419 lines
53 KiB
XML
419 lines
53 KiB
XML
<document id="DA535D364F96AF337012C186ED0CC290" ID-DOI="10.11646/zootaxa.3681.4.6" ID-GBIF-Dataset="cd4060b8-9845-4f8a-88d9-f50fa7fbfa21" ID-ISSN="1175-5326" ID-Zenodo-Dep="247302" ID-ZooBank="FD512749-AB7A-4EC5-9983-BDC648B78FC6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460640836473" checkinUser="plazi" docAuthor="Kunz, Tobias Saraiva & Borges-Martins, Márcio" docDate="2013" docId="A7629F4DFFECFFE6FF54FB35E385FE00" docLanguage="en" docName="zt03681p439.pdf" docOrigin="Zootaxa 3681 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tropidurus catalanensis Gudynas & Skuk 1983" docType="treatment" docVersion="8" lastPageNumber="432" masterDocId="5B5BE735FFE2FFF5FFC3FFC3E05DFFCF" masterDocTitle="A new microendemic species of Tropidurus (Squamata: Tropiduridae) from southern Brazil and revalidation of Tropidurus catalanensis Gudynas & Skuk, 1983" masterLastPageNumber="439" masterPageNumber="413" pageNumber="427" updateTime="1698335809894" updateUser="plazi">
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<mods:title id="84A1D161AEC733C011F4714586973C0C">A new microendemic species of Tropidurus (Squamata: Tropiduridae) from southern Brazil and revalidation of Tropidurus catalanensis Gudynas & Skuk, 1983</mods:title>
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<mods:namePart id="8C25E2FCB656D93E4FC24CAA5B99CED1">Kunz, Tobias Saraiva</mods:namePart>
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<mods:namePart id="1E66BB061977B913B7ECA121164824F5">Borges-Martins, Márcio</mods:namePart>
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<mods:title id="6DADA044898C024D743A2DE739F631B8">Zootaxa</mods:title>
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<mods:date id="E3E11ACC10C14A0D95A6615BCD54B973">2013</mods:date>
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<treatment id="A7629F4DFFECFFE6FF54FB35E385FE00" ID-DOI="http://doi.org/10.5281/zenodo.5632789" ID-GBIF-Taxon="119571242" ID-Zenodo-Dep="5632789" LSID="urn:lsid:plazi:treatment:A7629F4DFFECFFE6FF54FB35E385FE00" httpUri="http://treatment.plazi.org/id/A7629F4DFFECFFE6FF54FB35E385FE00" lastPageId="19" lastPageNumber="432" pageId="14" pageNumber="427">
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<subSubSection id="67D17DD0FFECFFFBFF54FB35E170FAFE" pageId="14" pageNumber="427" type="nomenclature">
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<paragraph id="2F742E5BFFECFFFBFF54FB35E3DDFADF" blockId="14.[151,896,1270,1329]" box="[151,896,1270,1296]" pageId="14" pageNumber="427">
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<heading id="743C9937FFECFFFBFF54FB35E3DDFADF" bold="true" box="[151,896,1270,1296]" fontSize="11" level="1" pageId="14" pageNumber="427" reason="1">
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<emphasis id="1DBFF249FFECFFFBFF54FB35E3DDFADF" bold="true" box="[151,896,1270,1296]" pageId="14" pageNumber="427">
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<taxonomicName id="E8CB55D8FFECFFFBFF54FB35E2BBFADF" authority="Gudynas & Skuk, 1983" authorityName="Gudynas & Skuk" authorityYear="1983" box="[151,742,1270,1296]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="catalanensis">
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<emphasis id="1DBFF249FFECFFFBFF54FB35E1E6FADF" bold="true" box="[151,443,1270,1296]" italics="true" pageId="14" pageNumber="427">Tropidurus catalanensis</emphasis>
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<bibRefCitation id="4B5A53AAFFECFFFBFE00FB35E2BBFADF" author="Gudynas" box="[451,742,1270,1296]" pageId="14" pageNumber="427" refString="Gudynas, E. & Skuk, G. (1983) A new species of the iguanid lizard genus Tropidurus from temperate South America (Lacertilia: Iguanidae). Centro Educativo Don Orione Contribuciones en Biologia, 10, 1 - 10." type="journal article" year="1983">Gudynas & Skuk, 1983</bibRefCitation>
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</taxonomicName>
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, revalidated
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="2F742E5BFFECFFFBFF54FADAE170FAFE" blockId="14.[151,896,1270,1329]" box="[151,301,1305,1329]" pageId="14" pageNumber="427">
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(
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<figureCitation id="B7F032DEFFECFFFBFF5CFADAE17BFAFE" box="[159,294,1305,1329]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStart-3="FIGURE 16" captionStart-4="FIGURE 17" captionStartId-0="15.[151,250,1327,1350]" captionStartId-1="16.[151,250,1051,1074]" captionStartId-2="17.[151,250,1050,1073]" captionStartId-3="17.[151,250,1975,1998]" captionStartId-4="18.[151,250,994,1017]" captionTargetBox-0="[196,1400,215,1287]" captionTargetBox-1="[151,1435,286,1030]" captionTargetBox-2="[151,1435,194,1028]" captionTargetBox-3="[151,1435,1124,1952]" captionTargetBox-4="[151,1435,194,900]" captionTargetId-0="figure@15.[170,1416,194,1306]" captionTargetId-1="figure@16.[151,1435,264,1030]" captionTargetId-2="figure@17.[151,1435,194,1029]" captionTargetId-3="figure@17.[151,1435,1124,1953]" captionTargetId-4="figure@18.[151,1435,194,973]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="17" captionTargetPageId-4="18" captionText-0="FIGURE 13. Dorsal (left) and ventral (right) views of the holotype (MNHN 3489; SVL 114.8 mm) of Tropidurus catalanensis." captionText-1="FIGURE 14. Tropidurus catalanensis from Iguaçu National Park, Foz do Iguaçu, Paraná, Brazil (not collected). Note the yellow coloration of chin, including infra and supralabials." captionText-2="FIGURE 15. Tropidurus catalanensis from Campos Novos, Santa Catarina, near the Uruguay riverbank (not collected)." captionText-3="FIGURE 16. Tropidurus catalanensis from São Francisco de Assis, Rio Grande do Sul. Adult male (not collected)." captionText-4="FIGURE 17. Lateral view of head of Tropidurus catalanensis (UFRGS 5324) from São Francisco de Assis, Rio Grande do Sul, Brazil. Freshly killed subadult male (head length 21.4 mm)." httpUri-0="https://zenodo.org/record/247314/files/figure.png" httpUri-1="https://zenodo.org/record/247315/files/figure.png" httpUri-2="https://zenodo.org/record/247316/files/figure.png" httpUri-3="https://zenodo.org/record/247317/files/figure.png" httpUri-4="https://zenodo.org/record/247318/files/figure.png" pageId="14" pageNumber="427">Figs. 13–17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="67D17DD0FFECFFFBFF54FAA2E266FA3D" pageId="14" pageNumber="427" type="reference_group">
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<paragraph id="2F742E5BFFECFFFBFF54FAA2E325FA59" blockId="14.[151,1436,1377,1522]" pageId="14" pageNumber="427">
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<treatmentCitationGroup id="0FDB0975FFECFFFBFF54FAA2E325FA59" pageId="14" pageNumber="427">
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<taxonomicName id="E8CB55D8FFECFFFBFF54FAA2E2CFFAB8" authority="Gudynas & Skuk 1983: 2" authorityName="Gudynas & Skuk" authorityPageNumber="2" authorityYear="1983" box="[151,658,1377,1399]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="catalanensis">
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<emphasis id="1DBFF249FFECFFFBFF54FAA2E1D6FAB8" box="[151,395,1377,1399]" italics="true" pageId="14" pageNumber="427">Tropidurus catalanensis</emphasis>
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<treatmentCitation id="AE6A084AFFECFFFBFE52FAA2E2CFFAB8" author="Gudynas" box="[401,658,1377,1399]" page="2" pageId="14" pageNumber="427" year="1983">
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<bibRefCitation id="4B5A53AAFFECFFFBFE52FAA2E226FAB8" author="Gudynas" box="[401,635,1377,1399]" pageId="14" pageNumber="427" refString="Gudynas, E. & Skuk, G. (1983) A new species of the iguanid lizard genus Tropidurus from temperate South America (Lacertilia: Iguanidae). Centro Educativo Don Orione Contribuciones en Biologia, 10, 1 - 10." type="journal article" year="1983">Gudynas & Skuk 1983</bibRefCitation>
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: 2
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</treatmentCitation>
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</taxonomicName>
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.
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<typeStatus id="F07090F9FFECFFFBFD5FFAA1E293FAB8" box="[668,718,1378,1399]" pageId="14" pageNumber="427">Type</typeStatus>
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locality: “from a hill
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<quantity id="E83383BEFFECFFFBFC6BFAA1E386FAB8" box="[936,987,1377,1399]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="427" unit="km" value="1.0">1 km</quantity>
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300 ESE of route Nº 30 and Catalán Grande stream (airline), Departamento de Artigas,
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<collectingCountry id="57DC6ECBFFECFFFBFDB9FA42E28AFA59" box="[634,727,1409,1430]" name="Uruguay" pageId="14" pageNumber="427">Uruguay</collectingCountry>
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”. MNHN 3489.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="2F742E5BFFECFFFBFF54FA5CE2E2FA7A" blockId="14.[151,1436,1377,1522]" box="[151,703,1439,1461]" pageId="14" pageNumber="427">
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<treatmentCitationGroup id="0FDB0975FFECFFFBFF54FA5CE2E2FA7A" box="[151,703,1439,1461]" pageId="14" pageNumber="427">
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<taxonomicName id="E8CB55D8FFECFFFBFF54FA5CE130FA7A" box="[151,365,1439,1461]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="torquatus">
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<emphasis id="1DBFF249FFECFFFBFF54FA5CE130FA7A" box="[151,365,1439,1461]" italics="true" pageId="14" pageNumber="427">Tropidurus torquatus</emphasis>
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</taxonomicName>
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||
—
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<treatmentCitation id="AE6A084AFFECFFFBFE44FA5CE207FA7B" author="Rodrigues" box="[391,602,1439,1461]" page="190" pageId="14" pageNumber="427" year="1987">
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<bibRefCitation id="4B5A53AAFFECFFFBFE44FA5CE277FA7B" author="Rodrigues" box="[391,554,1439,1461]" pageId="14" pageNumber="427" refString="Rodrigues, M. T. (1987) Sistematica, ecologia e zoogeografia dos Tropidurus do grupo torquatus ao sul do rio Amazonas (Sauria, Iguanidae). Arquivos de Zoologia, 31 (3), 105 - 230." type="journal article" year="1987">Rodrigues 1987</bibRefCitation>
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: 190
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</treatmentCitation>
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(
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<emphasis id="1DBFF249FFECFFFBFDAAFA63E2F2FA7A" box="[617,687,1440,1461]" italics="true" pageId="14" pageNumber="427">Partim</emphasis>
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).
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="2F742E5BFFECFFFBFF54FA7EE2FFFA1C" blockId="14.[151,1436,1377,1522]" box="[151,674,1469,1491]" pageId="14" pageNumber="427">
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<treatmentCitationGroup id="0FDB0975FFECFFFBFF54FA7EE2FFFA1C" box="[151,674,1469,1491]" pageId="14" pageNumber="427">
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<taxonomicName id="E8CB55D8FFECFFFBFF54FA7EE1ACFA1C" box="[151,497,1469,1491]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="subSpecies" species="torquatus" subSpecies="catalanensis">
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<emphasis id="1DBFF249FFECFFFBFF54FA7EE1ACFA1C" box="[151,497,1469,1491]" italics="true" pageId="14" pageNumber="427">Tropidurus torquatus catalanensis</emphasis>
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</taxonomicName>
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—
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<treatmentCitation id="AE6A084AFFECFFFBFDCFFA7EE2C0FA1C" author="Cei" box="[524,669,1469,1491]" page="315" pageId="14" pageNumber="427" year="1993">
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<bibRefCitation id="4B5A53AAFFECFFFBFDCFFA7EE237FA1C" author="Cei" box="[524,618,1469,1491]" pageId="14" pageNumber="427" refString="Cei, J. M. (1993) Reptiles del Noroeste, Nordeste y Este de la Argentina - Herpetofauna de las Selvas Subtropicales, Puna y Pampas. Monografia XIV, Museo Regionale di Scienze Naturali, Torino, Italia, 950 pp." type="book" year="1993">Cei 1993</bibRefCitation>
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: 315
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="2F742E5BFFECFFFBFF54FA1FE266FA3D" blockId="14.[151,1436,1377,1522]" box="[151,571,1500,1522]" pageId="14" pageNumber="427">
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||
<treatmentCitationGroup id="0FDB0975FFECFFFBFF54FA1FE266FA3D" box="[151,571,1500,1522]" pageId="14" pageNumber="427">
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<taxonomicName id="E8CB55D8FFECFFFBFF54FA1FE1D6FA3D" box="[151,395,1500,1522]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="catalanensis">
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<emphasis id="1DBFF249FFECFFFBFF54FA1FE1D6FA3D" box="[151,395,1500,1522]" italics="true" pageId="14" pageNumber="427">Tropidurus catalanensis</emphasis>
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</taxonomicName>
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||
—
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<treatmentCitation id="AE6A084AFFECFFFBFE66FA1FE26AFA3D" author="Cei" box="[421,567,1500,1522]" page="135" pageId="14" pageNumber="427" year="2003">
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<bibRefCitation id="4B5A53AAFFECFFFBFE66FA1FE259FA3E" author="Cei" box="[421,516,1500,1522]" pageId="14" pageNumber="427" refString="Cei, J. M. (2003) Specific supraocular scutellation patterns as significant diagnostic characters: a taxonomic inter and intrageneric " finger-print " in Lacertilia. Facena, 19, 129 - 135." type="journal article" year="2003">Cei 2003</bibRefCitation>
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: 135
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67D17DD0FFECFFFBFF54F9DCE3ABF9B0" pageId="14" pageNumber="427" type="materials_examined">
|
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<paragraph id="2F742E5BFFECFFFBFF54F9DCE3ABF9B0" blockId="14.[151,1436,1567,2024]" pageId="14" pageNumber="427">
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<emphasis id="1DBFF249FFECFFFBFF54F9DCE156F9F8" bold="true" box="[151,267,1567,1591]" pageId="14" pageNumber="427">
|
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<typeStatus id="F07090F9FFECFFFBFF54F9DCE15BF9F8" box="[151,262,1567,1591]" pageId="14" pageNumber="427" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Museo Nacional de Historia Natural,
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<collectingCountry id="57DC6ECBFFECFFFBFD00F9E3E378F9F7" box="[707,805,1568,1592]" name="Uruguay" pageId="14" pageNumber="427">Uruguay</collectingCountry>
|
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(
|
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<collectionCode id="49DAB69EFFECFFFBFCF6F9DCE3D3F9F8" LSID="urn:lsid:biocol.org:col:34988" box="[821,910,1567,1591]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="427">MNHN</collectionCode>
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||
) 3489; male, from a hill near route nº 30 and Arroyo Catalán Grande (airline), Departamento de Artigas,
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<collectingCountry id="57DC6ECBFFECFFFBFC86F987E3F6F993" box="[837,939,1604,1628]" name="Uruguay" pageId="14" pageNumber="427">Uruguay</collectingCountry>
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||
; collected
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||
<date id="5B75089BFFECFFFBFBE4F980E48AF994" box="[1063,1239,1603,1627]" pageId="14" pageNumber="427" value="1983-03-30">30 March 1983</date>
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||
by L. Amato, E. Gudynas and G. Skuk; by designation on the original description (
|
||
<figureCitation id="B7F032DEFFECFFFBFCBBF9A4E3B4F9B0" box="[888,1001,1639,1663]" captionStart="FIGURE 13" captionStartId="15.[151,250,1327,1350]" captionTargetBox="[196,1400,215,1287]" captionTargetId="figure@15.[170,1416,194,1306]" captionTargetPageId="15" captionText="FIGURE 13. Dorsal (left) and ventral (right) views of the holotype (MNHN 3489; SVL 114.8 mm) of Tropidurus catalanensis." httpUri="https://zenodo.org/record/247314/files/figure.png" pageId="14" pageNumber="427">Figure 13</figureCitation>
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||
).
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="67D17DD0FFECFFFBFF04F949E52DF8B4" pageId="14" pageNumber="427" type="diagnosis">
|
||
<paragraph id="2F742E5BFFECFFFBFF04F949E52DF8B4" blockId="14.[151,1436,1567,2024]" pageId="14" pageNumber="427">
|
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<emphasis id="1DBFF249FFECFFFBFF04F949E11CF96C" bold="true" box="[199,321,1674,1699]" pageId="14" pageNumber="427">Diagnosis.</emphasis>
|
||
<taxonomicName id="E8CB55D8FFECFFFBFE89F948E207F96C" box="[330,602,1675,1699]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="catalanensis">
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||
<emphasis id="1DBFF249FFECFFFBFE89F948E207F96C" box="[330,602,1675,1699]" italics="true" pageId="14" pageNumber="427">Tropidurus catalanensis</emphasis>
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||
</taxonomicName>
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||
is distinguished from all congeners, except those of the
|
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<taxonomicName id="E8CB55D8FFECFFFBFB22F94FE516F96C" box="[1249,1355,1676,1699]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1DBFF249FFECFFFBFB22F94FE516F96C" box="[1249,1355,1676,1699]" italics="true" pageId="14" pageNumber="427">torquatus</emphasis>
|
||
</taxonomicName>
|
||
group, by lacking a middorsal scale row and by having imbricate and keeled dorsal scales. From those of the
|
||
<taxonomicName id="E8CB55D8FFECFFFBFAF1F973E5C1F908" box="[1330,1436,1712,1735]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1DBFF249FFECFFFBFAF1F973E5C1F908" box="[1330,1436,1712,1735]" italics="true" pageId="14" pageNumber="427">torquatus</emphasis>
|
||
</taxonomicName>
|
||
group, except from
|
||
<taxonomicName id="E8CB55D8FFECFFFBFEB0F917E1A5F924" box="[371,504,1748,1771]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1DBFF249FFECFFFBFEB0F917E1A5F924" box="[371,504,1748,1771]" italics="true" pageId="14" pageNumber="427">T. torquatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E8CB55D8FFECFFFBFDEDF917E2F0F924" box="[558,685,1747,1771]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="imbituba" status="sp. nov.">
|
||
<emphasis id="1DBFF249FFECFFFBFDEDF917E2F0F924" box="[558,685,1747,1771]" italics="true" pageId="14" pageNumber="427">T. imbituba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="068C4F32FFECFFFBFD77F910E350F924" box="[692,781,1747,1771]" pageId="14" pageNumber="427" rank="species">
|
||
<emphasis id="1DBFF249FFECFFFBFD77F910E350F924" bold="true" box="[692,781,1747,1771]" pageId="14" pageNumber="427">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, it can be distinguished by having folds and pockets of the
|
||
<typeStatus id="F07090F9FFECFFFBFF54F93BE09AF8DF" box="[151,199,1784,1808]" pageId="14" pageNumber="427">type</typeStatus>
|
||
“A” (
|
||
<emphasis id="1DBFF249FFECFFFBFECFF93BE117F8C0" box="[268,330,1784,1807]" italics="true" pageId="14" pageNumber="427">sensu</emphasis>
|
||
<bibRefCitation id="4B5A53AAFFECFFFBFE92F934E255F8C0" author="Rodrigues" box="[337,520,1783,1807]" pageId="14" pageNumber="427" refString="Rodrigues, M. T. (1987) Sistematica, ecologia e zoogeografia dos Tropidurus do grupo torquatus ao sul do rio Amazonas (Sauria, Iguanidae). Arquivos de Zoologia, 31 (3), 105 - 230." type="journal article" year="1987">Rodrigues 1987</bibRefCitation>
|
||
). From
|
||
<taxonomicName id="E8CB55D8FFECFFFBFD9CF93BE282F8C0" box="[607,735,1783,1807]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="imbituba" status="sp. nov.">
|
||
<emphasis id="1DBFF249FFECFFFBFD9CF93BE282F8C0" box="[607,735,1783,1807]" italics="true" pageId="14" pageNumber="427">T. imbituba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="068C4F32FFECFFFBFD25F934E362F8C0" box="[742,831,1783,1807]" pageId="14" pageNumber="427" rank="species">
|
||
<emphasis id="1DBFF249FFECFFFBFD25F934E362F8C0" bold="true" box="[742,831,1783,1807]" pageId="14" pageNumber="427">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="E8CB55D8FFECFFFBFCB4F93BE3A0F8C0" box="[887,1021,1784,1807]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1DBFF249FFECFFFBFCB4F93BE3A0F8C0" box="[887,1021,1784,1807]" italics="true" pageId="14" pageNumber="427">T. torquatus</emphasis>
|
||
</taxonomicName>
|
||
, it differs in the yellow coloration of the chest and throat, more intense in adult males. From
|
||
<taxonomicName id="E8CB55D8FFECFFFBFCD4F8DFE3C5F8FC" box="[791,920,1819,1843]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="imbituba" status="sp. nov.">
|
||
<emphasis id="1DBFF249FFECFFFBFCD4F8DFE3C5F8FC" box="[791,920,1819,1843]" italics="true" pageId="14" pageNumber="427">T. imbituba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="068C4F32FFECFFFBFC62F8D8E3A1F8FC" box="[929,1020,1819,1843]" pageId="14" pageNumber="427" rank="species">
|
||
<emphasis id="1DBFF249FFECFFFBFC62F8D8E3A1F8FC" bold="true" box="[929,1020,1819,1843]" pageId="14" pageNumber="427">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
it also differs, with little overlap, in dorsal scale counts, ranging 104–130 (mean 116.9 ± 4.8) in males and 110–140 (mean 126.7 ± 6.8) in females, while in
|
||
<taxonomicName id="E8CB55D8FFECFFFBFF3AF8A7E125F8B4" box="[249,376,1891,1915]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="427" phylum="Chordata" rank="species" species="imbituba">
|
||
<emphasis id="1DBFF249FFECFFFBFF3AF8A7E125F8B4" box="[249,376,1891,1915]" italics="true" pageId="14" pageNumber="427">T. imbituba</emphasis>
|
||
</taxonomicName>
|
||
it ranges 99–113 (mean 104.9 ± 4.6) in males and 102–116 (mean 110.6 ± 3.7) in females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67D17DD0FFECFFE5FF04F845E34BF91E" lastPageId="16" lastPageNumber="429" pageId="14" pageNumber="427" type="description">
|
||
<paragraph id="2F742E5BFFECFFFBFF04F845E463F828" blockId="14.[151,1436,1567,2024]" pageId="14" pageNumber="427">
|
||
<emphasis id="1DBFF249FFECFFFBFF04F845E166F850" bold="true" box="[199,315,1926,1951]" pageId="14" pageNumber="427">Variation.</emphasis>
|
||
Head length 21–25% of SVL; tail length 58–64% of total length; largest male presented
|
||
<quantity id="E83383BEFFECFFFBFAE9F844E5C1F850" box="[1322,1436,1927,1951]" metricMagnitude="-1" metricUnit="m" metricValue="1.279" pageId="14" pageNumber="427" unit="mm" value="127.9">127.9 mm</quantity>
|
||
SVL; Smaller male with opaque and convoluted efferent ducts presented
|
||
<quantity id="E83383BEFFECFFFBFC1BF868E460F80C" box="[984,1085,1963,1987]" metricMagnitude="-2" metricUnit="m" metricValue="7.909999999999999" pageId="14" pageNumber="427" unit="mm" value="79.1">79.1 mm</quantity>
|
||
SVL; largest female presented
|
||
<quantity id="E83383BEFFECFFFBFF54F80CE0A6F828" box="[151,251,1999,2023]" metricMagnitude="-2" metricUnit="m" metricValue="9.879999999999999" pageId="14" pageNumber="427" unit="mm" value="98.8">98.8 mm</quantity>
|
||
SVL and the smaller female with oviductal eggs presented
|
||
<quantity id="E83383BEFFECFFFBFC54F80CE3A6F828" box="[919,1019,1999,2023]" metricMagnitude="-2" metricUnit="m" metricValue="7.27" pageId="14" pageNumber="427" unit="mm" value="72.7">72.7 mm</quantity>
|
||
SVL.
|
||
</paragraph>
|
||
<caption id="7BB47ED3FFEDFFFAFF54FAECE5C0FA8A" box="[151,1437,1327,1350]" httpUri="https://zenodo.org/record/247314/files/figure.png" pageId="15" pageNumber="428" targetBox="[196,1400,215,1287]" targetPageId="15">
|
||
<paragraph id="2F742E5BFFEDFFFAFF54FAECE5C0FA8A" blockId="15.[151,1437,1327,1350]" box="[151,1437,1327,1350]" pageId="15" pageNumber="428">
|
||
<emphasis id="1DBFF249FFEDFFFAFF54FAECE142FA8A" bold="true" box="[151,287,1327,1350]" pageId="15" pageNumber="428">FIGURE 13.</emphasis>
|
||
Dorsal (left) and ventral (right) views of the holotype (MNHN 3489; SVL 114.8 mm) of
|
||
<taxonomicName id="E8CB55D8FFEDFFFAFB61FAECE5C4FA8A" box="[1186,1433,1327,1349]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="428" phylum="Chordata" rank="species" species="catalanensis">Tropidurus catalanensis</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2F742E5BFFEDFFFAFF04FAB2E57CF91D" blockId="15.[151,1437,1393,2034]" pageId="15" pageNumber="428">
|
||
Frontal scales of the head generally convex in adults; 2–4 scales between the nasals; rostral contacting nasals in 37.3% of specimens; 10–15 scales between rostral and interparietal; 2–4 scale rows of enlarged supraoculars, generally three; 6–9 enlarged supraoculars in the first row (examples of the supraocular scuttelation patterns are in
|
||
<bibRefCitation id="4B5A53AAFFEDFFFAFF54FA1EE1C1FA3A" author="Gudynas" box="[151,412,1501,1525]" pageId="15" pageNumber="428" refString="Gudynas, E. & Skuk, G. (1983) A new species of the iguanid lizard genus Tropidurus from temperate South America (Lacertilia: Iguanidae). Centro Educativo Don Orione Contribuciones en Biologia, 10, 1 - 10." type="journal article" year="1983">Gudynas & Skuk 1983</bibRefCitation>
|
||
, p. 4, fig. 1;
|
||
<bibRefCitation id="4B5A53AAFFEDFFFAFDEEFA1EE2CAFA3A" author="Cei" box="[557,663,1501,1525]" pageId="15" pageNumber="428" refString="Cei, J. M. (1993) Reptiles del Noroeste, Nordeste y Este de la Argentina - Herpetofauna de las Selvas Subtropicales, Puna y Pampas. Monografia XIV, Museo Regionale di Scienze Naturali, Torino, Italia, 950 pp." type="book" year="1993">Cei 1993</bibRefCitation>
|
||
, p. 319, fig. 49A; and
|
||
<bibRefCitation id="4B5A53AAFFEDFFFAFC54FA1EE45DFA3A" author="Cei" box="[919,1024,1501,1525]" pageId="15" pageNumber="428" refString="Cei, J. M. (2003) Specific supraocular scutellation patterns as significant diagnostic characters: a taxonomic inter and intrageneric " finger-print " in Lacertilia. Facena, 19, 129 - 135." type="journal article" year="2003">Cei 2003</bibRefCitation>
|
||
, p. 135, fig. 2F); 4–7 supralabials to bellow the center of eye generally 5, followed by 3–7 (generally 4–5) small elongated scales to the commissure of mouth; supralabials contacting subocular in 60% of specimens, generally with 5th or 6th (rarely with both), and less frequently with 4th or 7th; 6–10 infralabials, generally 8–9; preauricular fringe with 4–9 spines, generally 7; 9–15 scales between the subocular and the preauricular fringe, generally 11–12; mental triangular or polygonal, followed by 2–5 (generally 3–4) angulated postmentals; first postmentals contacting one another in 13.8% of specimens; 2– 4 canthals, generally 3; 46–65 gular scales (mean 56.6 ± 3.8, n=216) (
|
||
<quantity id="E83383BEFFEDFFFAFC66F979E454F91D" box="[933,1033,1722,1746]" metricMagnitude="0" metricUnit="m" metricValue="1.4097" metricValueMax="1.6256" metricValueMin="1.1938" pageId="15" pageNumber="428" unit="in" value="55.5" valueMax="64.0" valueMin="47.0">47–64 in</quantity>
|
||
<bibRefCitation id="4B5A53AAFFEDFFFAFBD3F979E549F91D" author="Gudynas" box="[1040,1300,1722,1746]" pageId="15" pageNumber="428" refString="Gudynas, E. & Skuk, G. (1983) A new species of the iguanid lizard genus Tropidurus from temperate South America (Lacertilia: Iguanidae). Centro Educativo Don Orione Contribuciones en Biologia, 10, 1 - 10." type="journal article" year="1983">Gudynas & Skuk 1983</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2F742E5BFFEDFFFAFF04F91CE347F8AD" blockId="15.[151,1437,1393,2034]" pageId="15" pageNumber="428">
|
||
Two mite pockets under the oblique neck fold, the upper pocket is well developed and the lower one is poorly developed; antehumeral fold present; 2–3 axillary pockets, generally two; inguinal granular pocket present; 24–34 subdigital lamellae on the fourth toe (mean 29 ± 1.9, n=216) (
|
||
<quantity id="E83383BEFFEDFFFAFC80F8E5E3FAF8F1" box="[835,935,1830,1854]" metricMagnitude="-1" metricUnit="m" metricValue="7.366" metricValueMax="8.128" metricValueMin="6.604" pageId="15" pageNumber="428" unit="in" value="29.0" valueMax="32.0" valueMin="26.0">26–32 in</quantity>
|
||
<bibRefCitation id="4B5A53AAFFEDFFFAFC6FF8E5E4F2F8F1" author="Gudynas" box="[940,1199,1830,1854]" pageId="15" pageNumber="428" refString="Gudynas, E. & Skuk, G. (1983) A new species of the iguanid lizard genus Tropidurus from temperate South America (Lacertilia: Iguanidae). Centro Educativo Don Orione Contribuciones en Biologia, 10, 1 - 10." type="journal article" year="1983">Gudynas & Skuk 1983</bibRefCitation>
|
||
); 18–25 scales on the dorsum of tibia (mean 21.6 ± 1.3, n=217), usually 21–23.
|
||
</paragraph>
|
||
<paragraph id="2F742E5BFFEDFFE5FF04F8ACE504FF38" blockId="15.[151,1437,1393,2034]" lastBlockId="16.[151,1436,151,248]" lastPageId="16" lastPageNumber="429" pageId="15" pageNumber="428">
|
||
In the original description,
|
||
<bibRefCitation id="4B5A53AAFFEDFFFAFE3FF8ADE379F849" author="Gudynas" box="[508,804,1902,1926]" pageId="15" pageNumber="428" refString="Gudynas, E. & Skuk, G. (1983) A new species of the iguanid lizard genus Tropidurus from temperate South America (Lacertilia: Iguanidae). Centro Educativo Don Orione Contribuciones en Biologia, 10, 1 - 10." type="journal article" year="1983">Gudynas and Skuk (1983)</bibRefCitation>
|
||
stated that there were no significant sexual differences, thus, presenting all meristic and morphometric variables grouped for both sexes. Our data do not corroborate this statement in relation to dorsal scales and SAB. Males present 104–130 imbricate, mucronate and keeled dorsals (mean 116.9 ± 4.8, n=103), while females present 110–140 (mean 126.7 ± 6.8, n=110). Specimens from lowlands of southwestern Rio Grande do Sul and the vicinities of the mouth of the
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FC14FF5BE462FF7F" box="[983,1087,152,176]" name="Paraguay" pageId="16" pageNumber="429">Paraguay</collectingCountry>
|
||
River (northern
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FB3FFF54E531FF60" box="[1276,1388,151,175]" name="Argentina" pageId="16" pageNumber="429">Argentina</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FEC2FF7FE136FF1B" box="[257,363,188,212]" name="Paraguay" pageId="16" pageNumber="429">Paraguay</collectingCountry>
|
||
) present the lowest means (113.2 ±
|
||
<quantity id="E83383BEFFF2FFE5FCD0FF7FE308FF1B" box="[787,853,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="1.1683999999999999" pageId="16" pageNumber="429" unit="in" value="4.6">4.6 in</quantity>
|
||
males; 122.4 ±
|
||
<quantity id="E83383BEFFF2FFE5FBD7FF7FE40AFF1B" box="[1044,1111,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="1.1938" pageId="16" pageNumber="429" unit="in" value="4.7">4.7 in</quantity>
|
||
females) while those of the valleys of the upper
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FEB8FF23E180FF37" box="[379,477,224,248]" name="Uruguay" pageId="16" pageNumber="429">Uruguay</collectingCountry>
|
||
Basin present the highest means (119.8 ±
|
||
<quantity id="E83383BEFFF2FFE5FC74FF1CE3AAFF38" box="[951,1015,223,247]" metricMagnitude="-1" metricUnit="m" metricValue="1.0921999999999998" pageId="16" pageNumber="429" unit="in" value="4.3">4.3 in</quantity>
|
||
males; 130.4 ±
|
||
<quantity id="E83383BEFFF2FFE5FB69FF1CE4B7FF38" box="[1194,1258,223,247]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" pageId="16" pageNumber="429" unit="in" value="5.5">5.5 in</quantity>
|
||
females).
|
||
</paragraph>
|
||
<caption id="7BB47ED3FFF2FFE5FF54FBD8E2B7FB9F" httpUri="https://zenodo.org/record/247315/files/figure.png" pageId="16" pageNumber="429" targetBox="[151,1435,286,1030]" targetPageId="16">
|
||
<paragraph id="2F742E5BFFF2FFE5FF54FBD8E2B7FB9F" blockId="16.[151,1436,1051,1104]" pageId="16" pageNumber="429">
|
||
<emphasis id="1DBFF249FFF2FFE5FF54FBD8E17EFBFE" bold="true" box="[151,291,1051,1074]" pageId="16" pageNumber="429">FIGURE 14.</emphasis>
|
||
<taxonomicName id="E8CB55D8FFF2FFE5FEEFFBDFE278FBFD" box="[300,549,1052,1074]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="429" phylum="Chordata" rank="species" species="catalanensis">
|
||
<emphasis id="1DBFF249FFF2FFE5FEEFFBDFE278FBFD" box="[300,549,1052,1074]" italics="true" pageId="16" pageNumber="429">Tropidurus catalanensis</emphasis>
|
||
</taxonomicName>
|
||
from Iguaçu National Park, Foz do Iguaçu, Paraná, Brazil (not collected). Note the yellow coloration of chin, including infra and supralabials.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2F742E5BFFF2FFE5FF04FBBAE19AFA7E" blockId="16.[151,1436,1145,1998]" pageId="16" pageNumber="429">
|
||
<bibRefCitation id="4B5A53AAFFF2FFE5FF04FBBAE1B6FB5E" author="Gudynas" box="[199,491,1145,1169]" pageId="16" pageNumber="429" refString="Gudynas, E. & Skuk, G. (1983) A new species of the iguanid lizard genus Tropidurus from temperate South America (Lacertilia: Iguanidae). Centro Educativo Don Orione Contribuciones en Biologia, 10, 1 - 10." type="journal article" year="1983">Gudynas and Skuk (1983)</bibRefCitation>
|
||
presented vertebral and paravertebral scale counts, which ranged, respectively, 107– 131 and 110–142. Since lizards of the
|
||
<taxonomicName id="E8CB55D8FFF2FFE5FDA3FB5DE296FB7A" box="[608,715,1182,1205]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="429" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1DBFF249FFF2FFE5FDA3FB5DE296FB7A" box="[608,715,1182,1205]" italics="true" pageId="16" pageNumber="429">torquatus</emphasis>
|
||
</taxonomicName>
|
||
group lack a vertebral crest or enlarged vertebral scale rows, distinguishing vertebrals and paravertebrals is difficult, but these values are similar to what we found in our dorsal counts. SAB ranged 91–116 (mean 103.4 ± 5.4, n=97) in males, and 103–131 (mean 115.9 ± 6.6, n=105) in females (
|
||
<quantity id="E83383BEFFF2FFE5FF5CFACAE14FFAEE" box="[159,274,1289,1313]" metricMagnitude="0" metricUnit="m" metricValue="2.667" metricValueMax="3.0734" metricValueMin="2.2606" pageId="16" pageNumber="429" unit="in" value="105.0" valueMax="121.0" valueMin="89.0">89–121 in</quantity>
|
||
the original description, for males and females grouped).
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FC67FAC9E39EFAED" box="[932,963,1290,1314]" name="American Samoa" pageId="16" pageNumber="429">As</collectingCountry>
|
||
for dorsals, specimens from the lowlands of southwestern Rio Grande do Sul, northeastern
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FD00FAEEE36FFA8A" box="[707,818,1325,1349]" name="Argentina" pageId="16" pageNumber="429">Argentina</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FC12FAEDE464FA89" box="[977,1081,1326,1350]" name="Paraguay" pageId="16" pageNumber="429">Paraguay</collectingCountry>
|
||
present the lowest means (99.5 ±
|
||
<quantity id="E83383BEFFF2FFE5FF6DFA92E0ADFAA6" box="[174,240,1361,1385]" metricMagnitude="-2" metricUnit="m" metricValue="8.128" pageId="16" pageNumber="429" unit="in" value="3.2">3.2 in</quantity>
|
||
males, 110.5 ±
|
||
<quantity id="E83383BEFFF2FFE5FE65FA92E1BAFAA6" box="[422,487,1361,1385]" metricMagnitude="-2" metricUnit="m" metricValue="9.398" pageId="16" pageNumber="429" unit="in" value="3.7">3.7 in</quantity>
|
||
females), and those of the upper
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FCA6FA91E39AFAA5" box="[869,967,1362,1386]" name="Uruguay" pageId="16" pageNumber="429">Uruguay</collectingCountry>
|
||
Basin, the highest means (106.5 ±
|
||
<quantity id="E83383BEFFF2FFE5FA98FA92E5C1FAA6" box="[1371,1436,1361,1385]" metricMagnitude="-1" metricUnit="m" metricValue="1.143" pageId="16" pageNumber="429" unit="in" value="4.5">4.5 in</quantity>
|
||
males, 119.4 ±
|
||
<quantity id="E83383BEFFF2FFE5FE84FAB6E1D4FA42" box="[327,393,1397,1421]" metricMagnitude="-1" metricUnit="m" metricValue="1.4223999999999999" pageId="16" pageNumber="429" unit="in" value="5.6">5.6 in</quantity>
|
||
females). Ventrals 72–103 (mean 86.2 ± 6.1, n=214; the original description didn’t present variation of this character).
|
||
</paragraph>
|
||
<paragraph id="2F742E5BFFF2FFE5FF04FA7EE34BF91E" blockId="16.[151,1436,1145,1998]" pageId="16" pageNumber="429">
|
||
<emphasis id="1DBFF249FFF2FFE5FF04FA7EE103FA19" bold="true" box="[199,350,1469,1494]" pageId="16" pageNumber="429">Color in life.</emphasis>
|
||
Dorsal color is brown, gray or greenish, with an irregular pattern of black paravertebral patches and many yellow or cream spots in the dorsum of the trunk and limbs; yellow or cream spots are more numerous in males; black antehumeral band occupying 4–10 scale rows, separated from the opposite one by 3–35 scales (mean 15.7 ± 6.3); venter white or cream; adult males with femoral and preanal patches of black pigmented glandular scales; presence of midventral patches variable but rarely lacking in specimens larger than
|
||
<quantity id="E83383BEFFF2FFE5FB52F98EE4B3F9AA" box="[1169,1262,1613,1637]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="429" unit="mm" value="100.0">100 mm</quantity>
|
||
SVL; temporal region usually bluish; adults with black gular patches; chin yellow with dark reticulations enclosing yellow blotches; yellow pigment reaches the chest and sometimes the infra and supralabials; yellow pigment lacking in juveniles and usually less evident, or lacking, in females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="67D17DD0FFF2FFE5FF04F91EE3F9F802" pageId="16" pageNumber="429" type="distribution">
|
||
<paragraph id="2F742E5BFFF2FFE5FF04F91EE3F9F802" blockId="16.[151,1436,1145,1998]" pageId="16" pageNumber="429">
|
||
<emphasis id="1DBFF249FFF2FFE5FF04F91EE13DF939" bold="true" box="[199,352,1757,1782]" pageId="16" pageNumber="429">Distribution.</emphasis>
|
||
Northwestern
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FDDFF91DE222F939" box="[540,639,1758,1782]" name="Uruguay" pageId="16" pageNumber="429">Uruguay</collectingCountry>
|
||
, northeastern
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FCF3F91EE3FFF93A" box="[816,930,1757,1781]" name="Argentina" pageId="16" pageNumber="429">Argentina</collectingCountry>
|
||
, southern
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FBE8F91DE4CEF939" box="[1067,1171,1758,1782]" name="Paraguay" pageId="16" pageNumber="429">Paraguay</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FA92F91EE5C5F93A" box="[1361,1432,1757,1781]" name="Brazil" pageId="16" pageNumber="429">Brazil</collectingCountry>
|
||
, reaching southwestern São Paulo and southeastern Mato Grosso do Sul extremes. In northwestern
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FACBF8C1E537F8D5" box="[1288,1386,1794,1818]" name="Uruguay" pageId="16" pageNumber="429">Uruguay</collectingCountry>
|
||
and southwestern Rio Grande do Sul, the species is found on isolated rocky hills of low altitude regionally known as “cerros”. These hills are sparsely distributed along wide areas of
|
||
<emphasis id="1DBFF249FFF2FFE5FCBDF889E389F8AE" box="[894,980,1866,1889]" italics="true" pageId="16" pageNumber="429">campos</emphasis>
|
||
(grassland) of the Pampa Biome. In the remaining areas, species distribution follows the course of the main tributaries of the Paraná and
|
||
<collectingCountry id="57DC6ECBFFF2FFE5FB2BF8ADE517F849" box="[1256,1354,1902,1926]" name="Uruguay" pageId="16" pageNumber="429">Uruguay</collectingCountry>
|
||
Rivers (La Plata Basin). Along the valleys and canyons of these rivers, the species occurs mainly on isolated populations in rocky outcrops among areas originally covered by seasonal forests.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7BB47ED3FFF3FFE4FF54FBD9E504FBFF" box="[151,1369,1050,1073]" httpUri="https://zenodo.org/record/247316/files/figure.png" pageId="17" pageNumber="430" targetBox="[151,1435,194,1028]" targetPageId="17">
|
||
<paragraph id="2F742E5BFFF3FFE4FF54FBD9E504FBFF" blockId="17.[151,1369,1050,1073]" box="[151,1369,1050,1073]" pageId="17" pageNumber="430">
|
||
<emphasis id="1DBFF249FFF3FFE4FF54FBD9E142FBFF" bold="true" box="[151,287,1050,1073]" pageId="17" pageNumber="430">FIGURE 15.</emphasis>
|
||
<taxonomicName id="E8CB55D8FFF3FFE4FEE6FBD9E247FBFF" box="[293,538,1050,1072]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="430" phylum="Chordata" rank="species" species="catalanensis">
|
||
<emphasis id="1DBFF249FFF3FFE4FEE6FBD9E247FBFF" box="[293,538,1050,1072]" italics="true" pageId="17" pageNumber="430">Tropidurus catalanensis</emphasis>
|
||
</taxonomicName>
|
||
from Campos Novos, Santa Catarina, near the Uruguay riverbank (not collected).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7BB47ED3FFF3FFE4FF54F874E576F802" box="[151,1323,1975,1998]" httpUri="https://zenodo.org/record/247317/files/figure.png" pageId="17" pageNumber="430" targetBox="[151,1435,1124,1952]" targetPageId="17">
|
||
<paragraph id="2F742E5BFFF3FFE4FF54F874E576F802" blockId="17.[151,1323,1975,1998]" box="[151,1323,1975,1998]" pageId="17" pageNumber="430">
|
||
<emphasis id="1DBFF249FFF3FFE4FF54F874E142F803" bold="true" box="[151,287,1975,1998]" pageId="17" pageNumber="430">FIGURE 16.</emphasis>
|
||
<taxonomicName id="E8CB55D8FFF3FFE4FEE6F874E247F802" box="[293,538,1975,1997]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="17" pageNumber="430" phylum="Chordata" rank="species" species="catalanensis">
|
||
<emphasis id="1DBFF249FFF3FFE4FEE6F874E247F802" box="[293,538,1975,1997]" italics="true" pageId="17" pageNumber="430">Tropidurus catalanensis</emphasis>
|
||
</taxonomicName>
|
||
from São Francisco de Assis, Rio Grande do Sul. Adult male (not collected).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7BB47ED3FFF0FFE7FF54FC21E2A8FBD8" httpUri="https://zenodo.org/record/247318/files/figure.png" pageId="18" pageNumber="431" targetBox="[151,1435,194,900]" targetPageId="18">
|
||
<paragraph id="2F742E5BFFF0FFE7FF54FC21E2A8FBD8" blockId="18.[151,1436,994,1047]" pageId="18" pageNumber="431">
|
||
<emphasis id="1DBFF249FFF0FFE7FF54FC21E142FC37" bold="true" box="[151,287,994,1017]" pageId="18" pageNumber="431">FIGURE 17.</emphasis>
|
||
Lateral view of head of
|
||
<taxonomicName id="E8CB55D8FFF0FFE7FDD7FC20E354FC36" box="[532,777,995,1017]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="431" phylum="Chordata" rank="species" species="catalanensis">
|
||
<emphasis id="1DBFF249FFF0FFE7FDD7FC20E354FC36" box="[532,777,995,1017]" italics="true" pageId="18" pageNumber="431">Tropidurus catalanensis</emphasis>
|
||
</taxonomicName>
|
||
(UFRGS 5324) from São Francisco de Assis, Rio Grande do Sul, Brazil. Freshly killed subadult male (head length 21.4 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="67D17DD0FFF0FFE6FF04FB86E385FE00" lastPageId="19" lastPageNumber="432" pageId="18" pageNumber="431" type="discussion">
|
||
<paragraph id="2F742E5BFFF0FFE7FF04FB86E3F8FAB3" blockId="18.[151,1436,1092,2017]" pageId="18" pageNumber="431">
|
||
<emphasis id="1DBFF249FFF0FFE7FF04FB86E166FB92" bold="true" box="[199,315,1093,1117]" pageId="18" pageNumber="431">Remarks.</emphasis>
|
||
The distribution of
|
||
<taxonomicName id="E8CB55D8FFF0FFE7FDE7FB86E292FB93" box="[548,719,1092,1116]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="431" phylum="Chordata" rank="species" species="catalanensis">
|
||
<emphasis id="1DBFF249FFF0FFE7FDE7FB86E292FB93" box="[548,719,1092,1116]" italics="true" pageId="18" pageNumber="431">T. catalanensis</emphasis>
|
||
</taxonomicName>
|
||
in southern
|
||
<collectingCountry id="57DC6ECBFFF0FFE7FCA2FB87E3F8FB93" box="[865,933,1092,1116]" name="Brazil" pageId="18" pageNumber="431">Brazil</collectingCountry>
|
||
shows an interesting pattern, in association with river valleys and very restricted and localized populations, sometimes a hundred kilometers apart from other populations. The lack of gene flow between these populations is evidenced by some differences in meristic characters. For example, specimens from upstream of the Iguaçu river present, with high frequency, contact of the first post-mental with the opposite one (and, in this case, there is no contact of the gular scales with the mental scale). The frequency of this condition decreases downstream toward its mouth, and is rare in all other analyzed populations. Other variable condition is the contact between the nasals and the rostral scale, which is very frequent in some populations from southwestern Rio Grande do Sul (e.g. from São Francisco de Assis) and in those from Foz do Iguaçu (state of Paraná), but not frequent in other populations.
|
||
</paragraph>
|
||
<paragraph id="2F742E5BFFF0FFE6FF04FA4BE1D9FED3" blockId="18.[151,1436,1092,2017]" lastBlockId="19.[151,1436,151,463]" lastPageId="19" lastPageNumber="432" pageId="18" pageNumber="431">
|
||
Populations of the river valleys inhabit rocky outcrops with abundant crevices. This punctual distribution pattern, at least in the low meanders of the Southern Brazilian Plateau, seems to indicate that the current climatic condition is unfavorable for the species (see Behling
|
||
<emphasis id="1DBFF249FFF0FFE7FD2FFA12E378FA27" box="[748,805,1488,1512]" italics="true" pageId="18" pageNumber="431">et al.</emphasis>
|
||
2009 for evidences of the climate dynamics of southern
|
||
<collectingCountry id="57DC6ECBFFF0FFE7FF54FA37E086F9C3" box="[151,219,1524,1548]" name="Brazil" pageId="18" pageNumber="431">Brazil</collectingCountry>
|
||
during the Holocene). These rocky outcrops represent the only natural open habitats among these river valleys, originally covered with seasonal forests. In addition, these river valleys and canyons retain milder climatic conditions in relation to the higher portions of the plateau. Despite the intense deforestation of the seasonal forests in southern
|
||
<collectingCountry id="57DC6ECBFFF0FFE7FED9F9A3E13CF9B7" box="[282,353,1632,1656]" name="Brazil" pageId="18" pageNumber="431">Brazil</collectingCountry>
|
||
, these populations have not expanded their occupation beyond the limits of the outcrops, pending in favor of the argument that the current climatic condition is unfavorable for the species expansion, at least in the Southern Brazilian Plateau. This situation is quite contrasting with the alleged opportunism of these lizards that, according to
|
||
<bibRefCitation id="4B5A53AAFFF0FFE7FEFCF90FE24CF92B" author="Rodrigues" box="[319,529,1740,1764]" pageId="18" pageNumber="431" refString="Rodrigues, M. T. (1987) Sistematica, ecologia e zoogeografia dos Tropidurus do grupo torquatus ao sul do rio Amazonas (Sauria, Iguanidae). Arquivos de Zoologia, 31 (3), 105 - 230." type="journal article" year="1987">Rodrigues (1987)</bibRefCitation>
|
||
, being good colonizers, would be expanding its distribution favored by deforestations (at least for the
|
||
<taxonomicName id="E8CB55D8FFF0FFE7FE2EF932E20AF8C7" box="[493,599,1777,1800]" class="Reptilia" family="Tropiduridae" genus="Tropidurus" kingdom="Animalia" order="Squamata" pageId="18" pageNumber="431" phylum="Chordata" rank="species" species="torquatus">
|
||
<emphasis id="1DBFF249FFF0FFE7FE2EF932E20AF8C7" box="[493,599,1777,1800]" italics="true" pageId="18" pageNumber="431">torquatus</emphasis>
|
||
</taxonomicName>
|
||
populations in the state of São Paulo). In fact, there is evidence to believe that the opposite is occurring in southern
|
||
<collectingCountry id="57DC6ECBFFF0FFE7FD58F8D7E2BEF8E3" box="[667,739,1812,1836]" name="Brazil" pageId="18" pageNumber="431">Brazil</collectingCountry>
|
||
. Apparently, some populations have disappeared after the formation of reservoirs for hydroelectric dams on the Iguaçu River, in the state of Paraná (R.S. Bérnils, personal communication) and in the upper
|
||
<collectingCountry id="57DC6ECBFFF0FFE7FDDBF89EE227F8BA" box="[536,634,1885,1909]" name="Uruguay" pageId="18" pageNumber="431">Uruguay</collectingCountry>
|
||
Basin in the border of the states of Santa Catarina and Rio Grande do Sul (M. Borges-Martins, personal observation). Viability of populations of these saxicolous lizards could be conditioned to the maintenance of adequate habitats. In the surroundings of the Quebra-Queixo hydroelectric dam, on the Chapecó river (a large tributary of the
|
||
<collectingCountry id="57DC6ECBFFF0FFE7FD6CF80AE34CF82E" box="[687,785,1993,2017]" name="Uruguay" pageId="18" pageNumber="431">Uruguay</collectingCountry>
|
||
river), where rocks and crevices are very abundant, the species is also still abundant. For populations from southwestern Rio Grande do Sul, expansion of
|
||
<taxonomicName id="E8CB55D8FFF1FFE6FB29FF5BE53BFF7F" box="[1258,1382,152,176]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="432" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="1DBFF249FFF1FFE6FB29FF5BE53BFF7F" box="[1258,1382,152,176]" italics="true" pageId="19" pageNumber="432">Eucalyptus</emphasis>
|
||
</taxonomicName>
|
||
spp. plantations in the vicinities of the rocky outcrops could result in shading and affect the viability of some of these populations (personal observations; see
|
||
<bibRefCitation id="4B5A53AAFFF1FFE6FDA4FF1CE373FF38" author="Pillar" box="[615,814,223,248]" pageId="19" pageNumber="432" refString="Pillar, V. P., Muller, S. C., Castilhos, Z. M. S. & Jacques, A. V. A. (Eds.), (2009) Campos Sulinos - conservacao e uso sustentavel da biodiversidade. Ministerio do Meio Ambiente, Brasilia, 403 pp." type="book" year="2009">
|
||
Pillar
|
||
<emphasis id="1DBFF249FFF1FFE6FD6CFF22E2B1FF37" box="[687,748,224,248]" italics="true" pageId="19" pageNumber="432">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
for a discussion on the conservation of the southern Brazilian “
|
||
<emphasis id="1DBFF249FFF1FFE6FED2FEC6E13AFED3" box="[273,359,261,284]" italics="true" pageId="19" pageNumber="432">campos</emphasis>
|
||
”).
|
||
</paragraph>
|
||
<paragraph id="2F742E5BFFF1FFE6FF04FEEBE385FE00" blockId="19.[151,1436,151,463]" pageId="19" pageNumber="432">
|
||
It is worth considering that the mainly saxicolous habit of the populations from Southern
|
||
<collectingCountry id="57DC6ECBFFF1FFE6FB68FEE4E4AFFEF0" box="[1195,1266,295,319]" name="Brazil" pageId="19" pageNumber="432">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry id="57DC6ECBFFF1FFE6FB3EFEE4E531FEF0" box="[1277,1388,295,319]" name="Argentina" pageId="19" pageNumber="432">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="57DC6ECBFFF1FFE6FF54FE8EE0A4FEAA" box="[151,249,333,357]" name="Uruguay" pageId="19" pageNumber="432">Uruguay</collectingCountry>
|
||
(
|
||
<bibRefCitation id="4B5A53AAFFF1FFE6FECAFE8FE24CFEAB" author="Gudynas" box="[265,529,332,356]" pageId="19" pageNumber="432" refString="Gudynas, E. & Skuk, G. (1983) A new species of the iguanid lizard genus Tropidurus from temperate South America (Lacertilia: Iguanidae). Centro Educativo Don Orione Contribuciones en Biologia, 10, 1 - 10." type="journal article" year="1983">Gudynas & Skuk 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="4B5A53AAFFF1FFE6FDDDFE8FE2D5FEAB" author="Cei" box="[542,648,332,356]" pageId="19" pageNumber="432" refString="Cei, J. M. (1993) Reptiles del Noroeste, Nordeste y Este de la Argentina - Herpetofauna de las Selvas Subtropicales, Puna y Pampas. Monografia XIV, Museo Regionale di Scienze Naturali, Torino, Italia, 950 pp." type="book" year="1993">Cei 1993</bibRefCitation>
|
||
; this study) differs in what
|
||
<bibRefCitation id="4B5A53AAFFF1FFE6FC01FE8FE461FEAB" author="Cei" box="[962,1084,332,356]" pageId="19" pageNumber="432" refString="Cei, J. M. (1993) Reptiles del Noroeste, Nordeste y Este de la Argentina - Herpetofauna de las Selvas Subtropicales, Puna y Pampas. Monografia XIV, Museo Regionale di Scienze Naturali, Torino, Italia, 950 pp." type="book" year="1993">Cei (1993)</bibRefCitation>
|
||
observed for populations from Corrientes (
|
||
<collectingCountry id="57DC6ECBFFF1FFE6FEE1FEACE1C5FE48" box="[290,408,367,391]" name="Argentina" pageId="19" pageNumber="432">Argentina</collectingCountry>
|
||
), allegedly more arboreal. Still, the Atlas of
|
||
<bibRefCitation id="4B5A53AAFFF1FFE6FC05FEACE49BFE48" box="[966,1222,367,392]" pageId="19" pageNumber="432" refString="Alvarez, B. B., Aguirre, R. H., Cespedez, J. A., Hernando, A. B. & Tedesco, M. E. (2002) Atlas de Anfibios y Reptiles de las provincias de Corrientes, Chaco y Formosa, Argentina: Anuros, Cecilidos, Saurios, Anfisbenidos y Serpientes. Universidad Nacional del Nordeste, Corrientes, 160 pp." type="book">
|
||
Alvarez
|
||
<emphasis id="1DBFF249FFF1FFE6FBEEFEB2E430FE47" box="[1069,1133,368,392]" italics="true" pageId="19" pageNumber="432">et al.</emphasis>
|
||
(2002)
|
||
</bibRefCitation>
|
||
shows that these populations are also distributed exclusively along the Parana River and in the vicinities of the
|
||
<collectingCountry id="57DC6ECBFFF1FFE6FB28FE56E50EFE62" box="[1259,1363,405,429]" name="Paraguay" pageId="19" pageNumber="432">Paraguay</collectingCountry>
|
||
River mouth, both rivers of the La Plata Basin, as are the remaining populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |