA0FEA5" lastPageId="26" lastPageNumber="477" pageId="23" pageNumber="474">
<subSubSection id="4B85A5DF5861FF94FF2866BACE2EFE10" box="[151,533,438,465]" pageId="23" pageNumber="474" type="nomenclature">
<paragraph id="0320F6545861FF94FF2866BACE2EFE10" blockId="23.[151,533,438,501]" box="[151,533,438,465]" pageId="23" pageNumber="474">
<heading id="586841385861FF94FF2866BACE2EFE10" bold="true" box="[151,533,438,465]" fontSize="11" level="1" pageId="23" pageNumber="474" reason="1">
<emphasis id="31EB2A465861FF94FF2866BACE2EFE10" bold="true" box="[151,533,438,465]" pageId="23" pageNumber="474">
<taxonomicName id="C49F8DD75861FF94FF2866BACD45FE11" authorityName="Dupérré &amp; Tapia" authorityYear="2024" box="[151,382,439,465]" class="Arachnida" family="Paratropididae" genus="Paratropis" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="arenosa" status="sp. nov.">
<emphasis id="31EB2A465861FF94FF2866BACD45FE11" bold="true" box="[151,382,439,465]" italics="true" pageId="23" pageNumber="474">Paratropis arenosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2AD8973D5861FF94FE3A66BACE2EFE10" box="[389,533,438,465]" pageId="23" pageNumber="474" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4B85A5DF5861FF94FF2866D6CD50FE35" box="[151,363,475,501]" pageId="23" pageNumber="474" type="description">
<paragraph id="0320F6545861FF94FF2866D6CD50FE35" blockId="23.[151,533,438,501]" box="[151,363,475,501]" pageId="23" pageNumber="474">
<figureCitation id="9BA4EAD15861FF94FF2866D6CD2CFE35" box="[151,279,475,501]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStart-2="FIGURE 22" captionStartId-0="23.[151,250,1751,1775]" captionStartId-1="24.[151,248,1837,1861]" captionStartId-2="25.[151,250,1940,1964]" captionTargetBox-0="[179,1409,862,1715]" captionTargetBox-1="[170,1394,179,1800]" captionTargetBox-2="[161,1426,465,1909]" captionTargetId-0="figure-213@23.[166,1422,849,1728]" captionTargetId-1="figure-23@24.[153,1434,151,1813]" captionTargetId-2="figure-152@25.[151,1436,453,1917]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionText-0="FIGURE 20. Paratropis arenosa sp. nov. female ECFN 1323. A. Live, Las Antenas, Sucumbíos Province, dorso-lateral view. Scale bar: 5.0 mm." captionText-1="FIGURE 21 Paratropis arenosa sp. nov. female holotype.A. Habitus, dorsal view. B. Habitus, ventral view. Scale bars: 5.0 mm." captionText-2="FIGURE 22. Paratropis arenosa sp. nov. A. Female holotype, carapace dorsal view. B. Female holotype, sternum and endites ventral view. C. Spermathecae, dorsal view (holotype). D. Spermathecae, dorsal view (ECFN 7220). Scale bars:A, B = 1.0 mm; C, D = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13916181" figureDoi-1="http://doi.org/10.5281/zenodo.13916183" figureDoi-2="http://doi.org/10.5281/zenodo.13916185" httpUri-0="https://zenodo.org/record/13916181/files/figure.png" httpUri-1="https://zenodo.org/record/13916183/files/figure.png" httpUri-2="https://zenodo.org/record/13916185/files/figure.png" pageId="23" pageNumber="474">Figs 20–22</figureCitation>
,
<figureCitation id="9BA4EAD15861FF94FE9D66D6CD50FE35" box="[290,363,475,501]" captionStart="MAP 1" captionStartId="33.[151,208,1803,1827]" captionTargetBox="[182,1406,415,1776]" captionTargetId="figure-134@33.[169,1418,401,1789]" captionTargetPageId="33" captionText="MAP 1. Distribution of newly described Paratropis. Paratropis carcosita sp. nov. (yellow circle), Paratropis pukallucha sp. nov. (orange circle), Paratropis esmeraldas sp. nov. (blue circle), Paratropis kapak sp. nov. (purple circle), Paratropis pasochoa sp. nov. (yellow star), Paratropis arenosa sp. nov. (red inverted triangle) and, Paratropis cryptica sp. nov. (green inverted triangle). Provinces abbreviations: Azuay (AZ), Bolívar (BO), Carchi (CA), Cañar (CAN), Chimborazo (CH), Cotopaxi (CO), El Oro (EO), Esmeralsdas (ES), Guayas (GU), Imbabura (IM), Loja (LO), Los Ríos (LR), Manabí (MA), Morona Santiago (MS), Napo (NA), Orellana (OR), Pastaza (PA), Pichincha (PI), Santa Elena (SE), Santo Domingo de los Tsáchilas (SD), Sucumbíos (SU), Tugurahua (TU), Zamora Chinchipe (ZC)." figureDoi="http://doi.org/10.5281/zenodo.13921361" httpUri="https://zenodo.org/record/13921361/files/figure.png" pageId="23" pageNumber="474">Map 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4B85A5DF5861FF94FF28652EC8A7FD69" pageId="23" pageNumber="474" type="materials_examined">
<paragraph id="0320F6545861FF94FF28652EC8A7FD69" blockId="23.[151,1436,546,825]" pageId="23" pageNumber="474">
<emphasis id="31EB2A465861FF94FF28652ECD7CFDFC" bold="true" box="[151,327,546,573]" pageId="23" pageNumber="474">Type material.</emphasis>
<materialsCitation id="B3F7FC095861FF94FEEF652EC85AFDA1" collectedFrom="under pile of rocks" collectingDate="2019-05-22" collectingMethod="hand collected" collectionCode="QCAZ" collectorName="E. Tapia" elevation="826" location="Las Antenas" municipality="Lumbaqui" pageId="23" pageNumber="474" specimenCode="ECFN 1332" specimenCount="1" specimenCount-female="1" stateProvince="Sucumbios Province" typeStatus="holotype">
<specimenCount id="15993DDD5861FF94FEEF652ECD99FDFD" box="[336,418,547,573]" pageId="23" pageNumber="474" type="female">Female</specimenCount>
<typeStatus id="DC2448F65861FF94FE14652ECE36FDFD" box="[427,525,547,573]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
from
<collectingRegion id="C15B38B65861FF94FDEA652ECF7BFDFD" box="[597,832,547,573]" country="Ecuador" name="Sucumbios" pageId="23" pageNumber="474">Sucumbíos Province</collectingRegion>
,
<collectingMunicipality id="E3446C2E5861FF94FCF1652ECFFFFDFD" box="[846,964,547,573]" pageId="23" pageNumber="474">Lumbaqui</collectingMunicipality>
,
<location id="0640A08F5861FF94FC6E652EC867FDFD" LSID="urn:lsid:plazi:treatment:8B3647425861FF99FF2866BACDA0FEA5:0640A08F5861FF94FC6E652EC867FDFD" box="[977,1116,547,573]" county="Female" municipality="Lumbaqui" name="Las Antenas" pageId="23" pageNumber="474" stateProvince="Sucumbios Province">Las Antenas</location>
(
<geoCoordinate id="66AB90935861FF94FBD2652EC8C1FDFD" box="[1133,1274,547,573]" pageId="23" pageNumber="474">-00.036226°</geoCoordinate>
<geoCoordinate id="66AB90935861FF94FABC652FC9AFFDFD" box="[1283,1428,546,573]" pageId="23" pageNumber="474">–77.308526°</geoCoordinate>
)
<quantity id="C4675BB15861FF94FF28654ACCE4FDA1" box="[151,223,583,609]" metricMagnitude="2" metricUnit="m" metricValue="8.26" pageId="23" pageNumber="474" unit="m" value="826.0">
<elevation id="88B211675861FF94FF28654ACCE4FDA1" box="[151,223,583,609]" metricMagnitude="2" metricUnit="m" metricValue="8.26" pageId="23" pageNumber="474" unit="m" value="826.0">826 m</elevation>
</quantity>
,
<date id="7721D0945861FF94FF56654ACD44FDA1" box="[233,383,583,609]" pageId="23" pageNumber="474" value="2019-05-22">
<collectingDate id="6765297C5861FF94FF56654ACD44FDA1" box="[233,383,583,609]" pageId="23" pageNumber="474" value="2019-05-22">22 May 2019</collectingDate>
</date>
,
<collectingMethod id="DADE8E435861FF94FE36654ACE11FDA1" box="[393,554,583,609]" pageId="23" pageNumber="474">hand collected</collectingMethod>
<collectedFrom id="B083B5A95861FF94FD8F654ACF3AFDA1" box="[560,769,583,609]" pageId="23" pageNumber="474">under pile of rocks</collectedFrom>
,
<collectorName id="AE6A93825861FF94FCB4654ACF51FDA1" box="[779,874,583,609]" pageId="23" pageNumber="474">E. Tapia</collectorName>
,
<specimenCode id="53395E2F5861FF94FCCC654ACFC3FDA1" box="[883,1016,583,609]" collectionCode="ECFN" pageId="23" pageNumber="474">ECFN 1332</specimenCode>
(
<collectionCode id="658E6E915861FF94FBB9654AC862FDA1" box="[1030,1113,583,609]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="23" pageNumber="474" type="Museum">QCAZ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="B3F7FC095861FF94FBD5654ACD57FD69" collectedFrom="en talud de tierra bosque primario humedo" collectingDate="2018-02-24" collectionCode="ECFN" collectorName="E. Tapia" elevation="826" location="Las Antenas" pageId="23" pageNumber="474" specimenCode="ECFN 1323, ZMH-A0002999" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="DC2448F65861FF94FBD5654AC8E0FDA1" box="[1130,1243,583,609]" pageId="23" pageNumber="474" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="15993DDD5861FF94FB5B654AC93CFDA1" box="[1252,1287,583,609]" count="1" pageId="23" pageNumber="474" type="female">1♀</specimenCount>
,
<location id="0640A08F5861FF94FAAB654AC9A7FDA1" LSID="urn:lsid:plazi:treatment:8B3647425861FF99FF2866BACDA0FEA5:0640A08F5861FF94FAAB654AC9A7FDA1" box="[1300,1436,583,609]" name="Las Antenas" pageId="23" pageNumber="474">Las Antenas</location>
(
<geoCoordinate id="66AB90935861FF94FF206566CD0BFD45" box="[159,304,619,645]" pageId="23" pageNumber="474">–00.036226°</geoCoordinate>
<geoCoordinate id="66AB90935861FF94FE886567CDF2FD45" box="[311,457,618,645]" pageId="23" pageNumber="474">–77.308526°</geoCoordinate>
)
<quantity id="C4675BB15861FF94FE686566CE24FD45" box="[471,543,619,645]" metricMagnitude="2" metricUnit="m" metricValue="8.26" pageId="23" pageNumber="474" unit="m" value="826.0">
<elevation id="88B211675861FF94FE686566CE24FD45" box="[471,543,619,645]" metricMagnitude="2" metricUnit="m" metricValue="8.26" pageId="23" pageNumber="474" unit="m" value="826.0">826 m</elevation>
</quantity>
,
<date id="7721D0945861FF94FD956566CE84FD45" box="[554,703,619,645]" pageId="23" pageNumber="474" value="2018-02-24">
<collectingDate id="6765297C5861FF94FD956566CE84FD45" box="[554,703,619,645]" pageId="23" pageNumber="474" value="2018-02-24">24 Feb. 2018</collectingDate>
</date>
,
<collectedFrom id="B083B5A95861FF94FD766566C898FD45" box="[713,1187,619,645]" pageId="23" pageNumber="474">en talud de tierra bosque primario humedo</collectedFrom>
,
<collectorName id="AE6A93825861FF94FB126566C937FD45" box="[1197,1292,619,645]" pageId="23" pageNumber="474">E. Tapia</collectorName>
,
<specimenCode id="53395E2F5861FF94FAA96566C9A7FD45" box="[1302,1436,619,645]" collectionCode="ECFN" pageId="23" pageNumber="474">ECFN 1323</specimenCode>
(
<specimenCode id="53395E2F5861FF94FF206582CD58FD69" box="[159,355,655,681]" collectionCode="ZMH-A" pageId="23" pageNumber="474">ZMH-A0002999</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="B3F7FC095861FF94FEC96582C8A2FD69" box="[374,1177,655,681]" collectingDate="2014-02-11" collectionCode="QCAZ" collectorName="C. Rodriguez" location="Lago Agrio" pageId="23" pageNumber="474" specimenCode="ECFN 7220" specimenCount="1" specimenCount-female="1">
<specimenCount id="15993DDD5861FF94FEC96582CDA2FD69" box="[374,409,655,681]" count="1" pageId="23" pageNumber="474" type="female">1♀</specimenCount>
,
<location id="0640A08F5861FF94FE186582CE1CFD69" LSID="urn:lsid:plazi:treatment:8B3647425861FF99FF2866BACDA0FEA5:0640A08F5861FF94FE186582CE1CFD69" box="[423,551,655,681]" name="Lago Agrio" pageId="23" pageNumber="474">Lago Agrio</location>
(18N)
<date id="7721D0945861FF94FDC66582CEC1FD69" box="[633,762,655,681]" pageId="23" pageNumber="474" value="2014-02-11">
<collectingDate id="6765297C5861FF94FDC66582CEC1FD69" box="[633,762,655,681]" pageId="23" pageNumber="474" value="2014-02-11">2014/11/02</collectingDate>
</date>
,
<collectorName id="AE6A93825861FF94FCBA6582CFA5FD69" box="[773,926,655,681]" pageId="23" pageNumber="474">C. Rodriguez</collectorName>
,
<specimenCode id="53395E2F5861FF94FC176582C814FD69" box="[936,1071,655,681]" collectionCode="ECFN" pageId="23" pageNumber="474">ECFN 7220</specimenCode>
(
<collectionCode id="658E6E915861FF94FB816582C8AAFD69" box="[1086,1169,655,681]" collectionName="Ecuador, Quito, Pontificia Universidad Catolica del Ecuador, Catholic Zoology Museum" country="Ecuador" httpUri="http://grbio.org/cool/kwpg-nhbg" name="Museo de Zoologia, Pontificia Universidad Catolica del Ecuador" pageId="23" pageNumber="474" type="Museum">QCAZ</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4B85A5DF5861FF94FF7865BFC9BBFD31" pageId="23" pageNumber="474" type="diagnosis">
<paragraph id="0320F6545861FF94FF7865BFC9BBFD31" blockId="23.[151,1436,546,825]" pageId="23" pageNumber="474">
<emphasis id="31EB2A465861FF94FF7865BFCD79FD0C" bold="true" box="[199,322,690,716]" pageId="23" pageNumber="474">Diagnosis.</emphasis>
Females most resemble those of
<taxonomicName id="C49F8DD75861FF94FD1165BECF1FFD0C" authorityName="Dupérré &amp; Tapia" authorityYear="2024" box="[686,804,691,716]" class="Arachnida" family="Paratropididae" genus="Paratropis" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="cryptica" status="sp. nov.">
<emphasis id="31EB2A465861FF94FD1165BECF1FFD0C" box="[686,804,691,716]" italics="true" pageId="23" pageNumber="474">P. cryptica</emphasis>
</taxonomicName>
<emphasis id="31EB2A465861FF94FC9565BECFB9FD0D" bold="true" box="[810,898,691,717]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="2AD8973D5861FF94FC9565BECFB9FD0D" box="[810,898,691,717]" pageId="23" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but are distinguished by their spermathecal head being elongated, sinuous (
<figureCitation id="9BA4EAD15861FF94FE0365DACE7FFD31" box="[444,580,727,753]" captionStart="FIGURE 22" captionStartId="25.[151,250,1940,1964]" captionTargetBox="[161,1426,465,1909]" captionTargetId="figure-152@25.[151,1436,453,1917]" captionTargetPageId="25" captionText="FIGURE 22. Paratropis arenosa sp. nov. A. Female holotype, carapace dorsal view. B. Female holotype, sternum and endites ventral view. C. Spermathecae, dorsal view (holotype). D. Spermathecae, dorsal view (ECFN 7220). Scale bars:A, B = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916185" httpUri="https://zenodo.org/record/13916185/files/figure.png" pageId="23" pageNumber="474">Fig. 22C, D</figureCitation>
), while short and condensed spermathecal head in the latter (
<figureCitation id="9BA4EAD15861FF94FB5065DAC94FFD31" box="[1263,1396,726,753]" captionStart="FIGURE 25" captionStartId="28.[151,250,1943,1967]" captionTargetBox="[206,1380,193,1913]" captionTargetId="figure-23@28.[196,1392,181,1919]" captionTargetPageId="28" captionText="FIGURE 25. Paratropis cryptica sp. nov. female holotype. A. Carapace, dorsal view. B. Sternum and endites, ventral view. C. Spermathecae, dorsal view. D. Spermathecae, dorsal view (CASENT 470). E. Spermathecae, dorsal view (CASENT 448). F. Spermathecae, dorsal view (ECFN 1320). Scale bars: A, B = 1.0 mm, C–F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916191" httpUri="https://zenodo.org/record/13916191/files/figure.png" pageId="23" pageNumber="474">Fig. 25C–F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4B85A5DF5861FF94FF7865F6CFBDFCF9" pageId="23" pageNumber="474" type="etymology">
<paragraph id="0320F6545861FF94FF7865F6CFBDFCF9" blockId="23.[151,1436,546,825]" pageId="23" pageNumber="474">
<emphasis id="31EB2A465861FF94FF7865F6CD75FCD5" bold="true" box="[199,334,763,789]" pageId="23" pageNumber="474">Etymology.</emphasis>
The specific epithet is non-Latin adjective (invariable) taken from Spanish language meaning “with sand” in reference to the species covered with sand particles.
</paragraph>
</subSubSection>
<caption id="57E0A6DC5861FF94FF2861DACD6CF8D3" ID-DOI="http://doi.org/10.5281/zenodo.13916181" ID-Zenodo-Dep="13916181" httpUri="https://zenodo.org/record/13916181/files/figure.png" pageId="23" pageNumber="474" startId="23.[151,250,1751,1775]" targetBox="[179,1409,862,1715]" targetPageId="23" targetType="figure">
<paragraph id="0320F6545861FF94FF2861DACD6CF8D3" blockId="23.[151,1437,1751,1811]" pageId="23" pageNumber="474">
<emphasis id="31EB2A465861FF94FF2861DACD1BF92F" bold="true" box="[151,288,1751,1775]" pageId="23" pageNumber="474">FIGURE 20.</emphasis>
<taxonomicName id="C49F8DD75861FF94FE9861D5CDD7F92F" authorityName="Dupérré &amp; Tapia" authorityYear="2024" box="[295,492,1752,1775]" class="Arachnida" family="Paratropididae" genus="Paratropis" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="arenosa" status="sp. nov.">
<emphasis id="31EB2A465861FF94FE9861D5CDD7F92F" box="[295,492,1752,1775]" italics="true" pageId="23" pageNumber="474">Paratropis arenosa</emphasis>
</taxonomicName>
<emphasis id="31EB2A465861FF94FE4D61DACE78F92F" bold="true" box="[498,579,1751,1775]" pageId="23" pageNumber="474">
<taxonomicNameLabel id="2AD8973D5861FF94FE4D61DACE78F92F" box="[498,579,1751,1775]" pageId="23" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female ECFN 1323.
<emphasis id="31EB2A465861FF94FCA361DACF0EF92F" bold="true" box="[796,821,1751,1775]" pageId="23" pageNumber="474">A.</emphasis>
Live, Las Antenas, Sucumbíos Province, dorso-lateral view. Scale bar: 5.0 mm.
</paragraph>
</caption>
<subSubSection id="4B85A5DF5861FF99FF78604CC85EFF15" lastPageId="26" lastPageNumber="477" pageId="23" pageNumber="474" type="description">
<paragraph id="0320F6545861FF9AFF78604CC92DFE6D" blockId="23.[151,1437,1857,1992]" lastBlockId="25.[151,1437,150,429]" lastPageId="25" lastPageNumber="476" pageId="23" pageNumber="474">
<emphasis id="31EB2A465861FF94FF78604CCD62F89B" bold="true" box="[199,345,1857,1883]" pageId="23" pageNumber="474">Description.</emphasis>
Coloration (live female
<typeStatus id="DC2448F65861FF94FDD4604FCEF0F89C" box="[619,715,1858,1884]" pageId="23" pageNumber="474" type="paratype">paratype</typeStatus>
ECFN 1323). Female carapace dark brown, completely covered with soil particles; abdomen completely covered with soil particles; legs blackish, covered with soil particles (
<figureCitation id="9BA4EAD15861FF94FAD06068CCF7F863" captionStart="FIGURE 20" captionStartId="23.[151,250,1751,1775]" captionTargetBox="[179,1409,862,1715]" captionTargetId="figure-213@23.[166,1422,849,1728]" captionTargetPageId="23" captionText="FIGURE 20. Paratropis arenosa sp. nov. female ECFN 1323. A. Live, Las Antenas, Sucumbíos Province, dorso-lateral view. Scale bar: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13916181" httpUri="https://zenodo.org/record/13916181/files/figure.png" pageId="23" pageNumber="474">Fig. 20A</figureCitation>
).
<emphasis id="31EB2A465861FF94FF606084CDFBF863" bold="true" box="[223,448,1929,1955]" pageId="23" pageNumber="474">
(Female
<typeStatus id="DC2448F65861FF94FEF86084CD88F863" box="[327,435,1929,1955]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
):
</emphasis>
Total length: 13.82; carapace length: 5.90; carapace width: 6.51; abdomen length: 7.92. Coloration in ethanol. CEPHALOTHORAX: Carapace: reddish-brown, caput arched; fovea, slightly procurved (
<figureCitation id="9BA4EAD15861FF94FA9260A0C9B4F807" box="[1325,1423,1965,1991]" captionStart="FIGURE 22" captionStartId="25.[151,250,1940,1964]" captionTargetBox="[161,1426,465,1909]" captionTargetId="figure-152@25.[151,1436,453,1917]" captionTargetPageId="25" captionText="FIGURE 22. Paratropis arenosa sp. nov. A. Female holotype, carapace dorsal view. B. Female holotype, sternum and endites ventral view. C. Spermathecae, dorsal view (holotype). D. Spermathecae, dorsal view (ECFN 7220). Scale bars:A, B = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916185" httpUri="https://zenodo.org/record/13916185/files/figure.png" pageId="23" pageNumber="474">Fig. 22A</figureCitation>
). Chelicerae reddish-brown; promargin 11 teeth, retromargin 13 teeth. Labium sub-rectangular, 43 cuspules anteriorly (
<figureCitation id="9BA4EAD1586EFF9BFF20609ACD3DF871" box="[159,262,1943,1969]" captionStart="FIGURE 22" captionStartId="25.[151,250,1940,1964]" captionTargetBox="[161,1426,465,1909]" captionTargetId="figure-152@25.[151,1436,453,1917]" captionTargetPageId="25" captionText="FIGURE 22. Paratropis arenosa sp. nov. A. Female holotype, carapace dorsal view. B. Female holotype, sternum and endites ventral view. C. Spermathecae, dorsal view (holotype). D. Spermathecae, dorsal view (ECFN 7220). Scale bars:A, B = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916185" httpUri="https://zenodo.org/record/13916185/files/figure.png" pageId="24" pageNumber="475">Fig. 22B</figureCitation>
). Maxillae longer than wide, anterior prolateral lobe elongated, conical; 72 cuspules (
<figureCitation id="9BA4EAD1586EFF9BFB7B609AC910F871" box="[1220,1323,1943,1969]" captionStart="FIGURE 22" captionStartId="25.[151,250,1940,1964]" captionTargetBox="[161,1426,465,1909]" captionTargetId="figure-152@25.[151,1436,453,1917]" captionTargetPageId="25" captionText="FIGURE 22. Paratropis arenosa sp. nov. A. Female holotype, carapace dorsal view. B. Female holotype, sternum and endites ventral view. C. Spermathecae, dorsal view (holotype). D. Spermathecae, dorsal view (ECFN 7220). Scale bars:A, B = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916185" httpUri="https://zenodo.org/record/13916185/files/figure.png" pageId="24" pageNumber="475">Fig. 22B</figureCitation>
). Sternum length 2.96; width 3.51, oval; three pairs of oval sigilla (
<figureCitation id="9BA4EAD1586EFF9BFD4B60B6CF63F815" box="[756,856,1979,2005]" captionStart="FIGURE 22" captionStartId="25.[151,250,1940,1964]" captionTargetBox="[161,1426,465,1909]" captionTargetId="figure-152@25.[151,1436,453,1917]" captionTargetPageId="25" captionText="FIGURE 22. Paratropis arenosa sp. nov. A. Female holotype, carapace dorsal view. B. Female holotype, sternum and endites ventral view. C. Spermathecae, dorsal view (holotype). D. Spermathecae, dorsal view (ECFN 7220). Scale bars:A, B = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916185" httpUri="https://zenodo.org/record/13916185/files/figure.png" pageId="24" pageNumber="475">Fig. 22B</figureCitation>
). EYES: eight eyes on tubercle, elevated, height 0.91; AME: 0.21; ALE: 0.30: PLE:0.27; PME: 0.14; PME–PME 0.59. ABDOMEN. Dorsally dark gray, slightly encrusted with soil, with 5 transversal rows of thin spatulate setae on low tubercles; laterally gray fewer and smaller spatulate setae; ventrally light gray without spatulate setae (
<figureCitation id="9BA4EAD1586FFF9AFD7D67B6CF73FF15" box="[706,840,187,213]" captionStart="FIGURE 21" captionStartId="24.[151,248,1837,1861]" captionTargetBox="[170,1394,179,1800]" captionTargetId="figure-23@24.[153,1434,151,1813]" captionTargetPageId="24" captionText="FIGURE 21 Paratropis arenosa sp. nov. female holotype.A. Habitus, dorsal view. B. Habitus, ventral view. Scale bars: 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13916183" httpUri="https://zenodo.org/record/13916183/files/figure.png" pageId="25" pageNumber="476">Fig. 21A, B</figureCitation>
). LEGS: Uniformly reddish-brown, slightly encrusted with soil particles. Macrosetae present ventrally on tibia III–IV, metatarsi and tarsi II–IV. Spinules on leg I ventrally, metatarsus: 9; tarsus: 17; trichobothria: palpal tarsus: 8, tarsus I: 9, II: 7, III: 6, IV: 8/9; metatarsus I: 5, II: 4, III: 4, IV: 4; tarsal scopula absent. Leg measurements: I 17.33 (5.69/2.16/4.34/3.14/2.00); II 13.16 (4.28/2.11/2.60/2.46/1.71); III 12.82 (3.89/2.00/2.35/2.50/2.08); IV 18.33 (5.24/2.04/4.20/4.10/2.75); leg formula 4123. Tarsus I third claw present; tarsi II–IV without third claw; all paired claws with one tooth. GENITALIA. Two long spermathecae (5x longer than wide as wide); spermathecae head with elongated, sinuous vesicles, stalks elongated, sinuous (
<figureCitation id="9BA4EAD1586FFF9AFB1A669EC931FE6D" box="[1189,1290,403,429]" captionStart="FIGURE 22" captionStartId="25.[151,250,1940,1964]" captionTargetBox="[161,1426,465,1909]" captionTargetId="figure-152@25.[151,1436,453,1917]" captionTargetPageId="25" captionText="FIGURE 22. Paratropis arenosa sp. nov. A. Female holotype, carapace dorsal view. B. Female holotype, sternum and endites ventral view. C. Spermathecae, dorsal view (holotype). D. Spermathecae, dorsal view (ECFN 7220). Scale bars:A, B = 1.0 mm; C, D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13916185" httpUri="https://zenodo.org/record/13916185/files/figure.png" pageId="25" pageNumber="476">Fig. 22C</figureCitation>
).
</paragraph>
<caption id="57E0A6DC586EFF9BFF286020C9A6F884" ID-DOI="http://doi.org/10.5281/zenodo.13916183" ID-Zenodo-Dep="13916183" box="[151,1437,1837,1861]" httpUri="https://zenodo.org/record/13916183/files/figure.png" pageId="24" pageNumber="475" startId="24.[151,248,1837,1861]" targetBox="[170,1394,179,1800]" targetPageId="24" targetType="figure">
<paragraph id="0320F654586EFF9BFF286020C9A6F884" blockId="24.[151,1437,1837,1861]" box="[151,1437,1837,1861]" pageId="24" pageNumber="475">
<emphasis id="31EB2A46586EFF9BFF286020CD2CF885" bold="true" box="[151,279,1837,1861]" pageId="24" pageNumber="475">FIGURE 21</emphasis>
<taxonomicName id="C49F8DD7586EFF9BFEA26023CDE6F885" authorityName="Dupérré &amp; Tapia" authorityYear="2024" box="[285,477,1838,1861]" class="Arachnida" family="Paratropididae" genus="Paratropis" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="475" phylum="Arthropoda" rank="species" species="arenosa" status="sp. nov.">
<emphasis id="31EB2A46586EFF9BFEA26023CDE6F885" box="[285,477,1838,1861]" italics="true" pageId="24" pageNumber="475">Paratropis arenosa</emphasis>
</taxonomicName>
<emphasis id="31EB2A46586EFF9BFE5C6020CE09F885" bold="true" box="[483,562,1837,1861]" pageId="24" pageNumber="475">
<taxonomicNameLabel id="2AD8973D586EFF9BFE5C6020CE09F885" box="[483,562,1837,1861]" pageId="24" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
female holotype.
<emphasis id="31EB2A46586EFF9BFD5B6020CEC6F885" bold="true" box="[740,765,1837,1861]" pageId="24" pageNumber="475">A.</emphasis>
Habitus, dorsal view.
<emphasis id="31EB2A46586EFF9BFC656020CFCAF885" bold="true" box="[986,1009,1837,1861]" pageId="24" pageNumber="475">B.</emphasis>
Habitus, ventral view. Scale bars: 5.0 mm.
</paragraph>
</caption>
<caption id="57E0A6DC586FFF9AFF286099CD0CF834" ID-DOI="http://doi.org/10.5281/zenodo.13916185" ID-Zenodo-Dep="13916185" httpUri="https://zenodo.org/record/13916185/files/figure.png" pageId="25" pageNumber="476" startId="25.[151,250,1940,1964]" targetBox="[161,1426,465,1909]" targetPageId="25" targetType="figure">
<paragraph id="0320F654586FFF9AFF286099CD0CF834" blockId="25.[151,1436,1940,2036]" pageId="25" pageNumber="476">
<emphasis id="31EB2A46586FFF9AFF286099CD1BF86C" bold="true" box="[151,288,1940,1964]" pageId="25" pageNumber="476">FIGURE 22.</emphasis>
<taxonomicName id="C49F8DD7586FFF9AFE996098CDD0F86C" authorityName="Dupérré &amp; Tapia" authorityYear="2024" box="[294,491,1941,1964]" class="Arachnida" family="Paratropididae" genus="Paratropis" kingdom="Animalia" order="Araneae" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="arenosa" status="sp. nov.">
<emphasis id="31EB2A46586FFF9AFE996098CDD0F86C" box="[294,491,1941,1964]" italics="true" pageId="25" pageNumber="476">Paratropis arenosa</emphasis>
</taxonomicName>
<emphasis id="31EB2A46586FFF9AFE4E6099CE5BF86C" bold="true" box="[497,608,1940,1964]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="2AD8973D586FFF9AFE4E6099CE79F86C" box="[497,578,1940,1964]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Female holotype, carapace dorsal view.
<emphasis id="31EB2A46586FFF9AFC426099C82FF86C" bold="true" box="[1021,1044,1940,1964]" pageId="25" pageNumber="476">B.</emphasis>
Female holotype, sternum and endites ventral view.
<emphasis id="31EB2A46586FFF9AFEA060B5CD0CF810" bold="true" box="[287,311,1976,2000]" pageId="25" pageNumber="476">C.</emphasis>
Spermathecae, dorsal view (holotype).
<emphasis id="31EB2A46586FFF9AFD7960B5CEE4F810" bold="true" box="[710,735,1976,2000]" pageId="25" pageNumber="476">D.</emphasis>
Spermathecae, dorsal view (ECFN 7220). Scale bars: A, B = 1.0 mm; C, D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="0320F654586CFF99FF78679BC85EFF15" blockId="26.[151,1437,150,357]" pageId="26" pageNumber="477">
<emphasis id="31EB2A46586CFF99FF78679BCD04FF70" bold="true" box="[199,319,150,176]" pageId="26" pageNumber="477">Variation.</emphasis>
Female (ECFN 7220) carapace length: 7.50; abdomen length: 10.09; (ZMH-A0002999) carapace length: 8.19; abdomen length: 10.09. Female body length: 13.82–18.89,
<emphasis id="31EB2A46586CFF99FC7B67B7CFEBFF12" box="[964,976,186,210]" italics="true" pageId="26" pageNumber="477">x</emphasis>
17.05,
<emphasis id="31EB2A46586CFF99FB9767B6C872FF14" box="[1064,1097,187,212]" italics="true" pageId="26" pageNumber="477">n=</emphasis>
3.
</paragraph>
</subSubSection>
<subSubSection id="4B85A5DF586CFF99FF7867D3CE8CFF39" box="[199,695,222,249]" pageId="26" pageNumber="477" type="distribution">
<paragraph id="0320F654586CFF99FF7867D3CE8CFF39" blockId="26.[151,1437,150,357]" box="[199,695,222,249]" pageId="26" pageNumber="477">
<emphasis id="31EB2A46586CFF99FF7867D3CD5AFF38" bold="true" box="[199,353,222,248]" pageId="26" pageNumber="477">Distribution.</emphasis>
<collectingRegion id="C15B38B6586CFF99FED867D2CE74FF39" box="[359,591,223,249]" country="Ecuador" name="Sucumbios" pageId="26" pageNumber="477">Sucumbíos Province</collectingRegion>
(
<figureCitation id="9BA4EAD1586CFF99FDE267D2CE9DFF39" box="[605,678,223,249]" captionStart="MAP 1" captionStartId="33.[151,208,1803,1827]" captionTargetBox="[182,1406,415,1776]" captionTargetId="figure-134@33.[169,1418,401,1789]" captionTargetPageId="33" captionText="MAP 1. Distribution of newly described Paratropis. Paratropis carcosita sp. nov. (yellow circle), Paratropis pukallucha sp. nov. (orange circle), Paratropis esmeraldas sp. nov. (blue circle), Paratropis kapak sp. nov. (purple circle), Paratropis pasochoa sp. nov. (yellow star), Paratropis arenosa sp. nov. (red inverted triangle) and, Paratropis cryptica sp. nov. (green inverted triangle). Provinces abbreviations: Azuay (AZ), Bolívar (BO), Carchi (CA), Cañar (CAN), Chimborazo (CH), Cotopaxi (CO), El Oro (EO), Esmeralsdas (ES), Guayas (GU), Imbabura (IM), Loja (LO), Los Ríos (LR), Manabí (MA), Morona Santiago (MS), Napo (NA), Orellana (OR), Pastaza (PA), Pichincha (PI), Santa Elena (SE), Santo Domingo de los Tsáchilas (SD), Sucumbíos (SU), Tugurahua (TU), Zamora Chinchipe (ZC)." figureDoi="http://doi.org/10.5281/zenodo.13921361" httpUri="https://zenodo.org/record/13921361/files/figure.png" pageId="26" pageNumber="477">Map 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4B85A5DF586CFF99FF78660ECDA0FEA5" pageId="26" pageNumber="477" type="biology_ecology">
<paragraph id="0320F654586CFF99FF78660ECDA0FEA5" blockId="26.[151,1437,150,357]" pageId="26" pageNumber="477">
<emphasis id="31EB2A46586CFF99FF78660ECDB8FEDC" bold="true" box="[199,387,258,285]" pageId="26" pageNumber="477">Natural history.</emphasis>
Specimens were collected at
<quantity id="C4675BB1586CFF99FD7B660ECF32FEDD" box="[708,777,259,285]" metricMagnitude="2" metricUnit="m" metricValue="8.26" pageId="26" pageNumber="477" unit="m" value="826.0">826 m</quantity>
a.s.l. in an evergreen foothill forest of the Eastern Cordillera (BsPn03) (
<bibRefCitation id="670E8BA5586CFF99FEAF662ACE3BFE81" author="Guevara, J. E. &amp; Mogollon, H. &amp; Ceron, C. &amp; Josse, C." box="[272,512,295,321]" pageId="26" pageNumber="477" pagination="108 - 110" refId="ref11783" refString="Guevara, J. E., Mogollon, H., Ceron, C. &amp; Josse, C. (2013 b) Region Andes. Bosque siempreverde piemontano del Norte de la Cordillera Oriental de los Andes. In: Galeras, R., Guevara, J. E., Medina-Torres, B., Chinchero, M. A. &amp; Herrera, X. (Eds.), Sistema de Clasificacion de Ecosistemas del Ecuador Continental. Ministerio del Ambiente del Ecuador, Quito, pp. 108 - 110." type="book chapter" year="2013">
Guevara
<emphasis id="31EB2A46586CFF99FEC7662ACD8AFE81" box="[376,433,295,321]" italics="true" pageId="26" pageNumber="477">et al.</emphasis>
2013b
</bibRefCitation>
) and at
<quantity id="C4675BB1586CFF99FDE4662ACE98FE81" box="[603,675,295,321]" metricMagnitude="2" metricUnit="m" metricValue="3.01" pageId="26" pageNumber="477" unit="m" value="301.0">301 m</quantity>
in Aguarico-Putumayo-Caquetá lowland evergreen forest (BsTa01) (
<bibRefCitation id="670E8BA5586CFF99FF206646CDB5FEA5" author="Guevara, J. E. &amp; Pitman, N. &amp; Mogollon, H. &amp; Ceron, C. &amp; Palacios, W." box="[159,398,331,357]" pageId="26" pageNumber="477" pagination="175 - 177" refId="ref11882" refString="Guevara, J. E., Pitman, N., Mogollon, H., Ceron, C. &amp; Palacios, W. (2013 c) Region Amazonia. Bosque siempreverde de tierras bajas del Aguarico-Putumayo-Caqueta. In: Galeras, R., Guevara, J. E., Medina-Torres, B., Chinchero, M. A. &amp; Herrera, X. (Eds.), Sistema de Clasificacion de Ecosistemas del Ecuador Continental. Ministerio del Ambiente del Ecuador, Quito, pp. 175 - 177." type="book chapter" year="2013">
Guevara
<emphasis id="31EB2A46586CFF99FEB96646CD04FEA5" box="[262,319,331,357]" italics="true" pageId="26" pageNumber="477">et al.</emphasis>
2013c
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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