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<document ID-DOI="http://doi.org/10.5281/zenodo.4760453" ID-GBIF-Dataset="d92d447b-3342-41ca-bdcd-1175c037416f" ID-GBIF-Taxon="182073560" ID-ISSN="1854-0392" ID-Zenodo-Dep="4760453" ID-ZooBank="7F93F639-6206-40B9-A63B-071DA459E8ED" approvalRequired="1" approvalRequired_for_taxonomicNames="1" checkinTime="1620965460301" checkinUser="felipe" docAuthor="Mason, Denise &amp; Stark, Bill P." docDate="2015" docId="94578784B629FFA0264DFA90FBBAAC30" docLanguage="en" docName="Illiesia.11.04.29-40.pdf" docOrigin="Illiesia 11 (4)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Phanoperla wedda Zwick 1982" docType="treatment" docVersion="6" lastPageNumber="37" masterDocId="686EFFFCB621FFA82719FF84FFEAAB43" masterDocTitle="Notes On The Genus Phanoperla Banks From Sri Lanka And India (Plecoptera: Perlidae)" masterLastPageNumber="40" masterPageNumber="29" pageId="8" pageNumber="37" updateTime="1632020977812" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>Notes On The Genus Phanoperla Banks From Sri Lanka And India (Plecoptera: Perlidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mason, Denise</mods:namePart>
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi 39058, U. S. A. E-mail: dmason @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">dmason@mc.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Stark, Bill P.</mods:namePart>
<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi 39058, U. S. A. E-mail: stark @ mc. edu</mods:affiliation>
<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
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<mods:title>Illiesia</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>11</mods:number>
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<mods:number>4</mods:number>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760908</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4760453" ID-GBIF-Taxon="182073560" ID-Zenodo-Dep="4760453" LSID="urn:lsid:plazi:treatment:94578784B629FFA0264DFA90FBBAAC30" httpUri="http://treatment.plazi.org/id/94578784B629FFA0264DFA90FBBAAC30" lastPageNumber="37" pageId="8" pageNumber="37">
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<heading bold="true" box="[340,646,1300,1324]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="37" reason="2">
<taxonomicName authority="Zwick" authorityName="Zwick" authorityYear="1982" box="[340,646,1300,1324]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="wedda">
<emphasis bold="true" box="[340,646,1300,1324]" pageId="8" pageNumber="37">
<emphasis bold="true" box="[340,562,1300,1324]" italics="true" pageId="8" pageNumber="37">Phanoperla wedda</emphasis>
Zwick
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[422,565,1336,1360]" pageId="8" pageNumber="37" type="description">
<paragraph blockId="8.[422,565,1336,1360]" box="[422,565,1336,1360]" pageId="8" pageNumber="37">
(
<figureCitation box="[430,557,1336,1360]" captionStart="Figs" captionStartId="9.[189,241,1810,1834]" captionTargetBox="[232,1404,290,1728]" captionTargetId="figure-95@9.[232,1404,290,1728]" captionTargetPageId="9" captionText="Figs. 22-27. Phanoperla wedda egg. 22. Lateral aspect. 23. Collar end (s = stria). 24. Anchor and collar (at = anchor tentacle). 25. Lid and micropylar zone. 26. Anchor and collar, apical aspect. 27. Micropyles (m = micropyle)." figureDoi="http://doi.org/10.5281/zenodo.4760920" httpUri="https://zenodo.org/record/4760920/files/figure.png" pageId="8" pageNumber="37">Figs. 22-27</figureCitation>
)
</paragraph>
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<subSubSection box="[227,637,1400,1422]" pageId="8" pageNumber="37" type="reference_group">
<paragraph blockId="8.[227,797,1400,1487]" box="[227,637,1400,1422]" pageId="8" pageNumber="37">
<treatmentCitation author="Zwick, P." box="[227,634,1400,1422]" page="118" pageId="8" pageNumber="37" year="1982">
<taxonomicName authority="Zwick 1982 a: 118" authorityName="Zwick" authorityPageNumber="118" authorityYear="1982" box="[227,634,1400,1422]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="wedda">
<emphasis box="[227,418,1400,1421]" italics="true" pageId="8" pageNumber="37">Phanoperla wedda</emphasis>
<bibRefCitation author="Zwick, P." box="[440,634,1400,1422]" pageId="8" pageNumber="37" pagination="87 - 126" refId="ref5664" refString="Zwick, P. 1982 a. A revision of the Oriental stonefly genus Phanoperla (Plecoptera: Perlidae). Systematic Entomology, 7: 87 - 126." type="journal article" year="1982">Zwick 1982a:118</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="37" type="materials_examined">
<paragraph blockId="8.[227,797,1400,1487]" pageId="8" pageNumber="37">
<materialsCitation ID-GBIF-Occurrence="3111499321" country="Sri Lanka" county="Kandy District" location="Hasalaka Oya" municipality="Hasalaka Oya" pageId="8" pageNumber="37" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[658,758,1400,1422]" pageId="8" pageNumber="37">Holotype</typeStatus>
<specimenCount box="[779,797,1400,1422]" pageId="8" pageNumber="37" type="male"></specimenCount>
(United States National Museum),
<collectingMunicipality box="[625,793,1432,1454]" pageId="8" pageNumber="37">Hasalaka Oya</collectingMunicipality>
,
<collectingCounty box="[227,385,1465,1487]" pageId="8" pageNumber="37">Kandy District</collectingCounty>
,
<collectingCountry box="[395,471,1465,1487]" name="Sri Lanka" pageId="8" pageNumber="37">Ceylon</collectingCountry>
[Sri Lanka]
</materialsCitation>
</paragraph>
</subSubSection>
</treatmentCitationGroup>
<subSubSection pageId="8" pageNumber="37" type="materials_examined">
<paragraph blockId="8.[189,798,1526,1911]" pageId="8" pageNumber="37">
<emphasis bold="true" box="[189,428,1526,1550]" pageId="8" pageNumber="37">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111499314" collectingDate="1978-06-28" collectingDateMax="1978-07-04" collectingDateMin="1978-06-28" collectionCode="V" country="Sri Lanka" location="Matale District" municipality="Kibissa" pageId="8" pageNumber="37" specimenCount="1">
<emphasis bold="true" box="[439,602,1526,1550]" pageId="8" pageNumber="37">
<collectingCountry box="[439,596,1526,1550]" name="Sri Lanka" pageId="8" pageNumber="37">SRI LANKA</collectingCountry>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:94578784B629FFA0264DFA90FBBAAC30:19216049B629FFA0257DFA73FCF3AD4C" box="[612,793,1527,1551]" country="Sri Lanka" municipality="Kibissa" name="Matale District" pageId="8" pageNumber="37">Matale District</location>
,
<collectingMunicipality box="[189,277,1563,1587]" pageId="8" pageNumber="37">Kibissa</collectingMunicipality>
, 0.5 miles
<locationDeviation box="[413,667,1563,1587]" location="Sigiriya Jungle" pageId="8" pageNumber="37">W of Sigiriya Jungle</locationDeviation>
,
<date pageId="8" pageNumber="37" value="1978-06-28" valueMax="1978-07-04" valueMin="1978-06-28">
<collectingDate pageId="8" pageNumber="37" value="1978-06-28" valueMax="1978-07-04" valueMin="1978-06-28">28 June-4 July 1978</collectingDate>
</date>
, K.
<collectionCode box="[370,392,1599,1622]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="37" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111499318" collectingDate="1978-04-09" collectingDateMax="1978-04-10" collectingDateMin="1978-04-09" collectionCode="USNM, T" collectorName="Krombein &amp; M. D. Hubbard" country="Sri Lanka" location="Ratnapura District" municipality="Balihyloya" pageId="8" pageNumber="37" specimenCount="14" specimenCount-female="7" specimenCount-male="7">
<collectorName box="[420,541,1599,1623]" pageId="8" pageNumber="37">Krombein</collectorName>
,
<specimenCount box="[569,603,1599,1623]" pageId="8" pageNumber="37" type="male">7♂</specimenCount>
,
<specimenCount box="[634,667,1599,1623]" pageId="8" pageNumber="37" type="female">7♀</specimenCount>
(
<collectionCode box="[699,786,1599,1623]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="37" type="Museum">USNM</collectionCode>
).
<location LSID="urn:lsid:plazi:treatment:94578784B629FFA0264DFA90FBBAAC30:19216049B629FFA027A4F9E7FE4FAD38" box="[189,421,1635,1659]" country="Sri Lanka" municipality="Balihyloya" name="Ratnapura District" pageId="8" pageNumber="37">Ratnapura District</location>
,
<collectingMunicipality box="[441,569,1635,1659]" pageId="8" pageNumber="37">Balihyloya</collectingMunicipality>
,
<date box="[591,793,1635,1659]" pageId="8" pageNumber="37" value="1978-04-09" valueMax="1978-04-10" valueMin="1978-04-09">
<collectingDate box="[591,793,1635,1659]" pageId="8" pageNumber="37" value="1978-04-09" valueMax="1978-04-10" valueMin="1978-04-09">9-10 April 1978</collectingDate>
</date>
,
<collectorName box="[189,388,1671,1695]" pageId="8" pageNumber="37">M.D. Hubbard</collectorName>
,
<collectionCode box="[422,441,1672,1695]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="37">T</collectionCode>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111499315" collectingDate="1978-06-23" collectingDateMax="1978-06-26" collectingDateMin="1978-06-23" collectionCode="USNM, V" collectorName="Wijesinhe" country="Sri Lanka" location="Uggalkaltota" pageId="8" pageNumber="37" specimenCount="1" specimenCount-female="1">
<collectorName box="[475,594,1671,1695]" pageId="8" pageNumber="37">Wijesinhe</collectorName>
,
<specimenCount box="[627,661,1671,1695]" pageId="8" pageNumber="37" type="female">1♀</specimenCount>
(
<collectionCode box="[699,786,1671,1695]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="37" type="Museum">USNM</collectionCode>
).
<location LSID="urn:lsid:plazi:treatment:94578784B629FFA0264DFA90FBBAAC30:19216049B629FFA027A4F92FFEB2AD80" box="[189,344,1707,1731]" country="Sri Lanka" name="Uggalkaltota" pageId="8" pageNumber="37">Uggalkaltota</location>
,
<date box="[356,545,1707,1731]" pageId="8" pageNumber="37" value="1978-06-23" valueMax="1978-06-26" valueMin="1978-06-23">
<collectingDate box="[356,545,1707,1731]" pageId="8" pageNumber="37" value="1978-06-23" valueMax="1978-06-26" valueMin="1978-06-23">23-26 June 1978</collectingDate>
</date>
, K.
<collectionCode box="[583,605,1707,1730]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="37" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111499312" collectionCode="T" collectorName="Krombein, P. B &amp; Karunaratne" country="Sri Lanka" pageId="8" pageNumber="37" specimenCount="1">
<collectorName box="[617,793,1707,1731]" pageId="8" pageNumber="37">Krombein, P.B</collectorName>
.
<collectorName box="[189,340,1743,1766]" pageId="8" pageNumber="37">Karunaratne</collectorName>
,
<collectionCode box="[361,380,1743,1766]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="37">T</collectionCode>
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="3111499320" collectionCode="USNM" collectorName="Wijesinhe, L &amp; Jayawickrema, N &amp; Karanaratne" country="Sri Lanka" pageId="8" pageNumber="37" specimenCount="44" specimenCount-female="15" specimenCount-male="29">
<collectorName box="[400,558,1743,1767]" pageId="8" pageNumber="37">Wijesinhe, L</collectorName>
.
<collectorName box="[578,793,1743,1767]" pageId="8" pageNumber="37">Jayawickrema, N</collectorName>
.
<collectorName box="[189,337,1779,1802]" pageId="8" pageNumber="37">Karanaratne</collectorName>
,
<specimenCount box="[348,395,1779,1803]" pageId="8" pageNumber="37" type="male">29♂</specimenCount>
,
<specimenCount box="[409,456,1779,1803]" pageId="8" pageNumber="37" type="female">15♀</specimenCount>
(
<collectionCode box="[470,557,1779,1803]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="37" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="8.[189,798,1526,1911]" lastBlockId="8.[835,1444,228,1907]" pageId="8" pageNumber="37">
<emphasis bold="true" box="[189,241,1815,1839]" pageId="8" pageNumber="37">Egg.</emphasis>
Spindle shaped. Length ca. 364-381 μm, equatorial width ca. 270-280 μm (
<figureCitation box="[584,667,1851,1875]" captionStart="Figs" captionStartId="9.[189,241,1810,1834]" captionTargetBox="[232,1404,290,1728]" captionTargetId="figure-95@9.[232,1404,290,1728]" captionTargetPageId="9" captionText="Figs. 22-27. Phanoperla wedda egg. 22. Lateral aspect. 23. Collar end (s = stria). 24. Anchor and collar (at = anchor tentacle). 25. Lid and micropylar zone. 26. Anchor and collar, apical aspect. 27. Micropyles (m = micropyle)." figureDoi="http://doi.org/10.5281/zenodo.4760920" httpUri="https://zenodo.org/record/4760920/files/figure.png" pageId="8" pageNumber="37">Fig. 22</figureCitation>
). Collar ca. 25-29 μm long and ca. 102-114 μm wide, margin slightly flanged and irregularly incised (
<figureCitation box="[1316,1444,228,252]" captionStart="Figs" captionStartId="9.[189,241,1810,1834]" captionTargetBox="[232,1404,290,1728]" captionTargetId="figure-95@9.[232,1404,290,1728]" captionTargetPageId="9" captionText="Figs. 22-27. Phanoperla wedda egg. 22. Lateral aspect. 23. Collar end (s = stria). 24. Anchor and collar (at = anchor tentacle). 25. Lid and micropylar zone. 26. Anchor and collar, apical aspect. 27. Micropyles (m = micropyle)." figureDoi="http://doi.org/10.5281/zenodo.4760920" httpUri="https://zenodo.org/record/4760920/files/figure.png" pageId="8" pageNumber="37">Figs. 23-24</figureCitation>
μm), sides bearing a series of thick vertical ridges (ca.
<quantity box="[884,941,300,324]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="8" pageNumber="37" unit="in" value="10.0">10 in</quantity>
lateral aspect); ridges continue onto body of egg for ca. 142 μm, forming a series of narrow striae, subtended on each side by a row of punctations; broad sulci between striae without punctations (
<figureCitation box="[992,1166,444,468]" captionStart="Figs" captionStartId="9.[189,241,1810,1834]" captionTargetBox="[232,1404,290,1728]" captionTargetId="figure-95@9.[232,1404,290,1728]" captionTargetPageId="9" captionText="Figs. 22-27. Phanoperla wedda egg. 22. Lateral aspect. 23. Collar end (s = stria). 24. Anchor and collar (at = anchor tentacle). 25. Lid and micropylar zone. 26. Anchor and collar, apical aspect. 27. Micropyles (m = micropyle)." figureDoi="http://doi.org/10.5281/zenodo.4760920" httpUri="https://zenodo.org/record/4760920/files/figure.png" pageId="8" pageNumber="37">Figs. 22-23, 26</figureCitation>
). Mesal chorionic zone (ca. 178 μm wide) consists of rather widely spaced coarse pits which extend over micropylar row and become somewhat finer beyond micropyles (
<figureCitation captionStart="Figs" captionStartId="9.[189,241,1810,1834]" captionTargetBox="[232,1404,290,1728]" captionTargetId="figure-95@9.[232,1404,290,1728]" captionTargetPageId="9" captionText="Figs. 22-27. Phanoperla wedda egg. 22. Lateral aspect. 23. Collar end (s = stria). 24. Anchor and collar (at = anchor tentacle). 25. Lid and micropylar zone. 26. Anchor and collar, apical aspect. 27. Micropyles (m = micropyle)." figureDoi="http://doi.org/10.5281/zenodo.4760920" httpUri="https://zenodo.org/record/4760920/files/figure.png" pageId="8" pageNumber="37">Fig. 25</figureCitation>
). Lid covered with follicle cell impressions, most containing five pits. Micropylar orifices circular, without prominent sperm guides (
<figureCitation box="[1241,1323,660,684]" captionStart="Figs" captionStartId="9.[189,241,1810,1834]" captionTargetBox="[232,1404,290,1728]" captionTargetId="figure-95@9.[232,1404,290,1728]" captionTargetPageId="9" captionText="Figs. 22-27. Phanoperla wedda egg. 22. Lateral aspect. 23. Collar end (s = stria). 24. Anchor and collar (at = anchor tentacle). 25. Lid and micropylar zone. 26. Anchor and collar, apical aspect. 27. Micropyles (m = micropyle)." figureDoi="http://doi.org/10.5281/zenodo.4760920" httpUri="https://zenodo.org/record/4760920/files/figure.png" pageId="8" pageNumber="37">Fig. 27</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="37" type="discussion">
<paragraph blockId="8.[835,1444,228,1907]" pageId="8" pageNumber="37">
<emphasis bold="true" box="[835,973,695,719]" pageId="8" pageNumber="37">Comments.</emphasis>
The scanning electron micrographs provide a few subtle chorionic details not observed in
<bibRefCitation author="Zwick, P." box="[872,1063,768,792]" pageId="8" pageNumber="37" pagination="87 - 126" refId="ref5664" refString="Zwick, P. 1982 a. A revision of the Oriental stonefly genus Phanoperla (Plecoptera: Perlidae). Systematic Entomology, 7: 87 - 126." type="journal article" year="1982">Zwicks (1982a)</bibRefCitation>
description. These include, 1) the extension of chorionic striae (or “ribs”) onto the collar (
<figureCitation box="[921,991,840,864]" captionStart="Figs" captionStartId="4.[189,241,1814,1838]" captionTargetBox="[232,1404,290,1732]" captionTargetId="figure-85@4.[232,1404,290,1732]" captionTargetPageId="4" captionText="Figs. 7-12. Putative Phanoperla nana egg. 7. Lateral aspect. 8. Collar end. 9. Chorionic detail surrounding collar. 10. Lid and micropylar zone. 11. Collar and anchor. 12. Micropyles." figureDoi="http://doi.org/10.5281/zenodo.4760914" httpUri="https://zenodo.org/record/4760914/files/figure.png" pageId="8" pageNumber="37">Fig. 8</figureCitation>
), 2) the presence of FCIs on the lid, and 3) continuation of the fine punctations to the base of lid (not shown in Zwicks
<figureCitation box="[1285,1384,912,936]" captionStart="Figs" captionStartId="9.[189,241,1810,1834]" captionTargetBox="[232,1404,290,1728]" captionTargetId="figure-95@9.[232,1404,290,1728]" captionTargetPageId="9" captionText="Figs. 22-27. Phanoperla wedda egg. 22. Lateral aspect. 23. Collar end (s = stria). 24. Anchor and collar (at = anchor tentacle). 25. Lid and micropylar zone. 26. Anchor and collar, apical aspect. 27. Micropyles (m = micropyle)." figureDoi="http://doi.org/10.5281/zenodo.4760920" httpUri="https://zenodo.org/record/4760920/files/figure.png" pageId="8" pageNumber="37">Fig. 24c</figureCitation>
, but noted in his description as “…punctures, very fine along otherwise invisible suture of lid;…”). Although the egg appears distinctive among the Sri Lankan species, several species (e.g.
<taxonomicName authority="Sivec &amp; Stark 2010 b" authorityName="Sivec &amp; Stark" authorityYear="2010" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="huang">
<emphasis box="[1274,1373,1056,1080]" italics="true" pageId="8" pageNumber="37">P. huang</emphasis>
<bibRefCitation author="Sivec, I. &amp; B. P. Stark" pageId="8" pageNumber="37" pagination="98 - 112" refId="ref5475" refString="Sivec, I. &amp; B. P. Stark. 2010 b. Seven new species of Phanoperla from Vietnam and Thailand (Plecoptera: Perlidae). Illiesia, 6: 98 - 112." type="journal article" year="2010">Sivec &amp; Stark 2010b</bibRefCitation>
</taxonomicName>
;
<taxonomicName authority="Sivec &amp; Stark 2011" authorityName="Sivec &amp; Stark" authorityYear="2011" box="[1016,1408,1092,1116]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="magnaspina">
<emphasis box="[1016,1177,1092,1115]" italics="true" pageId="8" pageNumber="37">P. magnaspina</emphasis>
<bibRefCitation author="Sivec, I. &amp; B. P. Stark" box="[1185,1408,1092,1116]" pageId="8" pageNumber="37" pagination="264 - 279" refId="ref5512" refString="Sivec, I. &amp; B. P. Stark. 2011. New species of Neoperla Needham and Phanoperla Banks (Plecoptera: Perlidae) from the Philippine Islands. Illiesia, 7: 264 - 279." type="journal article" year="2011">Sivec &amp; Stark 2011</bibRefCitation>
</taxonomicName>
;
<taxonomicName authority="Sivec &amp; Stark 2010 b" authorityName="Sivec &amp; Stark" authorityYear="2010" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="uchida">
<emphasis italics="true" pageId="8" pageNumber="37">P. uchida</emphasis>
<bibRefCitation author="Sivec, I. &amp; B. P. Stark" box="[917,1158,1128,1152]" pageId="8" pageNumber="37" pagination="98 - 112" refId="ref5475" refString="Sivec, I. &amp; B. P. Stark. 2010 b. Seven new species of Phanoperla from Vietnam and Thailand (Plecoptera: Perlidae). Illiesia, 6: 98 - 112." type="journal article" year="2010">Sivec &amp; Stark 2010b</bibRefCitation>
</taxonomicName>
) from other areas have eggs with a mixture of similar chorionic striae and punctations (
<bibRefCitation author="Sivec, I. &amp; B. P. Stark" box="[1003,1283,1200,1224]" pageId="8" pageNumber="37" pagination="98 - 112" refId="ref5475" refString="Sivec, I. &amp; B. P. Stark. 2010 b. Seven new species of Phanoperla from Vietnam and Thailand (Plecoptera: Perlidae). Illiesia, 6: 98 - 112." type="journal article" year="2010">Sivec &amp; Stark 2010b</bibRefCitation>
;
<bibRefCitation author="Sivec, I. &amp; B. P. Stark" box="[1308,1366,1200,1224]" pageId="8" pageNumber="37" pagination="264 - 279" refId="ref5512" refString="Sivec, I. &amp; B. P. Stark. 2011. New species of Neoperla Needham and Phanoperla Banks (Plecoptera: Perlidae) from the Philippine Islands. Illiesia, 7: 264 - 279." type="journal article" year="2011">2011</bibRefCitation>
). The common occurrence of this egg pattern in other species groups raises the question of whether the two named species in the
<taxonomicName authorityName=": Zwick" authorityYear="1982" box="[1205,1288,1308,1331]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="testacea">
<emphasis box="[1205,1288,1308,1331]" italics="true" pageId="8" pageNumber="37">testacea</emphasis>
</taxonomicName>
-group with unknown females (
<taxonomicName authorityName="Kawai" authorityYear="1975" box="[1075,1209,1344,1368]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="ceylonica">
<emphasis box="[1075,1209,1344,1368]" italics="true" pageId="8" pageNumber="37">P. ceylonica</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Zwick" authorityYear="1982" box="[1229,1350,1344,1368]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="srilanka">
<emphasis box="[1229,1350,1344,1368]" italics="true" pageId="8" pageNumber="37">P. srilanka</emphasis>
</taxonomicName>
) might have eggs similar enough to those of
<taxonomicName authorityName="Zwick" authorityYear="1982" box="[1281,1380,1379,1403]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="wedda">
<emphasis box="[1281,1380,1379,1403]" italics="true" pageId="8" pageNumber="37">P. wedda</emphasis>
</taxonomicName>
(also included in this group) to go undetected. We have examined at least
<specimenCount box="[1050,1143,1452,1476]" pageId="8" pageNumber="37" type="egg">one egg</specimenCount>
from every female listed above with light microscopy and have examined several eggs (ca. 10) from each of
<specimenCount box="[1252,1378,1523,1548]" pageId="8" pageNumber="37" type="female">10 females</specimenCount>
with SEM. No significant variation of the chorionic pattern has been detected but additional samples are needed from more localities to test this hypothesis.
</paragraph>
<paragraph blockId="8.[835,1444,228,1907]" pageId="8" pageNumber="37">
The following key is modified from that of
<bibRefCitation author="Zwick, P." box="[835,1006,1739,1763]" pageId="8" pageNumber="37" pagination="87 - 126" refId="ref5664" refString="Zwick, P. 1982 a. A revision of the Oriental stonefly genus Phanoperla (Plecoptera: Perlidae). Systematic Entomology, 7: 87 - 126." type="journal article" year="1982">Zwick (1982a)</bibRefCitation>
but includes only males of species known from
<collectingCountry box="[996,1058,1775,1799]" name="India" pageId="8" pageNumber="37">India</collectingCountry>
and
<collectingCountry box="[1123,1242,1775,1799]" name="Sri Lanka" pageId="8" pageNumber="37">Sri Lanka</collectingCountry>
. The Sri Lankan Plecopteran fauna is completely endemic, consequently the initial couplet is based on geographic separation.
</paragraph>
</subSubSection>
</treatment>
</document>