treatments-xml/data/7E/3D/87/7E3D87A0880A8009FE60BBBC006BF426.xml
2024-06-21 12:41:40 +02:00

239 lines
30 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="DC5CCD3CD5BBF7A3A6C24A4B744BC376" ID-DOI="10.5281/zenodo.7666308" ID-ISSN="2305-2562" ID-Zenodo-Dep="7666308" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1677091000000" checkinUser="felipe" docAuthor="Redman, Guy T., Hamer, Michelle L. &amp; Barraclough, David A." docDate="2003" docId="7E3D87A0880A8009FE60BBBC006BF426" docLanguage="en" docName="AfrInvertebr.44.2.203277.pdf" docOrigin="African Invertebrates 44 (2)" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Harpagophora dittoktenus Attems 1914" docType="treatment" docVersion="2" lastPageNumber="225" masterDocId="8204FFD8881E801FFFECBF750212F119" masterDocTitle="Revision of the Harpagophoridae (Diplopoda, Spirostreptida) of southern Africa, including descriptions of five new species" masterLastPageNumber="277" masterPageNumber="203" pageNumber="223" updateTime="1699455364425" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="CFDF660EBF889EDB02B96173CD247F35" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F36B7057388EC33EBBCBBACACC286E3D">
<mods:title id="85055A15BBA81BF1BACD0BA6EA5ADD6F">Revision of the Harpagophoridae (Diplopoda, Spirostreptida) of southern Africa, including descriptions of five new species</mods:title>
</mods:titleInfo>
<mods:name id="1F620A142484CFDB8E78C649D2D593DA" type="personal">
<mods:role id="2E4478E269023B38FBB1EBF1D2B5210C">
<mods:roleTerm id="AF86B8601AE8FA5FB81B1DA8EF1341B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9C7CC77EE869A1E164B549096EBADB78">Redman, Guy T.</mods:namePart>
</mods:name>
<mods:name id="4DC9E903D26FA76DF8CAFB72E9949FF2" type="personal">
<mods:role id="A9D0757D17924222EF737B697BF79E2B">
<mods:roleTerm id="F45D49B3E3387433B4684519904CE920">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A42E0425A8510317FEED6C38CDBB010C">Hamer, Michelle L.</mods:namePart>
</mods:name>
<mods:name id="B9753D90AF8DDE9B0410B2C21ED0DC92" type="personal">
<mods:role id="AE82C96DC9235A91AB8AD9436F544608">
<mods:roleTerm id="4224D4E7426BA9FA3CEBCAB1B4B7C06E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09806BC937F23DDB4CFACB0A10D54EDD">Barraclough, David A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="ED5D844C1CC8052A0D5EBD3862F94873">text</mods:typeOfResource>
<mods:relatedItem id="752333D5D8240E7382FDE9E36755DA14" type="host">
<mods:titleInfo id="0821180B2AA9B72691AA6E49995CF671">
<mods:title id="1BC050CB24EA67BB3E0E9F3069BC3C3B">African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part id="9BC6A0E958EFD70A00E43993FCDBC976">
<mods:date id="B7C96CCD18D7F29388F12FA693110A45">2003</mods:date>
<mods:detail id="A76D573E84EED48FD3583E37F01750AC" type="pubDate">
<mods:number id="E52A2E0B63B6A3F814293BAE934B6A24">2003-12-31</mods:number>
</mods:detail>
<mods:detail id="D58FB159BECBD5FE83513F0F56CE76F1" type="volume">
<mods:number id="CF5581B507A60B83C44B52D617A4FE9E">44</mods:number>
</mods:detail>
<mods:detail id="FEB92A481796B091493A64895D4C9C1B" type="issue">
<mods:number id="ACE7CDC3B67F96F0C43D87569378A348">2</mods:number>
</mods:detail>
<mods:extent id="0A6975E27431004B74366D1A4CECD0F8" unit="page">
<mods:start id="B9F866958871EF88DC522EAB329D6DBF">203</mods:start>
<mods:end id="FF0C1F361DF324414006193EB6964AFD">277</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A210E55FE4F0D6890F4E3C435E48A913">journal article</mods:classification>
<mods:identifier id="5223CE3E0A14245E4967E0127C4164A9" type="DOI">10.5281/zenodo.7666308</mods:identifier>
<mods:identifier id="8AB73C74B6E22581BE4FE3417D71C067" type="ISSN">2305-2562</mods:identifier>
<mods:identifier id="C988B1F310DCC59001C53980827CBF9F" type="Zenodo-Dep">7666308</mods:identifier>
</mods:mods>
<treatment id="7E3D87A0880A8009FE60BBBC006BF426" LSID="urn:lsid:plazi:treatment:7E3D87A0880A8009FE60BBBC006BF426" httpUri="http://treatment.plazi.org/id/7E3D87A0880A8009FE60BBBC006BF426" lastPageId="22" lastPageNumber="225" pageId="20" pageNumber="223">
<subSubSection id="BE8E653D880A800BFE60BBBC0141F5FB" box="[396,851,1225,1250]" pageId="20" pageNumber="223" type="nomenclature">
<paragraph id="F62B36B6880A800BFE60BBBC0141F5FB" blockId="20.[396,851,1225,1250]" box="[396,851,1225,1250]" pageId="20" pageNumber="223">
<heading id="AD6381DA880A800BFE60BBBC0141F5FB" box="[396,851,1225,1250]" centered="true" fontSize="11" level="2" pageId="20" pageNumber="223" reason="3">
<taxonomicName id="31944D35880A800BFE60BBBC0141F5FB" ID-CoL="6LBXS" authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[396,851,1225,1250]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="223" phylum="Arthropoda" rank="species" species="dittoktenus">
<emphasis id="C4E0EAA4880A800BFE60BBBC00A4F5F8" box="[396,694,1225,1249]" italics="true" pageId="20" pageNumber="223">Harpagophora dittoktenus</emphasis>
Attems, 1914
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BE8E653D880A800BFDCFBB8100AEF414" box="[547,700,1268,1293]" pageId="20" pageNumber="223" type="description">
<paragraph id="F62B36B6880A800BFDCFBB8100AEF414" blockId="20.[144,987,1268,1352]" box="[547,700,1268,1293]" pageId="20" pageNumber="223">
<figureCitation id="6EAF2A33880A800BFDCFBB81007AF414" box="[547,616,1268,1293]" captionStart="Fig" captionStartId="1.[182,215,845,865]" captionTargetBox="[251,1061,203,829]" captionTargetId="figure-7@1.[232,1095,193,840]" captionTargetPageId="1" captionText="Fig. 1. Distribution of Harpagophora, Metaphora and Poratophilus species in southern Africa. G = Gauteng; KZN = KwaZulu-Natal; L = Lesotho; MP = Mpumalanga; S = Swaziland; NW Prov = North West Province; W Cape = Western Cape." figureDoi="http://doi.org/10.5281/zenodo.7666310" httpUri="https://zenodo.org/record/7666310/files/figure.png" pageId="20" pageNumber="223">Figs 1</figureCitation>
,
<figureCitation id="6EAF2A33880A800BFD9ABB8100AEF414" box="[630,700,1268,1293]" captionStart="Figs 4549" captionStartId="21.[182,221,1505,1525]" captionTargetBox="[237,1100,263,1451]" captionTargetId="figure-70@21.[234,1111,268,1455]" captionTargetPageId="21" captionText="Figs 4549. Harpagophora dittoktenus Attems, 1914. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="20" pageNumber="223">4549</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BE8E653D880A800BFF7CBA69015EF451" pageId="20" pageNumber="223" type="reference_group">
<paragraph id="F62B36B6880A800BFF7CBA69015EF451" blockId="20.[144,987,1268,1352]" pageId="20" pageNumber="223">
<taxonomicName id="31944D35880A800BFF7CBA69003DF429" ID-CoL="6LBXS" authority="Attems, 1914: 165" authorityName="Attems" authorityPageNumber="165" authorityYear="1914" box="[144,559,1308,1328]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="223" phylum="Arthropoda" rank="species" species="dittoktenus">
<emphasis id="C4E0EAA4880A800BFF7CBA690392F429" box="[144,384,1308,1328]" italics="true" pageId="20" pageNumber="223">Harpagophora dittoktenus</emphasis>
Attems, 1914: 165
</taxonomicName>
; Attems 1928;
<bibRefCitation id="92054B47880A800BFD55BA69015AF429" author="LAWRENCE, R. F." box="[697,840,1308,1328]" pageId="20" pageNumber="223" pagination="23 - 62" refId="ref35036" refString="LAWRENCE, R. F. 1965. New Spirostreptidae and Harpagophoridae (Diplopoda) from southern Africa. Memorias do Instituto de Investigacao Cientifica de Mocambique 7: 23 - 62." type="journal article" year="1965">Lawrence 1965</bibRefCitation>
;
<bibRefCitation id="92054B47880A800BFCBDBA6901C5F429" author="SCHUBART, O." box="[849,983,1308,1328]" pageId="20" pageNumber="223" pagination="154 - 172" refId="ref35110" refString="SCHUBART, O. 1966. Diplopoda III. Pselaphognatha, Opisthospermophora, Colobognatha, zugleich Addenda und allgemeine Bemerkungen. In: Hanstrom, B., Brink, P., &amp; Rudebeck, G., eds, South African Animal Life. Stockholm: Swedish Natural Science Research Council 12: 154 - 172." type="journal article" year="1966">Schubart 1966</bibRefCitation>
.
<taxonomicName id="31944D35880A800BFF7CBA410021F451" ID-CoL="6LBXS" authorityName="Attems" authorityYear="1914" box="[144,563,1332,1352]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="20" pageNumber="223" phylum="Arthropoda" rank="species" species="dittoktenus" subGenus="Harpagophorella">
<emphasis id="C4E0EAA4880A800BFF7CBA41030AF451" box="[144,280,1332,1352]" italics="true" pageId="20" pageNumber="223">Harpagophora</emphasis>
(
<emphasis id="C4E0EAA4880A800BFEC9BA4103D1F451" box="[293,451,1332,1352]" italics="true" pageId="20" pageNumber="223">Harpagophorella</emphasis>
)
<emphasis id="C4E0EAA4880A800BFE3CBA410021F451" box="[464,563,1332,1352]" italics="true" pageId="20" pageNumber="223">dittoktenus</emphasis>
</taxonomicName>
;
<bibRefCitation id="92054B47880A800BFDD3BA4100DEF451" author="DEMANGE, J. - M." box="[575,716,1332,1352]" pageId="20" pageNumber="223" pagination="561 - 564" refId="ref34494" refString="DEMANGE, J. - M. 1983. Donnees nouvelles sur la famille des Harpagophoridae (Myriapoda, Diplopoda). Bulletin de Museum National d'Histoire Naturelle Paris (4) 5: 561 - 564." type="journal article" year="1983">Demange 1983</bibRefCitation>
;
<bibRefCitation id="92054B47880A800BFD39BA41015AF451" author="HAMER, M." box="[725,840,1332,1352]" pageId="20" pageNumber="223" pagination="11 - 82" refId="ref34653" refString="HAMER, M. 1998. Checklist of southern African millipedes. Annals of the Natal Museum 39: 11 - 82." type="journal article" year="1998">Hamer 1998</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BE8E653D880A8009FF7CBA2C006BF426" lastPageId="22" lastPageNumber="225" pageId="20" pageNumber="223" type="materials_examined">
<paragraph id="F62B36B6880A800BFF7CBA2C0015F4AB" blockId="20.[144,1104,1369,1666]" pageId="20" pageNumber="223">
<materialsCitation id="46FC3CEB880A800BFF7CBA2C0011F4AB" country="South Africa" location="Ashton" pageId="20" pageNumber="223" specimenCount="4" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="292F8814880A800BFF7CBA2C02D5F46B" box="[144,199,1369,1394]" pageId="20" pageNumber="223">Type</typeStatus>
material (not examined):
<collectingCountry id="8E837626880A800BFE0ABA2C00A0F46B" box="[486,690,1369,1394]" name="South Africa" pageId="20" pageNumber="223">SOUTH AFRICA</collectingCountry>
: 1ơ. The
<typeStatus id="292F8814880A800BFCF4BA2C015AF46B" box="[792,840,1369,1394]" pageId="20" pageNumber="223">type</typeStatus>
locality was given only as Cape and the
<typeStatus id="292F8814880A800BFEB8BA0C0396F48B" box="[340,388,1401,1426]" pageId="20" pageNumber="223">type</typeStatus>
specimens are housed in MNHN. The
<typeStatus id="292F8814880A800BFCDFBA0C0171F48B" box="[819,867,1401,1426]" pageId="20" pageNumber="223">type</typeStatus>
locality is here set as
<collectingRegion id="3450F854880A800BFF7CBAEC033CF4AB" box="[144,302,1433,1458]" country="South Africa" name="Western Cape" pageId="20" pageNumber="223">Western Cape</collectingRegion>
,
<location id="F34B606D880A800BFED4BAEC039BF4AB" LSID="urn:lsid:plazi:treatment:7E3D87A0880A8009FE60BBBC006BF426:F34B606D880A800BFED4BAEC039BF4AB" box="[312,393,1433,1458]" country="South Africa" name="Ashton" pageId="20" pageNumber="223" stateProvince="Western Cape">Ashton</location>
[3320CC]
</materialsCitation>
.
</paragraph>
<paragraph id="F62B36B6880A800BFF7CBAB40642F4E3" blockId="20.[144,1104,1369,1666]" pageId="20" pageNumber="223">
<materialsCitation id="46FC3CEB880A800BFF7CBAB4065EF4E3" collectingDate="1900-08" collectionCode="SAMC" collectorName="W. F. Purcell" country="South Africa" county="Material" location="Robertson district" municipality="Ashton" pageId="20" pageNumber="223" specimenCode="SAMC 1672" specimenCount="1" stateProvince="Western Cape">
<collectingCounty id="1F4A4E3A880A800BFF7CBAB402E0F4C3" box="[144,242,1473,1498]" pageId="20" pageNumber="223">Material</collectingCounty>
examined:
<collectingCountry id="8E837626880A800BFE6CBAB40045F4C3" box="[384,599,1473,1498]" name="South Africa" pageId="20" pageNumber="223">SOUTH AFRICA</collectingCountry>
:
<emphasis id="C4E0EAA4880A800BFD89BAB4011AF4C0" box="[613,776,1473,1497]" italics="true" pageId="20" pageNumber="223">
<collectingRegion id="3450F854880A800BFD89BAB4011AF4C0" box="[613,776,1473,1497]" country="South Africa" name="Western Cape" pageId="20" pageNumber="223">Western Cape</collectingRegion>
</emphasis>
: 2ơ3^,
<collectingMunicipality id="164FACCC880A800BFC9BBAB401D8F4C3" box="[887,970,1473,1498]" pageId="20" pageNumber="223">Ashton</collectingMunicipality>
[3320CC],
<location id="F34B606D880A800BFF7CBA94034AF4E3" LSID="urn:lsid:plazi:treatment:7E3D87A0880A8009FE60BBBC006BF426:F34B606D880A800BFF7CBA94034AF4E3" box="[144,344,1505,1530]" country="South Africa" county="Material" municipality="Ashton" name="Robertson district" pageId="20" pageNumber="223" stateProvince="Western Cape">Robertson district</location>
,
<location id="F34B606D880A800BFE8DBA9403B8F4E3" LSID="urn:lsid:plazi:treatment:7E3D87A0880A8009FE60BBBC006BF426:F34B606D880A800BFE8DBA9403B8F4E3" box="[353,426,1505,1530]" country="South Africa" county="Material" municipality="Ashton" name="Karoo" pageId="20" pageNumber="223" stateProvince="Western Cape">Karoo</location>
, in earth under bushes,
<date id="822A1076880A800BFD44BA94011DF4E3" box="[680,783,1505,1530]" pageId="20" pageNumber="223" value="1900-08">
<collectingDate id="926EE99E880A800BFD44BA94011DF4E3" box="[680,783,1505,1530]" pageId="20" pageNumber="223" value="1900-08">viii.1900</collectingDate>
</date>
,
<collectorName id="5B615360880A800BFCF4BA9401B6F4E3" box="[792,932,1505,1530]" pageId="20" pageNumber="223">W. F. Purcell</collectorName>
(
<specimenCode id="A6329ECD880A800BFC5DBA940656F4E3" box="[945,1092,1505,1530]" collectionCode="SAMC" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="20" pageNumber="223" type="Museum">SAMC 1672</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="F62B36B6880A800BFF7CB97C0100F79B" blockId="20.[144,1104,1369,1666]" pageId="20" pageNumber="223">
<emphasis id="C4E0EAA4880A800BFF7CB97C0310F738" box="[144,258,1545,1569]" italics="true" pageId="20" pageNumber="223">Diagnosis</emphasis>
: Apex of larger femoral spine with two acute projections of unequal length, longer of these directed outwards (
<figureCitation id="6EAF2A33880A800BFDF2B95C0066F75B" box="[542,628,1577,1602]" captionStart="Figs 4549" captionStartId="21.[182,221,1505,1525]" captionTargetBox="[237,1100,263,1451]" captionTargetId="figure-70@21.[234,1111,268,1455]" captionTargetPageId="21" captionText="Figs 4549. Harpagophora dittoktenus Attems, 1914. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="20" pageNumber="223">Fig. 47</figureCitation>
). Spine branch shorter than pectinophore, narrow and finger-like with spines along its margins (
<figureCitation id="6EAF2A33880A800BFD0DB93C0127F77B" box="[737,821,1609,1634]" captionStart="Figs 4549" captionStartId="21.[182,221,1505,1525]" captionTargetBox="[237,1100,263,1451]" captionTargetId="figure-70@21.[234,1111,268,1455]" captionTargetPageId="21" captionText="Figs 4549. Harpagophora dittoktenus Attems, 1914. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="20" pageNumber="223">Fig. 49</figureCitation>
). Distal medial margin of telocoxite with distally directed blunt lobes (
<figureCitation id="6EAF2A33880A800BFD6FB91C0114F79B" box="[643,774,1641,1666]" captionStart="Figs 4549" captionStartId="21.[182,221,1505,1525]" captionTargetBox="[237,1100,263,1451]" captionTargetId="figure-70@21.[234,1111,268,1455]" captionTargetPageId="21" captionText="Figs 4549. Harpagophora dittoktenus Attems, 1914. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="20" pageNumber="223">Figs 47, 48</figureCitation>
).
</paragraph>
<caption id="A2EB663E880B800AFF5ABA9401D0F724" ID-DOI="http://doi.org/10.5281/zenodo.7666328" ID-Zenodo-Dep="7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="21" pageNumber="224" startId="21.[182,221,1505,1525]" targetBox="[237,1100,263,1451]" targetPageId="21">
<paragraph id="F62B36B6880B800AFF5ABA9401D0F724" blockId="21.[182,1142,1505,1597]" pageId="21" pageNumber="224">
Figs 4549.
<taxonomicName id="31944D35880B800AFEC4BA94008EF4EC" authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[296,668,1505,1525]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="21" pageNumber="224" phylum="Arthropoda" rank="species" species="dittoktenus">
<emphasis id="C4E0EAA4880B800AFEC4BA94000BF4EC" box="[296,537,1505,1525]" italics="true" pageId="21" pageNumber="224">Harpagophora dittoktenus</emphasis>
Attems, 1914
</taxonomicName>
. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch.
</paragraph>
</caption>
<paragraph id="F62B36B688088009FF7CBFBB0309F1FE" blockId="22.[144,1104,206,1343]" box="[144,283,206,231]" pageId="22" pageNumber="225">Description:</paragraph>
<paragraph id="F62B36B688088009FF7CBF830177F056" blockId="22.[144,1104,206,1343]" pageId="22" pageNumber="225">
<emphasis id="C4E0EAA488088009FF7CBF830306F017" box="[144,276,246,270]" italics="true" pageId="22" pageNumber="225">Dimensions</emphasis>
: Males, n = 2; females, n = 3. Body width 7.07.1, female 6.78.2; collum width 6.56.5, female 5.87.0; body length 65.075.0; female 65.080.0; leg length 3.73.7; female 3.23.8; antenna length 4.54.5; female 4.05.0.
</paragraph>
<paragraph id="F62B36B688088009FF7CBE23006BF076" blockId="22.[144,1104,206,1343]" box="[144,633,342,367]" pageId="22" pageNumber="225">
<emphasis id="C4E0EAA488088009FF7CBE230366F077" box="[144,372,342,366]" italics="true" pageId="22" pageNumber="225">Number of segments</emphasis>
: 4749; female 4551.
</paragraph>
<paragraph id="F62B36B688088009FF7CBE0300DFF0D6" blockId="22.[144,1104,206,1343]" pageId="22" pageNumber="225">
<emphasis id="C4E0EAA488088009FF7CBE0302CDF097" box="[144,223,374,398]" italics="true" pageId="22" pageNumber="225">Colour</emphasis>
: Head, clypeus and collum brown. Prozonites light brown with greyish tinge, mesozonites light brown, metazonites dark brown. Anal valve light brown with greyish tinge. Caudal spine, legs and antennae light brown.
</paragraph>
<paragraph id="F62B36B688088009FF7CBEA303A1F0F6" blockId="22.[144,1104,206,1343]" box="[144,435,470,495]" pageId="22" pageNumber="225">
<emphasis id="C4E0EAA488088009FF7CBEA30334F0F7" box="[144,294,470,494]" italics="true" pageId="22" pageNumber="225">First ozopore</emphasis>
: Segment 6.
</paragraph>
<paragraph id="F62B36B688088009FF7CBE83035CF336" blockId="22.[144,1104,206,1343]" pageId="22" pageNumber="225">
<emphasis id="C4E0EAA488088009FF7CBE8302F0F317" box="[144,226,502,526]" italics="true" pageId="22" pageNumber="225">Collum</emphasis>
: Angle of anterior and lateral margins approximately 100˚. One submarginal groove (
<figureCitation id="6EAF2A3388088009FF01BD630353F336" box="[237,321,534,559]" captionStart="Figs 4549" captionStartId="21.[182,221,1505,1525]" captionTargetBox="[237,1100,263,1451]" captionTargetId="figure-70@21.[234,1111,268,1455]" captionTargetPageId="21" captionText="Figs 4549. Harpagophora dittoktenus Attems, 1914. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="22" pageNumber="225">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="F62B36B688088009FF7CBD43019FF296" blockId="22.[144,1104,206,1343]" pageId="22" pageNumber="225">
<emphasis id="C4E0EAA488088009FF7CBD430311F357" box="[144,259,566,590]" italics="true" pageId="22" pageNumber="225">Gonopods</emphasis>
: Larger femoral spine straight and partially concealed under fold on aboral surface of telocoxite.Apex of larger femoral spine with two acute projections of unequal length, longer of these directed outward. Smaller femoral spine directed outwards (
<figureCitation id="6EAF2A3388088009FBF2BD0302CBF3B6" captionStart="Figs 4549" captionStartId="21.[182,221,1505,1525]" captionTargetBox="[237,1100,263,1451]" captionTargetId="figure-70@21.[234,1111,268,1455]" captionTargetPageId="21" captionText="Figs 4549. Harpagophora dittoktenus Attems, 1914. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="22" pageNumber="225">Figs 47, 48</figureCitation>
). Pectinophore with series of long spiniform projections emerging from hyaline plate and not forming part of margin. Spine branch shorter than pectinophore, narrow and finger-like with spines along its margins (
<figureCitation id="6EAF2A3388088009FD97BDA300DFF3F6" box="[635,717,726,751]" captionStart="Figs 4549" captionStartId="21.[182,221,1505,1525]" captionTargetBox="[237,1100,263,1451]" captionTargetId="figure-70@21.[234,1111,268,1455]" captionTargetPageId="21" captionText="Figs 4549. Harpagophora dittoktenus Attems, 1914. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="22" pageNumber="225">Fig. 49</figureCitation>
). Distal medial margin of telocoxite with distally directed blunt lobe. Lateral margin broadly rounded.(
<figureCitation id="6EAF2A3388088009FCB4BD8301C1F216" box="[856,979,758,783]" captionStart="Figs 4549" captionStartId="21.[182,221,1505,1525]" captionTargetBox="[237,1100,263,1451]" captionTargetId="figure-70@21.[234,1111,268,1455]" captionTargetPageId="21" captionText="Figs 4549. Harpagophora dittoktenus Attems, 1914. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="22" pageNumber="225">Figs 47, 48</figureCitation>
). Telocoxal spine absent, however medial margin of distal part of telocoxite acutely depressed at base of distal part of telocoxite, about a third of gonopod length from distal margin in general position of telocoxal spine (
<figureCitation id="6EAF2A3388088009FDF0BC23008FF276" box="[540,669,854,879]" captionStart="Figs 4549" captionStartId="21.[182,221,1505,1525]" captionTargetBox="[237,1100,263,1451]" captionTargetId="figure-70@21.[234,1111,268,1455]" captionTargetPageId="21" captionText="Figs 4549. Harpagophora dittoktenus Attems, 1914. 45. Oral view of first pair of male legs. 46. Lateral view of collum; arrow indicates anterior end of millipede. 47. Aboral view of gonopods (SAMC 1672). 48. Aboral view of gonopods from same sample. 49. Apical elements of telopodite. f1 = large femoral spine; ml = median lobe; p = pectinophore; s = spine branch." figureDoi="http://doi.org/10.5281/zenodo.7666328" httpUri="https://zenodo.org/record/7666328/files/figure.png" pageId="22" pageNumber="225">Figs 47, 48</figureCitation>
). Aboral surface of telocoxite with fold adjacent to telocoxal spine partially concealing larger femoral spine.
</paragraph>
<paragraph id="F62B36B688088009FF7CBCE30341F2D6" blockId="22.[144,1104,206,1343]" pageId="22" pageNumber="225">
<emphasis id="C4E0EAA488088009FF7CBCE30339F2B7" box="[144,299,918,942]" italics="true" pageId="22" pageNumber="225">Pre-anal ring</emphasis>
: Caudal spine short, just surpassing margin of anal valve and slightly upturned distally.
</paragraph>
<paragraph id="F62B36B688088009FF7CBCAB03C1F50E" blockId="22.[144,1104,206,1343]" pageId="22" pageNumber="225">
Distribution:The only known record of
<taxonomicName id="31944D3588088009FDD5BCAB00DDF2EF" authorityName="Attems" authorityYear="1914" box="[569,719,990,1015]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="225" phylum="Arthropoda" rank="species" species="dittoktenus">
<emphasis id="C4E0EAA488088009FDD5BCAB005FF2EF" box="[569,589,990,1014]" italics="true" pageId="22" pageNumber="225">H</emphasis>
.
<emphasis id="C4E0EAA488088009FDB4BCAB00DDF2EF" box="[600,719,990,1014]" italics="true" pageId="22" pageNumber="225">dittoktenus</emphasis>
</taxonomicName>
is from Ashton (Robertson District) in the
<collectingRegion id="3450F85488088009FF39BC8B0363F50E" box="[213,369,1022,1047]" country="South Africa" name="Western Cape" pageId="22" pageNumber="225">Western Cape</collectingRegion>
(
<figureCitation id="6EAF2A3388088009FE6CBC8B03D0F50E" box="[384,450,1022,1047]" captionStart="Fig" captionStartId="1.[182,215,845,865]" captionTargetBox="[251,1061,203,829]" captionTargetId="figure-7@1.[232,1095,193,840]" captionTargetPageId="1" captionText="Fig. 1. Distribution of Harpagophora, Metaphora and Poratophilus species in southern Africa. G = Gauteng; KZN = KwaZulu-Natal; L = Lesotho; MP = Mpumalanga; S = Swaziland; NW Prov = North West Province; W Cape = Western Cape." figureDoi="http://doi.org/10.5281/zenodo.7666310" httpUri="https://zenodo.org/record/7666310/files/figure.png" pageId="22" pageNumber="225">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="F62B36B688088009FF7CBB530199F586" blockId="22.[144,1104,206,1343]" pageId="22" pageNumber="225">
Remarks: The identity of the material examined was confirmed by comparison with the original illustration.According to the original description and the key by Attems (1928),
<taxonomicName id="31944D3588088009FF7CBB130339F567" authorityName="Attems" authorityYear="1914" box="[144,299,1126,1151]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="225" phylum="Arthropoda" rank="species" species="dittoktenus">
<emphasis id="C4E0EAA488088009FF7CBB1302B6F567" box="[144,164,1126,1150]" italics="true" pageId="22" pageNumber="225">H</emphasis>
.
<emphasis id="C4E0EAA488088009FF5CBB130339F567" box="[176,299,1126,1150]" italics="true" pageId="22" pageNumber="225">dittoktenus</emphasis>
</taxonomicName>
does not have telocoxal spines, but both the specimens examined had the marginal depression which could be interpreted as an obscure spine.
</paragraph>
<paragraph id="F62B36B688088009FF46BBD30350F5E6" blockId="22.[144,1104,206,1343]" pageId="22" pageNumber="225">
The gonopods of this species very closely resemble those of
<emphasis id="C4E0EAA488088009FCD3BBD301C9F5A7" box="[831,987,1190,1214]" italics="true" pageId="22" pageNumber="225">
<taxonomicName id="31944D3588088009FCD3BBD301C4F5A7" authorityName="Attems" authorityYear="1909" box="[831,982,1190,1214]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="225" phylum="Arthropoda" rank="species" species="alokopyga">H. alokopyga</taxonomicName>
,
</emphasis>
especially in terms of the telocoxite. The apical elements of the telopodite, however, distinguish the two species.
</paragraph>
<paragraph id="F62B36B688088009FF46BA73006BF426" blockId="22.[144,1104,206,1343]" pageId="22" pageNumber="225">
The origin of the name
<taxonomicName id="31944D3588088009FE5EBA73003FF407" authorityName="Attems" authorityYear="1914" box="[434,557,1286,1310]" class="Diplopoda" family="Harpagophoridae" genus="Harpagophora" kingdom="Animalia" order="Spirostreptida" pageId="22" pageNumber="225" phylum="Arthropoda" rank="species" species="dittoktenus">
<emphasis id="C4E0EAA488088009FE5EBA73003FF407" box="[434,557,1286,1310]" italics="true" pageId="22" pageNumber="225">dittoktenus</emphasis>
</taxonomicName>
is unclear and it has therefore not been modified to agree with the gender of the genus name.
</paragraph>
</subSubSection>
</treatment>
</document>