263 lines
40 KiB
XML
263 lines
40 KiB
XML
<document id="1BF8BB03DB6D1842A218227A0E9FB9ED" ID-CLB-Dataset="55565" ID-DOI="10.5281/zenodo.6540166" ID-GBIF-Dataset="e616c9be-941b-46d5-ae65-12f0e84dcf4a" ID-ISSN="1536-9307" ID-Zenodo-Dep="6540166" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638264341924" checkinUser="felipe" docAuthor="Kovařík, František & Šťáhlavský, František" docDate="2019" docId="DC014066B8406D1CFC5C359FFE4CD7F8" docLanguage="en" docName="Euscorpius.295.1-45.pdf" docOrigin="Euscorpius 295" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Reddyanus hofereki Kovařík & Šťáhlavský 2019, sp. n." docType="treatment" docVersion="6" lastPageNumber="14" masterDocId="2038381EB8496D13FF863174FFA0D41A" masterDocTitle="Revision of the genus Reddyanus from Southeast Asia, with description of five new species from Cambodia, Malaysia, Thailand and Vietnam (Scorpiones: Buthidae)" masterLastPageNumber="45" masterPageNumber="1" pageNumber="8" updateTime="1699471341931" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="4F98BC5FCF44FCF346562689F9ADDBBE">Revision of the genus Reddyanus from Southeast Asia, with description of five new species from Cambodia, Malaysia, Thailand and Vietnam (Scorpiones: Buthidae)</mods:title>
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<mods:name id="0472BF352B75E2FE2694740304073A41" type="personal">
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<mods:namePart id="985CD3ABB0A836EADB437B0BF9EA17EB">Kovařík, František</mods:namePart>
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<mods:affiliation id="88043D194B161EC50E5BDDE2754A7E73">Department of Zoology, Charles University, ViničnÁ 7, CZ- 128 44 Praha 2, Czech Republic;</mods:affiliation>
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<mods:namePart id="857561F26FA32B4EBC821F5E53993B88">Šťáhlavský, František</mods:namePart>
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<mods:affiliation id="001DB2298C93DB13640515486FE29182">Department of Zoology, Charles University, ViničnÁ 7, CZ- 128 44 Praha 2, Czech Republic;</mods:affiliation>
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<mods:nameIdentifier id="411DAF99456BE7B932CC0B2BAC6AB3E9" type="URL">www.scorpio.cz</mods:nameIdentifier>
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<mods:date id="915982D7B54E39F889F9EB81CD940379">2019</mods:date>
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<treatment id="DC014066B8406D1CFC5C359FFE4CD7F8" ID-DOI="http://doi.org/10.5281/zenodo.6540251" ID-GBIF-Taxon="195300217" ID-Zenodo-Dep="6540251" LSID="urn:lsid:zoobank.org:act:52EF458C-1BBD-4E4D-9076-4A778C052891" httpUri="http://treatment.plazi.org/id/DC014066B8406D1CFC5C359FFE4CD7F8" lastPageId="15" lastPageNumber="14" pageId="9" pageNumber="8">
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<subSubSection id="1CB2A2FBB8406D1AFC5C359FFAB5D11C" box="[986,1301,1259,1286]" pageId="9" pageNumber="8" type="nomenclature">
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<paragraph id="5417F170B8406D1AFC5C359FFAB5D11C" blockId="9.[814,1473,1066,2022]" box="[986,1301,1259,1286]" pageId="9" pageNumber="8">
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<taxonomicName id="93A88AF3B8406D1AFC5C359FFB6CD11C" ID-CoL="6X57V" authority="Kovařík & Šťáhlavský, 2019" authorityName="Kovařík & Šťáhlavský" authorityYear="2019" box="[986,1228,1259,1286]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="hofereki" status="sp. nov.">Reddyanus hofereki</taxonomicName>
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.
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<emphasis id="66DC2D62B8406D1AFB7B3598FAADD11C" bold="true" box="[1277,1293,1260,1286]" pageId="9" pageNumber="8">n</emphasis>
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.
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<subSubSection id="1CB2A2FBB8406D1AFC133479FAF0D17F" pageId="9" pageNumber="8" type="description">
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<paragraph id="5417F170B8406D1AFC133479FAC1D13C" blockId="9.[814,1473,1066,2022]" box="[917,1377,1293,1318]" pageId="9" pageNumber="8">
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(
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<figureCitation id="CC93EDF5B8406D1AFC1A3479FB99D13C" box="[924,1081,1293,1318]" captionStart-0="Figures 37–42" captionStart-1="Figures 43–48" captionStart-2="Figures 49–66" captionStart-3="Figures 67–70" captionStart-4="Figure 71" captionStartId-0="10.[115,192,1981,2003]" captionStartId-1="11.[115,192,1980,2002]" captionStartId-2="12.[113,190,1892,1914]" captionStartId-3="13.[115,192,2009,2031]" captionStartId-4="14.[116,184,1221,1243]" captionTargetBox-0="[119,1465,258,1929]" captionTargetBox-1="[113,1474,240,1956]" captionTargetBox-2="[121,1471,249,1859]" captionTargetBox-3="[118,1474,322,1929]" captionTargetBox-4="[114,1472,234,1207]" captionTargetId-0="figure-66@10.[113,1474,238,1957]" captionTargetId-1="figure-54@11.[113,1474,240,1958]" captionTargetId-2="figure-126@12.[113,1474,234,1865]" captionTargetId-3="figure-26@13.[113,1474,316,1948]" captionTargetId-4="figure-467@14.[114,1472,231,1207]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionTargetPageId-4="14" captionText-0="Figures 37–42: Reddyanus hofereki sp. n. Figures 37–38, 42. Male holotype in dorsal (37) and ventral (38) views, and telson lateral (42). Figures 39–41. Female paratype in dorsal (39) and ventral (40) views, and telson lateral (41). Scale bar: 10 mm (37–40)." captionText-1="Figures 43–48: Reddyanus hofereki sp. n. Figures 43–45. Male holotype, carapace and tergites I–III (43), coxosternal area and sternites (44–45). Figures 46–48. Female paratype, carapace and tergites I–III (46), coxosternal area and sternites (47–48)." captionText-2="Figures 49–66: Reddyanus hofereki sp. n. Figures 49–50, 61–63. Female paratype, pedipalp chela dorsal (49) and external (50) views, metasoma and telson in lateral (61), dorsal (62) and ventral (63) views. Figures 51–60, 64–66. Male holotype, pedipalp chela dorsal (51), external (52) and ventral (53) views, patella dorsal (54), external (55) and ventral (56) views, femur and trochanter internal (57), and dorsal (58) views, movable finger (59) and fixed finger (60) dentition, metasoma and telson in lateral (64), dorsal (65) and ventral (66) views. Trichobothrial pattern is indicated in Figures 52–55 and 57–58. Scale bar: 10 mm (61–66)." captionText-3="Figures 67–70. Reddyanus hofereki sp. n., male holotype, left legs IV–I, retrolateral aspect." captionText-4="Figure 71. Reddyanus hofereki sp. n., type locality." figureDoi-0="http://doi.org/10.5281/zenodo.6540180" figureDoi-1="http://doi.org/10.5281/zenodo.6540184" figureDoi-2="http://doi.org/10.5281/zenodo.6540186" figureDoi-3="http://doi.org/10.5281/zenodo.6540188" figureDoi-4="http://doi.org/10.5281/zenodo.6540190" httpUri-0="https://zenodo.org/record/6540180/files/figure.png" httpUri-1="https://zenodo.org/record/6540184/files/figure.png" httpUri-2="https://zenodo.org/record/6540186/files/figure.png" httpUri-3="https://zenodo.org/record/6540188/files/figure.png" httpUri-4="https://zenodo.org/record/6540190/files/figure.png" pageId="9" pageNumber="8">Figures 37–71</figureCitation>
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,
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<figureCitation id="CC93EDF5B8406D1AFBC2347AFBCED13C" box="[1092,1134,1294,1318]" captionStart="Figures 192–202" captionStartId="40.[113,190,1922,1944]" captionTargetBox="[125,1468,264,1882]" captionTargetId="figure-11@40.[113,1474,255,1897]" captionTargetPageId="40" captionText="Figures 192–202: Reddyanus, males, pedipalp trochanter and femur, patella, and chela dorsal, metasoma II–III in lateral views. Figure 192. R. feti, holotype. Figure 193. R. furai sp. n., holotype. Figure 194. R. hofereki sp. n., holotype. Figure 195. R. jendeki, holotype. Figure 196. R. krasenskyi, Java, Ujung Kulon. Figure 197. R. majkusi sp. n., holotype. Figure 198. R. neradi, holotype. Figure 199. R. petrzelkai, holotype. Figure 200. R. rolciki sp. n., holotype. Figure 201. R. schwotti sp. n., Thailand, Khao sok. Figure 202. R. zideki, paratype." figureDoi="http://doi.org/10.5281/zenodo.6540239" httpUri="https://zenodo.org/record/6540239/files/figure.png" pageId="9" pageNumber="8">194</figureCitation>
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,
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<figureCitation id="CC93EDF5B8406D1AFBFC3479FB04D13F" box="[1146,1188,1293,1317]" captionStart="Figures 203–214" captionStartId="41.[113,190,1922,1944]" captionTargetBox="[119,1452,261,1882]" captionTargetId="figure-9@41.[113,1474,234,1887]" captionTargetPageId="41" captionText="Figures 203–214: Reddyanus, males, telson lateral. Figure 203. R. feti, holotype. Figure 204. R. furai sp. n., holotype. Figure 205. R. hofereki sp. n., holotype. Figure 206. R. jendeki, holotype. Figure 207. R. krasenskyi, Java, Ujung Kulon. Figure 208. R. majkusi sp. n., holotype. Figure 209. R. neradi, holotype. Figure 210. R. petrzelkai, holotype. Figure 211. R. rolciki sp. n., holotype. Figure 212. R. schwotti sp. n., holotype. Figure 213. R. schwotti sp. n., paratype from Thailand, Khao sok. Figure 214. R. zideki, paratype." figureDoi="http://doi.org/10.5281/zenodo.6540241" httpUri="https://zenodo.org/record/6540241/files/figure.png" pageId="9" pageNumber="8">205</figureCitation>
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,
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<figureCitation id="CC93EDF5B8406D1AFB293479FB79D13F" box="[1199,1241,1293,1317]" captionStart="Figure 225" captionStartId="43.[116,184,2017,2039]" captionTargetBox="[111,1472,234,1993]" captionTargetId="figure-9@43.[111,1472,234,1993]" captionTargetPageId="43" captionText="Figure 225. Map showing confirmed distribution of Reddyanus spp. in Southeast Asia." figureDoi="http://doi.org/10.5281/zenodo.6540247" httpUri="https://zenodo.org/record/6540247/files/figure.png" pageId="9" pageNumber="8">225</figureCitation>
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,
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<tableCitation id="192AC4CBB8406D1AFB623479FA9DD13C" box="[1252,1341,1293,1318]" captionStart="Table 1" captionStartId="9.[115,171,989,1010]" captionTargetBox="[120,1443,244,968]" captionTargetId="graphics-483@9.[114,1472,234,972]" captionTargetPageId="9" captionText="Table 1. Comparative measurements of adults of Reddyanus furai sp. n. and R. hofereki sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/00D7A1F8B8406D1AFFF532A9FD82D00A" pageId="9" pageNumber="8" tableUuid="00D7A1F8B8406D1AFFF532A9FD82D00A">Tables 1</tableCitation>
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,
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<tableCitation id="192AC4CBB8406D1AFACC347AFAF8D13C" box="[1354,1368,1294,1318]" captionStart="Table 4" captionStartId="39.[113,169,674,695]" captionTargetId="graphics-578@39.[117,1471,234,657]" captionTargetPageId="39" captionText="Table 4. The diploid numbers, the postpachytene configuration (mit-information only about mitosis), the percentage of the total chromosome" httpUri="http://table.plazi.org/id/00D7A1F8B86E6D34FFF733D6FB8CD6CF" pageId="9" pageNumber="8" tableUuid="00D7A1F8B86E6D34FFF733D6FB8CD6CF">4</tableCitation>
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)
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</paragraph>
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<emphasis id="66DC2D62B8406D1AFCB3345AFAF0D17F" bold="true" pageId="9" pageNumber="8">http://zoobank.org/urn:lsid:zoobank.org:act:891C12CABF4D-4CAC-AAF4-DD4BFC0A20BE</emphasis>
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</paragraph>
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<subSubSection id="1CB2A2FBB8406D1AFCA834FAFA2DD27C" pageId="9" pageNumber="8" type="materials_examined">
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<paragraph id="5417F170B8406D1AFCA834FAFC63D1FC" blockId="9.[814,1473,1066,2022]" pageId="9" pageNumber="8">
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TYPE LOCALITY AND TYPE REPOSITORY.
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<collectingCountry id="2CBFB1E0B8406D1AFB6C34F9FAEED1BF" box="[1258,1358,1421,1445]" name="Malaysia" pageId="9" pageNumber="8">Malaysia</collectingCountry>
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,
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<collectingRegion id="966C3F92B8406D1AFAE634F9FA60D1BF" box="[1376,1472,1421,1445]" country="Malaysia" name="Kelantan" pageId="9" pageNumber="8">Kelantan</collectingRegion>
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<collectingCounty id="BD7689FCB8406D1AFCA834D9FC25D1DF" box="[814,901,1453,1477]" pageId="9" pageNumber="8">Dabong</collectingCounty>
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,
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<collectingMunicipality id="B4736B0AB8406D1AFC1734DAFB8FD1DC" box="[913,1071,1454,1478]" pageId="9" pageNumber="8">Gunung Stong</collectingMunicipality>
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<geoCoordinate id="319C97B7B8406D1AFB5434D9FAD1D1DF" box="[1234,1393,1453,1477]" degrees="101" direction="east" minutes="58" orientation="longitude" pageId="9" pageNumber="8" precision="1" seconds="41.4" value="101.978165">101°58'41.4"E</geoCoordinate>
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<quantity id="93505C95B8406D1AFAFA34D9FA60D1DF" box="[1404,1472,1453,1477]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="8" unit="m" value="125.0">
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</quantity>
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a. s. l.;
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<collectionCode id="32B969B5B8406D1AFCFA34BAFC1FD1FC" box="[892,959,1486,1510]" pageId="9" pageNumber="8">FKCP</collectionCode>
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</paragraph>
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<paragraph id="5417F170B8406D1AFCA8377AFA2DD27C" blockId="9.[814,1473,1066,2022]" pageId="9" pageNumber="8">
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TYPE MATERIAL.
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<materialsCitation id="E4C0FB2DB8406D1AFC583779FC3DD27F" ID-GBIF-Occurrence="3766411335" collectingDate="2016-08-30" collectionCode="FKCP" collectorName="D. Hoferek" country="Malaysia" elevation="125" latitude="5.342917" location="Gunung Stong" longLatPrecision="1" longitude="101.978165" municipality="Dabong" pageId="9" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Kelantan" typeStatus="holotype">
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<emphasis id="66DC2D62B8406D1AFC583779FBE6D23F" bold="true" box="[990,1094,1549,1573]" pageId="9" pageNumber="8">
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<collectingCountry id="2CBFB1E0B8406D1AFC583779FBE6D23F" box="[990,1094,1549,1573]" name="Malaysia" pageId="9" pageNumber="8">Malaysia</collectingCountry>
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</emphasis>
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,
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<collectingRegion id="966C3F92B8406D1AFBD23779FB14D23F" box="[1108,1204,1549,1573]" country="Malaysia" name="Kelantan" pageId="9" pageNumber="8">Kelantan</collectingRegion>
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<collectingMunicipality id="B4736B0AB8406D1AFB3D3779FAB2D23F" box="[1211,1298,1549,1573]" pageId="9" pageNumber="8">Dabong</collectingMunicipality>
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,
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<location id="5177A7ABB8406D1AFA98377AFA1CD23C" LSID="urn:lsid:plazi:treatment:DC014066B8406D1CFC5C359FFE4CD7F8:5177A7ABB8406D1AFA98377AFA1CD23C" box="[1310,1468,1550,1574]" country="Malaysia" latitude="5.342917" longLatPrecision="1" longitude="101.978165" municipality="Dabong" name="Gunung Stong" pageId="9" pageNumber="8" stateProvince="Kelantan">Gunung Stong</location>
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<geoCoordinate id="319C97B7B8406D1AFC413759FBC5D25F" box="[967,1125,1581,1605]" degrees="101" direction="east" minutes="58" orientation="longitude" pageId="9" pageNumber="8" precision="1" seconds="41.4" value="101.978165">101°58'41.4"E</geoCoordinate>
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<elevation id="DF851643B8406D1AFBF63759FB5AD25F" box="[1136,1274,1581,1605]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="9" pageNumber="8" unit="m" value="125.0">
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a. s. l.
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</elevation>
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,
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<collectingDate id="30522E58B8406D1AFA803759FA34D25F" box="[1286,1428,1581,1605]" pageId="9" pageNumber="8">
|
||
<date id="2016D7B0B8406D1AFA803759FAF6D25F" box="[1286,1366,1581,1605]" pageId="9" pageNumber="8" value="2016-08-30">30.VIII</date>
|
||
.2016
|
||
</collectingDate>
|
||
,
|
||
<specimenCount id="42AE3AF9B8406D1AFA19375AFA60D25C" box="[1439,1472,1582,1606]" pageId="9" pageNumber="8" type="male">1♂</specimenCount>
|
||
(
|
||
<typeStatus id="8B134FD2B8406D1AFCB03739FC37D27F" box="[822,919,1613,1637]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
)
|
||
</materialsCitation>
|
||
<materialsCitation id="E4C0FB2DB8406D1AFC223739FA2DD27C" ID-GBIF-Occurrence="3766411361" box="[932,1421,1613,1638]" pageId="9" pageNumber="8" typeStatus="paratype">
|
||
<specimenCount id="42AE3AF9B8406D1AFC223739FC1DD27F" box="[932,957,1613,1637]" pageId="9" pageNumber="8" type="female">1♀</specimenCount>
|
||
2juvs. (
|
||
<typeStatus id="8B134FD2B8406D1AFB92373AFB20D27C" box="[1044,1152,1614,1638]" pageId="9" pageNumber="8" type="paratype">paratypes</typeStatus>
|
||
), leg.
|
||
<collectorName id="F95D94A6B8406D1AFB46373AFA9AD27F" box="[1216,1338,1613,1637]" pageId="9" pageNumber="8">D. Hoferek</collectorName>
|
||
,
|
||
<collectionCode id="32B969B5B8406D1AFAC3373AFA28D27C" box="[1349,1416,1614,1638]" pageId="9" pageNumber="8">FKCP</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1CB2A2FBB8406D1AFCA837FAFA1BD2FC" pageId="9" pageNumber="8" type="etymology">
|
||
<paragraph id="5417F170B8406D1AFCA837FAFA1BD2FC" blockId="9.[814,1473,1066,2022]" pageId="9" pageNumber="8">
|
||
ETYMOLOGY. The species epithet is a patronym honoring David Hoferek, the collector of
|
||
<typeStatus id="8B134FD2B8406D1AFB0C37DAFB62D2DC" box="[1162,1218,1710,1734]" pageId="9" pageNumber="8">types</typeStatus>
|
||
of the new species. He is also the author of the photograph of
|
||
<typeStatus id="8B134FD2B8406D1AFB4A37BAFB5AD2FC" box="[1228,1274,1742,1766]" pageId="9" pageNumber="8">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="CC93EDF5B8406D1AFAD937B9FA0FD2FC" box="[1375,1455,1741,1766]" captionStart="Figure 71" captionStartId="14.[116,184,1221,1243]" captionTargetBox="[114,1472,234,1207]" captionTargetId="figure-467@14.[114,1472,231,1207]" captionTargetPageId="14" captionText="Figure 71. Reddyanus hofereki sp. n., type locality." figureDoi="http://doi.org/10.5281/zenodo.6540190" httpUri="https://zenodo.org/record/6540190/files/figure.png" pageId="9" pageNumber="8">Fig. 71</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1CB2A2FBB8406D1DFCA8367AFDCAD1C8" lastPageId="14" lastPageNumber="13" pageId="9" pageNumber="8" type="diagnosis">
|
||
<paragraph id="5417F170B8406D1DFCA8367AFDCAD1C8" blockId="9.[814,1473,1066,2022]" lastBlockId="14.[113,772,1274,1490]" lastPageId="14" lastPageNumber="13" pageId="9" pageNumber="8">
|
||
DIAGNOSIS. Adult from
|
||
<quantity id="93505C95B8406D1AFBBC367AFB2AD33F" box="[1082,1162,1806,1830]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="9" pageNumber="8" unit="mm" value="31.0">31 mm</quantity>
|
||
(female) to
|
||
<quantity id="93505C95B8406D1AFA983679FACED33F" box="[1310,1390,1805,1829]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="8" unit="mm" value="35.0">35 mm</quantity>
|
||
(male) long. Male has longer metasomal segments and telson. Legs and metasoma yellow, with brown spots. Base coloration of pedipalp yellow with dark spots, identical on femur and patella. Manus of pedipalps yellow with black spots, fingers black. Metasoma V and partly IV and telson black in male. First metasomal segment bears 10 carinae, second through fourth bear eight carinae, lateral carinae on metasoma II represented only by several granules in posterior part. Metasoma V bears five carinae which can be reduced. Terminal tubercle on second and third metasomal segments of male slightly enlarged. Telson elongate in male, shorter in female. Subaculear tooth large but not rounded, dorsally with 3–4 granules in two rows. Pectinal teeth number
|
||
<quantity id="93505C95B8476D1DFEE734CEFE36D1C8" box="[353,406,1466,1490]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="14" pageNumber="13" unit="in" value="11.0">11 in</quantity>
|
||
male,
|
||
<quantity id="93505C95B8476D1DFE5B34CEFDB4D1C8" box="[477,532,1466,1490]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="13" unit="in" value="10.0">10 in</quantity>
|
||
female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1CB2A2FBB8476D1CFFF7348EFD50D7B9" lastPageId="15" lastPageNumber="14" pageId="14" pageNumber="13" type="description">
|
||
<paragraph id="5417F170B8476D1DFFF7348EFD6AD288" blockId="14.[113,772,1529,2034]" pageId="14" pageNumber="13">
|
||
DESCRIPTION. The adults are
|
||
<quantity id="93505C95B8476D1DFE25348EFE4CD20B" box="[419,492,1530,1554]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="14" pageNumber="13" unit="mm" value="31.0">31 mm</quantity>
|
||
(female
|
||
<typeStatus id="8B134FD2B8476D1DFDCC348EFD08D208" box="[586,680,1530,1554]" pageId="14" pageNumber="13" type="paratype">paratype</typeStatus>
|
||
) –
|
||
<quantity id="93505C95B8476D1DFD41348DFF3AD22B" metricMagnitude="-2" metricUnit="m" metricValue="3.532" pageId="14" pageNumber="13" unit="mm" value="35.32">35.32 mm</quantity>
|
||
(male
|
||
<typeStatus id="8B134FD2B8476D1DFF6E376EFEE7D228" box="[232,327,1562,1586]" pageId="14" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
) long. The habitus is shown in
|
||
<figureCitation id="CC93EDF5B8476D1DFD18376EFF2FD248" captionStart="Figures 37–42" captionStartId="10.[115,192,1981,2003]" captionTargetBox="[119,1465,258,1929]" captionTargetId="figure-66@10.[113,1474,238,1957]" captionTargetPageId="10" captionText="Figures 37–42: Reddyanus hofereki sp. n. Figures 37–38, 42. Male holotype in dorsal (37) and ventral (38) views, and telson lateral (42). Figures 39–41. Female paratype in dorsal (39) and ventral (40) views, and telson lateral (41). Scale bar: 10 mm (37–40)." figureDoi="http://doi.org/10.5281/zenodo.6540180" httpUri="https://zenodo.org/record/6540180/files/figure.png" pageId="14" pageNumber="13">Figs. 37– 40</figureCitation>
|
||
. For position and distribution of trichobothria of pedipalps see
|
||
<figureCitation id="CC93EDF5B8476D1DFF1A372EFE80D26B" box="[156,288,1625,1650]" captionStart="Figures 49–66" captionStartId="12.[113,190,1892,1914]" captionTargetBox="[121,1471,249,1859]" captionTargetId="figure-126@12.[113,1474,234,1865]" captionTargetPageId="12" captionText="Figures 49–66: Reddyanus hofereki sp. n. Figures 49–50, 61–63. Female paratype, pedipalp chela dorsal (49) and external (50) views, metasoma and telson in lateral (61), dorsal (62) and ventral (63) views. Figures 51–60, 64–66. Male holotype, pedipalp chela dorsal (51), external (52) and ventral (53) views, patella dorsal (54), external (55) and ventral (56) views, femur and trochanter internal (57), and dorsal (58) views, movable finger (59) and fixed finger (60) dentition, metasoma and telson in lateral (64), dorsal (65) and ventral (66) views. Trichobothrial pattern is indicated in Figures 52–55 and 57–58. Scale bar: 10 mm (61–66)." figureDoi="http://doi.org/10.5281/zenodo.6540186" httpUri="https://zenodo.org/record/6540186/files/figure.png" pageId="14" pageNumber="13">Figs. 52–55</figureCitation>
|
||
, and 57–58. Sexual dimorphism: adult male with longer and narrow metasomal segments and telson.
|
||
</paragraph>
|
||
<paragraph id="5417F170B8476D1DFFF737EDFE3ED3E8" blockId="14.[113,772,1529,2034]" pageId="14" pageNumber="13">
|
||
<emphasis id="66DC2D62B8476D1DFFF737EDFF4CD2AB" bold="true" box="[113,236,1689,1713]" pageId="14" pageNumber="13">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="CC93EDF5B8476D1DFE8437EEFE2DD2A8" box="[258,397,1690,1714]" captionStart="Figures 37–42" captionStartId="10.[115,192,1981,2003]" captionTargetBox="[119,1465,258,1929]" captionTargetId="figure-66@10.[113,1474,238,1957]" captionTargetPageId="10" captionText="Figures 37–42: Reddyanus hofereki sp. n. Figures 37–38, 42. Male holotype in dorsal (37) and ventral (38) views, and telson lateral (42). Figures 39–41. Female paratype in dorsal (39) and ventral (40) views, and telson lateral (41). Scale bar: 10 mm (37–40)." figureDoi="http://doi.org/10.5281/zenodo.6540180" httpUri="https://zenodo.org/record/6540180/files/figure.png" pageId="14" pageNumber="13">Figs. 37–40</figureCitation>
|
||
). The base color is yellow with dark spots. The chelicera is yellow, strongly reticulated. The fingers of chelicerae are spotted. The carapace and mesosoma are with a pattern that forms three dark longitudinal strips. The tergits with two symmetrical yellow spots on posterior part. The ventral side of the mesosoma is reddish brown. Base coloration of pedipalp yellow with dark spots, identical on femur and patella. The legs are also yellow with large black spots. The metasoma I–III is yellowish or redish brown with black spots, metasoma IV–V is black in male and reddish black and spotted in female.
|
||
</paragraph>
|
||
<paragraph id="5417F170B8476D1DFCB6358DFC63D2CB" blockId="14.[816,1475,1273,2034]" pageId="14" pageNumber="13">
|
||
<emphasis id="66DC2D62B8476D1DFCB6358DFBECD108" bold="true" box="[816,1100,1273,1298]" pageId="14" pageNumber="13">Carapace and mesosoma</emphasis>
|
||
(
|
||
<figureCitation id="CC93EDF5B8476D1DFBDD358EFB7ED108" box="[1115,1246,1274,1298]" captionStart-0="Figures 37–42" captionStart-1="Figures 43–48" captionStartId-0="10.[115,192,1981,2003]" captionStartId-1="11.[115,192,1980,2002]" captionTargetBox-0="[119,1465,258,1929]" captionTargetBox-1="[113,1474,240,1956]" captionTargetId-0="figure-66@10.[113,1474,238,1957]" captionTargetId-1="figure-54@11.[113,1474,240,1958]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="Figures 37–42: Reddyanus hofereki sp. n. Figures 37–38, 42. Male holotype in dorsal (37) and ventral (38) views, and telson lateral (42). Figures 39–41. Female paratype in dorsal (39) and ventral (40) views, and telson lateral (41). Scale bar: 10 mm (37–40)." captionText-1="Figures 43–48: Reddyanus hofereki sp. n. Figures 43–45. Male holotype, carapace and tergites I–III (43), coxosternal area and sternites (44–45). Figures 46–48. Female paratype, carapace and tergites I–III (46), coxosternal area and sternites (47–48)." figureDoi-0="http://doi.org/10.5281/zenodo.6540180" figureDoi-1="http://doi.org/10.5281/zenodo.6540184" httpUri-0="https://zenodo.org/record/6540180/files/figure.png" httpUri-1="https://zenodo.org/record/6540184/files/figure.png" pageId="14" pageNumber="13">Figs. 37–48</figureCitation>
|
||
). The entire carapace is covered with large granules; carinae are absent. The anterior margin of the carapace is medially convex. The mesosoma bears one median carina and is strongly granulated. Tergite VII is pentacarinate. The pectinal tooth count is
|
||
<quantity id="93505C95B8476D1DFACA340EFA25D188" box="[1356,1413,1402,1426]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="14" pageNumber="13" unit="in" value="11.0">11 in</quantity>
|
||
male
|
||
<typeStatus id="8B134FD2B8476D1DFCB634EEFC2DD1A8" box="[816,909,1434,1458]" pageId="14" pageNumber="13" type="holotype">holotype</typeStatus>
|
||
and
|
||
<quantity id="93505C95B8476D1DFC5034EEFBB7D1A8" box="[982,1047,1434,1458]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="13" unit="in" value="10.0">10 in</quantity>
|
||
female
|
||
<typeStatus id="8B134FD2B8476D1DFB0734EEFB40D1A8" box="[1153,1248,1434,1458]" pageId="14" pageNumber="13" type="paratype">paratype</typeStatus>
|
||
. The
|
||
<specimenCount id="42AE3AF9B8476D1DFAA934EEFA62D1A8" box="[1327,1474,1434,1458]" pageId="14" pageNumber="13" type="subadult">two subadult</specimenCount>
|
||
<typeStatus id="8B134FD2B8476D1DFCB634CEFC36D1C8" box="[816,918,1466,1490]" pageId="14" pageNumber="13" type="paratype">paratypes</typeStatus>
|
||
have 10–11 pectinal tooth. The pectine marginal tips extend to the end of the third sternite in the male and to the third quorter of the third sternite in the female. The pectines have three marginal lamellae and 8–9 middle lamellae. The lamellae and fulcra bear numerous light setae. Sternites are smooth with two parallel furrows except sternite VII which bears four incomplete carinae and is smooth. The glabrous zone in the middle of the posterior margin of sternite V present in both sexes.
|
||
</paragraph>
|
||
<paragraph id="5417F170B8476D1CFCB637AEFD44D558" blockId="14.[816,1475,1273,2034]" lastBlockId="15.[113,772,235,739]" lastPageId="15" lastPageNumber="14" pageId="14" pageNumber="13">
|
||
<emphasis id="66DC2D62B8476D1DFCB637AEFBBFD2E8" bold="true" box="[816,1055,1754,1778]" pageId="14" pageNumber="13">Metasoma and telson</emphasis>
|
||
(
|
||
<figureCitation id="CC93EDF5B8476D1DFBAA37AEFB0BD2E8" box="[1068,1195,1754,1778]" captionStart="Figures 37–42" captionStartId="10.[115,192,1981,2003]" captionTargetBox="[119,1465,258,1929]" captionTargetId="figure-66@10.[113,1474,238,1957]" captionTargetPageId="10" captionText="Figures 37–42: Reddyanus hofereki sp. n. Figures 37–38, 42. Male holotype in dorsal (37) and ventral (38) views, and telson lateral (42). Figures 39–41. Female paratype in dorsal (39) and ventral (40) views, and telson lateral (41). Scale bar: 10 mm (37–40)." figureDoi="http://doi.org/10.5281/zenodo.6540180" httpUri="https://zenodo.org/record/6540180/files/figure.png" pageId="14" pageNumber="13">Figs. 41–42</figureCitation>
|
||
,
|
||
<figureCitation id="CC93EDF5B8476D1DFB3337ADFB5CD2EB" box="[1205,1276,1753,1777]" captionStart="Figures 49–66" captionStartId="12.[113,190,1892,1914]" captionTargetBox="[121,1471,249,1859]" captionTargetId="figure-126@12.[113,1474,234,1865]" captionTargetPageId="12" captionText="Figures 49–66: Reddyanus hofereki sp. n. Figures 49–50, 61–63. Female paratype, pedipalp chela dorsal (49) and external (50) views, metasoma and telson in lateral (61), dorsal (62) and ventral (63) views. Figures 51–60, 64–66. Male holotype, pedipalp chela dorsal (51), external (52) and ventral (53) views, patella dorsal (54), external (55) and ventral (56) views, femur and trochanter internal (57), and dorsal (58) views, movable finger (59) and fixed finger (60) dentition, metasoma and telson in lateral (64), dorsal (65) and ventral (66) views. Trichobothrial pattern is indicated in Figures 52–55 and 57–58. Scale bar: 10 mm (61–66)." figureDoi="http://doi.org/10.5281/zenodo.6540186" httpUri="https://zenodo.org/record/6540186/files/figure.png" pageId="14" pageNumber="13">61–66</figureCitation>
|
||
). The first segment bears 10 carinae. The metasoma II–IV bear eight carinae, lateral carinae on metasoma II represented only by several granules in posterior part. The fifth segment bears five carinae from which the dorsal in male are reduced. Ventral carina is present on the telson in male and ventral and lateral in female. Surfaces between carinae are obviously smooth, with small, sparse granules that become more numerous on metasoma I– II in female. Terminal tubercle on the second and the third metasomal segments of both sexes only very slightly enlarged. The telson elongate in male, shorter in female. Subaculear tooth not rounded, dorsally with 3–4 granules in two rows.
|
||
</paragraph>
|
||
<paragraph id="5417F170B8466D1CFFF7303EFDD9D639" blockId="15.[113,772,235,739]" pageId="15" pageNumber="14">
|
||
<emphasis id="66DC2D62B8466D1CFFF7303EFF40D578" bold="true" box="[113,224,330,354]" pageId="15" pageNumber="14">Pedipalps</emphasis>
|
||
(
|
||
<figureCitation id="CC93EDF5B8466D1CFF70303FFE20D579" box="[246,384,331,355]" captionStart="Figures 49–66" captionStartId="12.[113,190,1892,1914]" captionTargetBox="[121,1471,249,1859]" captionTargetId="figure-126@12.[113,1474,234,1865]" captionTargetPageId="12" captionText="Figures 49–66: Reddyanus hofereki sp. n. Figures 49–50, 61–63. Female paratype, pedipalp chela dorsal (49) and external (50) views, metasoma and telson in lateral (61), dorsal (62) and ventral (63) views. Figures 51–60, 64–66. Male holotype, pedipalp chela dorsal (51), external (52) and ventral (53) views, patella dorsal (54), external (55) and ventral (56) views, femur and trochanter internal (57), and dorsal (58) views, movable finger (59) and fixed finger (60) dentition, metasoma and telson in lateral (64), dorsal (65) and ventral (66) views. Trichobothrial pattern is indicated in Figures 52–55 and 57–58. Scale bar: 10 mm (61–66)." figureDoi="http://doi.org/10.5281/zenodo.6540186" httpUri="https://zenodo.org/record/6540186/files/figure.png" pageId="15" pageNumber="14">Figs. 49–60</figureCitation>
|
||
). The pedipalps are very sparsely hirsute. The femur bears five and patella seven granulated carina. The chela is with four granulated carinae in both sexes. The chela and patella are slightly and femur is densely granulated. Movable finger with six and fixed finger with seven rows of granules, both with several accessory granules and 6 external and 5 internal accessory granules.
|
||
</paragraph>
|
||
<paragraph id="5417F170B8466D1CFFF7335FFD49D6D9" blockId="15.[113,772,235,739]" pageId="15" pageNumber="14">
|
||
<emphasis id="66DC2D62B8466D1CFFF7335FFF06D659" bold="true" box="[113,166,555,579]" pageId="15" pageNumber="14">Legs</emphasis>
|
||
(
|
||
<figureCitation id="CC93EDF5B8466D1CFF32335FFE94D659" box="[180,308,555,579]" captionStart="Figures 67–70" captionStartId="13.[115,192,2009,2031]" captionTargetBox="[118,1474,322,1929]" captionTargetId="figure-26@13.[113,1474,316,1948]" captionTargetPageId="13" captionText="Figures 67–70. Reddyanus hofereki sp. n., male holotype, left legs IV–I, retrolateral aspect." figureDoi="http://doi.org/10.5281/zenodo.6540188" httpUri="https://zenodo.org/record/6540188/files/figure.png" pageId="15" pageNumber="14">Figs. 67–70</figureCitation>
|
||
). The legs without tibial spurs. The tibia and tarsomeres of legs with setae not arranged into bristle combs on dorsal surfaces. Tarsomere II of leg IV with two sparse rows of <20 spiniform setae on ventral surface. The femur bears 3–4 and patella, 5 carinae; both femur and patella are granulated.
|
||
</paragraph>
|
||
<paragraph id="5417F170B8466D1CFFF733BFFE08D6F9" blockId="15.[113,772,235,739]" box="[113,424,715,739]" pageId="15" pageNumber="14">
|
||
<emphasis id="66DC2D62B8466D1CFFF733BFFEB8D6F9" bold="true" box="[113,280,715,739]" pageId="15" pageNumber="14">Measurements</emphasis>
|
||
. See
|
||
<tableCitation id="192AC4CBB8466D1CFED433BFFE02D6F9" box="[338,418,715,739]" captionStart="Table 1" captionStartId="9.[115,171,989,1010]" captionTargetBox="[120,1443,244,968]" captionTargetId="graphics-483@9.[114,1472,234,972]" captionTargetPageId="9" captionText="Table 1. Comparative measurements of adults of Reddyanus furai sp. n. and R. hofereki sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/00D7A1F8B8406D1AFFF532A9FD82D00A" pageId="15" pageNumber="14" tableUuid="00D7A1F8B8406D1AFFF532A9FD82D00A">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5417F170B8466D1CFFF7327FFD50D7B9" blockId="15.[113,771,779,931]" pageId="15" pageNumber="14">
|
||
AFFINITIES. The described features distinguish
|
||
<taxonomicName id="93A88AF3B8466D1CFDD4327FFD63D739" authority="Kovařík & Šťáhlavský, 2019" authorityName="Kovařík & Šťáhlavský" authorityYear="2019" box="[594,707,779,803]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="hofereki" status="sp. nov.">
|
||
<emphasis id="66DC2D62B8466D1CFDD4327FFD63D739" box="[594,707,779,803]" italics="true" pageId="15" pageNumber="14">R. hofereki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="7DEF9019B8466D1CFD4F327FFCA3D738" box="[713,771,779,803]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="66DC2D62B8466D1CFD4F327FFD42D739" bold="true" box="[713,738,779,803]" pageId="15" pageNumber="14">sp</emphasis>
|
||
.
|
||
<emphasis id="66DC2D62B8466D1CFD68327FFD5DD739" bold="true" box="[750,765,779,803]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from all other species of the genus. They are recounted in the key. Morphologically closest species is
|
||
<taxonomicName id="93A88AF3B8466D1CFD8E323FFD31D779" baseAuthorityName="KovarIk" baseAuthorityYear="1998" box="[520,657,843,867]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="krasenskyi">
|
||
<emphasis id="66DC2D62B8466D1CFD8E323FFD31D779" box="[520,657,843,867]" italics="true" pageId="15" pageNumber="14">R. krasenskyi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="966C3F92B8466D1CFD48323FFD5FD778" box="[718,767,843,866]" country="Indonesia" name="Jawa Timur" pageId="15" pageNumber="14">Java</collectingRegion>
|
||
, which differs mainly by the darker pedipalp patella and more elongate metasomal segments in male (
|
||
<figureCitation id="CC93EDF5B8466D1CFD8F32FFFDCAD7B9" box="[521,618,907,931]" captionStart="Figures 192–202" captionStartId="40.[113,190,1922,1944]" captionTargetBox="[125,1468,264,1882]" captionTargetId="figure-11@40.[113,1474,255,1897]" captionTargetPageId="40" captionText="Figures 192–202: Reddyanus, males, pedipalp trochanter and femur, patella, and chela dorsal, metasoma II–III in lateral views. Figure 192. R. feti, holotype. Figure 193. R. furai sp. n., holotype. Figure 194. R. hofereki sp. n., holotype. Figure 195. R. jendeki, holotype. Figure 196. R. krasenskyi, Java, Ujung Kulon. Figure 197. R. majkusi sp. n., holotype. Figure 198. R. neradi, holotype. Figure 199. R. petrzelkai, holotype. Figure 200. R. rolciki sp. n., holotype. Figure 201. R. schwotti sp. n., Thailand, Khao sok. Figure 202. R. zideki, paratype." figureDoi="http://doi.org/10.5281/zenodo.6540239" httpUri="https://zenodo.org/record/6540239/files/figure.png" pageId="15" pageNumber="14">Figs. 194</figureCitation>
|
||
versus 196).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1CB2A2FBB8466D1CFFF732BFFE4CD7F8" box="[113,492,970,995]" pageId="15" pageNumber="14" type="distribution">
|
||
<paragraph id="5417F170B8466D1CFFF732BFFE4CD7F8" blockId="15.[113,492,970,995]" box="[113,492,970,995]" pageId="15" pageNumber="14">
|
||
DISTRIBUTION.
|
||
<collectingCountry id="2CBFB1E0B8466D1CFE9432BFFED4D7F9" box="[274,372,971,995]" name="Malaysia" pageId="15" pageNumber="14">Malaysia</collectingCountry>
|
||
(
|
||
<figureCitation id="CC93EDF5B8466D1CFE0432BFFE40D7F8" box="[386,480,970,995]" captionStart="Figure 225" captionStartId="43.[116,184,2017,2039]" captionTargetBox="[111,1472,234,1993]" captionTargetId="figure-9@43.[111,1472,234,1993]" captionTargetPageId="43" captionText="Figure 225. Map showing confirmed distribution of Reddyanus spp. in Southeast Asia." figureDoi="http://doi.org/10.5281/zenodo.6540247" httpUri="https://zenodo.org/record/6540247/files/figure.png" pageId="15" pageNumber="14">Fig. 225</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |