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<document id="6CC04F3AE332A86152AAE4515919B0C2" ID-ISSN="1175­5334" approvalRequired="298" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="51" approvalRequired_for_textStreams="166" approvalRequired_for_treatments="80" checkinTime="1627004795516" checkinUser="felipe" docAuthor="Oláh, J. &amp; Johanson, K. A." docDate="2008" docId="BC22C3221702A92D989DFCD47918F8FE" docLanguage="en" docName="zt01802p248.pdf" docOrigin="Zootaxa 1802" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.2:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="2" docTitle="Calosopsyche domingensis" docType="treatment" docVersion="3" lastPageNumber="52" masterDocId="401BBB5A1731A91E980AFFCB7843FFFC" masterDocTitle="Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)" masterLastPageNumber="248" masterPageNumber="1" pageNumber="52" updateTime="1651296210969" updateUser="GgImagineBatch" zenodo-license-document="CLOSED">
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<mods:title id="DB894FF90AF028453B2A32BDB91F41AC">Generic review of Hydropsychinae, with description of Schmidopsyche, new genus, 3 new genus clusters, 8 new species groups, 4 new species clades, 12 new species clusters and 62 new species from the Oriental and Afrotropical regions (Trichoptera: Hydropsychidae)</mods:title>
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<mods:namePart id="D526E44F47668C80ED4A69A9C663F2E9">Oláh, J.</mods:namePart>
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<mods:namePart id="12D3150F0974023DB7726DAF5C6DAE2D">Johanson, K. A.</mods:namePart>
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<mods:date id="B3FA765CD545C71E6BC2BE220C614DCF">2008</mods:date>
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<mods:number id="4DC1F834FE7932D96AE052677B53738F">2008-06-16</mods:number>
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<mods:number id="DB21BB5873D27BD48434E6C8312B25D7">1802</mods:number>
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<mods:identifier id="2CA294416AB222C93261A4B43689C27D" type="ISSN">1175­5334</mods:identifier>
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<treatment id="BC22C3221702A92D989DFCD47918F8FE" ID-DOI="http://doi.org/10.5281/zenodo.5125629" ID-Zenodo-Dep="5125629" LSID="urn:lsid:plazi:treatment:BC22C3221702A92D989DFCD47918F8FE" httpUri="http://treatment.plazi.org/id/BC22C3221702A92D989DFCD47918F8FE" lastPageNumber="52" pageId="51" pageNumber="52">
<subSubSection id="7C9121BF1702A92D989DFCD47A00FCC5" box="[151,579,799,825]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="343472341702A92D989DFCD47A00FCC5" blockId="51.[151,579,799,825]" box="[151,579,799,825]" pageId="51" pageNumber="52">
<heading id="6F7CC5581702A92D989DFCD47A00FCC5" bold="true" box="[151,579,799,825]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
<taxonomicName id="F38B09B71702A92D989DFCD47A00FCC5" ID-CoL="Q3MV" authority="(Banks)" baseAuthorityName="Banks" box="[151,579,799,825]" class="Insecta" family="Hydropsychidae" genus="Calosopsyche" kingdom="Animalia" order="Trichoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="domingensis">
<emphasis id="06FFAE261702A92D989DFCD47A00FCC5" bold="true" box="[151,579,799,825]" pageId="51" pageNumber="52">
<emphasis id="06FFAE261702A92D989DFCD47999FCC5" bold="true" box="[151,474,799,825]" italics="true" pageId="51" pageNumber="52">Calosopsyche domingensis</emphasis>
(Banks)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF1702A92D989DFCBB7BABFC1B" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="343472341702A92D989DFCBB7A34FC7B" blockId="51.[151,1000,880,999]" box="[151,631,880,903]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="149B551A1702A92D989DFCBB7A34FC7B" box="[151,631,880,903]" pageId="51" pageNumber="52">
<treatmentCitation id="B52A54251702A92D989DFCBB7A30FC7B" author="Banks N." box="[151,627,880,903]" page="398" pageId="51" pageNumber="52" year="1941">
<taxonomicName id="F38B09B71702A92D989DFCBB7A30FC7B" authority="Banks, 1941: 398" authorityName="Banks" authorityPageNumber="398" authorityYear="1941" box="[151,627,880,903]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="domingensis">
<emphasis id="06FFAE261702A92D989DFCBB79EDFC7B" box="[151,430,880,903]" italics="true" pageId="51" pageNumber="52">Hydropsyche domingensis</emphasis>
<bibRefCitation id="501A0FC51702A92D99BFFCBB7A30FC7B" author="Banks N." box="[437,627,880,903]" pageId="51" pageNumber="52" pagination="385 - 402" refId="ref106558" refString="Banks N. (1941) New neuropteroid insects from the Antilles. Memorias de la Sociedad Cubana de Historia Natural, 15, 385 - 402." type="journal article" year="1941">Banks, 1941: 398</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="343472341702A92D989DFC5B7BD6FC5B" blockId="51.[151,1000,880,999]" box="[151,917,912,935]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="149B551A1702A92D989DFC5B7BD6FC5B" box="[151,917,912,935]" pageId="51" pageNumber="52">
<taxonomicName id="F38B09B71702A92D989DFC5B7A1AFC5B" ID-CoL="6VQLG" authority="(Banks, 1941)" baseAuthorityName="Banks" baseAuthorityYear="1941" box="[151,601,912,935]" class="Insecta" family="Hydropsychidae" genus="Plectropsyche" kingdom="Animalia" order="Trichoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="domingensis">
<emphasis id="06FFAE261702A92D989DFC5B79FAFC5B" box="[151,441,912,935]" italics="true" pageId="51" pageNumber="52">Plectropsyche domingensis</emphasis>
(
<bibRefCitation id="501A0FC51702A92D99CDFC5B7A12FC5B" author="Banks N." box="[455,593,912,935]" pageId="51" pageNumber="52" pagination="385 - 402" refId="ref106558" refString="Banks N. (1941) New neuropteroid insects from the Antilles. Memorias de la Sociedad Cubana de Historia Natural, 15, 385 - 402." type="journal article" year="1941">Banks, 1941</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="B52A54251702A92D9A6FFC5B7BD2FC5B" author="Ross, H. H. &amp; Unzicker, J. D." box="[613,913,912,935]" page="308" pageId="51" pageNumber="52" year="1977">
<bibRefCitation id="501A0FC51702A92D9A6FFC5B7BD2FC5B" author="Ross, H. H. &amp; Unzicker, J. D." box="[613,913,912,935]" pageId="51" pageNumber="52" pagination="298 - 312" refId="ref111855" refString="Ross, H. H. &amp; Unzicker, J. D. (1977) The relationships of the genera of American Hydropsychinae as indicated by phallic structures (Trichoptera, Hydropsychidae). Journal of the Georgia Entomological Society, 12 (4), 298 - 312." type="journal article" year="1977">Ross &amp; Unzicker 1977: 308</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="343472341702A92D989DFC7B7B7EFC3B" blockId="51.[151,1000,880,999]" box="[151,829,944,967]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="149B551A1702A92D989DFC7B7B7EFC3B" box="[151,829,944,967]" pageId="51" pageNumber="52">
<taxonomicName id="F38B09B71702A92D989DFC7B7A7EFC3B" authority="Banks, 1941" authorityName="Banks" authorityYear="1941" box="[151,573,944,967]" class="Insecta" family="Hydropsychidae" genus="Hydropsyche" kingdom="Animalia" order="Trichoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="domingensis">
<emphasis id="06FFAE261702A92D989DFC7B79EDFC3B" box="[151,430,944,967]" italics="true" pageId="51" pageNumber="52">Hydropsyche domingensis</emphasis>
<bibRefCitation id="501A0FC51702A92D99BFFC7B7A7EFC3B" author="Banks N." box="[437,573,944,967]" pageId="51" pageNumber="52" pagination="385 - 402" refId="ref106558" refString="Banks N. (1941) New neuropteroid insects from the Antilles. Memorias de la Sociedad Cubana de Historia Natural, 15, 385 - 402." type="journal article" year="1941">Banks, 1941</bibRefCitation>
</taxonomicName>
;
<treatmentCitation id="B52A54251702A92D9A42FC7B7B7AFC3B" author="Botosaneanu, L." box="[584,825,944,967]" page="17" pageId="51" pageNumber="52" year="1996">
<bibRefCitation id="501A0FC51702A92D9A42FC7B7B7AFC3B" author="Botosaneanu, L." box="[584,825,944,967]" pageId="51" pageNumber="52" pagination="5 - 26" refId="ref106807" refString="Botosaneanu, L. (1996) Caddis flies (Trichoptera) from the Dominican Republic (West Indies). II. All families except Hydroptilidae; with general observations for Hispaniola. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie, 66, 5 - 26." type="journal article" year="1996">Botosaneanu 1996: 17</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="343472341702A92D989DFC1B7BABFC1B" blockId="51.[151,1000,880,999]" box="[151,1000,976,999]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="149B551A1702A92D989DFC1B7BABFC1B" box="[151,1000,976,999]" pageId="51" pageNumber="52">
<taxonomicName id="F38B09B71702A92D989DFC1B7A14FC1B" ID-CoL="Q3MV" authority="(Banks, 1941)" baseAuthorityName="Banks" baseAuthorityYear="1941" box="[151,599,976,999]" class="Insecta" family="Hydropsychidae" genus="Calosopsyche" kingdom="Animalia" order="Trichoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="domingensis">
<emphasis id="06FFAE261702A92D989DFC1B79F5FC1B" box="[151,438,976,999]" italics="true" pageId="51" pageNumber="52">Calosopsyche domingensis</emphasis>
(
<bibRefCitation id="501A0FC51702A92D99CFFC1B7A0CFC1B" author="Banks N." box="[453,591,976,999]" pageId="51" pageNumber="52" pagination="385 - 402" refId="ref106558" refString="Banks N. (1941) New neuropteroid insects from the Antilles. Memorias de la Sociedad Cubana de Historia Natural, 15, 385 - 402." type="journal article" year="1941">Banks, 1941</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="B52A54251702A92D9A68FC1B7BA7FC1B" author="Flint, O. S., Jr. &amp; Holzenthal, R. W. &amp; Harris, S. C." box="[610,996,976,999]" page="75" pageId="51" pageNumber="52" year="1999">
<bibRefCitation id="501A0FC51702A92D9A68FC1B7BA7FC1B" author="Flint, O. S., Jr. &amp; Holzenthal, R. W. &amp; Harris, S. C." box="[610,996,976,999]" pageId="51" pageNumber="52" pagination="73 - 84" refId="ref107255" refString="Flint, O. S., Jr., Holzenthal, R. W. &amp; Harris, S. C. (1999) Nomenclatural and systematic changes in the Neotropical caddisflies (Insecta: Trichoptera). Insecta Mundi, 13 (1 - 2), 73 - 84." type="journal article" year="1999">Flint, Holzenthal &amp; Harris 1999: 75</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7C9121BF1702A92D989DFBDC7918F8FE" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="343472341702A92D989DFBDC7A47FBCE" blockId="51.[151,1436,1047,1794]" box="[151,516,1047,1074]" pageId="51" pageNumber="52">
<materialsCitation id="84E378691702A92D989DFBDC7A42FBCE" box="[151,513,1047,1074]" location="Santo Domingo" pageId="51" pageNumber="52" specimenCount="1" stateProvince="Santo Domingo" typeStatus="holotype">
<emphasis id="06FFAE261702A92D989DFBDC797FFBCD" bold="true" box="[151,316,1047,1073]" pageId="51" pageNumber="52">
<typeStatus id="EB30CC961702A92D989DFBDC7897FBCD" box="[151,212,1047,1073]" pageId="51" pageNumber="52">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="F64FBCD61702A92D994EFBD37A42FBCE" box="[324,513,1048,1074]" country="Dominican Republic" name="Santo Domingo" pageId="51" pageNumber="52">Santo Domingo</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="343472341702A92D98CCFB8B7918F8FE" blockId="51.[151,1436,1047,1794]" pageId="51" pageNumber="52">
Remarks: As is seen from the frequent generic transfers, the status of this species was highly instable. Even in the cladistic analysis of
<bibRefCitation id="501A0FC51702A92D9A19FBA37A89FB7E" author="Schefter, P. W." box="[531,714,1128,1154]" pageId="51" pageNumber="52" pagination="133 - 154" refId="ref111953" refString="Schefter, P. W. (2005) Re-evaluation of genera in the subfamily Hydropsychinae (Trichoptera: Hydropsychidae). Aquatic Insects, 27 (2), 133 - 154." type="journal article" year="2005">Schefter (2005)</bibRefCitation>
it was suggested and isolated into a distinct genus. However, based upon defined primary generic characters, the species represents a true
<taxonomicName id="F38B09B71702A92D9C21FB5B7C8DFB55" box="[1067,1230,1168,1193]" class="Insecta" family="Hydropsychidae" genus="Calosopsyche" kingdom="Animalia" order="Trichoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="06FFAE261702A92D9C21FB5B7C8DFB55" box="[1067,1230,1168,1193]" italics="true" pageId="51" pageNumber="52">Calosopsyche</emphasis>
</taxonomicName>
, having identical primary generic characters with the other members of the genus. We have found some mistakes in the character matrix used in the cladistic analysis by
<bibRefCitation id="501A0FC51702A92D9AA9FB2B7B22FB06" author="Schefter, P. W." box="[675,865,1248,1274]" pageId="51" pageNumber="52" pagination="133 - 154" refId="ref111953" refString="Schefter, P. W. (2005) Re-evaluation of genera in the subfamily Hydropsychinae (Trichoptera: Hydropsychidae). Aquatic Insects, 27 (2), 133 - 154." type="journal article" year="2005">Schefter (2005)</bibRefCitation>
. These mistakes were mainly caused by sexual dimorphism. The pretarsal claw is asymmetrical, not symmetrical. Like in other genera, or other species, even inside the genus
<taxonomicName id="F38B09B71702A92D9956FAFB79BDFAB5" box="[348,510,1328,1353]" class="Insecta" family="Hydropsychidae" genus="Calosopsyche" kingdom="Animalia" order="Trichoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="06FFAE261702A92D9956FAFB79BDFAB5" box="[348,510,1328,1353]" italics="true" pageId="51" pageNumber="52">Calosopsyche</emphasis>
</taxonomicName>
, asymmetry of the pretarsal claws is frequently sex linked. Even the degree of asymmetry varies greatly. Asymmetry involving laterally flanked setal bundle may occur only in the fore-leg, fore- and mid-leg, or in all legs in the male. This may, or may not be combined with identical variation in the female. The hind wing crossvein m-cu, representing one of the most important primary generic characters, is well visible in both
<taxonomicName id="F38B09B71702A92D9974FA1B7BFBFA16" authority="Flint &amp; Bueno-Soria" authorityName="Flint &amp; Bueno-Soria" box="[382,952,1488,1514]" class="Insecta" family="Hydropsychidae" genus="Calosopsyche" kingdom="Animalia" order="Trichoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="continentalis">
<emphasis id="06FFAE261702A92D9974FA1B7AFDFA15" box="[382,702,1488,1513]" italics="true" pageId="51" pageNumber="52">Calosopsyche continentalis</emphasis>
Flint &amp; Bueno-Soria
</taxonomicName>
, and
<taxonomicName id="F38B09B71702A92D9BFFFA1B7DD4FA16" authority="(Banks)" baseAuthorityName="Banks" box="[1013,1431,1488,1514]" class="Insecta" family="Hydropsychidae" genus="Calosopsyche" kingdom="Animalia" order="Trichoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="domingensis">
<emphasis id="06FFAE261702A92D9BFFFA1B7D73FA15" box="[1013,1328,1488,1513]" italics="true" pageId="51" pageNumber="52">Calosopsyche domingensis</emphasis>
(Banks)
</taxonomicName>
, like in all other species in the genus. Among the tertiary generic characters, the
<taxonomicName id="F38B09B71702A92D9C4BFA337C8DF9EE" authorityName="M. Zang &amp; R.H. Petersen" authorityYear="1989" box="[1089,1230,1528,1554]" class="Agaricomycetes" family="Phallaceae" genus="Endophallus" kingdom="Fungi" order="Phallales" pageId="51" pageNumber="9" phylum="Basidiomycota" rank="genus">endophallus</taxonomicName>
is long, not short as listed in the character matrix. Even the 3 setose processes are present, but shortened, forming almost a single, united, lobe. A careful examination shows that 3 separate setose surfaces are distinguishable, among them, the preanal appendages. After correcting the character matrix of
<bibRefCitation id="501A0FC51702A92D9BE0F9BB7CE0F976" author="Schefter, P. W." box="[1002,1187,1648,1674]" pageId="51" pageNumber="52" pagination="133 - 154" refId="ref111953" refString="Schefter, P. W. (2005) Re-evaluation of genera in the subfamily Hydropsychinae (Trichoptera: Hydropsychidae). Aquatic Insects, 27 (2), 133 - 154." type="journal article" year="2005">Schefter (2005)</bibRefCitation>
for these characters, one of us (
<collectionCode id="529AEAF11702A92D991AF953790AF94E" box="[272,329,1688,1714]" pageId="51" pageNumber="52">KAJ</collectionCode>
) re-ran the analysis in
<collectionCode id="529AEAF11702A92D9A59F9537AE1F94E" box="[595,674,1688,1714]" country="India" httpUri="http://grbio.org/cool/i1x1-g5cx" name="Punjab Agricultural University" pageId="51" pageNumber="52">PAUP</collectionCode>
*. The many more shortest trees obtained were 2 steps shorter and the strict consensus tree did not improve resolution. Furthermore, the two
<emphasis id="06FFAE261702A92D9C1EF90B7C86F925" box="[1044,1221,1728,1753]" italics="true" pageId="51" pageNumber="52">Calaosopsyche</emphasis>
species were still distantly related.
</paragraph>
</subSubSection>
</treatment>
</document>
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