145 lines
22 KiB
XML
145 lines
22 KiB
XML
<document id="4AA9F8D5803F434FEF82AF1703F6FC04" ID-ISSN="11755334" approvalRequired="298" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="51" approvalRequired_for_textStreams="166" approvalRequired_for_treatments="80" checkinTime="1627004795516" checkinUser="felipe" docAuthor="Oláh, J. & Johanson, K. A." docDate="2008" docId="BC22C3221740A96C989DFE1C793AFED6" docLanguage="en" docName="zt01802p248.pdf" docOrigin="Zootaxa 1802" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.2:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="2" docTitle="Hydropsyche kiogupa Olah & Schefter 2008, new species" docType="treatment" docVersion="2" lastPageNumber="115" masterDocId="401BBB5A1731A91E980AFFCB7843FFFC" masterDocTitle="Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)" masterLastPageNumber="248" masterPageNumber="1" pageNumber="114" updateTime="1651296210969" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="C484C8240D0E9C4F31AB6811318A7B65">Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)</mods:title>
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<mods:namePart id="1E0E62FD2EAB9530A137DEA251B4F70A">Oláh, J.</mods:namePart>
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<mods:namePart id="E1EF5F202C6590CDDB4177F8A691E025">Johanson, K. A.</mods:namePart>
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<mods:typeOfResource id="808247AF92839BAC34F625794722D4A1">text</mods:typeOfResource>
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<mods:title id="2069A28CCC38C0979333F661704322F8">Zootaxa</mods:title>
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<mods:part id="591B8F1CC8FCCEF335A5DEA6A37D673E">
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<mods:date id="5105E413043FBC862F7B4EE1B965060C">2008</mods:date>
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<mods:number id="9CD929C5DF843664991AA4CA8B4A35CD">2008-06-16</mods:number>
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<mods:number id="D5820FDD2060DE151020834B8B2323A6">1802</mods:number>
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<mods:identifier id="67FFEE3430C97CCDFFD01EF7CBE70AEA" type="ISSN">11755334</mods:identifier>
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<treatment id="BC22C3221740A96C989DFE1C793AFED6" LSID="urn:lsid:plazi:treatment:BC22C3221740A96C989DFE1C793AFED6" httpUri="http://treatment.plazi.org/id/BC22C3221740A96C989DFE1C793AFED6" lastPageId="114" lastPageNumber="115" pageId="113" pageNumber="114">
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<subSubSection id="7C9121BF1740A96F989DFE1C7B4EFE0D" box="[151,781,471,497]" pageId="113" pageNumber="114" type="nomenclature">
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<paragraph id="343472341740A96F989DFE1C7B4EFE0D" blockId="113.[151,781,471,538]" box="[151,781,471,497]" pageId="113" pageNumber="114">
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<heading id="6F7CC5581740A96F989DFE1C7B4EFE0D" bold="true" box="[151,781,471,497]" fontSize="11" level="1" pageId="113" pageNumber="114" reason="1">
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<taxonomicName id="F38B09B71740A96F989DFE1C7A32FE0D" authority="Olah & Schefter" authorityName="Olah & Schefter" authorityYear="2008" box="[151,625,471,497]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="kiogupa" status="new species">
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<emphasis id="06FFAE261740A96F989DFE1C79DDFE0D" bold="true" box="[151,414,471,497]" italics="true" pageId="113" pageNumber="114">Hydropsyche kiogupa</emphasis>
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Oláh & Schefter
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</taxonomicName>
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,
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7C9121BF1740A96F989DFDCB7C3AF81E" pageId="113" pageNumber="114" type="description">
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<paragraph id="343472341740A96F989DFDCB7970FDE6" blockId="113.[151,781,471,538]" box="[151,307,512,538]" pageId="113" pageNumber="114">
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<figureCitation id="ACB06EB11740A96F989DFDCB7970FDE6" box="[151,307,512,538]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 183–187</figureCitation>
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</paragraph>
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<paragraph id="343472341740A96F989DFD9B7A46FC7E" blockId="113.[151,1436,592,2018]" pageId="113" pageNumber="114">
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This species is most similar to
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<taxonomicName id="F38B09B71740A96F9A06FD9B7B43FD96" authority="Banks" authorityName="Banks" box="[524,768,592,618]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="pallipenne">
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<emphasis id="06FFAE261740A96F9A06FD9B7AEDFD95" box="[524,686,592,617]" italics="true" pageId="113" pageNumber="114">H. pallipenne</emphasis>
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Banks
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</taxonomicName>
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from
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<collectingCountry id="4C9C32A41740A96F9B46FD9B7BF4FD96" box="[844,951,592,618]" name="Malaysia" pageId="113" pageNumber="114">Malaysia</collectingCountry>
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(
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<collectingRegion id="F64FBCD61740A96F9BC3FD9B7C60FD96" box="[969,1059,592,618]" country="Malaysia" name="Pahang" pageId="113" pageNumber="114">Pahang</collectingRegion>
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) and
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<collectingCountry id="4C9C32A41740A96F9C6DFD9B7C8EFD96" box="[1127,1229,592,618]" name="Pakistan" pageId="113" pageNumber="114">Pakistan</collectingCountry>
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, from which it is easily distinguished by its characteristically S-shaped gonocoxites. Moreover, the phallotremal sclerites are robust, not tapering and slender; and the sclerous endothecal process is bilobed, with a small, dorsobasal lobe. Male. Body and wings brown. Forewing membranes light brown with dark brown maculations, especially on veins. Sternum V with protuberance. Maxillary palp formula I-II-(III, IV)-V, segment V as long as segments I–IV together. Head dorsum brown, with 9 brown warts. Swollen setal wart present on proepisternum. Pretarsal claws asymmetrical, laterally flanked by setal bundle. Spur formula 244. Both protibiae with posteroapical spur equal to anteroapical spur.
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</paragraph>
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<paragraph id="343472341740A96F98CFFC5B7C65FC06" blockId="113.[151,1436,592,2018]" pageId="113" pageNumber="114">Wings. Venation typical for the genus. Forewing crossveins m-cu and cu separate. Sc and R, and Cu2 and A run separately before costa. Hind wing Sc and R meeting before costa at crossvein r. Stem of M and Cu1 running closely. Median cell closed. Fork 1 present. Forewing length 9.3 mm.</paragraph>
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<paragraph id="343472341740A96F98CFFBC37C3AF81E" blockId="113.[151,1436,592,2018]" pageId="113" pageNumber="114">
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Male genitalia. Abdominal segment IX fused annularly; median keel slightly narrowing in dorsal view (
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<figureCitation id="ACB06EB11740A96F9895FBFB7949FBB6" box="[159,266,1072,1098]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 184</figureCitation>
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); dorsal and lateral surfaces granulous; anterior margin arciform, ventrum short; dorsum slightly longer than ventrum (
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<figureCitation id="ACB06EB11740A96F999EFB9379B8FB8E" box="[404,507,1112,1138]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 183</figureCitation>
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). Antecosta well developed into double line, narrowing dorsally before margin, broadest ventrally; external groove of antecostal suture present. Apical lobe on posterolateral margins sharply triangular (
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<figureCitation id="ACB06EB11740A96F9910FB6379C1FB3E" box="[282,386,1192,1218]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 183</figureCitation>
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). Posterior spine rows continuous, but scarce, except on apical lobes (
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<figureCitation id="ACB06EB11740A96F9CC8FB637D69FB3E" box="[1218,1322,1192,1218]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 183</figureCitation>
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). Depression between segments IX and X very short and deep, oblique anteriorly, not vertical. Body of segment X large, dorsal margin short, ventral margin long (
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<figureCitation id="ACB06EB11740A96F9AC6FB337B70FAEE" box="[716,819,1272,1298]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 183</figureCitation>
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); subquadrangular in dorsal view (
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<figureCitation id="ACB06EB11740A96F9CC6FB337D70FAEE" box="[1228,1331,1272,1298]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 184</figureCitation>
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). Lateral setose areas (preanal appendages) forming compact ovoid area with pale background centrally on lateral surfaces, and distal setae scattered toward apical margin of segment X. Apicoventral setose lobes forming pair of stout, short, digitiform processes with apical setae; directed posteromesad. Dorsal interlobular gap small, diamond-shaped, encircled by apicoventral setose lobes (
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<figureCitation id="ACB06EB11740A96F9B11FA537BC7FA4E" box="[795,900,1432,1458]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 184</figureCitation>
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). Apicodorsal setose lobes fusing into setose ridge, or crest; elongated dorsally, shifted anteriorly to short, deep intersegmental depression; setae arranged in lateral rows (
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<figureCitation id="ACB06EB11740A96F9945FA2379F5F9FE" box="[335,438,1512,1538]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 184</figureCitation>
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). Smooth cavity forming small, shallow depression, occupying basal part of segment X. Short, wide, darkly pigmented transverse suture present on each side, running ventrally from bottom of intersegmental depression (
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<figureCitation id="ACB06EB11740A96F99D3F9F37A03F9AE" box="[473,576,1592,1618]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 183</figureCitation>
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); visible in dorsal view as prominent suture crossing segment X in intersegmental depression. Longitudinal sutures encircling ventral part of shallow cavity. Coxopodites reaching apex of segment X; strongly sinuous, slightly dilating distally from mid-length; weakly curving mesad in ventral view (
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<figureCitation id="ACB06EB11740A96F9919F97B793FF936" box="[275,380,1712,1738]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 185</figureCitation>
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). Harpagones parallel sided and curving dorsally along their lengths, slightly capitate in lateral view (
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<figureCitation id="ACB06EB11740A96F993AF91379D6F90E" box="[304,405,1752,1778]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 183</figureCitation>
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); curving mediad in ventral view (
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<figureCitation id="ACB06EB11740A96F9B2DF9137BCDF90E" box="[807,910,1752,1778]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 185</figureCitation>
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). Phallic apparatus simply bent, with slightly developed ventrad curving basal part; horizontal section of phallotheca straight, dilating apically.
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<taxonomicName id="F38B09B71740A96F9D03F8CB7DD8F8E6" authorityName="M. Zang & R.H. Petersen" authorityYear="1989" box="[1289,1435,1792,1818]" class="Agaricomycetes" family="Phallaceae" genus="Endophallus" kingdom="Fungi" order="Phallales" pageId="113" pageNumber="9" phylum="Basidiomycota" rank="genus">Endophallus</taxonomicName>
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forming narrow tube attached to dorsal wall of phallotheca; reaching bend of phallic apparatus. Phallotremal sclerites well separated into pair of stout, claw-like processes with blunt apex; falciform in lateral view; pincer-shaped in ventral view (
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<figureCitation id="ACB06EB11740A96F99ECF8B37A0CF86E" box="[486,591,1912,1938]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 187</figureCitation>
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). Pair of sclerous endothecal processes arising laterally on dominating bilobed phallotremal sclerites; basodorsal lobe small, visible in lateral view (
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<figureCitation id="ACB06EB11740A96F9C30F86B7CE7F846" box="[1082,1188,1952,1978]" captionStart="FIGURES 183–187" captionStartId="114.[151,269,1509,1533]" captionTargetBox="[179,1413,358,1467]" captionTargetId="figure-191@114.[151,1436,324,1486]" captionTargetPageId="114" captionText="FIGURES 183–187. Hydropsyche kiogupa Oláh & Schefter, new species, holotype, male. 183 — genitalia, lateral; 184 — genitalia, dorsal; 185 — gonocoxite, ventral; 186 — phallus, lateral; 187 — phallus apex, ventral." pageId="113" pageNumber="114">Fig. 186</figureCitation>
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). Sclerotized ventral process of phallotheca shorter than phallotremal sclerites, with slightly bilobed apex.
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</paragraph>
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</subSubSection>
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<subSubSection id="7C9121BF1743A96C98CFFF5C79DEFF26" pageId="114" pageNumber="115" type="materials_examined">
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<paragraph id="343472341743A96C98CFFF5C79DEFF26" blockId="114.[151,1436,151,298]" pageId="114" pageNumber="115">
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<materialsCitation id="84E378691743A96C98CFFF5C79DEFF26" collectingDate="1961-07-15" collectionCode="ROM" collectorName="F. Schmid" country="India" location="Ukhrul" pageId="114" pageNumber="115" specimenCount="1" stateProvince="Manipur" typeStatus="holotype">
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<emphasis id="06FFAE261743A96C98CFFF5C7939FF4D" bold="true" box="[197,378,151,177]" pageId="114" pageNumber="115">
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<typeStatus id="EB30CC961743A96C98CFFF5C7974FF4D" box="[197,311,151,177]" pageId="114" pageNumber="115">Holotype</typeStatus>
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male
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</emphasis>
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||
:
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<emphasis id="06FFAE261743A96C9983FF5C79AAFF4D" bold="true" box="[393,489,151,177]" pageId="114" pageNumber="115">
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||
<collectingCountry id="4C9C32A41743A96C9983FF5C79A7FF4D" box="[393,484,151,177]" name="India" pageId="114" pageNumber="115">INDIA</collectingCountry>
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||
:
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||
</emphasis>
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||
<collectingRegion id="F64FBCD61743A96C99FAFF537A14FF4E" box="[496,599,152,178]" country="India" name="Manipur" pageId="114" pageNumber="115">Manipur</collectingRegion>
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||
,
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<location id="315424EF1743A96C9A68FF537AFBFF4E" LSID="urn:lsid:plazi:treatment:BC22C3221740A96C989DFE1C793AFED6:315424EF1743A96C9A68FF537AFBFF4E" box="[610,696,152,178]" country="India" name="Ukhrul" pageId="114" pageNumber="115" stateProvince="Manipur">Ukhrul</location>
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,
|
||
<date id="403554F41743A96C9ACEFF537B0AFF4E" box="[708,841,152,178]" pageId="114" pageNumber="115" value="1961-07-15">
|
||
<collectingDate id="5071AD1C1743A96C9ACEFF537B0AFF4E" box="[708,841,152,178]" pageId="114" pageNumber="115" value="1961-07-15">15.vii.1961</collectingDate>
|
||
</date>
|
||
[
|
||
<collectorName id="997E17E21743A96C9B52FF537B96FF4E" box="[856,981,152,178]" pageId="114" pageNumber="115">F. Schmid</collectorName>
|
||
], (
|
||
<collectionCode id="529AEAF11743A96C9BFAFF537C74FF4E" box="[1008,1079,152,178]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="114" pageNumber="115" type="Museum">ROM</collectionCode>
|
||
, pinned with cleared abdomen in plastic genital vial).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7C9121BF1743A96C98CFFF2C793AFED6" pageId="114" pageNumber="115" type="etymology">
|
||
<paragraph id="343472341743A96C98CFFF2C793AFED6" blockId="114.[151,1436,151,298]" pageId="114" pageNumber="115">
|
||
<emphasis id="06FFAE261743A96C98CFFF2C7912FEFD" bold="true" box="[197,337,231,257]" pageId="114" pageNumber="115">Etymology.</emphasis>
|
||
<taxonomicName id="F38B09B71743A96C9956FF2379F9FEFD" authorityName="Olah & Schefter" authorityYear="2008" box="[348,442,232,257]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="kiogupa">
|
||
<emphasis id="06FFAE261743A96C9956FF2379F9FEFD" box="[348,442,232,257]" italics="true" pageId="114" pageNumber="115">kiogupa</emphasis>
|
||
</taxonomicName>
|
||
, from “kiogupa”, meaning double in Sanskrit, referring to the double-curving, Sshaped gonocoxite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |