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<document id="6C6B4A1FA53F4C65FAA2BF2AF3AEBE53" ID-ISSN="1175­5334" approvalRequired="298" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="51" approvalRequired_for_textStreams="166" approvalRequired_for_treatments="80" checkinTime="1627004795516" checkinUser="felipe" docAuthor="Oláh, J. &amp; Johanson, K. A." docDate="2008" docId="BC22C32217E9A9C5989DF8967C70FB8E" docLanguage="en" docName="zt01802p248.pdf" docOrigin="Zootaxa 1802" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.2:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="2" docTitle="Cheumatopsyche fianara Olah &amp; Johanson 2008, new species" docType="treatment" docVersion="2" lastPageNumber="220" masterDocId="401BBB5A1731A91E980AFFCB7843FFFC" masterDocTitle="Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)" masterLastPageNumber="248" masterPageNumber="1" pageNumber="217" updateTime="1651296210969" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="D8F3769BE0E499B4A25177F4F1D9631D">Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)</mods:title>
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<mods:namePart id="708EAE54DFFA2615CFDC3FBE9FF31B46">Oláh, J.</mods:namePart>
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<mods:namePart id="CD78A08B0815ACFB9C3A366CDD83E3E5">Johanson, K. A.</mods:namePart>
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<mods:date id="BD4B346756271FB07286738FAC3CE1AF">2008</mods:date>
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<subSubSection id="7C9121BF17E9A9C6989DF8967B0BF88B" box="[151,840,1885,1911]" pageId="216" pageNumber="217" type="nomenclature">
<paragraph id="3434723417E9A9C6989DF8967B0BF88B" blockId="216.[151,840,1885,1951]" box="[151,840,1885,1911]" pageId="216" pageNumber="217">
<heading id="6F7CC55817E9A9C6989DF8967B0BF88B" bold="true" box="[151,840,1885,1911]" fontSize="11" level="1" pageId="216" pageNumber="217" reason="1">
<emphasis id="06FFAE2617E9A9C6989DF8967B0BF88B" bold="true" box="[151,840,1885,1911]" pageId="216" pageNumber="217">
<taxonomicName id="F38B09B717E9A9C6989DF8967AEFF88B" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[151,684,1885,1911]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="fianara" status="new species">
<emphasis id="06FFAE2617E9A9C6989DF8967985F88B" bold="true" box="[151,454,1885,1911]" italics="true" pageId="216" pageNumber="217">Cheumatopsyche fianara</emphasis>
Oláh &amp; Johanson
</taxonomicName>
,
<taxonomicNameLabel id="1DCC135D17E9A9C69AB2F8967B0BF88B" box="[696,840,1885,1911]" pageId="216" pageNumber="217" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="7C9121BF17E9A9C5989DF84E79FDFCCE" lastPageId="219" lastPageNumber="220" pageId="216" pageNumber="217" type="description">
<paragraph id="3434723417E9A9C6989DF84E7970F863" blockId="216.[151,840,1885,1951]" box="[151,307,1925,1951]" pageId="216" pageNumber="217">
<figureCitation id="ACB06EB117E9A9C6989DF84E7970F863" box="[151,307,1925,1951]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="216" pageNumber="217">Fig. 363366</figureCitation>
</paragraph>
<paragraph id="3434723417E9A9C6989DF81E7DD7F813" blockId="216.[151,1428,2005,2031]" box="[151,1428,2005,2031]" pageId="216" pageNumber="217">
This species is similar to
<taxonomicName id="F38B09B717E9A9C699B4F81E7B44F813" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" box="[446,775,2005,2031]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="216" pageNumber="217" phylum="Arthropoda" rank="species" species="jaoriva">
<emphasis id="06FFAE2617E9A9C699B4F81E7A76F812" box="[446,565,2005,2030]" italics="true" pageId="216" pageNumber="217">C. jaoriva</emphasis>
Oláh &amp; Johanson
</taxonomicName>
and is distinguished by the anterior margins of segment
</paragraph>
<caption id="60F422BC17E8A9C7989DFF567CB7FF48" box="[151,1268,157,181]" pageId="217" pageNumber="218" startId="217.[151,241,157,181]">
<paragraph id="3434723417E8A9C7989DFF567CB7FF48" blockId="217.[151,1426,157,1978]" box="[151,1268,157,181]" pageId="217" pageNumber="218">
<emphasis id="06FFAE2617E8A9C7989DFF567940FF49" bold="true" box="[151,259,157,181]" pageId="217" pageNumber="218">TABLE 4</emphasis>
. Combination of wing characters for species in the
<taxonomicName id="F38B09B717E8A9C79B20FF557C17FF49" authorityName="J Olah &amp; KA Johanson" authorityYear="2008" box="[810,1108,158,181]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="mahakaya">
<emphasis id="06FFAE2617E8A9C79B20FF557C17FF49" box="[810,1108,158,181]" italics="true" pageId="217" pageNumber="218">Cheumatopsyche mahakaya</emphasis>
</taxonomicName>
species group.
</paragraph>
</caption>
<paragraph id="3434723417E8A9C79895FF197C68FE88" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">Species Forewing Forew- Distance Distance Distance Distance between hind membrane ing between between between hind wing crossveins r and s in colour length forewing forewing Cu2 wing Sc and R relation to length of r (mm) crossveins m- and A1 at in relation to cu and cu wing margin length of r</paragraph>
<paragraph id="3434723417E8A9C79895FE437C68FDFB" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">
<taxonomicName id="F38B09B717E8A9C79895FE43794DFE63" authorityName="J Olah &amp; KA Johanson" authorityYear="2008" box="[159,270,392,415]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="ambala">
<emphasis id="06FFAE2617E8A9C79895FE43794DFE63" box="[159,270,392,415]" italics="true" pageId="217" pageNumber="218">C. ambala</emphasis>
</taxonomicName>
uniformly 11.0 tangential far Sc apical to R r apical to s by a distance of coloured by a distance 2.0 times length of r brown of 1.0 times length of r
</paragraph>
<paragraph id="3434723417E8A9C79895FDD17DD1FDCD" blockId="217.[151,1426,157,1978]" box="[159,1426,538,561]" pageId="217" pageNumber="218">
<emphasis id="06FFAE2617E8A9C79895FDD17970FDCD" box="[159,307,538,561]" italics="true" pageId="217" pageNumber="218">C. amdakara-</emphasis>
uniformly 6.0 m-cu apical to far tangential r apical to s by a distance of
</paragraph>
<paragraph id="3434723417E8A9C79895FDF67AD0FD40" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">
<emphasis id="06FFAE2617E8A9C79895FDF6789AFDA8" box="[159,217,573,596]" italics="true" pageId="217" pageNumber="218">maya</emphasis>
coloured cu by a dis- 1.0 times length of r medium brown tance of 1.6 times length of m-cu
</paragraph>
<paragraph id="3434723417E8A9C79895FD1B7C68FCB3" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">
<taxonomicName id="F38B09B717E8A9C79895FD1B796FFD1B" authorityName="J Olah &amp; KA Johanson" authorityYear="2008" box="[159,300,720,743]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="balaromta">
<emphasis id="06FFAE2617E8A9C79895FD1B796FFD1B" box="[159,300,720,743]" italics="true" pageId="217" pageNumber="218">C. balaromta</emphasis>
</taxonomicName>
uniformly 7.5 tangential far Sc apical to R r apical to s by a distance of coloured by a distance 1.0 times length of r medium brown of 2.0 times length of r
</paragraph>
<paragraph id="3434723417E8A9C79895FCA97C68FC1D" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">
<taxonomicName id="F38B09B717E8A9C79895FCA97955FC85" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[159,278,866,889]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="bidikala">
<emphasis id="06FFAE2617E8A9C79895FCA97955FC85" box="[159,278,866,889]" italics="true" pageId="217" pageNumber="218">C. bidikala</emphasis>
</taxonomicName>
uniformly 6.5 tangential close Sc apical to R r apical to s by a distance of coloured pale by a distance 2.0 times length of r brown, thick of 1.0 times pterostigma length of r
</paragraph>
<paragraph id="3434723417E8A9C79895FC3E7AD0FB6B" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">
<taxonomicName id="F38B09B717E8A9C79895FC3E7946FBF0" authorityName="J Olah &amp; KA Johanson" authorityYear="2008" box="[159,261,1013,1036]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="bolosa">
<emphasis id="06FFAE2617E8A9C79895FC3E7946FBF0" box="[159,261,1013,1036]" italics="true" pageId="217" pageNumber="218">C. bolosa</emphasis>
</taxonomicName>
uniformly 11.0 m-cu apical to close Sc apical to R r apical to s by a distance of coloured dark cu by a dis- by a distance 2.0 times length of r brown, thick tance of 1.0 of 1.0 times pterostigma times length of length of r m-cu
</paragraph>
<paragraph id="3434723417E8A9C79895FB617DD1FB3D" blockId="217.[151,1426,157,1978]" box="[159,1426,1194,1217]" pageId="217" pageNumber="218">
<emphasis id="06FFAE2617E8A9C79895FB617950FB3D" box="[159,275,1194,1217]" italics="true" pageId="217" pageNumber="218">
<taxonomicName id="F38B09B717E8A9C79895FB617953FB3D" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[159,272,1194,1217]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="fianara">C. fianara</taxonomicName>
,
</emphasis>
pale brown, 11.0 tangential close Sc apical to R r apical to s by a distance of
</paragraph>
<paragraph id="3434723417E8A9C79895FB067C68FAD6" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">new species darker cloud by a distance 1.0 times length of r apicad of 1.0 times length of r</paragraph>
<paragraph id="3434723417E8A9C79895FAF67AD0FA23" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">
<taxonomicName id="F38B09B717E8A9C79895FAF67948FAA8" box="[159,267,1341,1364]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="jaoriva">
<emphasis id="06FFAE2617E8A9C79895FAF67948FAA8" box="[159,267,1341,1364]" italics="true" pageId="217" pageNumber="218">C. jaoriva</emphasis>
</taxonomicName>
uniformly 12.0 m-cu basal to far Sc apical to R r apical to s by a distance of coloured cu by a dis- by a distance 2.0 times length of r medium tance of 0.6 of 0.5 times brown, thick times length of length of r pterostigma m-cu
</paragraph>
<paragraph id="3434723417E8A9C79895FA387981F968" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">
<taxonomicName id="F38B09B717E8A9C79895FA38796FF9F6" authorityName="J Olah &amp; KA Johanson" authorityYear="2008" box="[159,300,1523,1546]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="mahakaya">
<emphasis id="06FFAE2617E8A9C79895FA38796FF9F6" box="[159,300,1523,1546]" italics="true" pageId="217" pageNumber="218">C. mahakaya</emphasis>
</taxonomicName>
uniformly 10.0 tangential far Sc apical to R r apical to s by a distance of coloured dark by a distance 1.0 times length of r brown, cor- of 1.0 times neous length of r pterostigma
</paragraph>
<paragraph id="3434723417E8A9C79895F9637C68F8DB" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">
<taxonomicName id="F38B09B717E8A9C79895F963796AF943" authorityName="J Olah &amp; KA Johanson" authorityYear="2008" box="[159,297,1704,1727]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="mandrara">
<emphasis id="06FFAE2617E8A9C79895F963796AF943" box="[159,297,1704,1727]" italics="true" pageId="217" pageNumber="218">C. mandrara</emphasis>
</taxonomicName>
brown, darker 9.5 tangential far Sc apical to R r apical to s by a distance of patches on by a distance 1.5 times length of r crossveins of 1.0 times length of r
</paragraph>
<paragraph id="3434723417E8A9C79895F8F07DD1F8AE" blockId="217.[151,1426,157,1978]" box="[159,1426,1851,1874]" pageId="217" pageNumber="218">
<emphasis id="06FFAE2617E8A9C79895F8F0796AF8AE" box="[159,297,1851,1874]" italics="true" pageId="217" pageNumber="218">
<taxonomicName id="F38B09B717E8A9C79895F8F07966F8AE" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[159,293,1851,1874]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="217" pageNumber="218" phylum="Arthropoda" rank="species" species="masiposa">C. masiposa</taxonomicName>
,
</emphasis>
brown, clouds 9.0 tangential far Sc apical to R r apical to s by a distance of
</paragraph>
<paragraph id="3434723417E8A9C79895F8967C68F846" blockId="217.[151,1426,157,1978]" pageId="217" pageNumber="218">new species apicad, thick by a distance 1.0 times length of r pterostigma of 1.5 times length of r</paragraph>
<paragraph id="3434723417EBA9C49895FF6B7C68FEE3" blockId="218.[159,1426,160,468]" pageId="218" pageNumber="219">
<emphasis id="06FFAE2617EBA9C49895FF6B7955FF4B" box="[159,278,160,183]" italics="true" pageId="218" pageNumber="219">
<taxonomicName id="F38B09B717EBA9C49895FF6B7951FF4B" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[159,274,160,183]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="rantsoa">C. rantsoa</taxonomicName>
,
</emphasis>
brown pattern 9.0 tangential met Sc apical to R r apical to s by a distance of new species apicad and on by a distance 2.0 times length of r crossveins of 0.3 times length of r
</paragraph>
<paragraph id="3434723417EBA9C49895FEF97AD0FE28" blockId="218.[159,1426,160,468]" pageId="218" pageNumber="219">
<taxonomicName id="F38B09B717EBA9C49895FEF9794BFEB5" authorityName="J Olah &amp; KA Johanson" authorityYear="2008" box="[159,264,306,329]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="tanana">
<emphasis id="06FFAE2617EBA9C49895FEF9794BFEB5" box="[159,264,306,329]" italics="true" pageId="218" pageNumber="219">C. tanana</emphasis>
</taxonomicName>
uniformly 8.0 m-cu apical to far Sc apical to R r apical to s by a distance of coloured pale cu by a dis- by a distance 1.0 times length of r brown tance of 1.0 of 1.0 times times length of length of r m-cu
</paragraph>
<paragraph id="3434723417EBA9C4989DFDD67B7BFD7B" blockId="218.[151,1436,541,807]" pageId="218" pageNumber="219">IX being more regularly bow-shaped, not shifted posterad and not so deeply incised anterodorsad in lateral view; the tip of segment X is perfectly rounded; the basal half of the coxopodites are slender; and the harpagones lack basal a tooth and is straight, not curving.</paragraph>
<paragraph id="3434723417EBA9C498CFFD5E7B53FCDB" blockId="218.[151,1436,541,807]" pageId="218" pageNumber="219">Male. Body and wings brown, with brown pubescence. Maxilary palp formula I-(III, IV)-II-V, segment V as long as sum of segments IIV. Head dorsum brown with brown setal warts, indiscernible on the pinned animal. Swollen setal wart absent on proepisternum. Pretarsal claws asymmetrical, laterally flanked by setal bundle. Protibia with 2 spurs, posteroapical spur smallest.</paragraph>
<caption id="60F422BC17EBA9C4989DF9D57BFCF9A4" pageId="218" pageNumber="219" startId="218.[151,269,1566,1590]" targetBox="[171,1380,857,1514]" targetPageId="218">
<paragraph id="3434723417EBA9C4989DF9D57BFCF9A4" blockId="218.[151,1436,1566,1624]" pageId="218" pageNumber="219">
<emphasis id="06FFAE2617EBA9C4989DF9D5793AF9CA" bold="true" box="[151,377,1566,1590]" pageId="218" pageNumber="219">FIGURES 363366.</emphasis>
<taxonomicName id="F38B09B717EBA9C4998AF9D47B13F9C9" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[384,848,1566,1590]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="218" pageNumber="219" phylum="Arthropoda" rank="species" species="fianara" status="new species">
<emphasis id="06FFAE2617EBA9C4998AF9D47AC8F9CA" box="[384,651,1567,1590]" italics="true" pageId="218" pageNumber="219">Cheumatopsyche fianara</emphasis>
Oláh &amp; Johanson
</taxonomicName>
,
<taxonomicNameLabel id="1DCC135D17EBA9C49B51F9D57B9CF9C9" box="[859,991,1566,1589]" pageId="218" pageNumber="219" rank="species">new species</taxonomicNameLabel>
, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral.
</paragraph>
</caption>
<paragraph id="3434723417EBA9C498CFF9477B40F8E2" blockId="218.[151,1437,1676,2022]" pageId="218" pageNumber="219">Wings. Forewing crossveins m-cu and cu tangential. Cu2 and A1 running very closely. Wing membranes uniformly brown, without pronounced pattern. Hind wing Sc and R confluent apically of crossvein r at distance equal to length of crossvein. Crossvein r located apically of crossvein s by distance equal to length of crossvein. Fork 1 present. Forewing length 11.0 mm.</paragraph>
<paragraph id="3434723417EBA9C598CFF8E779FDFCCE" blockId="218.[151,1437,1676,2022]" lastBlockId="219.[151,1437,152,1138]" lastPageId="219" lastPageNumber="220" pageId="218" pageNumber="219">
Male genitalia. Abdominal segment IX fused annularly, tergum shorter than sternum (
<figureCitation id="ACB06EB117EBA9C49CC8F8E77D69F8BA" box="[1218,1322,1836,1862]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="218" pageNumber="219">Fig. 363</figureCitation>
); anterior margins of segment IX almost regular bow-shaped; central section undulating; dorsal section slightly more drawn in lateral aspect (
<figureCitation id="ACB06EB117EBA9C499A4F8B77A57F86A" box="[430,532,1916,1942]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="218" pageNumber="219">Fig. 363</figureCitation>
). Apical lobe on posterolateral margins nearly triangular, located above articulation cavity of gonocoxites (
<figureCitation id="ACB06EB117EBA9C49A08F86F7A29F842" box="[514,618,1956,1982]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="218" pageNumber="219">Fig. 363</figureCitation>
). Spine row on posterior margins of segment IX complete, not intermittent; spines heterogenous, irregularly bending on laterocaudal lobes of dorsum; spines short, stiff, straight on apical lobes. Intersegmental step between segments IX and X filled, occupied by pair of setose protuberances, appearing as setose laterocaudal lobes. Body of segment X, long, nearly triangular, with dorsad curving, blunt apex (
<figureCitation id="ACB06EB117EAA9C5995FFF2379F8FEFE" box="[341,443,232,258]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 363</figureCitation>
); simple, lobe-shaped in dorsal aspect (
<figureCitation id="ACB06EB117EAA9C59B8CFF237BAEFEFE" box="[902,1005,232,258]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 364</figureCitation>
); ventrum slightly bellied (
<figureCitation id="ACB06EB117EAA9C59D22FF237DCCFEFE" box="[1320,1423,232,258]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 363</figureCitation>
). Apicoventral lobes fusing to segment X along mesal margin, dominating distal part of segment X (
<figureCitation id="ACB06EB117EAA9C59D2DFEDB7DCDFED6" box="[1319,1422,272,298]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 363</figureCitation>
); delineated by setae; rounded; dorsum flat in dorsal view (
<figureCitation id="ACB06EB117EAA9C59B3EFEF37BD8FEAE" box="[820,923,312,338]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 364</figureCitation>
). Smooth, mesal lobe reduced. Dorsal interlobular gap narrow, triangular. Apicoventral lobe apices orienting dorsad (
<figureCitation id="ACB06EB117EAA9C59C2EFEAB7CD3FE86" box="[1060,1168,352,378]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 363</figureCitation>
); separated by dorsal, interlobular gap; shape highly dependent on observation angle. Unusual, setose structure present on basal, dorsolateral areas of segment X. Setose structures forming pair of well-developed protuberances, representing laterocaudal lobes. Transverse sutures invisible. Longitudinal sutures running along bottom of segment X as continuation of apicoventral setose lobes (
<figureCitation id="ACB06EB117EAA9C59A9BFDCB7AB9FDE6" box="[657,762,512,538]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 363</figureCitation>
). Setose areas (superior or preanal appendages) forming pair of elevating, circular, flap-like setal warts located apically from middle of segment X. Coxopodites reaching apex of segment X; broad at bases followed by long, slender section, abruptly broadening apicad, slightly bending posterad (
<figureCitation id="ACB06EB117EAA9C5997AFDB37994FD6E" box="[368,471,632,658]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 363</figureCitation>
); straight in ventral view (
<figureCitation id="ACB06EB117EAA9C59B01FDB37B32FD6E" box="[779,881,632,658]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 365</figureCitation>
). Harpagones with short, broad triangular basis in lateral view (
<figureCitation id="ACB06EB117EAA9C59958FD6B79F9FD46" box="[338,442,672,698]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 363</figureCitation>
); continuing into long, upwardly orienting, straight, thin digitate process (
<figureCitation id="ACB06EB117EAA9C59D2DFD6B7DCCFD46" box="[1319,1423,672,698]" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 363</figureCitation>
). Phallothecal phallobase enlarged, straight, horizontal shaft parallel-sided, slightly constricted mid-length (
<figureCitation id="ACB06EB117EAA9C59D67FD037884FCF6" captionStart="FIGURES 363366" captionStartId="218.[151,269,1566,1590]" captionTargetBox="[171,1380,857,1514]" captionTargetId="figure-369@218.[151,1436,828,1543]" captionTargetPageId="218" captionText="FIGURES 363366. Cheumatopsyche fianara Oláh &amp; Johanson, new species, holotype, male. 363 — genitalia, lateral; 364 — genitalia, dorsal; 365 — gonocoxite, ventral; 366 — phallus, lateral." pageId="219" pageNumber="220">Fig. 366</figureCitation>
); chitinesed endothecal process circular; about half as wide as shaft. Phallotremmal sclerites broad and rounded in lateral aspect.
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF17EAA9C598CFFCF4794FFBDE" pageId="219" pageNumber="220" type="materials_examined">
<paragraph id="3434723417EAA9C598CFFCF47C93FC7E" blockId="219.[151,1437,152,1138]" pageId="219" pageNumber="220">
<materialsCitation id="84E3786917EAA9C598CFFCF47C93FC7E" collectingDate="1990-03-20" collectingDateMax="1990-03-24" collectingDateMin="1990-03-20" collectionCode="NMNH" collectorName="W. E. Steiner" country="Madagascar" elevation="900" location="Ranomafana" pageId="219" pageNumber="220" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="06FFAE2617EAA9C598CFFCF47A37FCA5" bold="true" box="[197,628,831,857]" pageId="219" pageNumber="220">
<typeStatus id="EB30CC9617EAA9C598CFFCF47974FCA5" box="[197,311,831,857]" pageId="219" pageNumber="220">Holotype</typeStatus>
male:
<collectingCountry id="4C9C32A417EAA9C59999FCF47A2CFCA5" box="[403,623,831,857]" name="Madagascar" pageId="219" pageNumber="220">MADAGASCAR</collectingCountry>
:
</emphasis>
<collectingRegion id="F64FBCD617EAA9C59A75FC8B7BC8FCA6" box="[639,907,832,858]" country="Madagascar" name="Fianarantsoa" pageId="219" pageNumber="220">Fianarantsoa Province</collectingRegion>
,
<quantity id="F373DFD117EAA9C59B90FC8B7B99FCA6" box="[922,986,832,858]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="219" pageNumber="220" unit="km" value="7.0">7 km</quantity>
W
<location id="315424EF17EAA9C59C06FC8B7CE0FCA6" LSID="urn:lsid:plazi:treatment:BC22C32217E9A9C5989DF8967C70FB8E:315424EF17EAA9C59C06FC8B7CE0FCA6" box="[1036,1187,832,858]" country="Madagascar" name="Ranomafana" pageId="219" pageNumber="220" stateProvince="Fianarantsoa">Ranomafana</location>
,
<quantity id="F373DFD117EAA9C59CB9FC8B7D40FCA6" box="[1203,1283,832,858]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="219" pageNumber="220" unit="m" value="900.0">
<elevation id="BFA6950717EAA9C59CB9FC8B7D40FCA6" box="[1203,1283,832,858]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="219" pageNumber="220" unit="m" value="900.0">900 m</elevation>
</quantity>
, black light, mountain rainforest near river and stream,
<date id="403554F417EAA9C59A81FCA37B76FC7E" box="[651,821,872,898]" pageId="219" pageNumber="220" value="1990-03-20" valueMax="1990-03-24" valueMin="1990-03-20">
<collectingDate id="5071AD1C17EAA9C59A81FCA37B76FC7E" box="[651,821,872,898]" pageId="219" pageNumber="220" value="1990-03-20" valueMax="1990-03-24" valueMin="1990-03-20">2024.iii.1990</collectingDate>
</date>
[
<collectorName id="997E17E217EAA9C59B4FFCA37B9EFC7E" box="[837,989,872,898]" pageId="219" pageNumber="220">W.E. Steiner</collectorName>
] - (
<collectionCode id="529AEAF117EAA9C59C0FFCA37C20FC7E" box="[1029,1123,872,898]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="219" pageNumber="220" type="Museum">NMNH</collectionCode>
, pinned).
</materialsCitation>
</paragraph>
<paragraph id="3434723417EAA9C598CFFC5B794FFBDE" blockId="219.[151,1437,152,1138]" pageId="219" pageNumber="220">
<materialsCitation id="84E3786917EAA9C598CFFC5B7B66FC56" box="[197,805,912,938]" collectingDate="1990-03-20" collectingDateMax="1990-03-24" collectingDateMin="1990-03-20" collectionCode="NMNH" collectorName="W. E. Steiner" country="Madagascar" elevation="900" location="Ranomafana" pageId="219" pageNumber="220" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<typeStatus id="EB30CC9617EAA9C598CFFC5B7978FC56" box="[197,315,912,938]" pageId="219" pageNumber="220">Paratypes</typeStatus>
: same data as holotype -
<specimenCount id="228DB9BD17EAA9C59A55FC5B7AEDFC56" box="[607,686,912,938]" pageId="219" pageNumber="220" type="male">1 male</specimenCount>
(
<collectionCode id="529AEAF117EAA9C59AB7FC5B7B5EFC56" box="[701,797,912,938]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="219" pageNumber="220" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E3786917EAA9C59B3BFC5B7D01FC56" box="[817,1346,912,938]" collectingDate="1990-01-20" collectingDateMax="1990-01-31" collectingDateMin="1990-01-20" collectionCode="NMNH" collectorName="W. E. Steiner" country="Madagascar" elevation="900" location="Ranomafana" pageId="219" pageNumber="220" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
ditto, except
<date id="403554F417EAA9C59BC3FC5B7C20FC56" box="[969,1123,912,938]" pageId="219" pageNumber="220" value="1990-01-20" valueMax="1990-01-31" valueMin="1990-01-20">
<collectingDate id="5071AD1C17EAA9C59BC3FC5B7C20FC56" box="[969,1123,912,938]" pageId="219" pageNumber="220" value="1990-01-20" valueMax="1990-01-31" valueMin="1990-01-20">2031.i.1990</collectingDate>
</date>
-
<specimenCount id="228DB9BD17EAA9C59C71FC5B7C89FC56" box="[1147,1226,912,938]" pageId="219" pageNumber="220" type="male">1 male</specimenCount>
(
<collectionCode id="529AEAF117EAA9C59CD3FC5B7D7AFC56" box="[1241,1337,912,938]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="219" pageNumber="220" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E3786917EAA9C59D44FC5B78A5FC2E" collectingDate="1990-01-20" collectingDateMax="1990-01-31" collectingDateMin="1990-01-20" collectionCode="OPC" collectorName="W. E. Steiner" country="Madagascar" elevation="900" isTailEnumeration="true" location="Ranomafana" pageId="219" pageNumber="220" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
<specimenCount id="228DB9BD17EAA9C59D44FC5B7DDFFC56" box="[1358,1436,912,938]" pageId="219" pageNumber="220" type="male">1 male</specimenCount>
(
<collectionCode id="529AEAF117EAA9C59895FC73789DFC2E" box="[159,222,952,978]" pageId="219" pageNumber="220">OPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E3786917EAA9C598FEFC737AB4FC2E" box="[244,759,952,978]" collectingDate="1990-02-01" collectingDateMax="1990-02-09" collectingDateMin="1990-02-01" collectionCode="NMNH" collectorName="W. E. Steiner" country="Madagascar" elevation="900" isTailEnumeration="true" location="Ranomafana" pageId="219" pageNumber="220" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
ditto, except
<date id="403554F417EAA9C59984FC737A50FC2E" box="[398,531,952,978]" pageId="219" pageNumber="220" value="1990-02-01" valueMax="1990-02-09" valueMin="1990-02-01">
<collectingDate id="5071AD1C17EAA9C59984FC737A50FC2E" box="[398,531,952,978]" pageId="219" pageNumber="220" value="1990-02-01" valueMax="1990-02-09" valueMin="1990-02-01">19.ii.1990</collectingDate>
</date>
-
<specimenCount id="228DB9BD17EAA9C59A24FC737A3DFC2E" box="[558,638,952,978]" pageId="219" pageNumber="220" type="male">1 male</specimenCount>
(
<collectionCode id="529AEAF117EAA9C59A85FC737AACFC2E" box="[655,751,952,978]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="219" pageNumber="220" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E3786917EAA9C59B0EFC737CBEFC2E" box="[772,1277,952,978]" collectingDate="1990-03-08" collectingDateMax="1990-03-13" collectingDateMin="1990-03-08" collectionCode="OPC" collectorName="W. E. Steiner" country="Madagascar" elevation="900" isTailEnumeration="true" location="Ranomafana" pageId="219" pageNumber="220" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
ditto, except
<date id="403554F417EAA9C59B94FC737C79FC2E" box="[926,1082,952,978]" pageId="219" pageNumber="220" value="1990-03-08" valueMax="1990-03-13" valueMin="1990-03-08">
<collectingDate id="5071AD1C17EAA9C59B94FC737C79FC2E" box="[926,1082,952,978]" pageId="219" pageNumber="220" value="1990-03-08" valueMax="1990-03-13" valueMin="1990-03-08">813.iii.1990</collectingDate>
</date>
-
<specimenCount id="228DB9BD17EAA9C59C5FFC737CE6FC2E" box="[1109,1189,952,978]" pageId="219" pageNumber="220" type="male">1 male</specimenCount>
(
<collectionCode id="529AEAF117EAA9C59CBCFC737CB6FC2E" box="[1206,1269,952,978]" pageId="219" pageNumber="220">OPC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E3786917EAA9C59D00FC737A43FC06" collectingDate="1990-10-30" collectingDateMax="1990-03-13" collectingDateMin="1990-03-08" collectionCode="NMNH" collectorName="W. E. Steiner" country="Madagascar" elevation="900" isTailEnumeration="true" location="Ranomafana" pageId="219" pageNumber="220" specimenCount="1" specimenCount-male="1" stateProvince="Fianarantsoa" typeStatus="paratype">
ditto, except
<date id="403554F417EAA9C5989DFC2B794FFC06" box="[151,268,992,1018]" pageId="219" pageNumber="220" value="1990-10-30">
<collectingDate id="5071AD1C17EAA9C5989DFC2B794FFC06" box="[151,268,992,1018]" pageId="219" pageNumber="220" value="1990-10-30">30.x.1990</collectingDate>
</date>
-
<specimenCount id="228DB9BD17EAA9C59925FC2B79C0FC06" box="[303,387,992,1018]" pageId="219" pageNumber="220" type="male">1 male</specimenCount>
(
<collectionCode id="529AEAF117EAA9C59992FC2B79BBFC06" box="[408,504,992,1018]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="219" pageNumber="220" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="84E3786917EAA9C59A1BFC2B7944FBDE" collectingDate="1990-10-18" collectingDateMax="1990-03-13" collectingDateMin="1990-03-08" collectionCode="NMNH" collectorName="W. E. Steiner &amp; C. Kremen &amp; Daniels" country="Madagascar" elevation="900" isTailEnumeration="true" location="Ranomafana" pageId="219" pageNumber="220" specimenCount="2" specimenCount-male="2" stateProvince="Fianarantsoa" typeStatus="paratype">
ditto, except
<date id="403554F417EAA9C59ABEFC2B7B69FC06" box="[692,810,992,1018]" pageId="219" pageNumber="220" value="1990-10-18">
<collectingDate id="5071AD1C17EAA9C59ABEFC2B7B69FC06" box="[692,810,992,1018]" pageId="219" pageNumber="220" value="1990-10-18">18.x.1990</collectingDate>
</date>
[
<collectorName id="997E17E217EAA9C59B37FC2B7B99FC06" box="[829,986,992,1018]" pageId="219" pageNumber="220">W.E. Steiner</collectorName>
,
<collectorName id="997E17E217EAA9C59BE1FC2B7C33FC06" box="[1003,1136,992,1018]" pageId="219" pageNumber="220">C. Kremen</collectorName>
&amp; P.
<collectorName id="997E17E217EAA9C59CC8FC2B7D5DFC06" box="[1218,1310,992,1018]" pageId="219" pageNumber="220">Daniels</collectorName>
] -
<specimenCount id="228DB9BD17EAA9C59D42FC2B7DDFFC06" box="[1352,1436,992,1018]" pageId="219" pageNumber="220" type="male">2 male</specimenCount>
(
<collectionCode id="529AEAF117EAA9C59895FBC378BCFBDE" box="[159,255,1032,1058]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="219" pageNumber="220" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF17EAA9C598CFFBE47A43FBB6" box="[197,512,1071,1098]" pageId="219" pageNumber="220" type="distribution">
<paragraph id="3434723417EAA9C598CFFBE47A43FBB6" blockId="219.[151,1437,152,1138]" box="[197,512,1071,1098]" pageId="219" pageNumber="220">
<emphasis id="06FFAE2617EAA9C598CFFBE47926FBB5" bold="true" box="[197,357,1071,1097]" pageId="219" pageNumber="220">Distribution.</emphasis>
<collectingCountry id="4C9C32A417EAA9C59966FBFB79BFFBB6" box="[364,508,1072,1098]" name="Madagascar" pageId="219" pageNumber="220">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF17EAA9C598CFFB9C7C70FB8E" box="[197,1075,1111,1138]" pageId="219" pageNumber="220" type="etymology">
<paragraph id="3434723417EAA9C598CFFB9C7C70FB8E" blockId="219.[151,1437,152,1138]" box="[197,1075,1111,1138]" pageId="219" pageNumber="220">
<emphasis id="06FFAE2617EAA9C598CFFB9C7912FB8D" bold="true" box="[197,337,1111,1137]" pageId="219" pageNumber="220">Etymology.</emphasis>
<taxonomicName id="F38B09B717EAA9C59953FB9379ECFB8D" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[345,431,1112,1137]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="219" pageNumber="220" phylum="Arthropoda" rank="species" species="fianara">
<emphasis id="06FFAE2617EAA9C59953FB9379ECFB8D" box="[345,431,1112,1137]" italics="true" pageId="219" pageNumber="220">fianara</emphasis>
</taxonomicName>
, derived from the
<typeStatus id="EB30CC9617EAA9C59A82FB937AF9FB8E" box="[648,698,1112,1138]" pageId="219" pageNumber="220">type</typeStatus>
locality,
<collectingRegion id="F64FBCD617EAA9C59B2DFB937C6CFB8E" box="[807,1071,1112,1138]" country="Madagascar" name="Fianarantsoa" pageId="219" pageNumber="220">Fianarantsoa Province</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>