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<mods:title id="A8994E481ECE949CABFCDC9E800DD9F8">Reassessment of the status of some European and Asian Melitaea taxa described as subspecies of Melitaea phoebe ([Denis &amp; Schiffermüller], 1775), with designations of lectotypes where appropriate (Lepidoptera: Nymphalidae)</mods:title>
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<mods:namePart id="776484F6BC5C2B9678AD6DFFF0C8E843">Russell, Peter J. C.</mods:namePart>
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<mods:namePart id="8957104EF28B74F94BADCB3F1B5A3E46">Lukhtanov, Vladimir A.</mods:namePart>
<mods:affiliation id="6703B1AA3B6FE50E43C7EE3B4CF49180">Department of Karyosystematics, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, 199034 Saint Petersburg, Russia.</mods:affiliation>
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<mods:namePart id="21E620732D62F7CB68255D7576D0F672">Tennent, W. John</mods:namePart>
<mods:affiliation id="B04C0FAF1D7EED5E667BD34BD8C383E2">Scientific Associate, Division of Insects, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K. &amp; Honorary Associate, Oxford University Museum of Natural History, Parks Road, Oxford OX 1 3 PW, U. K</mods:affiliation>
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<treatment id="DB6B87B0FFF0FFEEE1FD03D1FCA1F88F" ID-DOI="http://doi.org/10.5281/zenodo.6580855" ID-GBIF-Taxon="195626728" ID-Zenodo-Dep="6580855" LSID="urn:lsid:plazi:treatment:DB6B87B0FFF0FFEEE1FD03D1FCA1F88F" httpUri="http://treatment.plazi.org/id/DB6B87B0FFF0FFEEE1FD03D1FCA1F88F" lastPageNumber="33" pageId="8" pageNumber="33">
<subSubSection id="1BD8652DFFF0FFEEE1FD03D1FDD7FC66" box="[199,628,906,933]" pageId="8" pageNumber="33" type="nomenclature">
<paragraph id="537D36A6FFF0FFEEE1FD03D1FDD7FC66" blockId="8.[151,1437,150,2013]" box="[199,628,906,933]" pageId="8" pageNumber="33">
<taxonomicName id="94C24D25FFF0FFEEE1FD03D1FDD7FC66" authority="Gross &amp; Ebert, 1975" authorityName=", Gross &amp; Ebert" authorityYear="1975" box="[199,628,906,933]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="phoebe" subSpecies="abbas">
<emphasis id="61B6EAB4FFF0FFEEE1FD03D1FE2AFC66" bold="true" box="[199,393,906,933]" italics="true" pageId="8" pageNumber="33">M. phoebe abbas</emphasis>
<bibRefCitation id="37534B57FFF0FFEEE0B503D0FDD7FC66" author="Gross, F. J. &amp; Ebert, G." box="[399,628,906,933]" pageId="8" pageNumber="33" pagination="8 - 45" refId="ref9178" refString="Gross, F. J. &amp; Ebert, G. (1975) Beitrage zur Kenntis der Rhopaloceren Irans. 2. Beitrag. Neue Taxa der Pieridae, Satyridae und Nymphalidae. Journal of the Entomological Society of Iran Supplement, 1, 8 - 45." type="journal article" year="1975">Gross &amp; Ebert, 1975</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="1BD8652DFFF0FFEEE34003D1FCA1F88F" pageId="8" pageNumber="33" type="discussion">
<paragraph id="537D36A6FFF0FFEEE34003D1FDCDFBF7" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="33">
[TL:
<quantity id="943A9B43FFF0FFEEE38903D0FD5AFC67" box="[691,761,906,933]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="33" unit="km" value="50.0">50 km</quantity>
NW Ardekan, Tange-Surkh,
<collectingCountry id="2BD57636FFF0FFEEE50603D1FBCFFC66" box="[1084,1132,907,933]" name="Iran" pageId="8" pageNumber="33">Iran</collectingCountry>
,
<quantity id="943A9B43FFF0FFEEE54D03D0FB73FC66" box="[1143,1232,906,933]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="8" pageNumber="33" unit="m" value="2250.0">2250 m</quantity>
]: junior subjective synonym of
<taxonomicName id="94C24D25FFF0FFEEE01903F4FDD6FC0B" authority="Higgins, 1941" authorityName="Higgins" authorityYear="1941" box="[291,629,942,968]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="enoch">
<emphasis id="61B6EAB4FFF0FFEEE01903F4FE73FC0B" box="[291,464,942,968]" italics="true" pageId="8" pageNumber="33">Melitaea enoch</emphasis>
<bibRefCitation id="37534B57FFF0FFEEE0EC03F4FDD6FC0B" author="Higgins, L. G." box="[470,629,942,968]" pageId="8" pageNumber="33" pagination="175 - 365" refId="ref9372" refString="Higgins, L. G. (1941) An illustrated catalogue of the Palearctic Melitaea (Lep. Rhopalocera). Transactions of the Royal Entomological Society of London, 91, 175 - 365. https: // doi. org / 10.1111 / j. 1365 - 2311.1941. tb 01045. x" type="journal article" year="1941">Higgins, 1941</bibRefCitation>
</taxonomicName>
(
<emphasis id="61B6EAB4FFF0FFEEE3BF03F4FD4FFC0B" bold="true" box="[645,748,942,968]" pageId="8" pageNumber="33">syn. nov.</emphasis>
). The synonymy of
<taxonomicName id="94C24D25FFF0FFEEE2F403F4FBB2FC0B" authorityName="Gross &amp; Ebert" authorityYear="1975" box="[974,1041,942,968]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="abbas">
<emphasis id="61B6EAB4FFF0FFEEE2F403F4FBB2FC0B" box="[974,1041,942,968]" italics="true" pageId="8" pageNumber="33">abbas</emphasis>
</taxonomicName>
with
<taxonomicName id="94C24D25FFF0FFEEE56A03F5FB1BFC0B" authorityName="Higgins" authorityYear="1941" box="[1104,1208,942,968]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="enoch">
<emphasis id="61B6EAB4FFF0FFEEE56A03F5FB1BFC0B" box="[1104,1208,942,968]" italics="true" pageId="8" pageNumber="33">M. enoch</emphasis>
</taxonomicName>
is based on the very close similarity of the wing morphology of the
<specimenCount id="45C4FD2FFFF0FFEEE3990389FCE3FC2F" box="[675,832,978,1005]" pageId="8" pageNumber="33" type="generic" typeStatus="holotypes">two holotypes</specimenCount>
(see figs 17A &amp; B and 16A &amp; B, respectively) and the fact that
<bibRefCitation id="37534B57FFF0FFEEE1C303ACFE1CFBD2" author="Toth, J. P. &amp; Varga, Z. &amp; Bereczki, J." box="[249,447,1014,1041]" pageId="8" pageNumber="33" pagination="106 - 115" refId="ref11405" refString="Toth, J. P., Varga, Z. &amp; Bereczki, J. (2016) Long-term survival and diversification of an endemic Melitaea species in mountains of Iran and adjacent areas. Journal of Zoological Systematics and Evolutionary Research, 54 (2), 106 - 115. https: // doi. org / 10.1111 / jzs. 12122" type="journal article" year="2016">
Tóth
<emphasis id="61B6EAB4FFF0FFEEE00F03ADFEC4FBD3" box="[309,359,1014,1040]" italics="true" pageId="8" pageNumber="33">et al</emphasis>
. (2016)
</bibRefCitation>
could find no genetic differences between the taxon
<taxonomicName id="94C24D25FFF0FFEEE52803ACFBF6FBD3" authorityName="Gross &amp; Ebert" authorityYear="1975" box="[1042,1109,1014,1040]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="abbas">
<emphasis id="61B6EAB4FFF0FFEEE52803ACFBF6FBD3" box="[1042,1109,1014,1040]" italics="true" pageId="8" pageNumber="33">abbas</emphasis>
</taxonomicName>
from the northern (where
<taxonomicName id="94C24D25FFF0FFEEE44403ADFF79FBF7" authorityName="Higgins" authorityYear="1941" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="enoch">
<emphasis id="61B6EAB4FFF0FFEEE44403ADFF79FBF7" italics="true" pageId="8" pageNumber="33">M. enoch</emphasis>
</taxonomicName>
is found) and southern populations.
</paragraph>
<paragraph id="537D36A6FFF0FFEEE1FD0464FCA2FA5F" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="33">
The original description (
<bibRefCitation id="37534B57FFF0FFEEE0DA0464FD49FB9B" author="Gross, F. J. &amp; Ebert, G." box="[480,746,1086,1113]" pageId="8" pageNumber="33" pagination="8 - 45" refId="ref9178" refString="Gross, F. J. &amp; Ebert, G. (1975) Beitrage zur Kenntis der Rhopaloceren Irans. 2. Beitrag. Neue Taxa der Pieridae, Satyridae und Nymphalidae. Journal of the Entomological Society of Iran Supplement, 1, 8 - 45." type="journal article" year="1975">Gross &amp; Ebert 1975: 44</bibRefCitation>
) indicated the presence of a
<typeStatus id="8C798804FFF0FFEEE51E0464FB25FB9B" box="[1060,1158,1086,1112]" pageId="8" pageNumber="33" type="holotype">holotype</typeStatus>
male, an
<typeStatus id="8C798804FFF0FFEEE5CA0464FAE9FB9B" box="[1264,1354,1086,1112]" pageId="8" pageNumber="33" type="allotype">allotype</typeStatus>
female and
<specimenCount id="45C4FD2FFFF0FFEEE1FC0438FEB2FBBF" box="[198,273,1122,1148]" pageId="8" pageNumber="33" type="male">9 male</specimenCount>
<typeStatus id="8C798804FFF0FFEEE0220439FE24FBBE" box="[280,391,1123,1149]" pageId="8" pageNumber="33" type="paratype">paratypes</typeStatus>
. The
<typeStatus id="8C798804FFF0FFEEE0F90438FD86FBBF" box="[451,549,1122,1148]" pageId="8" pageNumber="33" type="holotype">holotype</typeStatus>
(
<figureCitation id="CBF92A23FFF0FFEEE30E0438FD7CFBBF" box="[564,735,1122,1149]" captionStart="FIGURES 1720" captionStartId="9.[151,264,1710,1734]" captionTargetBox="[151,1436,381,1687]" captionTargetId="figure-121@9.[151,1436,381,1687]" captionTargetPageId="9" captionText="FIGURES 1720. 17, Holotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © R. Trusch. 18, Allotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © Alipanah. 19, Holotype of Melitaea ornata adversaria Korb, Stradomsky &amp; Kuznetsov, 2015. 20, Paratype of Melitaea ornata reliquiae Korb, Stradomsky &amp; Kuznetsov, 2015. Figs 19, 20 © S.K. Korb, B.V. Stradomsky &amp; G.V. Kuznetsov.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577646" httpUri="https://zenodo.org/record/6577646/files/figure.png" pageId="8" pageNumber="33">Figs 17A, B, C</figureCitation>
) and
<specimenCount id="45C4FD2FFFF0FFEEE2270438FCD8FBBF" box="[797,891,1122,1148]" pageId="8" pageNumber="33" type="male">six male</specimenCount>
<typeStatus id="8C798804FFF0FFEEE2B80439FC4EFBBE" box="[898,1005,1123,1149]" pageId="8" pageNumber="33" type="paratype">paratypes</typeStatus>
are present in the Staatliches Museum für Naturkunde Karlsruhe (
<bibRefCitation id="37534B57FFF0FFEEE0F104DCFCB7FB62" author="Hesselbarth, G. &amp; van Oorschot, H. &amp; Wagener, S." box="[459,788,1158,1185]" pageId="8" pageNumber="33" pagination="1 - 847" refId="ref9287" refString="Hesselbarth, G., van Oorschot, H. &amp; Wagener, S. (1995) Die Tagfalter der Turkei unter Berucksichtigung der angrenzenden Lander, Sigbert Wagener, Bocholt, Germany, 1, 1 - 757, 2, 758 - 1354, 3, 1 - 847." type="journal article" year="1995">
Hesselbarth
<emphasis id="61B6EAB4FFF0FFEEE36304DDFD2FFB63" box="[601,652,1158,1184]" italics="true" pageId="8" pageNumber="33">et al</emphasis>
. 1995:1031
</bibRefCitation>
) and an
<typeStatus id="8C798804FFF0FFEEE24204DCFC71FB63" box="[888,978,1158,1184]" pageId="8" pageNumber="33" type="allotype">allotype</typeStatus>
female (
<figureCitation id="CBF92A23FFF0FFEEE50D04DCFB45FB63" box="[1079,1254,1158,1185]" captionStart="FIGURES 1720" captionStartId="9.[151,264,1710,1734]" captionTargetBox="[151,1436,381,1687]" captionTargetId="figure-121@9.[151,1436,381,1687]" captionTargetPageId="9" captionText="FIGURES 1720. 17, Holotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © R. Trusch. 18, Allotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © Alipanah. 19, Holotype of Melitaea ornata adversaria Korb, Stradomsky &amp; Kuznetsov, 2015. 20, Paratype of Melitaea ornata reliquiae Korb, Stradomsky &amp; Kuznetsov, 2015. Figs 19, 20 © S.K. Korb, B.V. Stradomsky &amp; G.V. Kuznetsov.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577646" httpUri="https://zenodo.org/record/6577646/files/figure.png" pageId="8" pageNumber="33">Figs 18A, B, C</figureCitation>
) and a
<typeStatus id="8C798804FFF0FFEEE40604DDFA3FFB62" box="[1340,1436,1159,1185]" pageId="8" pageNumber="33" type="paratype">paratype</typeStatus>
male are present in the Hayk Mirzayans Insect Museum, Iranian Research Institute of Plant Protection,
<collectingRegion id="9106F844FFF0FFEEE40604F0FA3AFB07" box="[1340,1433,1194,1220]" country="Iran" name="Tehran" pageId="8" pageNumber="33">Teheran</collectingRegion>
,
<collectingCountry id="2BD57636FFF0FFEEE1AD0495FF6BFB2A" box="[151,200,1231,1257]" name="Iran" pageId="8" pageNumber="33">Iran</collectingCountry>
. This was confirmed by
<bibRefCitation id="37534B57FFF0FFEEE0E40494FD77FB2B" author="Alipanah, H." box="[478,724,1230,1256]" pageId="8" pageNumber="33" pagination="191 - 210" refId="ref8644" refString="Alipanah, H. (2016) List of the Lepidoptera type material deposited in the Hayk Mirzayans Insect Museum (HMIM), Iranian Research Institute of Plant Protection (IRIPP) (Insecta: Lepidoptera). SHILAP Revista de Lepidopterologia, 44 (174), 191 - 210." type="journal article" year="2016">Alipanah (2016: 200)</bibRefCitation>
, who stated that Tange-Surkh is a misspelling of Tang-e Sorkh and Ardekan is currently known as Sepidân. Of the remaining
<specimenCount id="45C4FD2FFFF0FFEEE24A04A9FC4BFACF" box="[880,1000,1266,1293]" pageId="8" pageNumber="33" type="male">two males</specimenCount>
, one is deposited in the Zoologisches Forschungsmuseum Alexander Koenig in Bonn,
<collectingCountry id="2BD57636FFF0FFEEE3F3054CFC92FAF3" box="[713,817,1302,1328]" name="Germany" pageId="8" pageNumber="33">Germany</collectingCountry>
and the other in Staatliches Museum für Naturkunde Karlsruhe (SMNK),
<collectingCountry id="2BD57636FFF0FFEEE0460560FE45FA97" box="[380,486,1338,1364]" name="Germany" pageId="8" pageNumber="33">Germany</collectingCountry>
. Recent research by
<bibRefCitation id="37534B57FFF0FFEEE3F30560FC26FA97" author="Toth, J. P. &amp; Bereczki, J. &amp; Varga, Z. &amp; Rota, J. &amp; Sramko, G. &amp; Wahlberg, N." box="[713,901,1338,1365]" pageId="8" pageNumber="33" pagination="749 - 757" refId="ref11329" refString="Toth, J. P., Bereczki, J., Varga, Z., Rota, J., Sramko, G. &amp; Wahlberg, N. (2014) Relationships within the Melitaea phoebe species group (Lepidoptera: Nymphalidae): new insights from molecular and morphometric information. Systematic Entomology, 39, 749 - 757. https: // doi. org / 10.1111 / syen. 12083" type="journal article" year="2014">
Tóth
<emphasis id="61B6EAB4FFF0FFEEE23E0561FC97FA97" box="[772,820,1338,1364]" italics="true" pageId="8" pageNumber="33">et al</emphasis>
. (2014
</bibRefCitation>
,
<bibRefCitation id="37534B57FFF0FFEEE2AA0560FC6FFA96" author="Toth, J. P. &amp; Varga, Z. &amp; Bereczki, J." box="[912,972,1338,1365]" pageId="8" pageNumber="33" pagination="106 - 115" refId="ref11405" refString="Toth, J. P., Varga, Z. &amp; Bereczki, J. (2016) Long-term survival and diversification of an endemic Melitaea species in mountains of Iran and adjacent areas. Journal of Zoological Systematics and Evolutionary Research, 54 (2), 106 - 115. https: // doi. org / 10.1111 / jzs. 12122" type="journal article" year="2016">2016</bibRefCitation>
), indicated that the taxon
<taxonomicName id="94C24D25FFF0FFEEE5D20560FA88FA97" authorityName="Gross &amp; Ebert" authorityYear="1975" box="[1256,1323,1338,1364]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="species" species="abbas">
<emphasis id="61B6EAB4FFF0FFEEE5D20560FA88FA97" box="[1256,1323,1338,1364]" italics="true" pageId="8" pageNumber="33">abbas</emphasis>
</taxonomicName>
should be elevated to species level. There was no label on the pin of the
<typeStatus id="8C798804FFF0FFEEE26C0504FC13FABB" box="[854,944,1374,1400]" pageId="8" pageNumber="33" type="allotype">allotype</typeStatus>
to indicate its status, this has been rectified and its current specific identification noted (
<figureCitation id="CBF92A23FFF0FFEEE3BC05D8FD57FA5F" box="[646,756,1410,1436]" captionStart="FIGURES 1720" captionStartId="9.[151,264,1710,1734]" captionTargetBox="[151,1436,381,1687]" captionTargetId="figure-121@9.[151,1436,381,1687]" captionTargetPageId="9" captionText="FIGURES 1720. 17, Holotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © R. Trusch. 18, Allotype of Melitaea phoebe abbas, Gross &amp; Ebert, 1975. © Alipanah. 19, Holotype of Melitaea ornata adversaria Korb, Stradomsky &amp; Kuznetsov, 2015. 20, Paratype of Melitaea ornata reliquiae Korb, Stradomsky &amp; Kuznetsov, 2015. Figs 19, 20 © S.K. Korb, B.V. Stradomsky &amp; G.V. Kuznetsov.A = upperside, B = underside, C = labels on specimen pin." figureDoi="http://doi.org/10.5281/zenodo.6577646" httpUri="https://zenodo.org/record/6577646/files/figure.png" pageId="8" pageNumber="33">Fig. 18 C</figureCitation>
).
</paragraph>
<paragraph id="537D36A6FFF0FFEEE1FD05FCFCA1F88F" blockId="8.[151,1437,150,2013]" pageId="8" pageNumber="33">
There is some muddle in the figures of this species under the name
<taxonomicName id="94C24D25FFF0FFEEE28C05FDFBC8FA03" box="[950,1131,1446,1472]" class="Insecta" family="Nymphalidae" genus="Melitaea" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="33" phylum="Arthropoda" rank="subSpecies" species="telona" subSpecies="abbas">
<emphasis id="61B6EAB4FFF0FFEEE28C05FDFBC8FA03" box="[950,1131,1446,1472]" italics="true" pageId="8" pageNumber="33">M. telona abbas</emphasis>
</taxonomicName>
presented by
<bibRefCitation id="37534B57FFF0FFEEE43C05FDFEC4FA27" author="Van Oorschot, H. &amp; Coutsis, J. G." pageId="8" pageNumber="33" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot and Coutsis (2014</bibRefCitation>
: plate 13 figs 11, 12, 13 &amp; 16); their figure 16 correctly shows the
<typeStatus id="8C798804FFF0FFEEE56C0590FB1BFA27" box="[1110,1208,1482,1508]" pageId="8" pageNumber="33" type="holotype">holotype</typeStatus>
in SMNK; however the captions for the specimens (numbers 14631465) in figures 1113 indicate firstly that they are all female and secondly that they are housed in the SMNK (
<bibRefCitation id="37534B57FFF0FFEEE3A60648FBE1F9EE" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[668,1090,1554,1581]" pageId="8" pageNumber="33" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 167)</bibRefCitation>
; figure 13 does appear to be a female but figures 11 and 12 appear to be males. There are no female
<typeStatus id="8C798804FFF0FFEEE2A3066DFBA7F992" box="[921,1028,1591,1617]" pageId="8" pageNumber="33" type="paratype">paratypes</typeStatus>
among the specimens in SMNK, the
<typeStatus id="8C798804FFF0FFEEE1AD0600FF52F9B7" box="[151,241,1626,1652]" pageId="8" pageNumber="33" type="allotype">allotype</typeStatus>
and only female is in the
<collectingRegion id="9106F844FFF0FFEEE32A0600FDC9F9B7" box="[528,618,1626,1652]" country="Iran" name="Tehran" pageId="8" pageNumber="33">Teheran</collectingRegion>
museum (see above) and was taken at the same locality but on
<date id="277C1066FFF0FFEEE4160600FA3FF9B7" box="[1324,1436,1626,1652]" pageId="8" pageNumber="33" value="1972-06-16">16.6.1972</date>
(see our fig. 19C). The
<specimenCount id="45C4FD2FFFF0FFEEE0A30625FE5CF95B" box="[409,511,1662,1689]" pageId="8" pageNumber="33" type="male">two male</specimenCount>
specimens illustrated were taken apparently on
<date id="277C1066FFF0FFEEE52C0624FB0AF95B" box="[1046,1193,1662,1689]" pageId="8" pageNumber="33" value="1972-06-16">16 June 1972</date>
and
<date id="277C1066FFF0FFEEE5E40624FA3FF95A" box="[1246,1436,1662,1689]" pageId="8" pageNumber="33" value="1975-06-12" valueMax="1975-06-15" valueMin="1975-06-12">1215 June 1975</date>
(
<bibRefCitation id="37534B57FFF0FFEEE1A506F8FD9EF97E" author="Van Oorschot, H. &amp; Coutsis, J. G." box="[159,573,1698,1725]" pageId="8" pageNumber="33" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014: 167)</bibRefCitation>
. These dates exactly match those on two of the labels on the pins of
<typeStatus id="8C798804FFF0FFEEE40606F9FA3FF97E" box="[1340,1436,1699,1725]" pageId="8" pageNumber="33" type="paratype">paratype</typeStatus>
specimens in SMNK, but those specimens do not appear to match the specimens figured by
<bibRefCitation id="37534B57FFF0FFEEE5AB069DFF7FF8C7" author="Van Oorschot, H. &amp; Coutsis, J. G." pageId="8" pageNumber="33" refId="ref11693" refString="Van Oorschot, H. &amp; Coutsis, J. G. (2014) The genus Melitaea Fabricius, 1807 (Lepidoptera, Nymphalidae, Nymphalinae) Taxonomy and systematics with special reference to the male genitalia. Tshikolovets publications, Pardubice, Czech Republic, 360 pp." type="book" year="2014">van Oorschot &amp; Coutsis (2014</bibRefCitation>
: plate 13 figs 11 &amp; 12), the
<specimenCount id="45C4FD2FFFF0FFEEE31706B1FD01F8C7" box="[557,674,1770,1797]" pageId="8" pageNumber="33" type="male">two males</specimenCount>
illustrated have differently orientated antennae and wing sections missing which do not match any specimens in the collection of
<typeStatus id="8C798804FFF0FFEEE2520755FC70F8EA" box="[872,979,1807,1833]" pageId="8" pageNumber="33" type="paratype">paratypes</typeStatus>
in the SMNK. Neither John Coutsis nor the present authors are able to explain this discrepancy.
</paragraph>
</subSubSection>
</treatment>
</document>